180 lines
26 KiB
XML
180 lines
26 KiB
XML
<document id="50325E6106015EF872BEF0CF6793F519" ID="10.11646/zootaxa.4196.1.1" ID-DOI="10.11646/zootaxa.4196.1.1" ID-GBIF-Dataset="b7651a7b-df24-4347-9585-0633b88dba51" ID-ISSN="1175-5326" ID-Zenodo-Dep="167648" ID-ZooBank="026D88BD-86C0-4811-814A-CB5CACB3104F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479712233041" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2016" docId="8D3787848912FFDB21CA3F36FCA2FDA5" docLanguage="en" docName="zootaxa.4196.1.1.pdf" docOrigin="Zootaxa 4196 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zeanillus pallidus Broun" docType="treatment" docVersion="8" lastPageNumber="17" masterDocId="710EFFFC891DFFCB215D3B54FFBCFFFA" masterDocTitle="A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution" masterLastPageNumber="37" masterPageNumber="1" pageNumber="16" updateTime="1698393955637" updateUser="plazi">
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<mods:title id="95272D1CA91D5163FA9AF971E5E5D521">A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution</mods:title>
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<mods:name id="1256A76E16B848AD81AB7049DEAC62E9" type="personal">
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<mods:roleTerm id="94825A8092044F2AFAD945B8A94339E3">Author</mods:roleTerm>
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<mods:namePart id="69DB759512C6220D87AB48AF25162BE6">Sokolov, Igor M.</mods:namePart>
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<mods:typeOfResource id="D85D42B5605E14AE27542AFA5A6516B1">text</mods:typeOfResource>
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<mods:title id="010D646AA1F3C699218BBF00053E8838">Zootaxa</mods:title>
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<mods:part id="FE89ACFA19C2332887358D2D7AF9DBE1">
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<mods:date id="6DF439FE2C956C14D94FB8EF7848AD75">2016</mods:date>
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<mods:number id="B85DCDA4576367F4D8272DEE37E9D089">4196</mods:number>
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<mods:number id="585D9F4E77D62241FCE5D588B4DE2D23">1</mods:number>
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<mods:classification id="D113A6D1A79BCA27A8031BEBD4582764">journal article</mods:classification>
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<mods:identifier id="F1BAABD64623F796E17378527A30ABCB" type="DOI">10.11646/zootaxa.4196.1.1</mods:identifier>
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<mods:identifier id="B2E747168DD64A99F56EFF530634D6AC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="321D08380C3750881CE2C1C2D1408495" type="Zenodo-Dep">167648</mods:identifier>
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<treatment id="8D3787848912FFDB21CA3F36FCA2FDA5" ID-DOI="http://doi.org/10.5281/zenodo.6089676" ID-GBIF-Taxon="124814318" ID-Zenodo-Dep="6089676" LSID="urn:lsid:plazi:treatment:8D3787848912FFDB21CA3F36FCA2FDA5" httpUri="http://treatment.plazi.org/id/8D3787848912FFDB21CA3F36FCA2FDA5" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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<subSubSection id="4D8465198912FFC421CA3F36FBF1FB64" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph id="052136928912FFC421CA3F36FE66FB87" blockId="15.[151,1101,1122,1182]" box="[151,474,1122,1149]" pageId="15" pageNumber="16">
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<heading id="5E6981FE8912FFC421CA3F36FE66FB87" bold="true" box="[151,474,1122,1149]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
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<taxonomicName id="C29E4D118912FFC421CA3F36FE66FB87" ID-CoL="8MF5D" authority="Broun" authorityName="Broun" box="[151,474,1122,1149]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pallidus">
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<emphasis id="37EAEA808912FFC421CA3F36FE66FB87" bold="true" box="[151,474,1122,1149]" pageId="15" pageNumber="16">
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<emphasis id="37EAEA808912FFC421CA3F36FECCFB86" bold="true" box="[151,368,1122,1148]" italics="true" pageId="15" pageNumber="16">Zeanillus pallidus</emphasis>
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(Broun)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="052136928912FFC421CA3FD1FBF1FB64" blockId="15.[151,1101,1122,1182]" box="[151,1101,1157,1182]" pageId="15" pageNumber="16">
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<figureCitation id="9DA52A178912FFC421CA3FD1FF5CFB64" box="[151,224,1157,1182]" captionStart="FIGURE 1" captionStartId="8.[151,250,1917,1939]" captionTargetBox="[172,1423,199,1885]" captionTargetId="figure@8.[164,1430,193,1891]" captionTargetPageId="8" captionText="FIGURE 1. SEM illustrations of head, dorsal aspect, of Zeanillus species. A— Z. phyllobius; B— Z. punctigerus; C— Z. pallidus; D— Z. pellucidus; E— Z. lescheni; F— Z. nanus. Legend: cl—clypeus; cs—clypeal seta; fcc—fronto-clypeal carina; fs—frontal seta; lb—labrum; mp 3 — maxillary palpomere 3; mp 4 — maxillary palpomere 4; pos—postorbital seta; ssa—anterior supraorbital seta; ssp—posterior supraorbital seta. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/257540/files/figure.png" pageId="15" pageNumber="16">Figs. 1</figureCitation>
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C, 2B, 3C–D, 4C, 5C, 6C, 7D, 8B, 10DIO, 11D, 12M–P, 14D, 15D, 16E, 18A
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</paragraph>
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</subSubSection>
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<subSubSection id="4D8465198912FFC421CA3F9DFF40FAE6" pageId="15" pageNumber="16" type="reference_group">
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<paragraph id="052136928912FFC421CA3F9DFF40FAE6" blockId="15.[151,1437,1225,1308]" pageId="15" pageNumber="16">
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<treatmentCitationGroup id="258E11BC8912FFC421CA3F9DFF40FAE6" pageId="15" pageNumber="16">
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<taxonomicName id="C29E4D118912FFC421CA3F9DFE4EFB25" ID-CoL="8L2MQ" authority="Broun, 1884: 228" authorityName="Broun" authorityPageNumber="228" authorityYear="1884" box="[151,498,1225,1247]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pallidus">
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<emphasis id="37EAEA808912FFC421CA3F9DFE8BFB25" box="[151,311,1225,1247]" italics="true" pageId="15" pageNumber="16">Anillus pallidus</emphasis>
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<treatmentCitation id="843F10838912FFC420603F9EFE4EFB25" author="Broun" box="[317,498,1225,1247]" page="228" pageId="15" pageNumber="16" year="1884">
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<bibRefCitation id="610F4B638912FFC420603F9EFE03FB25" author="Broun" box="[317,447,1225,1247]" pageId="15" pageNumber="16" refString="Broun, T. (1884) On a decade of new Feroniidae. New Zealand Journal of Science, 2, 226 - 228." type="journal article" year="1884">Broun, 1884</bibRefCitation>
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: 228
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</treatmentCitation>
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</taxonomicName>
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. Type locality: «Sea Beach, Taieri» (original citation).
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<taxonomicName id="C29E4D118912FFC421CA3FBCFDA8FB04" ID-CoL="8L2MP" authority="Broun, 1914: 152" authorityName="Broun" authorityPageNumber="152" authorityYear="1914" box="[151,532,1256,1278]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="marginatus">
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<emphasis id="37EAEA808912FFC421CA3FBCFEE4FB04" box="[151,344,1256,1278]" italics="true" pageId="15" pageNumber="16">Anillus marginatus</emphasis>
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<treatmentCitation id="843F10838912FFC420033FBDFDA8FB04" author="Broun" box="[350,532,1256,1278]" page="152" pageId="15" pageNumber="16" year="1914">
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<bibRefCitation id="610F4B638912FFC420033FBDFE5CFB04" author="Broun" box="[350,480,1256,1278]" pageId="15" pageNumber="16" refString="Broun, T. (1914) Descriptions of new genera and species of Coleoptera, Part III. New Zealand Institute Bulletin, 1, 143 - 266." type="journal article" year="1914">Broun, 1914</bibRefCitation>
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: 152
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</treatmentCitation>
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</taxonomicName>
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. Type locality: «Invercargill» (original citation). Synonymy established by
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<treatmentCitation id="843F10838912FFC4244D3FBDFF44FAE6" author="Moore" page="402" pageId="15" pageNumber="16" year="1980">
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<bibRefCitation id="610F4B638912FFC4244D3FBDFF44FAE6" author="Moore" pageId="15" pageNumber="16" refString="Moore, B. P. (1980) A synopsis of the New Zealand Anillini (Coleoptera: Carabidae: Bembidiinae), with descriptions of new genera and species. New Zealand Journal of Zoology, 7, 399 - 406. http: // dx. doi. org / 10.1080 / 03014223.1980.10423793" type="journal article" year="1980">Moore (1980: 402)</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="4D8465198912FFC421CA3E1DFE72F9EC" pageId="15" pageNumber="16" type="materials_examined">
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<paragraph id="052136928912FFC421CA3E1DFE46FA34" blockId="15.[151,1437,1353,2026]" pageId="15" pageNumber="16">
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<materialsCitation id="B5F63CCF8912FFC421CA3E1DFEEAFA50" ID-GBIF-Occurrence="1341350632" location="Flagstaff-Whare Flat Road" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Otago">
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<emphasis id="37EAEA808912FFC421CA3E1DFE3FFA98" bold="true" box="[151,387,1353,1378]" pageId="15" pageNumber="16">Examined material.</emphasis>
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More than one hundred specimens (15 dissected) from the following localities:
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<collectingRegion id="C75AF8708912FFC425A33E1EFAFBFA98" box="[1278,1351,1354,1378]" country="New Zealand" name="Otago" pageId="15" pageNumber="16">Otago</collectingRegion>
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: Symes Road at Old Man Range,
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<locationDeviation id="14932FB08912FFC420933E39FD17FA7F" box="[462,683,1389,1414]" pageId="15" pageNumber="16">
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<quantity id="C2669B778912FFC420933E39FDB1FA7C" box="[462,525,1389,1414]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="km" value="10.0">10km</quantity>
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S Beaumont
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</locationDeviation>
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,
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<location id="004160498912FFC423E63E39FC4CFA7C" LSID="urn:lsid:plazi:treatment:8D3787848912FFDB21CA3F36FCA2FDA5:004160498912FFC423E63E39FC4CFA7C" box="[699,1008,1389,1414]" name="Flagstaff-Whare Flat Road" pageId="15" pageNumber="16" stateProvince="Otago">Flagstaff-Whare Flat Road</location>
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near Dunedin, Taieri Mouth, Picnic Gulley, Akatore
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</materialsCitation>
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;
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<materialsCitation id="B5F63CCF8912FFC420373EC5FE46FA34" ID-GBIF-Occurrence="1341350617" location="Southland" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Southland">
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||
<collectingRegion id="C75AF8708912FFC420373EC5FE5CFA50" box="[362,480,1425,1450]" country="New Zealand" name="Southland" pageId="15" pageNumber="16">Southland</collectingRegion>
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: Otanomomo Reserve, Kaka Point Scenic Reserve, Titpua near Marbel Bush, Thomsons Bush in Invercargill.
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</materialsCitation>
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</paragraph>
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<paragraph id="052136928912FFC4219A3E8DFE72F9EC" blockId="15.[151,1437,1353,2026]" pageId="15" pageNumber="16">
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<emphasis id="37EAEA808912FFC4219A3E8DFEE1FA08" bold="true" box="[199,349,1497,1522]" pageId="15" pageNumber="16">Recognition.</emphasis>
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Adults of this species (
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<figureCitation id="9DA52A178912FFC4233B3E8DFD09FA08" box="[614,693,1497,1522]" captionStart="FIGURE 11" captionStartId="20.[151,250,1887,1909]" captionTargetBox="[198,1372,193,1855]" captionTargetId="figure@20.[198,1374,193,1857]" captionTargetPageId="20" captionText="FIGURE 11. Digital images of habitus of Zeanillus species, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Coopers Knob); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Road); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge): J— Z. nanus (NZ, Otago, Pisa Range). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/167659/files/figure.png" pageId="15" pageNumber="16">Fig. 11</figureCitation>
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D) are distinguished from other congeners by the characters listed in the subgeneric diagnosis.
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</paragraph>
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</subSubSection>
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<subSubSection id="4D8465198912FFDB219A3D75FCA2FDA5" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
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<paragraph id="052136928912FFC4219A3D75FB47F9C0" blockId="15.[151,1437,1353,2026]" box="[199,1275,1569,1594]" pageId="15" pageNumber="16">
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<emphasis id="37EAEA808912FFC4219A3D75FEE5F9C0" bold="true" box="[199,345,1569,1594]" pageId="15" pageNumber="16">Description.</emphasis>
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Size. Medium for genus (SBL range
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<quantity id="C2669B778912FFC423A13D75FC1DF9C3" box="[764,929,1569,1594]" metricMagnitude="-3" metricUnit="m" metricValue="1.73" metricValueMax="1.84" metricValueMin="1.62" pageId="15" pageNumber="16" unit="mm" value="1.73" valueMax="1.84" valueMin="1.62">1.62–1.84 mm</quantity>
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, mean 1.73±
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<quantity id="C2669B778912FFC425733D75FB19F9C3" box="[1070,1189,1569,1594]" metricMagnitude="-5" metricUnit="m" metricValue="5.4" pageId="15" pageNumber="16" unit="mm" value="0.054">0.054 mm</quantity>
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, n=20).
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</paragraph>
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<paragraph id="052136928912FFC4219A3D11FE4DF95C" blockId="15.[151,1437,1353,2026]" pageId="15" pageNumber="16">Habitus. Body form subdepressed, subparallel, moderately elongate (WE/SBL 0.34±0.006), head of moderate width for genus compared to pronotum (WH/WPm 0.77±0.018), pronotum of moderate width in comparison to elytra (WPm/WE 0.76±0.014).</paragraph>
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<paragraph id="052136928912FFC4219A3DE5FCA1F933" blockId="15.[151,1437,1353,2026]" box="[199,797,1713,1738]" pageId="15" pageNumber="16">Color. Body color testaceous, appendages testaceous.</paragraph>
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<paragraph id="052136928912FFC4219A3D81FC57F8CC" blockId="15.[151,1437,1353,2026]" pageId="15" pageNumber="16">Prothorax. Pronotum slightly long (LP/LE 0.39±0.012) and moderately elongate (WPm/LP 1.29±0.028), with lateral margins rectilinear and markedly constricted posteriorly (WPm/WPp 1.97±0.076). Width between anterior angles much greater than between posterior angles (WPa/WPp 1.51±0.069).</paragraph>
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<paragraph id="052136928912FFC4219A3C15FEA1F858" blockId="15.[151,1437,1353,2026]" pageId="15" pageNumber="16">Elytra. Moderately depressed, comparatively long (LE/SBL 0.51±0.011) and slightly wide (WE/LE 0.67±0.015). Lateral margins slightly divergent at basal fourth, subparallel at middle and evenly rounded to apex in apical third.</paragraph>
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<paragraph id="052136928912FFDB219A3CF9FD02FF2E" blockId="15.[151,1437,1353,2026]" lastBlockId="16.[151,1437,151,608]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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Male genitalia. Median lobe (
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<figureCitation id="9DA52A178912FFC4237B3CF9FDC6F83C" box="[550,634,1965,1990]" captionStart="FIGURE 12" captionStartId="21.[151,250,1424,1446]" captionTargetBox="[155,1428,199,1393]" captionTargetId="figure@21.[150,1436,193,1403]" captionTargetPageId="21" captionText="FIGURE 12. Line drawings of aedeagus of Zeanillus species. Z. punctigerus (NZ, Canterbury, Sharplin Falls): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. nunni (NZ, Otago, Evansdale): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. pallidus (NZ, Otago, Taieri Mouth): M—median lobe, right lateral aspect, N—apical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ao—apical orifice; bo—basal orifice; ds—dorsal sclerites, of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167660/files/figure.png" pageId="15" pageNumber="16">Fig. 12</figureCitation>
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M) with straight ventral margin and semicircular short apex. Apical orifice area without ostial fields. Sclerites of internal sac united basally, of moderate length, much shorter than the length between apical and basal orifices. Left paramere without apical constriction (
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<figureCitation id="9DA52A17890DFFDB250E3BC3FB1BFF4A" box="[1107,1191,151,176]" captionStart="FIGURE 12" captionStartId="21.[151,250,1424,1446]" captionTargetBox="[155,1428,199,1393]" captionTargetId="figure@21.[150,1436,193,1403]" captionTargetPageId="21" captionText="FIGURE 12. Line drawings of aedeagus of Zeanillus species. Z. punctigerus (NZ, Canterbury, Sharplin Falls): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. nunni (NZ, Otago, Evansdale): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. pallidus (NZ, Otago, Taieri Mouth): M—median lobe, right lateral aspect, N—apical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ao—apical orifice; bo—basal orifice; ds—dorsal sclerites, of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167660/files/figure.png" pageId="16" pageNumber="17">Fig. 12</figureCitation>
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O). Ring sclerite with short and markedly transverse handle (
|
||
<figureCitation id="9DA52A17890DFFDB23153BE8FD26FF2E" box="[584,666,188,213]" captionStart="FIGURE 14" captionStartId="23.[151,250,1033,1055]" captionTargetBox="[165,1404,198,1020]" captionTargetId="figure@23.[162,1414,193,1024]" captionTargetPageId="23" captionText="FIGURE 14. Line drawings of ring sclerite of Zeanillus species, male genitalia, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Orari Gorge); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Rd); D— Z. pallidus (NZ, Southland, Thomsons Bush); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. nanus (NZ, Otago, Pisa Range); G— Z. montivagus (NZ, Otago, North Rough Ridge); H— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); I— Z. lescheni (NZ, Otago, Waikaia Forest); J— Z. carltoni (NZ, Otago, Mount Watkin). Legend: hd—handle of ring sclerite. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167662/files/figure.png" pageId="16" pageNumber="17">Fig. 14</figureCitation>
|
||
D).
|
||
</paragraph>
|
||
<paragraph id="05213692890DFFDB219A3B8BFC8CFEBA" blockId="16.[151,1437,151,608]" pageId="16" pageNumber="17">
|
||
Female internal genitalia. Goncoxite 2 with short ensiform setae (
|
||
<figureCitation id="9DA52A17890DFFDB22AD3B8BFBF5FF02" box="[1008,1097,223,248]" captionStart="FIGURE 15" captionStartId="24.[151,250,1566,1588]" captionTargetBox="[159,1415,184,1535]" captionTargetId="figure@24.[150,1436,175,1545]" captionTargetPageId="24" captionText="FIGURE 15. Line drawings of right ovipositor sclerite, lateral aspect, of Zeanillus species. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Woodside Glen); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge); J— Z. nanus (NZ, Otago, Balclutha). Legend: es—ensiform seta; gc 1 — gonocoxite 1; gc 2 — gonocoxite 2; lt—laterotergite; ns—nematiform seta. Scale bar = 0.05 mm." httpUri="https://zenodo.org/record/167663/files/figure.png" pageId="16" pageNumber="17">Fig. 15</figureCitation>
|
||
D). Spermatheca sclerotized, elongate, with two compartments (
|
||
<figureCitation id="9DA52A17890DFFDB231D3A50FD27FEE7" box="[576,667,260,285]" captionStart="FIGURE 16" captionStartId="25.[151,250,1600,1622]" captionTargetBox="[171,1418,211,1571]" captionTargetId="figure@25.[151,1435,193,1579]" captionTargetPageId="25" captionText="FIGURE 16. Line drawings of spermatheca, spermathecal duct, and spermathecal gland of Zeanillus species. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Woodside Glen); D— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); E— Z. pallidus (NZ, Otago, Taieri Mouth); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. montivagus (NZ, Otago, North Rough Ridge); H— Z. nanus (NZ, Otago, Balclutha). Legend: sd—spermathecal duct; sg—spermathecal gland; sp—spermatheca. Scale bar = 0.05 mm." httpUri="https://zenodo.org/record/167664/files/figure.png" pageId="16" pageNumber="17">Fig. 16</figureCitation>
|
||
E). Spermathecal duct short and without coils. Attachments of spermathecal duct and gland to spermatheca close together.
|
||
</paragraph>
|
||
<paragraph id="05213692890DFFDB219A3A18FE1DFE57" blockId="16.[151,1437,151,608]" pageId="16" pageNumber="17">
|
||
<emphasis id="37EAEA80890DFFDB219A3A18FDB4FE9F" bold="true" box="[199,520,332,357]" pageId="16" pageNumber="17">Geographical distribution.</emphasis>
|
||
This species inhabits coastal plains of the eastern
|
||
<collectingRegion id="C75AF870890DFFDB250A3A18FB75FE9F" box="[1111,1225,332,357]" country="New Zealand" name="Southland" pageId="16" pageNumber="17">Southland</collectingRegion>
|
||
and South
|
||
<collectingRegion id="C75AF870890DFFDB240D3A18FA24FE9E" box="[1360,1432,332,356]" country="New Zealand" name="Otago" pageId="16" pageNumber="17">Otago</collectingRegion>
|
||
, from Invercargill up to Dunedin, also goes upstream along the Clutha River almost to Alexandra in Central
|
||
<collectingRegion id="C75AF870890DFFDB24053A24FA21FE72" box="[1368,1437,368,392]" country="New Zealand" name="Otago" pageId="16" pageNumber="17">Otago</collectingRegion>
|
||
(18A, green diamonds).
|
||
</paragraph>
|
||
<paragraph id="05213692890DFFDB219A3AE3FB3BFE2A" blockId="16.[151,1437,151,608]" box="[199,1159,439,464]" pageId="16" pageNumber="17">
|
||
<emphasis id="37EAEA80890DFFDB219A3AE3FEF3FE2A" bold="true" box="[199,335,439,464]" pageId="16" pageNumber="17">Way of life.</emphasis>
|
||
Specimens were collected in broadleaf/podocarp and
|
||
<taxonomicName id="C29E4D11890DFFDB22F13AECFB91FE2A" box="[940,1069,440,464]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="37EAEA80890DFFDB22F13AECFB91FE2A" box="[940,1069,440,464]" italics="true" pageId="16" pageNumber="17">Nothofagus</emphasis>
|
||
</taxonomicName>
|
||
forests.
|
||
</paragraph>
|
||
<paragraph id="05213692890DFFDB219A3A88FCA2FDA5" blockId="16.[151,1437,151,608]" pageId="16" pageNumber="17">
|
||
<emphasis id="37EAEA80890DFFDB219A3A88FECCFE0F" bold="true" box="[199,368,476,501]" pageId="16" pageNumber="17">Relationships.</emphasis>
|
||
The structure of male genitalia suggest the relatedness to the representatives of
|
||
<taxonomicName id="C29E4D11890DFFDB25A43A88FADDFE0E" box="[1273,1377,476,500]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="37EAEA80890DFFDB25A43A88FADDFE0E" box="[1273,1377,476,500]" italics="true" pageId="16" pageNumber="17">Zeanillus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2CD957FB890DFFDB24343A89FA20FE0E" box="[1385,1436,477,500]" pageId="16" pageNumber="17" sensu="stricto">s.str.</taxonomicNameLabel>
|
||
The absence of postorbital seta is a character shared with the members of
|
||
<emphasis id="37EAEA80890DFFDB22B13954FBCCFDE2" box="[1004,1136,512,536]" italics="true" pageId="16" pageNumber="17">Otagonillus</emphasis>
|
||
. Based on these facts and distributional data it is postulated that
|
||
<taxonomicName id="C29E4D11890DFFDB231B3971FD02FDC6" box="[582,702,548,572]" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pallidus">
|
||
<emphasis id="37EAEA80890DFFDB231B3971FD02FDC6" box="[582,702,548,572]" italics="true" pageId="16" pageNumber="17">Z. pallidus</emphasis>
|
||
</taxonomicName>
|
||
is the sister taxon to the members of
|
||
<taxonomicName id="C29E4D11890DFFDB25383970FB71FDC6" box="[1125,1229,548,572]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="37EAEA80890DFFDB25383970FB71FDC6" box="[1125,1229,548,572]" italics="true" pageId="16" pageNumber="17">Zeanillus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2CD957FB890DFFDB25893971FAB4FDC6" box="[1236,1288,549,572]" pageId="16" pageNumber="17" sensu="stricto">s.str.</taxonomicNameLabel>
|
||
and shared a common ancestry with the representatives of
|
||
<emphasis id="37EAEA80890DFFDB23CE391CFCABFD9A" box="[659,791,584,608]" italics="true" pageId="16" pageNumber="17">Otagonillus</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |