257 lines
34 KiB
XML
257 lines
34 KiB
XML
<document id="75C35B4FD16312DB4CBE034AA431B46C" ID-DOI="10.11646/zootaxa.3750.5.7" ID-GBIF-Dataset="d35e62c7-975b-4dff-892e-e8e5c926820e" ID-ISSN="1175-5326" ID-Zenodo-Dep="217852" ID-ZooBank="22C6D62A-D6CA-4BBE-AD0D-29174C2A2769" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460711129622" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2013" docId="7505414FFF8D736DFF29FA0EFC0746E1" docLanguage="en" docName="zt03750p568.pdf" docOrigin="Zootaxa 3750 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alloraphes peruanus Franz" docType="treatment" docVersion="6" lastPageNumber="564" masterDocId="893C3937FF837362FFBEFFC2FFB14304" masterDocTitle="Revision of the Neotropical genus Alloraphes Franz (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="568" masterPageNumber="549" pageNumber="563" updateTime="1698357997849" updateUser="plazi">
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<mods:titleInfo id="C261B7A92F90F8F3604609EA137C643E">
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<mods:title id="183CF3CBD1167D2BB88B7C538026362A">Revision of the Neotropical genus Alloraphes Franz (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:name id="4CC2983CE7390854D6446B2433A13A58" type="personal">
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<mods:roleTerm id="68249A9D4D40DDA6D8BDB643A07CCC79">Author</mods:roleTerm>
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<mods:namePart id="3210B37AED5225618B32EF6872ECBB03">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="E5903CCB4739C2E09082E38CE604A555">text</mods:typeOfResource>
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<mods:title id="C8E888A5896D38AAFF984A995B23388F">Zootaxa</mods:title>
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<mods:part id="78B062A20C6EE43D9EF5EE624F8B4D62">
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<mods:date id="B02133118430031EEF709F4349CF3B6B">2013</mods:date>
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<mods:number id="7A3CC25B5E23A0B189D4F0788D2056CD">3750</mods:number>
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<mods:detail id="18CCE52ECAE3EB954ACE7BA4287B8A99" type="issue">
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<mods:number id="8156D5009E31FCD37B258530AECD937A">5</mods:number>
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<mods:classification id="C3764D49DCA4171BA715A4BE6AE036EF">journal article</mods:classification>
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<mods:identifier id="00476534ED2AD82B9802DE10C0F87FBE" type="DOI">10.11646/zootaxa.3750.5.7</mods:identifier>
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<mods:identifier id="74BEE133DD787FF05D33A5044A8B5E47" type="ISSN">1175-5326</mods:identifier>
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<treatment id="7505414FFF8D736DFF29FA0EFC0746E1" ID-DOI="http://doi.org/10.5281/zenodo.6154531" ID-GBIF-Taxon="119585167" ID-Zenodo-Dep="6154531" LSID="urn:lsid:plazi:treatment:7505414FFF8D736DFF29FA0EFC0746E1" httpUri="http://treatment.plazi.org/id/7505414FFF8D736DFF29FA0EFC0746E1" lastPageId="15" lastPageNumber="564" pageId="14" pageNumber="563">
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<subSubSection id="B5B6A3D2FF8D736CFF29FA0EFD214548" pageId="14" pageNumber="563" type="nomenclature">
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<paragraph id="FD13F059FF8D736CFF29FA0EFE5746E2" blockId="14.[151,497,1484,1543]" box="[151,486,1484,1510]" pageId="14" pageNumber="563">
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<heading id="A65B4735FF8D736CFF29FA0EFE5746E2" bold="true" box="[151,486,1484,1510]" fontSize="11" level="1" pageId="14" pageNumber="563" reason="1">
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<taxonomicName id="3AAC8BDAFF8D736CFF29FA0EFE5746E2" ID-CoL="BYZJ" authority="Franz" authorityName="Franz" box="[151,486,1484,1510]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="563" phylum="Arthropoda" rank="species" species="peruanus">
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<emphasis id="CFD82C4BFF8D736CFF29FA0EFE5746E2" bold="true" box="[151,486,1484,1510]" pageId="14" pageNumber="563">
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<emphasis id="CFD82C4BFF8D736CFF29FA0EFE2546E2" bold="true" box="[151,404,1484,1510]" italics="true" pageId="14" pageNumber="563">Alloraphes peruanus</emphasis>
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Franz
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="FD13F059FF8D736CFF29FA2CFE404503" blockId="14.[151,497,1484,1543]" box="[151,497,1518,1543]" pageId="14" pageNumber="563">
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(
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<figureCitation id="6597ECDCFF8D736CFF21FA2CFF5F4503" box="[159,238,1518,1543]" captionStart="FIGURES 1 – 12" captionStartId="3.[151,264,1879,1902]" captionTargetBox="[217,1363,203,1830]" captionTargetId="figure@3.[209,1368,195,1845]" captionTargetPageId="3" captionText="FIGURES 1 – 12. Dorsal (1 – 9, 11 – 12) and laterodorsal (10) habitus of Alloraphes Franz. Alloraphes jamaicae Franz, holotype male (1); A. chiapasensis Franz, holotype male (2); A. yucatani Franz, holotype male (3); A. lenkoi Franz, paratype male (4) and paratype female (5); A. peruanus Franz, holotype male (6) and paratype female (7; but see the Remarks section in the redescription); A. dentatus sp. n., holotype male (8); A. peckorum sp. n., holotype male (9 – 10); A. myrmecophilus Franz, stat. rest., paratype male (11);? Alloraphes magnus Franz, holotype female (12)." httpUri="https://zenodo.org/record/217853/files/figure.png" pageId="14" pageNumber="563">Figs. 6</figureCitation>
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,
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<figureCitation id="6597ECDCFF8D736CFF44FA2CFEF24503" box="[250,323,1518,1543]" captionStart="FIGURES 27 – 43" captionStartId="8.[151,264,1654,1677]" captionTargetBox="[156,1426,184,1626]" captionTargetId="figure@8.[151,1436,167,1633]" captionTargetPageId="8" captionText="FIGURES 27 – 43. Aedeagus (27 – 34, 36 – 43) and female genitalia (35) of Alloraphes Franz in ventral (27, 29, 31, 33, 35, 36, 38, 40, 42) and lateral (28, 30, 32, 34, 37, 39, 41, 43) views. Alloraphes jamaicae Franz, holotype (27 – 28); A. chiapasensis Franz, holotype, parameres missing (29 – 30); A. yucatani Franz, holotype (31 – 32); A. lenkoi Franz, paratypes (33 – 35); A. peruanus Franz, holotype (36 – 37); A. dentatus sp. n., holotype (38 – 39); A. peckorum sp. n., holotype (40 – 41); A. myrmecophilus Franz, stat. rest., paratype (42 – 43). Abbreviations: ap, apical projection; bpa, basal pumping apparatus; ov, egg; sp, spermatheca." httpUri="https://zenodo.org/record/217858/files/figure.png" pageId="14" pageNumber="563">36–37</figureCitation>
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,
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<figureCitation id="6597ECDCFF8D736CFEF0FA2CFE264503" box="[334,407,1518,1543]" captionStart="FIGURES 44 – 60" captionStartId="10.[151,264,1803,1826]" captionTargetBox="[162,1411,295,1754]" captionTargetId="figure@10.[151,1436,283,1765]" captionTargetPageId="10" captionText="FIGURES 44 – 60. Aedeagus (44 – 51, 53 – 60) and female genitalia (52) of Alloraphes Franz in ventral (44, 46, 48, 50, 52, 53, 55, 57, 59) and lateral (45, 47, 49, 51, 54, 56, 58, 60) views. Alloraphes jamaicae Franz, holotype (44 – 45); A. chiapasensis Franz, holotype, paramere detached (46 – 47); A. yucatani Franz, holotype (48 – 49); A. lenkoi Franz, paratypes (50 – 52); A. peruanus Franz, holotype (53 – 54); A. dentatus sp. n., holotype (55 – 56); A. peckorum sp. n., holotype (57 – 58); A. myrmecophilus Franz, stat. rest., paratype (59 – 60). Abbreviations: ap, apical projection; bpa, basal pumping apparatus; ov, egg; sp, spermatheca." httpUri="https://zenodo.org/record/217859/files/figure.png" pageId="14" pageNumber="563">53–54</figureCitation>
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,
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<figureCitation id="6597ECDCFF8D736CFE1CFA2CFE704503" box="[418,449,1518,1543]" captionStart="FIGURES 61 – 67" captionStartId="11.[151,264,1809,1832]" captionTargetBox="[161,1423,953,1744]" captionTargetId="figure@11.[151,1436,939,1757]" captionTargetPageId="11" captionText="FIGURES 61 – 67. Original sets of labels pinned under historical type specimens. Alloraphes jamaicae Franz, holotype (61); A. chiapasensis Franz, holotype (62); A. yucatani Franz, holotype (63); A. lenkoi Franz, paratype (64); A. peruanus Franz, holotype (65); A. myrmecophilus Franz, stat. rest., paratype (66);? Alloraphes magnus Franz, holotype (67)." httpUri="https://zenodo.org/record/217860/files/figure.png" pageId="14" pageNumber="563">65</figureCitation>
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,
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<figureCitation id="6597ECDCFF8D736CFE72FA2CFE584503" box="[460,489,1518,1543]" captionStart="FIGURE 68" captionStartId="12.[151,250,1290,1312]" captionTargetBox="[151,1418,167,1268]" captionTargetId="figure@12.[151,1436,167,1268]" captionTargetPageId="12" captionText="FIGURE 68. Distribution of Alloraphes: a, A. jamaicae Franz; b, A. chiapasensis Franz; c, A. yucatani Franz; d, A. lenkoi Franz; e, A. peruanus Franz; f, A. dentatus sp. n.; g, A. peckorum sp. n.; h, A. myrmecophilus Franz stat. rest.; i, A. bolivarensis Franz; j,? Alloraphes magnus Franz." httpUri="https://zenodo.org/record/217861/files/figure.png" pageId="14" pageNumber="563">68</figureCitation>
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)
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</paragraph>
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<paragraph id="FD13F059FF8D736CFF29F9F4FD214548" blockId="14.[151,656,1590,1613]" box="[151,656,1590,1613]" pageId="14" pageNumber="563">
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<taxonomicName id="3AAC8BDAFF8D736CFF29F9F4FD9B4548" authority="Franz, 1980a: 214" authorityName="Franz" authorityPageNumber="214" authorityYear="1980" box="[151,554,1590,1613]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="563" phylum="Arthropoda" rank="species" species="peruanus">
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<emphasis id="CFD82C4BFF8D736CFF29F9F4FEDB4549" box="[151,362,1590,1613]" italics="true" pageId="14" pageNumber="563">Alloraphes peruanus</emphasis>
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Franz, 1980a: 214
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</taxonomicName>
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, Fig. 200.
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</paragraph>
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</subSubSection>
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<subSubSection id="B5B6A3D2FF8D736CFF29F9BBFD9F44B6" pageId="14" pageNumber="563" type="materials_examined">
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<paragraph id="FD13F059FF8D736CFF29F9BBFAA745FA" blockId="14.[151,1436,1657,1970]" pageId="14" pageNumber="563">
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<emphasis id="CFD82C4BFF8D736CFF29F9BBFD074596" bold="true" box="[151,694,1657,1682]" pageId="14" pageNumber="563">
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<typeStatus id="22174EFBFF8D736CFF29F9BBFF634596" box="[151,210,1657,1682]" pageId="14" pageNumber="563">Type</typeStatus>
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material.
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<typeStatus id="22174EFBFF8D736CFEF0F9BBFE0E4596" box="[334,447,1657,1682]" pageId="14" pageNumber="563" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="85BBB0C9FF8D736CFE73F9BBFDB64596" box="[461,519,1657,1682]" name="Peru" pageId="14" pageNumber="563">Peru</collectingCountry>
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(San Martín):
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</emphasis>
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♂, three labels (
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<figureCitation id="6597ECDCFF8D736CFCCBF9BBFC7A4596" box="[885,971,1657,1682]" captionStart="FIGURES 61 – 67" captionStartId="11.[151,264,1809,1832]" captionTargetBox="[161,1423,953,1744]" captionTargetId="figure@11.[151,1436,939,1757]" captionTargetPageId="11" captionText="FIGURES 61 – 67. Original sets of labels pinned under historical type specimens. Alloraphes jamaicae Franz, holotype (61); A. chiapasensis Franz, holotype (62); A. yucatani Franz, holotype (63); A. lenkoi Franz, paratype (64); A. peruanus Franz, holotype (65); A. myrmecophilus Franz, stat. rest., paratype (66);? Alloraphes magnus Franz, holotype (67)." httpUri="https://zenodo.org/record/217860/files/figure.png" pageId="14" pageNumber="563">Fig. 65</figureCitation>
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): "Umg. Tarapoto /
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<collectingCountry id="85BBB0C9FF8D736CFB11F9B8FB544595" box="[1199,1253,1658,1681]" name="Peru" pageId="14" pageNumber="563">Peru</collectingCountry>
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, lg. Franz" with "SA133" on reverse side [white, printed; reverse handwritten in blue ink]; "
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<taxonomicName id="3AAC8BDAFF8D736CFBB4F95FFB3445B1" box="[1034,1157,1693,1717]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="563" phylum="Arthropoda" rank="genus">
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<emphasis id="CFD82C4BFF8D736CFBB4F95FFB3445B1" box="[1034,1157,1693,1717]" italics="true" pageId="14" pageNumber="563">Alloraphes</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="3AAC8BDAFF8D736CFB1FF95CFAB845B1" box="[1185,1289,1694,1717]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="563" phylum="Arthropoda" rank="species" species="peruanus">
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<emphasis id="CFD82C4BFF8D736CFB1FF95CFAB845B1" box="[1185,1289,1694,1717]" italics="true" pageId="14" pageNumber="563">peruanus</emphasis>
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</taxonomicName>
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m. / det. H. Franz" [white, handwritten and printed]; "
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<typeStatus id="22174EFBFF8D736CFDD0F900FD0645DE" box="[622,695,1730,1754]" pageId="14" pageNumber="563">Typus</typeStatus>
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" [red, handwritten] (NHMW).
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<typeStatus id="22174EFBFF8D736CFBA9F900FB3545DE" box="[1047,1156,1730,1754]" pageId="14" pageNumber="563" type="paratype">Paratypes</typeStatus>
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(2 ♀; identity uncertain, see the Remarks section below): same data as
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<typeStatus id="22174EFBFF8D736CFD22F927FCB045FA" box="[668,769,1765,1790]" pageId="14" pageNumber="563" type="holotype">holotype</typeStatus>
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, except for yellow "
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<typeStatus id="22174EFBFF8D736CFC5EF924FBE345FA" box="[992,1106,1766,1790]" pageId="14" pageNumber="563" type="paratype">paratypus</typeStatus>
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" label (NHMW).
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</paragraph>
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<paragraph id="FD13F059FF8D736CFF79F8CBFD9F44B6" blockId="14.[151,1436,1657,1970]" pageId="14" pageNumber="563">
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<emphasis id="CFD82C4BFF8D736CFF79F8CBFE104426" bold="true" box="[199,417,1801,1826]" pageId="14" pageNumber="563">Revised diagnosis.</emphasis>
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Male: frons and vertex confluent, without impression; pronotum with evenly arcuate antebasal transverse groove and without median pit; elytra unmodified; aedeagus in ventral view with nearly symmetrical apex of median lobe, long apical projections and narrow apices of parameres, one bearing two thin and short setae, the other one with a dense row of several thin and short setae. Female diagnostic characters unknown (see the Remarks section).
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</paragraph>
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</subSubSection>
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<subSubSection id="B5B6A3D2FF8C736DFF79FF55FD7440CC" pageId="15" pageNumber="564" type="description">
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<paragraph id="FD13F059FF8C736DFF79FF55FF7B43D0" blockId="15.[151,1436,151,1509]" pageId="15" pageNumber="564">
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<emphasis id="CFD82C4BFF8C736DFF79FF55FEC543B4" bold="true" box="[199,372,151,176]" pageId="15" pageNumber="564">Redescription.</emphasis>
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Body of male (
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<figureCitation id="6597ECDCFF8C736DFD92FF55FDC343B4" box="[556,626,151,176]" captionStart="FIGURES 1 – 12" captionStartId="3.[151,264,1879,1902]" captionTargetBox="[217,1363,203,1830]" captionTargetId="figure@3.[209,1368,195,1845]" captionTargetPageId="3" captionText="FIGURES 1 – 12. Dorsal (1 – 9, 11 – 12) and laterodorsal (10) habitus of Alloraphes Franz. Alloraphes jamaicae Franz, holotype male (1); A. chiapasensis Franz, holotype male (2); A. yucatani Franz, holotype male (3); A. lenkoi Franz, paratype male (4) and paratype female (5); A. peruanus Franz, holotype male (6) and paratype female (7; but see the Remarks section in the redescription); A. dentatus sp. n., holotype male (8); A. peckorum sp. n., holotype male (9 – 10); A. myrmecophilus Franz, stat. rest., paratype male (11);? Alloraphes magnus Franz, holotype female (12)." httpUri="https://zenodo.org/record/217853/files/figure.png" pageId="15" pageNumber="564">Fig. 6</figureCitation>
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) strongly convex, brown, covered with light brown vestiture; BL
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<quantity id="3A545DBCFF8C736DFAD5FF5AFF7743D0" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="15" pageNumber="564" unit="mm" value="0.83">0.83 mm</quantity>
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.
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</paragraph>
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<paragraph id="FD13F059FF8C736DFF79FF1DFC83428C" blockId="15.[151,1436,151,1509]" pageId="15" pageNumber="564">
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Head broadest at strikingly large, strongly convex and coarsely faceted eyes, HL
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||
<quantity id="3A545DBCFF8C736DFBC2FF22FB5443F3" box="[1148,1253,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="564" unit="mm" value="0.13">0.13 mm</quantity>
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, HW
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<quantity id="3A545DBCFF8C736DFA93FF22FA2943F3" box="[1325,1432,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="564" unit="mm" value="0.2">0.20 mm</quantity>
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||
; tempora absent, eyes adjacent to occipital constriction; vertex and frons confluent, uniformly and weakly convex, anterior part of frons slightly flattened; supraantennal tubercles feebly marked. Punctures on frons and vertex inconspicuous, barely discernible; setae in the only known specimen almost entirely missing. Only rudiments of antennae preserved (left scape and right scape and pedicel).
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||
</paragraph>
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<paragraph id="FD13F059FF8C736DFF79FE57FE644164" blockId="15.[151,1436,151,1509]" pageId="15" pageNumber="564">
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Pronotum only slightly elongate, broadest in anterior fourth; PL
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||
<quantity id="3A545DBCFF8C736DFC21FE56FBB442A8" box="[927,1029,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="15" pageNumber="564" unit="mm" value="0.23">0.23 mm</quantity>
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||
, PW
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||
<quantity id="3A545DBCFF8C736DFBFAFE56FB1A42A8" box="[1092,1195,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="564" unit="mm" value="0.21">0.21 mm</quantity>
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||
. Anterior margin and lateral margins in anterior third strongly rounded; sides distinctly but slightly narrowing caudad, pronotum distinctly constricted near posterior third; posterior margin slightly arcuate; ante-basal transverse groove distinct and evenly arcuate, without median pit and indistinctly deepened at each end. Punctures on pronotal disc inconspicuous, fine and sparse; setae on disc not preserved, on sides of pronotum visible remains of moderately long and disordered bristles.
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||
</paragraph>
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||
<paragraph id="FD13F059FF8C736DFF79FDAEFA2841F4" blockId="15.[151,1436,151,1509]" pageId="15" pageNumber="564">
|
||
Elytra more convex than pronotum, oval and unmodified, broadest slightly anterior to middle; EL
|
||
<quantity id="3A545DBCFF8C736DFA91FDAEFA264180" box="[1327,1431,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="15" pageNumber="564" unit="mm" value="0.48">0.48 mm</quantity>
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, EW
|
||
<quantity id="3A545DBCFF8C736DFF75FD52FE8241A3" box="[203,307,656,680]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="15" pageNumber="564" unit="mm" value="0.38">0.38 mm</quantity>
|
||
, EI 1.27; sub-humeral lines sharply marked and as long as 0.6 EL. Punctures on elytra distinct but small, shallow and with diffused margins, separated by spaces comparable to puncture diameters; setae on the only known specimen only partly preserved, relatively long, sparse and distinctly suberect.
|
||
<collectingCountry id="85BBB0C9FF8C736DFBE0FD15FB2741F4" box="[1118,1174,727,752]" name="India" pageId="15" pageNumber="564">Hind</collectingCountry>
|
||
wings well developed.
|
||
</paragraph>
|
||
<paragraph id="FD13F059FF8C736DFF79FD3FFD214011" blockId="15.[151,1436,151,1509]" box="[199,656,764,789]" pageId="15" pageNumber="564">Legs long and slender; all tibiae straight.</paragraph>
|
||
<paragraph id="FD13F059FF8C736DFF79FCDDFB0C40A0" blockId="15.[151,1436,151,1509]" pageId="15" pageNumber="564">
|
||
Aedeagus (
|
||
<figureCitation id="6597ECDCFF8C736DFEF8FCDDFE61403C" box="[326,464,799,824]" captionStart="FIGURES 27 – 43" captionStartId="8.[151,264,1654,1677]" captionTargetBox="[156,1426,184,1626]" captionTargetId="figure@8.[151,1436,167,1633]" captionTargetPageId="8" captionText="FIGURES 27 – 43. Aedeagus (27 – 34, 36 – 43) and female genitalia (35) of Alloraphes Franz in ventral (27, 29, 31, 33, 35, 36, 38, 40, 42) and lateral (28, 30, 32, 34, 37, 39, 41, 43) views. Alloraphes jamaicae Franz, holotype (27 – 28); A. chiapasensis Franz, holotype, parameres missing (29 – 30); A. yucatani Franz, holotype (31 – 32); A. lenkoi Franz, paratypes (33 – 35); A. peruanus Franz, holotype (36 – 37); A. dentatus sp. n., holotype (38 – 39); A. peckorum sp. n., holotype (40 – 41); A. myrmecophilus Franz, stat. rest., paratype (42 – 43). Abbreviations: ap, apical projection; bpa, basal pumping apparatus; ov, egg; sp, spermatheca." httpUri="https://zenodo.org/record/217858/files/figure.png" pageId="15" pageNumber="564">Figs. 36–37</figureCitation>
|
||
,
|
||
<figureCitation id="6597ECDCFF8C736DFE60FCDDFD97403C" box="[478,550,799,824]" captionStart="FIGURES 44 – 60" captionStartId="10.[151,264,1803,1826]" captionTargetBox="[162,1411,295,1754]" captionTargetId="figure@10.[151,1436,283,1765]" captionTargetPageId="10" captionText="FIGURES 44 – 60. Aedeagus (44 – 51, 53 – 60) and female genitalia (52) of Alloraphes Franz in ventral (44, 46, 48, 50, 52, 53, 55, 57, 59) and lateral (45, 47, 49, 51, 54, 56, 58, 60) views. Alloraphes jamaicae Franz, holotype (44 – 45); A. chiapasensis Franz, holotype, paramere detached (46 – 47); A. yucatani Franz, holotype (48 – 49); A. lenkoi Franz, paratypes (50 – 52); A. peruanus Franz, holotype (53 – 54); A. dentatus sp. n., holotype (55 – 56); A. peckorum sp. n., holotype (57 – 58); A. myrmecophilus Franz, stat. rest., paratype (59 – 60). Abbreviations: ap, apical projection; bpa, basal pumping apparatus; ov, egg; sp, spermatheca." httpUri="https://zenodo.org/record/217859/files/figure.png" pageId="15" pageNumber="564">53–54</figureCitation>
|
||
) slender and lightly sclerotized; AeL
|
||
<quantity id="3A545DBCFF8C736DFC67FCDDFBF34033" box="[985,1090,799,824]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="564" unit="mm" value="0.25">0.25 mm</quantity>
|
||
; in ventral view apical part of median lobe nearly symmetrical and with broad, shallow and asymmetrical emargination of apical margin; apical projections long; parameres in lateral view narrowing only in short and apical part which is curved dorsally, one bearing two thin and short setae, the other one with a dense row of several thin and short setae.
|
||
</paragraph>
|
||
<paragraph id="FD13F059FF8C736DFF79FC6DFD7440CC" blockId="15.[151,1436,151,1509]" box="[199,709,943,968]" pageId="15" pageNumber="564">Female. Unknown (see the Remarks section).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5B6A3D2FF8C736DFF79FC16FD3240E9" box="[199,643,980,1005]" pageId="15" pageNumber="564" type="distribution">
|
||
<paragraph id="FD13F059FF8C736DFF79FC16FD3240E9" blockId="15.[151,1436,151,1509]" box="[199,643,980,1005]" pageId="15" pageNumber="564">
|
||
<emphasis id="CFD82C4BFF8C736DFF79FC16FED140E9" bold="true" box="[199,352,980,1005]" pageId="15" pageNumber="564">Distribution.</emphasis>
|
||
Northern
|
||
<collectingCountry id="85BBB0C9FF8C736DFE6DFC17FDB740E8" box="[467,518,981,1004]" name="Peru" pageId="15" pageNumber="564">Peru</collectingCountry>
|
||
(
|
||
<figureCitation id="6597ECDCFF8C736DFDA8FC16FDD840E9" box="[534,617,980,1005]" captionStart="FIGURE 68" captionStartId="12.[151,250,1290,1312]" captionTargetBox="[151,1418,167,1268]" captionTargetId="figure@12.[151,1436,167,1268]" captionTargetPageId="12" captionText="FIGURE 68. Distribution of Alloraphes: a, A. jamaicae Franz; b, A. chiapasensis Franz; c, A. yucatani Franz; d, A. lenkoi Franz; e, A. peruanus Franz; f, A. dentatus sp. n.; g, A. peckorum sp. n.; h, A. myrmecophilus Franz stat. rest.; i, A. bolivarensis Franz; j,? Alloraphes magnus Franz." httpUri="https://zenodo.org/record/217861/files/figure.png" pageId="15" pageNumber="564">Fig. 68</figureCitation>
|
||
e).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5B6A3D2FF8C736DFF79FC35FC0746E1" pageId="15" pageNumber="564" type="discussion">
|
||
<paragraph id="FD13F059FF8C736DFF79FC35FC0746E1" blockId="15.[151,1436,151,1509]" pageId="15" pageNumber="564">
|
||
<emphasis id="CFD82C4BFF8C736DFF79FC35FE8A4714" bold="true" box="[199,315,1015,1040]" pageId="15" pageNumber="564">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="22174EFBFF8C736DFECAFC3AFE154714" box="[372,420,1016,1040]" pageId="15" pageNumber="564">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="3AAC8BDAFF8C736DFDAFFC3AFD294714" box="[529,664,1016,1040]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="perunaus">
|
||
<emphasis id="CFD82C4BFF8C736DFDAFFC3AFD294714" box="[529,664,1016,1040]" italics="true" pageId="15" pageNumber="564">A. perunaus</emphasis>
|
||
</taxonomicName>
|
||
is composed of a
|
||
<typeStatus id="22174EFBFF8C736DFCDAFC35FC774714" box="[868,966,1015,1040]" pageId="15" pageNumber="564" type="holotype">holotype</typeStatus>
|
||
male (partly damaged, with missing right elytron and most of the antennae) and three
|
||
<typeStatus id="22174EFBFF8C736DFD1BFBDFFCA04731" box="[677,785,1053,1077]" pageId="15" pageNumber="564" type="paratype">paratypes</typeStatus>
|
||
(one male and two females). The
|
||
<typeStatus id="22174EFBFF8C736DFB13FBDFFABF4731" box="[1197,1294,1053,1077]" pageId="15" pageNumber="564" type="paratype">paratype</typeStatus>
|
||
male is not conspecific with
|
||
<taxonomicName id="3AAC8BDAFF8C736DFEE2FB82FE57475C" box="[348,486,1088,1112]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="peruanus">
|
||
<emphasis id="CFD82C4BFF8C736DFEE2FB82FE57475C" box="[348,486,1088,1112]" italics="true" pageId="15" pageNumber="564">A. peruanus</emphasis>
|
||
</taxonomicName>
|
||
and is fixed below as a
|
||
<typeStatus id="22174EFBFF8C736DFCB5FBFDFCDC475C" box="[779,877,1087,1112]" pageId="15" pageNumber="564" type="holotype">holotype</typeStatus>
|
||
of a new species,
|
||
<taxonomicName id="3AAC8BDAFF8C736DFBF7FB82FB7A475C" box="[1097,1227,1088,1112]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="dentatus" status="sp. nov.">
|
||
<emphasis id="CFD82C4BFF8C736DFBF7FB82FB7A475C" box="[1097,1227,1088,1112]" italics="true" pageId="15" pageNumber="564">A. dentatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D4EB9130FF8C736DFB6BFBFDFAA6475C" box="[1237,1303,1087,1112]" pageId="15" pageNumber="564" rank="species">
|
||
<emphasis id="CFD82C4BFF8C736DFB6BFBFDFAA6475C" bold="true" box="[1237,1303,1087,1112]" pageId="15" pageNumber="564">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<taxonomicName id="3AAC8BDAFF8C736DFA9FFB82FF4E4778" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="peruanus">
|
||
<emphasis id="CFD82C4BFF8C736DFA9FFB82FF4E4778" italics="true" pageId="15" pageNumber="564">Alloraphes peruanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3AAC8BDAFF8C736DFE8BFBA6FE024778" box="[309,435,1124,1148]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="dentatus">
|
||
<emphasis id="CFD82C4BFF8C736DFE8BFBA6FE024778" box="[309,435,1124,1148]" italics="true" pageId="15" pageNumber="564">A. dentatus</emphasis>
|
||
</taxonomicName>
|
||
are externally highly similar but differ clearly in several characters.
|
||
<taxonomicName id="3AAC8BDAFF8C736DFB0CFBA6FA2D4778" box="[1202,1436,1124,1148]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="peruanus">
|
||
<emphasis id="CFD82C4BFF8C736DFB0CFBA6FA2D4778" box="[1202,1436,1124,1148]" italics="true" pageId="15" pageNumber="564">Alloraphes peruanus</emphasis>
|
||
</taxonomicName>
|
||
has much larger eyes than
|
||
<taxonomicName id="3AAC8BDAFF8C736DFE70FB4AFDFE47A4" box="[462,591,1160,1184]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="dentatus">
|
||
<emphasis id="CFD82C4BFF8C736DFE70FB4AFDFE47A4" box="[462,591,1160,1184]" italics="true" pageId="15" pageNumber="564">A. dentatus</emphasis>
|
||
</taxonomicName>
|
||
, more uniformly convex vertex, which in
|
||
<taxonomicName id="3AAC8BDAFF8C736DFB82FB4AFB0C47A4" box="[1084,1213,1160,1184]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="dentatus">
|
||
<emphasis id="CFD82C4BFF8C736DFB82FB4AFB0C47A4" box="[1084,1213,1160,1184]" italics="true" pageId="15" pageNumber="564">A. dentatus</emphasis>
|
||
</taxonomicName>
|
||
is noticeably more convex in middle than on sides, and the pronotal ante-basal transverse groove is more evenly arcuate, while in
|
||
<taxonomicName id="3AAC8BDAFF8C736DFA3AFB6EFF4747EC" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="dentatus">
|
||
<emphasis id="CFD82C4BFF8C736DFA3AFB6EFF4747EC" italics="true" pageId="15" pageNumber="564">A. dentatus</emphasis>
|
||
</taxonomicName>
|
||
it is slightly bi-arcuate with a median pit. In my opinion it is not possible to unambiguously assign the two remaining
|
||
<typeStatus id="22174EFBFF8C736DFEAFFB37FEC04609" box="[273,369,1269,1293]" pageId="15" pageNumber="564" type="paratype">paratype</typeStatus>
|
||
females (one of them shown in
|
||
<figureCitation id="6597ECDCFF8C736DFD5BFB36FC984608" box="[741,809,1268,1293]" captionStart="FIGURES 1 – 12" captionStartId="3.[151,264,1879,1902]" captionTargetBox="[217,1363,203,1830]" captionTargetId="figure@3.[209,1368,195,1845]" captionTargetPageId="3" captionText="FIGURES 1 – 12. Dorsal (1 – 9, 11 – 12) and laterodorsal (10) habitus of Alloraphes Franz. Alloraphes jamaicae Franz, holotype male (1); A. chiapasensis Franz, holotype male (2); A. yucatani Franz, holotype male (3); A. lenkoi Franz, paratype male (4) and paratype female (5); A. peruanus Franz, holotype male (6) and paratype female (7; but see the Remarks section in the redescription); A. dentatus sp. n., holotype male (8); A. peckorum sp. n., holotype male (9 – 10); A. myrmecophilus Franz, stat. rest., paratype male (11);? Alloraphes magnus Franz, holotype female (12)." httpUri="https://zenodo.org/record/217853/files/figure.png" pageId="15" pageNumber="564">Fig. 7</figureCitation>
|
||
) to either
|
||
<taxonomicName id="3AAC8BDAFF8C736DFC1FFB36FB9B4608" box="[929,1066,1268,1292]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="peruanus">
|
||
<emphasis id="CFD82C4BFF8C736DFC1FFB36FB9B4608" box="[929,1066,1268,1292]" italics="true" pageId="15" pageNumber="564">A. peruanus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="3AAC8BDAFF8C736DFBECFB36FB634608" box="[1106,1234,1268,1292]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="dentatus">
|
||
<emphasis id="CFD82C4BFF8C736DFBECFB36FB634608" box="[1106,1234,1268,1292]" italics="true" pageId="15" pageNumber="564">A. dentatus</emphasis>
|
||
</taxonomicName>
|
||
. They both differ from males of
|
||
<taxonomicName id="3AAC8BDAFF8C736DFE80FADAFE774634" box="[318,454,1304,1328]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="peruanus">
|
||
<emphasis id="CFD82C4BFF8C736DFE80FADAFE774634" box="[318,454,1304,1328]" italics="true" pageId="15" pageNumber="564">A. peruanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3AAC8BDAFF8C736DFE40FADAFDCF4634" box="[510,638,1304,1328]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="dentatus">
|
||
<emphasis id="CFD82C4BFF8C736DFE40FADAFDCF4634" box="[510,638,1304,1328]" italics="true" pageId="15" pageNumber="564">A. dentatus</emphasis>
|
||
</taxonomicName>
|
||
in distinctly broader elytra and smaller eyes; the characters associated with the vertex and pronotum are ambiguous. It seems more probable that the females are conspecific with
|
||
<taxonomicName id="3AAC8BDAFF8C736DFA3AFAFEFF4E467C" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="peruanus">
|
||
<emphasis id="CFD82C4BFF8C736DFA3AFAFEFF4E467C" italics="true" pageId="15" pageNumber="564">A. peruanus</emphasis>
|
||
</taxonomicName>
|
||
and not with
|
||
<taxonomicName id="3AAC8BDAFF8C736DFE23FAA2FDAC467C" box="[413,541,1376,1400]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="564" phylum="Arthropoda" rank="species" species="dentatus">
|
||
<emphasis id="CFD82C4BFF8C736DFE23FAA2FDAC467C" box="[413,541,1376,1400]" italics="true" pageId="15" pageNumber="564">A. dentatus</emphasis>
|
||
</taxonomicName>
|
||
, based on additional collecting data given by Franz (1980a). The
|
||
<typeStatus id="22174EFBFF8C736DFAB7FA9DFADA467C" box="[1289,1387,1375,1400]" pageId="15" pageNumber="564" type="holotype">holotype</typeStatus>
|
||
and two
|
||
<typeStatus id="22174EFBFF8C736DFF76FA47FE824699" box="[200,307,1413,1437]" pageId="15" pageNumber="564" type="paratype">paratypes</typeStatus>
|
||
were collected in Juan Guerra, in leaf litter of a relatively dry forest,
|
||
<date id="8912D699FF8C736DFBFEFA46FB0B4699" box="[1088,1210,1412,1437]" pageId="15" pageNumber="564" value="1968-09-28">28.ix.1968</date>
|
||
, while one
|
||
<typeStatus id="22174EFBFF8C736DFA85FA47FA2A4699" box="[1339,1435,1413,1437]" pageId="15" pageNumber="564" type="paratype">paratype</typeStatus>
|
||
in Santana, in forest leaf litter,
|
||
<date id="8912D699FF8C736DFE49FA65FDC346C4" box="[503,626,1447,1472]" pageId="15" pageNumber="564" value="1968-09-27">27.ix.1968</date>
|
||
. These two localities are in different parts of Tarapoto and the
|
||
<typeStatus id="22174EFBFF8C736DFA82FA6BFA2D46C5" box="[1340,1436,1449,1473]" pageId="15" pageNumber="564" type="paratype">paratype</typeStatus>
|
||
from Santana is most likely the male described below as a new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |