352 lines
39 KiB
XML
352 lines
39 KiB
XML
<document id="F1D86543734CD5CDBBF1AAD3C9EB384A" ID-DOI="10.1649/0010-065X-70.3.611" ID-Zenodo-Dep="5339686" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630184562620" checkinUser="diego" docAuthor="Owens, Brittany E. & Carlton, Christopher E." docDate="2016" docId="193887C2FFAF9977FEE6FC0EFB24FA1D" docLanguage="en" docName="0010-065X-70.3.611.pdf" docOrigin="The Coleopterists Bulletin 70 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-70.3.611" docStyle="DocumentStyle:A0A70CFE30726088A421CCB226FF86C7.4:TheColeopBull.2010-2016.journal_article.1cover" docStyleId="A0A70CFE30726088A421CCB226FF86C7" docStyleName="TheColeopBull.2010-2016.journal_article.1cover" docStyleVersion="4" docTitle="Pseudactium Casey 1908" docType="treatment" docVersion="3" lastPageNumber="633" masterDocId="E501FFBAFFA5997CFFEDFFBBFF82FFA5" masterDocTitle="Collecting in the Museum: New Species, Taxonomic Changes, and Range Extensions of Euplectite Pselaphinae (Coleoptera: Staphylinidae) of the Southeastern United States" masterLastPageNumber="633" masterPageNumber="611" pageNumber="611" updateTime="1699178805792" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="CD13A6F31436FFE8B555A1604EE72FFB">Collecting in the Museum: New Species, Taxonomic Changes, and Range Extensions of Euplectite Pselaphinae (Coleoptera: Staphylinidae) of the Southeastern United States</mods:title>
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<mods:namePart id="76E5A157602F7DD6313330FD459824F1">Owens, Brittany E.</mods:namePart>
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<mods:namePart id="399157E33A83D93D9A04920916DF2A83">Carlton, Christopher E.</mods:namePart>
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<mods:title id="0622AC4DF0EBC099FCB71AFEE5EF1D1E">The Coleopterists Bulletin</mods:title>
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<mods:part id="7D6188B15C9281F0BE390240E6381E50">
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<mods:date id="7A0D5F8041D03998AA93468548F82FF3">2016</mods:date>
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<mods:number id="57FFB37FB6B472B2A89BCD90408CF759">2016-09-30</mods:number>
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<mods:number id="923017074BDDA31F40F85D753579B0A1">70</mods:number>
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<mods:url id="EBECFE4616737C5E92799D5CD5015DDB">http://dx.doi.org/10.1649/0010-065x-70.3.611</mods:url>
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<mods:identifier id="FFF6283DE5431C0C8D36828B491969C6" type="DOI">10.1649/0010-065X-70.3.611</mods:identifier>
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<mods:identifier id="E3AB51AD842A6ADA72EA0EE7F0B6E455" type="Zenodo-Dep">5339686</mods:identifier>
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<subSubSection id="D98B655FFFAF9976FEE6FC0EFD88FC6F" box="[267,522,949,970]" pageId="10" type="nomenclature">
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<paragraph id="912E36D4FFAF9976FEE6FC0EFD88FC6F" blockId="10.[267,522,949,970]" box="[267,522,949,970]" pageId="10">
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<taxonomicName id="56914D57FFAF9976FEE6FC0EFD88FC6F" ID-CoL="8W2RB" authority="Casey, 1908" authorityName="Casey" authorityYear="1908" box="[267,522,949,970]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="genus">
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<emphasis id="A3E5EAC6FFAF9976FEE6FC0EFD88FC6F" bold="true" box="[267,522,949,970]" pageId="10">
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<emphasis id="A3E5EAC6FFAF9976FEE6FC0EFE0BFC6F" bold="true" box="[267,393,949,970]" italics="true" pageId="10">Pseudactium</emphasis>
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<bibRefCitation id="F5004B25FFAF9976FE7CFC0EFD88FC6F" author="Casey & Remarks on some new Pselaphidae & The Canadian Entomologist" box="[401,522,949,970]" pageId="10" pagination="279 - 281" refId="ref17979" refString="Casey, T. L. 1908. Remarks on some new Pselaphidae. The Canadian Entomologist 40: 271 - 272, 279 - 281." type="journal article" year="1908">Casey, 1908</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D98B655FFFAF9977FF42FC64FE02FE44" lastPageId="11" pageId="10" type="diagnosis">
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<paragraph id="912E36D4FFAF9976FF42FC64FCE8FD95" blockId="10.[151,638,991,1729]" lastBlockId="10.[685,1172,194,1729]" pageId="10">
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<emphasis id="A3E5EAC6FFAF9976FF42FC64FE28FC51" bold="true" box="[175,426,991,1012]" pageId="10">Diagnosis. Males. Head:</emphasis>
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Rounded trapezoidal, tempora broadly rounded behind ocular canthus to cervical constriction, vertexal foveae setose, frontal sulcus broad, shallow, delimited by foveae; antennal acetabula remote, antennomeres 1–8 unmodified, antennomeres 9–11 forming loose club; labrum of normal form for tribe, bearing pair of sensory pegs, mandibles broadly sickle-shaped, maxillae and labium typical for subfamily, labial palpi minute, 2-segmented, maxillary palpi unmodified, palpomere 1 minute and obliquely joined to palpomere 2, palpomere 2 pedunculate, narrower in basal 1/3, slightly expanded and rounded distally, palpomere 3 subtriangular, palpomere 4 widest, unmodified, bearing apical palpal cone; gula simple and slightly rounded anteriorly, paired gular foveae present; eyes present and well-developed, comprised of about 30 facets.
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<emphasis id="A3E5EAC6FFAF9976FF10FA18FECDFA1D" bold="true" box="[253,335,1443,1464]" pageId="10">Thorax:</emphasis>
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Pronotum cordate, disc evenly convex, antebasal transverse sulcus present, delimited by lateral antebasal foveae, longitudinal median sulcus absent, basolateral fovea present; prosternum slightly convex, lateral procoxal foveae present; mesoventrite with paired lateral and median mesoventral foveae, paired mesocoxal foveae present, mesocoxae contiguous; lateral metaventral foveae present, metacoxae conjunct; legs variously modified; elytra with sutural fovea and striae, antebasal discal foveae and subhumeral foveae present, discal striae very strong basally; lateroapical cleft present; wings present or absent.
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<emphasis id="A3E5EAC6FFAF9976FC8BFF66FC4DFF57" bold="true" box="[870,975,221,242]" pageId="10">Abdomen:</emphasis>
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First visible tergite with small basolateral foveae; ventrite II bearing large basolateral foveae; ventrite III bearing lateral processes; ventrites IV–VI unmodified and afoveate; ventrite VII of males tripartite, lateral sclerites triangular, penial plate ovate and hinged basally.
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<emphasis id="A3E5EAC6FFAF9976FD40FEC7FC9AFE34" bold="true" box="[685,792,380,401]" pageId="10">Aedeagus:</emphasis>
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Strongly asymmetrical, composed of basal bulb bearing 2 large distal processes, 1 usually elongate and originating along ventrolateral or ventral margin of phallobase and the shorter often digitate and originating along the lateral margin of phallobase or along dorsal aspect of elongate process.
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</paragraph>
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<paragraph id="912E36D4FFAF9976FD28FD8DFCF1FD10" blockId="10.[685,1172,194,1729]" pageId="10">
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<emphasis id="A3E5EAC6FFAF9976FD28FD8DFC9CFDEE" bold="true" box="[709,798,566,587]" pageId="10">Females.</emphasis>
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Externally similar to males except eyes smaller with fewer facets, secondary sexual modifications to the legs absent, and ventrite VII not modified into penial plate (revision,
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<bibRefCitation id="F5004B25FFAF9976FBA1FD3DFCEAFD10" author="Carlton" pageId="10" pagination="171 - 190" refId="ref17745" refString="Carlton, C. E., and D. S. Chandler. 1994. Revision of the Nearctic genus Pseudactium Casey (Coleoptera: Pselaphidae: Euplectinae). The Coleopterists Bulletin 48: 171 - 190." type="journal article" year="1994">Carlton and Chandler 1994</bibRefCitation>
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).
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</paragraph>
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<paragraph id="912E36D4FFAF9976FD28FD00FCE0FA38" blockId="10.[685,1172,194,1729]" pageId="10">
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<emphasis id="A3E5EAC6FFAF9976FD28FD00FC81FD75" bold="true" box="[709,771,699,720]" pageId="10">Notes.</emphasis>
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Members of the genus
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<taxonomicName id="56914D57FFAF9976FBF4FD00FB11FD75" authorityName="Casey" authorityYear="1908" box="[1049,1171,699,720]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="genus">
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<emphasis id="A3E5EAC6FFAF9976FBF4FD00FB11FD75" box="[1049,1171,699,720]" italics="true" pageId="10">Pseudactium</emphasis>
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</taxonomicName>
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exhibit an interesting biogeographic pattern and have been identified as possible indicators of biodiversity in old-growth forests (
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<bibRefCitation id="F5004B25FFAF9976FC23FCB1FD5FFC9F" author="Carlton" pageId="10" pagination="171 - 190" refId="ref17745" refString="Carlton, C. E., and D. S. Chandler. 1994. Revision of the Nearctic genus Pseudactium Casey (Coleoptera: Pselaphidae: Euplectinae). The Coleopterists Bulletin 48: 171 - 190." type="journal article" year="1994">Carlton and Chandler 1994</bibRefCitation>
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). Known species almost exclusively inhabit mature deciduous forests in the eastern
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<collectingCountry id="E9867644FFAF9976FBABFCFBFBE6FCF0" box="[1094,1124,832,853]" name="United States of America" pageId="10">US</collectingCountry>
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and occur south of Pleistocene glacial maxima. A number of species and collections are known from restricted localities within this range.
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<taxonomicName id="56914D57FFAF9976FBF6FC34FB91FC1A" authority="Carlton and Chandler, 1994" authorityName="Carlton and Chandler" authorityYear="1994" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="copelandi">
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<emphasis id="A3E5EAC6FFAF9976FBF6FC34FC8BFC1A" italics="true" pageId="10">Pseudactium copelandi</emphasis>
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<bibRefCitation id="F5004B25FFAF9976FCE2FC11FB91FC1A" author="Carlton" box="[783,1043,938,959]" pageId="10" pagination="171 - 190" refId="ref17745" refString="Carlton, C. E., and D. S. Chandler. 1994. Revision of the Nearctic genus Pseudactium Casey (Coleoptera: Pselaphidae: Euplectinae). The Coleopterists Bulletin 48: 171 - 190." type="journal article" year="1994">Carlton and Chandler, 1994</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="56914D57FFAF9976FBF1FC11FC54FC7C" authority="Carlton and Chandler, 1994" authorityName="Carlton and Chandler" authorityYear="1994" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="pecki">
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<emphasis id="A3E5EAC6FFAF9976FBF1FC11FD5FFC7C" italics="true" pageId="10">Pseudactium pecki</emphasis>
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<bibRefCitation id="F5004B25FFAF9976FD09FC7FFC54FC7C" author="Carlton" box="[740,982,964,985]" pageId="10" pagination="171 - 190" refId="ref17745" refString="Carlton, C. E., and D. S. Chandler. 1994. Revision of the Nearctic genus Pseudactium Casey (Coleoptera: Pselaphidae: Euplectinae). The Coleopterists Bulletin 48: 171 - 190." type="journal article" year="1994">Carlton and Chandler, 1994</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="56914D57FFAF9976FC33FC7FFC22FC51" authority="Carlton and Chandler, 1994" authorityName="Carlton and Chandler" authorityYear="1994" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="steevesi">
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<emphasis id="A3E5EAC6FFAF9976FC33FC7FFB16FC7C" box="[990,1172,964,985]" italics="true" pageId="10">Pseudactium steevesi</emphasis>
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<bibRefCitation id="F5004B25FFAF9976FD40FC64FC22FC51" author="Carlton" box="[685,928,991,1012]" pageId="10" pagination="171 - 190" refId="ref17745" refString="Carlton, C. E., and D. S. Chandler. 1994. Revision of the Nearctic genus Pseudactium Casey (Coleoptera: Pselaphidae: Euplectinae). The Coleopterists Bulletin 48: 171 - 190." type="journal article" year="1994">Carlton and Chandler, 1994</bibRefCitation>
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</taxonomicName>
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, and
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<taxonomicName id="56914D57FFAF9976FC22FC64FB11FC51" authority="Owens & Carlton, 2016" authorityName="Owens & Carlton" authorityYear="2016" box="[975,1171,991,1012]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="jaitlynum" status="sp. nov.">
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<emphasis id="A3E5EAC6FFAF9976FC22FC64FB11FC51" box="[975,1171,991,1012]" italics="true" pageId="10">Pseudactium jaitlynum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B8D657BDFFAF9976FD40FC41FCA3FBAA" box="[685,801,1018,1039]" pageId="10" rank="species">new species</taxonomicNameLabel>
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appear endemic to localities in northern
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<collectingRegion id="5355F836FFAF9976FD3CFBAFFCA7FB8C" box="[721,805,1044,1065]" country="United States of America" name="Alabama" pageId="10">Alabama</collectingRegion>
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, and
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<taxonomicName id="56914D57FFAF9976FCB8FBAFFCE0FBE1" authority="Carlton and Chandler, 1994" authorityName="Carlton and Chandler" authorityYear="1994" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="magazinensis">
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<emphasis id="A3E5EAC6FFAF9976FCB8FBAFFBCBFB8C" box="[853,1097,1044,1065]" italics="true" pageId="10">Pseudactium magazinensis</emphasis>
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<bibRefCitation id="F5004B25FFAF9976FBBDFBAFFCE0FBE1" author="Carlton" pageId="10" pagination="171 - 190" refId="ref17745" refString="Carlton, C. E., and D. S. Chandler. 1994. Revision of the Nearctic genus Pseudactium Casey (Coleoptera: Pselaphidae: Euplectinae). The Coleopterists Bulletin 48: 171 - 190." type="journal article" year="1994">Carlton and Chandler, 1994</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="56914D57FFAF9976FC7DFB94FD59FBFB" authority="Carlton, 1995" authorityName="Carlton" authorityYear="1995" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="ursum">
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<emphasis id="A3E5EAC6FFAF9976FC7DFB94FBC6FBE1" box="[912,1092,1071,1092]" italics="true" pageId="10">Pseudactium ursum</emphasis>
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<bibRefCitation id="F5004B25FFAF9976FBA7FB94FD59FBFB" author="Carlton" pageId="10" pagination="170 - 173" refId="ref17667" refString="Carlton, C. E. 1995. A new species of Pseudactium Casey from Arkansas (Coleoptera: Staphylinidae: Pselaphinae). The Coleopterists Bulletin 49: 170 - 173." type="journal article" year="1995">Carlton, 1995</bibRefCitation>
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</taxonomicName>
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are known only from the type localities in
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<collectingRegion id="5355F836FFAF9976FD40FBDFFC86FBDC" box="[685,772,1124,1145]" country="United States of America" name="Arkansas" pageId="10">Arkansas</collectingRegion>
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. The existence of some species and disjunct populations in
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<collectingRegion id="5355F836FFAF9976FCD6FBC4FC1CFB31" box="[827,926,1151,1172]" country="United States of America" name="Tennessee" pageId="10">Tennessee</collectingRegion>
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,
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<collectingRegion id="5355F836FFAF9976FC44FBC4FBA5FB31" box="[937,1063,1151,1172]" country="United States of America" name="Pennsylvania" pageId="10">Pennsylvania</collectingRegion>
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, and
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<collectingRegion id="5355F836FFAF9976FBB1FBC4FB16FB31" box="[1116,1172,1151,1172]" country="United States of America" name="Texas" pageId="10">Texas</collectingRegion>
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may indicate sites of past refugia and long-term persistence of stable forest habitats.
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<taxonomicName id="56914D57FFAF9976FBF6FB0FFBA8FB46" authority="Carlton and Chandler, 1994" authorityName="Carlton and Chandler" authorityYear="1994" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="stannardi">
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<emphasis id="A3E5EAC6FFAF9976FBF6FB0FFC87FB46" italics="true" pageId="10">Pseudactium stannardi</emphasis>
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<bibRefCitation id="F5004B25FFAF9976FCFEFB75FBA8FB46" author="Carlton" box="[787,1066,1230,1251]" pageId="10" pagination="171 - 190" refId="ref17745" refString="Carlton, C. E., and D. S. Chandler. 1994. Revision of the Nearctic genus Pseudactium Casey (Coleoptera: Pselaphidae: Euplectinae). The Coleopterists Bulletin 48: 171 - 190." type="journal article" year="1994">Carlton and Chandler, 1994</bibRefCitation>
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</taxonomicName>
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is known only from the
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<typeStatus id="4E2A8876FFAF9976FCD5FB52FC08FB5B" box="[824,906,1257,1278]" pageId="10" type="holotype">holotype</typeStatus>
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that was collected in a forested loess soil habitat in northwestern
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<collectingRegion id="5355F836FFAF9976FBDBFAB8FBE7FABD" box="[1078,1125,1283,1304]" country="United States of America" name="Iowa" pageId="10">Iowa</collectingRegion>
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east of the
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<collectingRegion id="5355F836FFAF9976FD00FAA5FCC2FA96" box="[749,832,1310,1331]" country="United States of America" name="Missouri" pageId="10">Missouri</collectingRegion>
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River. This represents a significant exception to the general distributional pattern of the genus. Attempts by the second author to collect additional specimens at or near the type locality during 2015 failed.
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</paragraph>
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<paragraph id="912E36D4FFAF9977FD28FA18FE02FE44" blockId="10.[685,1172,194,1729]" lastBlockId="11.[173,659,194,481]" lastPageId="11" pageId="10">
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Herein we describe one new species and report range extensions for
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<taxonomicName id="56914D57FFAF9976FC98FA06FD5EFA48" authority="Casey, 1908" authorityName="Casey" authorityYear="1908" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="carolinae">
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<emphasis id="A3E5EAC6FFAF9976FC98FA06FBCFFA77" box="[885,1101,1469,1490]" italics="true" pageId="10">Pseudactium carolinae</emphasis>
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<bibRefCitation id="F5004B25FFAF9976FBB9FA06FD5EFA48" author="Casey & Remarks on some new Pselaphidae & The Canadian Entomologist" pageId="10" pagination="279 - 281" refId="ref17979" refString="Casey, T. L. 1908. Remarks on some new Pselaphidae. The Canadian Entomologist 40: 271 - 272, 279 - 281." type="journal article" year="1908">Casey, 1908</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="56914D57FFAF9976FCF3FA63FB16FA48" authority="(Park, 1962)" baseAuthorityName="Park" baseAuthorityYear="1962" box="[798,1172,1496,1517]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="mendicum">
|
||
<emphasis id="A3E5EAC6FFAF9976FCF3FA63FB84FA48" box="[798,1030,1496,1517]" italics="true" pageId="10">Pseudactium mendicum</emphasis>
|
||
(
|
||
<bibRefCitation id="F5004B25FFAF9976FBF6FA63FB0FFA48" author="Park" box="[1051,1165,1496,1517]" pageId="10" pagination="150 - 156" refId="ref18830" refString="Park, O. 1962. A review of the genus Ramecia (Coleoptera: Pselaphidae). The Ohio Journal of Science 64: 150 - 156." type="journal article" year="1962">Park, 1962</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from forested areas in the eastern
|
||
<collectingCountry id="E9867644FFAF9976FC1DFA49FB93F9A2" box="[1008,1041,1522,1543]" name="United States of America" pageId="10">US</collectingCountry>
|
||
.
|
||
<taxonomicName id="56914D57FFAF9976FBF6FA49FC87F987" authority="Owens & Carlton, 2016" authorityName="Owens & Carlton" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="jaitlynum" status="sp. nov.">
|
||
<emphasis id="A3E5EAC6FFAF9976FBF6FA49FC87F987" italics="true" pageId="10">Pseudactium jaitlynum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B8D657BDFFAF9976FCFDF9B6FC07F987" box="[784,901,1549,1570]" pageId="10" rank="species">new species</taxonomicNameLabel>
|
||
is described from a single specimen collected from a mature forest in Madison Co.,
|
||
<collectingRegion id="5355F836FFAF9976FD36F9F9FCB0F9F2" box="[731,818,1602,1623]" country="United States of America" name="Alabama" pageId="10">Alabama</collectingRegion>
|
||
. It is the fourth species known exclusively from the northern part of that state. Litter sifting in mature forests in north-central and southeastern
|
||
<collectingRegion id="5355F836FFAF9976FD1AF929FCD0F902" box="[759,850,1682,1703]" country="United States of America" name="Louisiana" pageId="10">Louisiana</collectingRegion>
|
||
has documented two populations of
|
||
<taxonomicName id="56914D57FFAF9976FD38F917FCD1F964" authorityName="Casey" authorityYear="1908" box="[725,851,1708,1729]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="carolinae">
|
||
<emphasis id="A3E5EAC6FFAF9976FD38F917FCD1F964" box="[725,851,1708,1729]" italics="true" pageId="10">P. carolinae</emphasis>
|
||
</taxonomicName>
|
||
and a single population of
|
||
<taxonomicName id="56914D57FFAE9977FF40FF79FEA5FF72" baseAuthorityName="Park" baseAuthorityYear="1962" box="[173,295,194,215]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="mendicum">
|
||
<emphasis id="A3E5EAC6FFAE9977FF40FF79FEA5FF72" box="[173,295,194,215]" italics="true" pageId="11">P. mendicum</emphasis>
|
||
</taxonomicName>
|
||
, constituting new records of the species for the state. A single collection record of
|
||
<taxonomicName id="56914D57FFAE9977FF40FF4CFEA0FEA9" authorityName="Casey" authorityYear="1908" box="[173,290,247,268]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="carolinae">
|
||
<emphasis id="A3E5EAC6FFAE9977FF40FF4CFEA0FEA9" box="[173,290,247,268]" italics="true" pageId="11">P. carolinae</emphasis>
|
||
</taxonomicName>
|
||
from Fayette Co.,
|
||
<collectingRegion id="5355F836FFAE9977FE11FF4CFD0DFEA9" box="[508,655,247,268]" country="United States of America" name="West Virginia" pageId="11">West Virginia</collectingRegion>
|
||
, and a single collection record from Great Smoky Mountains National Park in Sevier Co.,
|
||
<collectingRegion id="5355F836FFAE9977FF40FEFCFE92FEF9" box="[173,272,327,348]" country="United States of America" name="Tennessee" pageId="11">Tennessee</collectingRegion>
|
||
, are also
|
||
<emphasis id="A3E5EAC6FFAE9977FE93FEFCFDB9FEF9" bold="true" box="[382,571,327,348]" pageId="11">new state records</emphasis>
|
||
. Further targeting of known mature and old growth forest sites will likely yield new records throughout the eastern
|
||
<collectingCountry id="E9867644FFAE9977FF15FE2CFE94FE09" box="[248,278,407,428]" name="United States of America" pageId="11">US</collectingCountry>
|
||
for a number of
|
||
<taxonomicName id="56914D57FFAE9977FE25FE2CFD0DFE09" box="[456,655,407,428]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="A3E5EAC6FFAE9977FE25FE2CFDC2FE09" box="[456,576,407,428]" italics="true" pageId="11">Pseudactium</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, and may shed light on the age and localized persistence of such habitats.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D98B655FFFAE9977FF04FDB1FB2BFE26" pageId="11" type="key">
|
||
<paragraph id="912E36D4FFAE9977FF04FDB1FDB3FDFE" blockId="11.[233,599,522,603]" pageId="11">
|
||
<emphasis id="A3E5EAC6FFAE9977FF04FDB1FDD5FDBA" bold="true" box="[233,599,522,545]" pageId="11">KEY TO MALES OF THE SPECIES OF</emphasis>
|
||
<taxonomicName id="56914D57FFAE9977FED9FD9CFD89FD99" ID-CoL="8W2RB" authority="CASEY" authorityName="Casey" authorityYear="1908" box="[308,523,551,574]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A3E5EAC6FFAE9977FED9FD9CFE3DFD99" bold="true" box="[308,447,551,574]" italics="true" pageId="11">PSEUDACTIUM</emphasis>
|
||
<emphasis id="A3E5EAC6FFAE9977FE2AFD9CFD89FD99" bold="true" box="[455,523,551,574]" pageId="11">CASEY</emphasis>
|
||
</taxonomicName>
|
||
(Modified from
|
||
<bibRefCitation id="F5004B25FFAE9977FE41FDFDFDA9FDFE" author="Carlton" box="[428,555,582,603]" pageId="11" pagination="170 - 173" refId="ref17667" refString="Carlton, C. E. 1995. A new species of Pseudactium Casey from Arkansas (Coleoptera: Staphylinidae: Pselaphinae). The Coleopterists Bulletin 49: 170 - 173." type="journal article" year="1995">Carlton 1995</bibRefCitation>
|
||
)
|
||
</paragraph>
|
||
<keyStep id="2A652E71FFAE9977FF40FDCBFD11FCE2" pageId="11">
|
||
<paragraph id="912E36D4FFAE9977FF40FDCBFD11FD70" blockId="11.[173,660,624,1729]" pageId="11">
|
||
<keyLead id="2A6095E1FFAE9977FF40FDCBFD11FD70" pageId="11">
|
||
<key id="E50061E5FFAE9977FF40FDCBFD11FD70" pageId="11">1. Mesotrochanter and (or) metatrochanter modified, bearing acute spine or blunt tubercle, or distal angle prolonged beyond junction with femur...................................... 2</key>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FF40FD58FD11FCE2" blockId="11.[173,660,624,1729]" pageId="11">
|
||
<keyLead id="2A6095E1FFAE9977FF40FD58FD11FCE2" pageId="11">1′. Neither mesotrochanters nor metatrochanters bearing spines or tubercles, convex along ventral margin or at most arcuate-carinate............................................ 7</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="2A652E71FFAE9977FF40FCEEFD10FB89" pageId="11">
|
||
<paragraph id="912E36D4FFAE9977FF40FCEEFD11FC1F" blockId="11.[173,660,624,1729]" pageId="11">
|
||
<keyLead id="2A6095E1FFAE9977FF40FCEEFD11FC1F" pageId="11">2. Lateral processes of ventrite III low and rounded or carinate, not obviously projecting beyond lateral margins of abdomen in dorsal view............................................... 3</keyLead>
|
||
</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FF40FC7CFD10FB89" blockId="11.[173,660,624,1729]" pageId="11">
|
||
<keyLead id="2A6095E1FFAE9977FF40FC7CFD10FB89" pageId="11">
|
||
2′. Lateral processes of ventrite III large, rounded triangular, obviously projecting beyond lateral margins of abdomen in dorsal view ..............
|
||
<taxonomicName id="56914D57FFAE9977FE71FBACFD10FB89" ID-CoL="4NHGQ" authority="(Brendel)" baseAuthorityName="Brendel" baseAuthorityYear="1893" box="[412,658,1047,1068]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="parabolicum">
|
||
<emphasis id="A3E5EAC6FFAE9977FE71FBACFD10FB89" bold="true" box="[412,658,1047,1068]" pageId="11">
|
||
<emphasis id="A3E5EAC6FFAE9977FE71FBACFE2AFB89" bold="true" box="[412,424,1047,1068]" italics="true" pageId="11">P</emphasis>
|
||
.
|
||
<emphasis id="A3E5EAC6FFAE9977FE58FBACFDADFB89" bold="true" box="[437,559,1047,1068]" italics="true" pageId="11">parabolicum</emphasis>
|
||
(Brendel)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="2A652E71FFAE9977FF40FB82FD11FA88" pageId="11">
|
||
<paragraph id="912E36D4FFAE9977FF40FB82FD11FB1C" blockId="11.[173,660,624,1729]" pageId="11">
|
||
<keyLead id="2A6095E1FFAE9977FF40FB82FD11FB1C" pageId="11">3. Metatrochanter bearing an acute spine or blunt tubercle along ventral margin; ventral margin of mesotrochanter convex, angulate, or bearing an acute spine before distal angle ............................................................. 5</keyLead>
|
||
</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FF40FB7DFD11FA88" blockId="11.[173,660,624,1729]" pageId="11">
|
||
<keyLead id="2A6095E1FFAE9977FF40FB7DFD11FA88" pageId="11">3′. Metatrochanter simple, ventral margin evenly convex; ventral margin of mesotrochanter concave, distal angle projecting beyond junction with femur..............................................4</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="2A652E71FFAE9977FF40FA81FD11F921" pageId="11">
|
||
<paragraph id="912E36D4FFAE9977FF40FA81FD11FA72" blockId="11.[173,660,624,1729]" pageId="11">
|
||
<keyLead id="2A6095E1FFAE9977FF40FA81FD11FA72" pageId="11">
|
||
4. Metacoxa unmodified, bearing at most a rounded flange anterior to trochanteral articulation; distal processes of aedeagus fused near phallobase, neither bearing distal digits....................
|
||
<taxonomicName id="56914D57FFAE9977FE51FA1CFD11FA72" ID-CoL="6WD98" authority="Carlton and Chandler" authorityName="Carlton and Chandler" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="alexanderi">
|
||
<emphasis id="A3E5EAC6FFAE9977FE51FA1CFD11FA72" bold="true" pageId="11">
|
||
<emphasis id="A3E5EAC6FFAE9977FE51FA1CFDBFFA19" bold="true" box="[444,573,1447,1468]" italics="true" pageId="11">P. alexanderi</emphasis>
|
||
Carlton and Chandler
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FF40FA5EFD11F921" blockId="11.[173,660,624,1729]" pageId="11">
|
||
<keyLead id="2A6095E1FFAE9977FF40FA5EFD11F921" pageId="11">
|
||
4′. Metacoxa bearing a minute spine anterior to trochanteral articulation; shorter distal process of aedeagus arising midway along dorsal aspect of elongate process, bearing a cluster of digits and tubercles near apex.........
|
||
<taxonomicName id="56914D57FFAE9977FEB4F9D4FD11F921" ID-CoL="4NHGR" authority="Carlton and Chandler" authorityName="Carlton and Chandler" authorityYear="1994" box="[345,659,1647,1668]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="pecki">
|
||
<emphasis id="A3E5EAC6FFAE9977FEB4F9D4FD11F921" bold="true" box="[345,659,1647,1668]" pageId="11">
|
||
<emphasis id="A3E5EAC6FFAE9977FEB4F9D4FE27F921" bold="true" box="[345,421,1647,1668]" italics="true" pageId="11">P. pecki</emphasis>
|
||
Carlton and Chandler
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph id="912E36D4FFAE9977FF40F92AFB2BFF56" blockId="11.[173,660,624,1729]" lastBlockId="11.[706,1193,195,1464]" pageId="11">
|
||
<keyLead id="2A6095E1FFAE9977FF40F92AFD10F964" pageId="11">
|
||
<keyStep id="2A652E71FFAE9977FF40F92AFD10F964" pageId="11">5. Ventral margin of mesotrochanter evenly convex or angulate near middle; lateral</keyStep>
|
||
</keyLead>
|
||
processes of ventrite III small, not reaching lateral margins of abdomen.......................6
|
||
</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FD2FFEBBFB2BFE26" blockId="11.[706,1193,195,1464]" pageId="11">
|
||
5′. Ventral margin of mesotrochanter bearing an acute spine just before distal angle; lateral processes of ventrite III longer, reaching or slightly surpassing lateral margins of the abdomen ..........................
|
||
<taxonomicName id="56914D57FFAE9977FC18FED5FB2BFE26" ID-CoL="4NHGG" authority="Casey" authorityName="Casey" authorityYear="1908" box="[1013,1193,366,387]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="carolinae">
|
||
<emphasis id="A3E5EAC6FFAE9977FC18FED5FB2BFE26" bold="true" box="[1013,1193,366,387]" pageId="11">
|
||
<emphasis id="A3E5EAC6FFAE9977FC18FED5FBE4FE26" bold="true" box="[1013,1126,366,387]" italics="true" pageId="11">P. carolinae</emphasis>
|
||
Casey
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D98B655FFFAE9977FD2FFE2BFB24FA1D" pageId="11" type="description">
|
||
<paragraph id="912E36D4FFAE9977FD2FFE2BFB2BFDB6" blockId="11.[706,1193,195,1464]" pageId="11">
|
||
6. Ventral margin of mesotrochanter angulate near middle; lateral processes of ventrite III smaller, less than ½ as long as ventrite III, barely visible.................
|
||
<taxonomicName id="56914D57FFAE9977FC1CFE59FB2BFDB6" authority="Carlton and Chandler" authorityName="Carlton and Chandler" authorityYear="1994" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="steevesi">
|
||
<emphasis id="A3E5EAC6FFAE9977FC1CFE59FB2BFDB6" bold="true" pageId="11">
|
||
<emphasis id="A3E5EAC6FFAE9977FC1CFE59FBD0FE52" bold="true" box="[1009,1106,482,503]" italics="true" pageId="11">P. steevesi</emphasis>
|
||
Carlton and Chandler
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FD2FFD9BFB2AFDC9" blockId="11.[706,1193,195,1464]" pageId="11">
|
||
6′. Ventral margin of mesotrochanter convex; lateral processes of ventrite III larger, 2/3 length of ventrite III...
|
||
<taxonomicName id="56914D57FFAE9977FC0CFDECFB2AFDC9" authority="(Park)" baseAuthorityName="Park" baseAuthorityYear="1962" box="[993,1192,599,620]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="mendicum">
|
||
<emphasis id="A3E5EAC6FFAE9977FC0CFDECFB2AFDC9" bold="true" box="[993,1192,599,620]" pageId="11">
|
||
<emphasis id="A3E5EAC6FFAE9977FC0CFDECFBE2FDC9" bold="true" box="[993,1120,599,620]" italics="true" pageId="11">P. mendicum</emphasis>
|
||
(Park)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FD2FFDC2FB2BFD0F" blockId="11.[706,1193,195,1464]" pageId="11">7. Lateral processes of ventrite III larger, at least ½ length of ventrite III......................8</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FD2FFD0CFB2AFCA6" blockId="11.[706,1193,195,1464]" pageId="11">7′. Lateral processes of ventrite III minute, consisting of low, setose timidities near postlateral angle of ventrite III..................9</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FD2FFCAAFB2AFCF9" blockId="11.[706,1193,195,1464]" pageId="11">
|
||
8. Each gena with 40 or more capitate setae .................
|
||
<taxonomicName id="56914D57FFAE9977FC47FC97FB2AFCF9" authority="Carlton and Chandler" authorityName="Carlton and Chandler" authorityYear="1994" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="magazinensis">
|
||
<emphasis id="A3E5EAC6FFAE9977FC47FC97FB2AFCF9" bold="true" pageId="11">
|
||
<emphasis id="A3E5EAC6FFAE9977FC47FC97FBCCFCE4" bold="true" box="[938,1102,812,833]" italics="true" pageId="11">P. magazinensis</emphasis>
|
||
Carlton and Chandler
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FD2FFCD1FB2BFC13" blockId="11.[706,1193,195,1464]" pageId="11">
|
||
8′. Capitate setae on each gena of head 30 or fewer .......................
|
||
<taxonomicName id="56914D57FFAE9977FC3DFC3EFB2BFC13" authority="Carlton and Chandler" authorityName="Carlton and Chandler" authorityYear="1994" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="copelandi">
|
||
<emphasis id="A3E5EAC6FFAE9977FC3DFC3EFB2BFC13" bold="true" pageId="11">
|
||
<emphasis id="A3E5EAC6FFAE9977FC3DFC3EFBCFFC3F" bold="true" box="[976,1101,901,922]" italics="true" pageId="11">P. copelandi</emphasis>
|
||
Carlton and Chandler
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FD2FFC78FB2BFB8F" blockId="11.[706,1193,195,1464]" pageId="11">
|
||
9. Capitate setae on each gena of head 40 or more; protrochanters arcuate along ventral margin ......................
|
||
<taxonomicName id="56914D57FFAE9977FC35FC41FB2BFB8F" authority="Carlton and Chandler" authorityName="Carlton and Chandler" authorityYear="1994" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="stannardi">
|
||
<emphasis id="A3E5EAC6FFAE9977FC35FC41FB2BFB8F" bold="true" pageId="11">
|
||
<emphasis id="A3E5EAC6FFAE9977FC35FC41FBD2FBAA" bold="true" box="[984,1104,1018,1039]" italics="true" pageId="11">P. stannardi</emphasis>
|
||
Carlton and Chandler
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FD2FFB83FB2BFB26" blockId="11.[706,1193,195,1464]" pageId="11">9′. Capitate setae on each gena of head 20 or fewer; protrochanters variable along ventral margin......................................................10</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FD2FFB2AFB2AFAB5" blockId="11.[706,1193,195,1464]" pageId="11">
|
||
10. Aedeagus with digitate accessory process arising from dorsal aspect of elongate process near base, distally bearing numerous small and two larger digits clustered at apex ........................
|
||
<taxonomicName id="56914D57FFAE9977FC29FB40FB2AFAB5" authority="(LeConte)" baseAuthorityName="LeConte" box="[964,1192,1275,1296]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="arcuatum">
|
||
<emphasis id="A3E5EAC6FFAE9977FC29FB40FB2AFAB5" bold="true" box="[964,1192,1275,1296]" pageId="11">
|
||
<emphasis id="A3E5EAC6FFAE9977FC29FB40FBB9FAB5" bold="true" box="[964,1083,1275,1296]" italics="true" pageId="11">P. arcuatum</emphasis>
|
||
(LeConte)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="912E36D4FFAE9977FD2FFAA5FB24FA1D" blockId="11.[706,1193,195,1464]" pageId="11">
|
||
10′. Aedeagus with digitate accessory process arising near right side of phallobase, bearing lateral row of two larger and many smaller digits, distally (
|
||
<figureCitation id="09AA2A51FFAE9977FC0BFAD5FBAAFA26" box="[998,1064,1390,1411]" captionStart="Fig" captionStartId="12.[175,206,1710,1729]" captionTargetBox="[183,562,991,1674]" captionTargetId="figure-354@12.[151,638,977,1687]" captionTargetPageId="12" captionText="Fig. 11. Pseudactium jaitlynum, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5339696" httpUri="https://zenodo.org/record/5339696/files/figure.png" pageId="11">Fig. 11</figureCitation>
|
||
)................... .............
|
||
<emphasis id="A3E5EAC6FFAE9977FCB3FA33FB24FA1D" bold="true" pageId="11">
|
||
<taxonomicName id="56914D57FFAE9977FCB3FA33FB26FA38" authority="Owens and Carlton" authorityName="Owens and Carlton" authorityYear="2016" box="[862,1188,1416,1437]" class="Insecta" family="Staphylinidae" genus="Pseudactium" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="jaitlynum" status="sp. nov.">
|
||
<emphasis id="A3E5EAC6FFAE9977FCB3FA33FC51FA38" bold="true" box="[862,979,1416,1437]" italics="true" pageId="11">P. jaitlynum</emphasis>
|
||
Owens and Carlton
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B8D657BDFFAE9977FBDCFA18FB24FA1D" box="[1073,1190,1443,1464]" pageId="11" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |