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<document id="AA289935615CD8E0B3634BFBE489F94C" ENCODING="UTF-8" ID-GBIF-Dataset="892cdbaf-7871-4cfb-a1ca-075d71882e41" checkinTime="1263711157264" checkinUser="donat" docAuthor="Ward, P. S." docDate="2009" docId="174E1FEE3CDC7C8DB62D64F369D6B35E" docLanguage="en" docName="22935.xml" docOrigin="Journal of Hymenoptera Research 18" docSource="http://antbase.org/ants/publications/22935/22935.pdf" docTitle="Tetraponera variegata Ward, 2009, stat. n." docType="treatment" docVersion="11" lastPageNumber="301" masterDocId="7C2A329B41C2F7BE72B2F1BED5CC0575" masterDocTitle="The ant genus Tetraponera in the Afrotropical region: the T. grandidieri group (Hymenoptera: Formicidae)." masterLastPageNumber="304" masterPageNumber="285" pageNumber="300" updateTime="1701311923478" updateUser="plazi">
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<mods:title id="C7208F84CF0B84F5DFD011593AE0E587">The ant genus Tetraponera in the Afrotropical region: the T. grandidieri group (Hymenoptera: Formicidae).</mods:title>
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<mods:namePart id="F8A52690A0E56EC0F382854AEB59CDC2">Ward, P. S.</mods:namePart>
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<mods:title id="937BC0619764B63DBAFF888CE5AF87FB">Journal of Hymenoptera Research</mods:title>
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<mods:date id="2628DF63C56641487BECEB7E7B7BD49B">2009</mods:date>
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<mods:classification id="E5BCABD288459D594E63F613AA68F83B">journal article</mods:classification>
<mods:identifier id="C078DC9CA4101A8F90C79D4E267D55D4" type="HNS-Pub">22935</mods:identifier>
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<treatment id="174E1FEE3CDC7C8DB62D64F369D6B35E" ID-DOI="http://doi.org/10.5281/zenodo.6219042" ID-GBIF-Taxon="100116650" ID-Zenodo-Dep="6219042" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E" httpUri="http://treatment.plazi.org/id/174E1FEE3CDC7C8DB62D64F369D6B35E" lastPageNumber="301" pageNumber="300">
<subSubSection id="DD1F3360672F9BB2B3947D256B62D5BC" type="nomenclature">
<paragraph id="6097B1692101CDB3F4F83C2D57468FC6" pageNumber="300">
<taxonomicName id="3E4FECBEB7E1F0B780AD921E4AE8EA50" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:261542" class="Insecta" family="Formicidae" genus="Tetraponera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera variegata" order="Hymenoptera" pageNumber="300" phylum="Arthropoda" rank="species" species="variegata">Tetraponera variegata</taxonomicName>
(Forel 1895)
<taxonomicNameLabel id="3D021ECAFE2CF13E1A94AF791954780E">stat. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C20FE2BD847DC955F910099A0B4DE139" type="description">
<paragraph id="952CDB8952A3F4EFEDCC82790B67E4DA" pageNumber="300">(Figs 21-22, 24)</paragraph>
</subSubSection>
<subSubSection id="4C7105B80919EC627B3554DE472180B7" type="reference_group">
<paragraph id="F992762B940576078FC8A5EA8E0530E8" pageNumber="300">
<materialsCitation id="9CB4F3D91D24E2395D78B6783D55AB2E" ID-GBIF-Occurrence="923898910" collectionCode="MHNG" collectorName="Sikora" country="Madagascar" location="Centr Madag" specimenCode="CASENT 0101045, CASENT 0101046" typeStatus="Syntypes">
Sima Grandidieri
<taxonomicName id="F17186C2D8BCDFFBE018DD17F7CE7806" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:152379" class="Insecta" family="Formicidae" genus="Tetraponera" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera grandidieri variegata (Forel)" order="Hymenoptera" pageNumber="300" phylum="Arthropoda" rank="subSpecies" species="grandidieri" species.completionEvidence="missing" species.completionRound="1" subSpecies="variegata">var. variegata Forel</taxonomicName>
1895: 487.
<typeStatus id="282E11B613CE2FABD6F93EED0955EE89">Syntypes</typeStatus>
, 2 workers, &quot;
<location id="2B9C074AF817115386D3A97BA00400E4" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:2B9C074AF817115386D3A97BA00400E4" country="Madagascar" name="Centr Madag">Centr Madag</location>
.&quot; (
<collectorName id="C8EE306F8DE647C46D2C7D87DCD697B4">Sikora</collectorName>
) (
<collectionCode id="DFA080C8CBEE4412839419D6BC514719" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
) [examined] [Imaged on AntWeb:
<specimenCode id="DEC0E4DECE07F422E3BFC6A9FC2BCD56">CASENT0101045</specimenCode>
,
<specimenCode id="649B63804B6A8F8D4C61CCA157E06B49">CASENT0101046</specimenCode>
].
<taxonomicNameLabel id="C6E2B1DE06C2730A189E23859A8A210F">
<typeStatus id="5D08986651CB8DF5D52C4965429C92ED">Syn. n</typeStatus>
.
</taxonomicNameLabel>
<typeStatus id="8F5A611E46612FECE2F7BB0F3F9674E0">One syntype</typeStatus>
(
<specimenCode id="CCFAD5346DCD4355AD057B5539C535E3">CASENT0101046</specimenCode>
)
<typeStatus id="319CE2A47BDF0C53E471A3847BFFD56F">here designated lectotype</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph id="D1C59DCE7558AFBB604DABC159CAEBDC" pageNumber="301">
<taxonomicName id="DB50191E3688267FE4E5A2E017D39546" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:152379" class="Insecta" family="Formicidae" genus="Tetraponera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera grandidieri variegata (Forel)" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="variety" species="grandidieri" variety="variegata">
<pageStartToken id="C8B108CAC8B6FC056A12EC08D1CA6BEC" pageNumber="301">Tetraponera</pageStartToken>
<taxonomicName id="5CE489578FB86AB3144607546FF6E29D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:152379" class="Insecta" family="Formicidae" genus="Tetraponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera grandidieri variegata (Forel)" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="variety" species="grandidieri" variety="variegata">grandidieri var. variegata (Forel)</taxonomicName>
</taxonomicName>
; Wheeler 1922: 1014. Combination in
<taxonomicName id="6AD8C3C9FE6CD242E860C52752F1D168" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24859" class="Insecta" family="Formicidae" genus="Tetraponera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera Smith" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="genus">Tetraponera</taxonomicName>
.
</paragraph>
<paragraph id="10D0378799D8267BB90FA6D9E6F1564A" pageNumber="301">
<taxonomicName id="D60D7A1C3ECBCE227AFE0DCEC6058686" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:152379" class="Insecta" family="Formicidae" genus="Tetraponera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera grandidieri variegata (Forel)" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="variety" species="grandidieri" variety="variegata">
Tetraponera
<taxonomicName id="C3BDADDF530342F738C399ED325F61A1" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:152379" class="Insecta" family="Formicidae" genus="Tetraponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera grandidieri variegata (Forel)" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="variety" species="grandidieri" variety="variegata">grandidieri var. variegata (Forel)</taxonomicName>
</taxonomicName>
; Santschi 1926: 27. Description of queen.
</paragraph>
</subSubSection>
<subSubSection id="D227307DCAD503E41F925880F19FB6FC" type="materials_examined">
<paragraph id="432442672C5EAA505DB0A473D223EF1C" pageNumber="301">
<materialsCitation id="261F2790EC4E7BB14EBB122C2A790E3C" ID-GBIF-Occurrence="923898906" collectionCode="CASC, MHNG, NHMB, PSWC" collectorName="Fisher, B. L." country="MADAGASCAR" location="Antananarivo">
Material Examined.-(
<collectionCode id="1711E23625441BD864838BA55EACB9DD" collectionName="USA, California, San Francisco, California Academy of Sciences">CASC</collectionCode>
,
<collectionCode id="0FD41030A4D95D43455B2E4746A1804E" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
,
<collectionCode id="AB6515F2BA6A2B7691C4B36DE899DC98" collectionName="Switzerland, Basel, Naturhistorisches Museum">NHMB</collectionCode>
,
<collectionCode id="0BCE878C8CDFF381FC51363E6C22BF6C" collectionName="PSWC">PSWC</collectionCode>
)
<collectingCountry id="C937EBBB6EC684FCA2E87938098C58C8">MADAGASCAR</collectingCountry>
<location id="099C922CC1607690812CE37B00BED19E" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:099C922CC1607690812CE37B00BED19E" country="MADAGASCAR" name="Antananarivo">Antananarivo</location>
:
<location id="3225D613E788F6FF8D09AEDD3878FE81" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:3225D613E788F6FF8D09AEDD3878FE81" country="MADAGASCAR" name="3 km 41 ° NE Andranomay, 11.5 km 147 ° SSE Anjozorobe">3 km 41° NE Andranomay, 11.5 km 147° SSE Anjozorobe</location>
, 1300 m (
<collectorName id="E32765FC098633B44E61FFC466F25452">Fisher, B. L.</collectorName>
; et al)
</materialsCitation>
;
<materialsCitation id="2FC569E7AAC0BDD002E5DFBBE4E0A7F5" ID-GBIF-Occurrence="923898909" collectorName="Irwin, M. E." country="Madagascar" location="Antsiranana">
<location id="4237A0A0524F96C39F0853D68BDE820A" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:4237A0A0524F96C39F0853D68BDE820A" country="Madagascar" name="Antsiranana">Antsiranana</location>
:
<location id="DDB0CD63C77FCD683D1F07CE41390FA5" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:DDB0CD63C77FCD683D1F07CE41390FA5" country="Madagascar" name="PN Marojejy">PN Marojejy</location>
, 488 m (
<collectorName id="6EF20EC1A328D49ACBA046EF0E23F912">Irwin, M. E.</collectorName>
)
</materialsCitation>
;
<materialsCitation id="9C7CF7C536F7FC600B863BBDE648A795" ID-GBIF-Occurrence="923898911" collectorName="Steiner, W. E." country="Madagascar" location="Fianarantsoa">
<location id="876084640B91B68A2FD6AF667DC7C332" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:876084640B91B68A2FD6AF667DC7C332" country="Madagascar" name="Fianarantsoa">Fianarantsoa</location>
:
<location id="2DC1422D76A08FE0AF51C1F1CF8BF17B" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:2DC1422D76A08FE0AF51C1F1CF8BF17B" country="Madagascar" name="7 km W Ranomafana">7 km W Ranomafana</location>
, 1100 m (
<collectorName id="9FED463B09C106CC193F543935141E5C">Steiner, W. E.</collectorName>
)
</materialsCitation>
;
<materialsCitation id="F920C971DB58A88FFE661CA33094D7F3" ID-GBIF-Occurrence="923898908" collectorName="Irwin, M. E., Schlinger, E. I." country="Madagascar" location="Ranomafana National Park">
<location id="4CE983F4EA471DF43F219FA14A7E8664" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:4CE983F4EA471DF43F219FA14A7E8664" country="Madagascar" name="Ranomafana National Park">Ranomafana National Park</location>
,
<location id="FA3A20EC1D87108B4F9A84F1A36F88B4" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:FA3A20EC1D87108B4F9A84F1A36F88B4" country="Madagascar" name="Talatakely">Talatakely</location>
, 850 m (
<collectorName id="150D638CA0AA1163432E02A41B837FAB">Irwin, M. E.</collectorName>
;
<collectorName id="D7BDD6559E3DEA53AFF056EC30054FB9">Schlinger, E. I.</collectorName>
)
</materialsCitation>
;
<materialsCitation id="4CFA1DA7990F02B7BB4E89A6F08DA2D9" ID-GBIF-Occurrence="923898907" collectorName="Fisher, В. L.; et al." country="Madagascar" location="Kalambatritra">
RS
<location id="A361468DE55AC0C0E0E1EEAFAC79F37F" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:A361468DE55AC0C0E0E1EEAFAC79F37F" country="Madagascar" name="Kalambatritra">Kalambatritra</location>
,
<location id="DBBCAAD702D5103E445B21FA39D36988" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:DBBCAAD702D5103E445B21FA39D36988" country="Madagascar" name="Ampanihy">Ampanihy</location>
, 1269 m (
<collectorName id="C2E7436FD8A977F49F9A766DF690122C">Fisher, В. L.; et al.</collectorName>
)
</materialsCitation>
;
<materialsCitation id="684F81F7143F666B1DE4107AE53E3BF8" ID-GBIF-Occurrence="923898914" collectorName="Pauly, A." country="Madagascar" location="Toamasina">
<location id="82230AA92C089F4CB53E2586860FFFC8" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:82230AA92C089F4CB53E2586860FFFC8" country="Madagascar" name="Toamasina">Toamasina</location>
:
<location id="701CE77CC70FF241CEF1190F4D483FC5" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:701CE77CC70FF241CEF1190F4D483FC5" country="Madagascar" name="Moramanga">Moramanga</location>
(
<location id="CDF60D898A87A452502B7095D81DA56C" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:CDF60D898A87A452502B7095D81DA56C" country="Madagascar" name="Descarpentriesi Morarano-Chrome">Descarpentriesi Morarano-Chrome</location>
(
<collectorName id="C67E4DC08E2B34CD53C5D74C40E06793">Pauly, A.</collectorName>
)
</materialsCitation>
;
<materialsCitation id="985409CFF88B66AF2636AA8833A7E176" ID-GBIF-Occurrence="923898915" collectorName="Fisher, B. L.; et al" country="Madagascar" location="Toliara">
<location id="FD44465E843EB792549DC7FBA44A43C4" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:FD44465E843EB792549DC7FBA44A43C4" country="Madagascar" name="Toliara">Toliara</location>
:
<location id="6548F381EEF0AD1EDFF596AB46C3E8B7" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:6548F381EEF0AD1EDFF596AB46C3E8B7" country="Madagascar" name="Foret Ivohibe">
<normalizedToken id="C2D2A829F92E338FCBB4D8FA635DE988" originalValue="Forêt">Foret</normalizedToken>
Ivohibe
</location>
, 650 m ((
<collectorName id="3631847B935DA1FFE57483B32B021428">Fisher, B. L.; et al</collectorName>
.)
</materialsCitation>
;
<materialsCitation id="80EF2FBBAF29FB5D4CFC6BC11D0E28AC" ID-GBIF-Occurrence="923898912" collectorName="Sikora" country="Madagascar" location="Centr Madag">
province unknown: &quot;
<location id="DD80701F15A4EFDD4EC5AA63AA9F9AB1" LSID="urn:lsid:plazi:treatment:174E1FEE3CDC7C8DB62D64F369D6B35E:DD80701F15A4EFDD4EC5AA63AA9F9AB1" country="Madagascar" name="Centr Madag">Centr Madag</location>
.&quot; (
<collectorName id="5E82A08250664920E11DC80FEF102AA7">Sikora</collectorName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="6798869C2786CF7F997D3798AA24605C" pageNumber="301">Worker measurements (n = 6). HW</paragraph>
</subSubSection>
<subSubSection id="EA60FB07940AC53E77FFA290DA16826E" type="description">
<paragraph id="B020301795063F0DF8BE0CB25CF5B52A" pageNumber="301">1.15-1.36, HL 1.36-1.62, LHT 1.39-1.59, CI 0.80-0.85, FCI 0.13-0.17, REL 0.29-0.32, REL2 0.35-0.38, SI 0.76-0.81, FI 0.30-0.32, PLI 0.49-0.55, PWI 0.41-0.46.</paragraph>
</subSubSection>
<subSubSection id="7D9EF0C918260E581C23BF92B10BAEFD" type="diagnosis">
<paragraph id="EE81FA2CF3C6B8D54785D922E7905B78" pageNumber="301">
Worker diagnosis. Similar to
<taxonomicName id="08A18D0F54B45FDA1BFBA1ACCC193A0C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37337" class="Insecta" family="Formicidae" genus="Tetraponera" genus.bestMatchDistance="418" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera grandidieri (Forel)" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="species" species="grandidieri">T. grandidieri</taxonomicName>
(q.v.), but larger on average. Basal margin of mandible lacking tooth; anterior clypeal margin broadly convex and crenulate, directed forward; metanotal spiracle visible in lateral view of mesosoma; dorsal face of propodeum broadly convex in posterior view; standing pilosity and appressed pubescence generally sparse; integument mostly sublucid, with fine coriarious /puncticulate sculpture; mesosoma, petiole and postpetiole orange-brown, head and gaster a contrasting blackish brown, legs with a black band on the distal portions of the femora.
</paragraph>
</subSubSection>
<subSubSection id="8497008D718152C5E6F49B38FE022867" type="discussion">
<paragraph id="A49D5684EC505D95A5966B4B84E8C983" pageNumber="301">
Comments.
<taxonomicName id="B2A412E98C3024A0D3D20015A91DFA45" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:261542" class="Insecta" family="Formicidae" genus="Tetraponera" genus.bestMatchDistance="458" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera variegata" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="species" species="variegata">T. variegata</taxonomicName>
can be distinguished from related species by the bicolored body and black banded legs (Fig. 22).
<taxonomicName id="7EB6D70636F21E71805291FC192304D7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37337" class="Insecta" family="Formicidae" genus="Tetraponera" genus.bestMatchDistance="294" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera grandidieri (Forel)" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="species" species="grandidieri">T. grandidieri</taxonomicName>
lacks black banding on the legs and, although the body is often bicolored, only the head is dark, not the head and gaster (as in
<taxonomicName id="769F81955485FCC45E18628CA52F8390" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:261542" class="Insecta" family="Formicidae" genus="Tetraponera" genus.bestMatchDistance="513" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera variegata" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="species" species="variegata">T. variegata</taxonomicName>
). Although such color differences might appear to be a weak basis for treating
<taxonomicName id="2FCFD4C604DC95AECCEB59CF19A62490" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:261542" class="Insecta" family="Formicidae" genus="Tetraponera" genus.bestMatchDistance="531" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera variegata" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="species" species="variegata">T. variegata</taxonomicName>
as a species distinct from
<taxonomicName id="7C8CE327D1539DE61185E30301AEBEF9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37337" class="Insecta" family="Formicidae" genus="Tetraponera" genus.bestMatchDistance="236" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera grandidieri (Forel)" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="species" species="grandidieri">T. grandidieri</taxonomicName>
, the two forms have been collected sympatrically at several sites (PN Marojejy, PN Ranomafana,
<normalizedToken id="655FC46AC294C52A368AA9BA5AB06AC9" originalValue="Forêt">Foret</normalizedToken>
Ivohibe) without showing any signs of intergradation. One other species,
<taxonomicName id="9FCA531A040A05D74F34185781EE8C64" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:260972" class="Insecta" family="Formicidae" genus="Tetraponera" genus.bestMatchDistance="6284" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera hespera WArd" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="species" species="hespera">T. hespera</taxonomicName>
, from northwestern Madagascar, exhibits black leg banding in most populations but in that species the body is unicolored light yellow or orange-brown.
</paragraph>
<paragraph id="5B42F42B7DCE389B14DBB8F9E51504B2" pageNumber="301">
I have designated a
<typeStatus id="CD67CA836EDCC8C75F4FFFEE86B52F23">lectotype</typeStatus>
for
<taxonomicName id="7D67A9720E223D491828D9BD3ED3F6EE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:261542" class="Insecta" family="Formicidae" genus="Tetraponera" genus.bestMatchDistance="608" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera variegata" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="species" species="variegata">T. variegata</taxonomicName>
since there is a dealate queen in
<collectionCode id="A053019805B01A00FAB116734A9EF80A" collectionName="Switzerland, Basel, Naturhistorisches Museum">NHMB</collectionCode>
(Basel) from Moramanga (leg. Descarpentries) labeled, incorrectly, as a
<taxonomicName id="A68AEFFF4B6349DAFF1E15726873D513" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:261542" class="Insecta" family="Formicidae" genus="Tetraponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera variegata" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="species" species="variegata">variegata</taxonomicName>
&quot;type&quot;. This specimen has no status as a type, but it reflects the practice of earlier myrmecologists of designating &quot;type specimens&quot; for queens and maies when they were described later than the worker caste of the same species.
</paragraph>
</subSubSection>
<subSubSection id="976E2B5DF78F308A88E0A1A40980A3EB" type="distribution">
<paragraph id="D1539EB58E3FAE1F50388315AE61A7CB" pageNumber="301">
Distribution and biology.
<taxonomicName id="894034D49634114C85EB09960F0AEAE8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:261542" class="Insecta" family="Formicidae" genus="Tetraponera" genus.bestMatchDistance="685" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera variegata" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="species" species="variegata">T. variegata</taxonomicName>
is known from several widely scattered locations in the rainforest zone of eastern Madagascar (Fig. 24). Its range broadly overlaps the distributions of Г.
<taxonomicName id="479598189F58CC441BC534428EA04E58" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37337" class="Insecta" family="Formicidae" genus="Tetraponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera grandidieri (Forel)" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="species" species="grandidieri">
grandidieri,
<taxonomicName id="CCDF5B3D1654C1651977DF1F3D2D16B1" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:260975" class="Insecta" family="Formicidae" genus="Tetraponera" genus.bestMatchDistance="4700" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera inermis Ward" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="species" species="inermis">T. inermis</taxonomicName>
</taxonomicName>
and
<taxonomicName id="14F23412569C5C0C142CDB25E4512554" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:260977" class="Insecta" family="Formicidae" genus="Tetraponera" genus.bestMatchDistance="2449" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tetraponera merita Ward" order="Hymenoptera" pageNumber="301" phylum="Arthropoda" rank="species" species="merita">T. merita</taxonomicName>
. Specimens have been collected in Malaise traps and foraging on vegetation. Up to this point no nests have been found.
</paragraph>
</subSubSection>
</treatment>
</document>