326 lines
45 KiB
XML
326 lines
45 KiB
XML
<document id="27DD0845BDE060678A9BF45376C125DC" ID-DOI="10.11646/zootaxa.3946.3.2" ID-GBIF-Dataset="d57d5862-5fc5-480c-84c9-3b598b597197" ID-ISSN="1175-5326" ID-Zenodo-Dep="244741" ID-ZooBank="24E2F41D-89BF-473F-ACE0-ED951BCB2699" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461034391373" checkinUser="plazi" docAuthor="Grinang, Jongkar & Ng, Peter K. L." docDate="2015" docId="03DF87F3FFC26E73FF4FFE3FD03D2FF6" docLanguage="en" docName="zt03946p346.pdf" docOrigin="Zootaxa 3946 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Terrathelphusa mas Grinang & Ng, 2015, n. sp." docType="treatment" docVersion="9" lastPageNumber="344" masterDocId="FFE6FF8BFFC86E7DFFD8FF92D3052E7E" masterDocTitle="The identity of the semiterrestrial crab Terrathelphusa kuchingensis (Nobili, 1901) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of four new species from southwestern Sarawak, Borneo, Malaysia" masterLastPageNumber="346" masterPageNumber="331" pageNumber="341" updateTime="1698574262036" updateUser="plazi">
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<mods:titleInfo id="7D3A8EDD6868B5A730A3743562487F3A">
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<mods:title id="1AADC033151638C91A00FC09B4F19C17">The identity of the semiterrestrial crab Terrathelphusa kuchingensis (Nobili, 1901) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of four new species from southwestern Sarawak, Borneo, Malaysia</mods:title>
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<mods:name id="04A074072E5C3D01EE83FA37800DD9D9" type="personal">
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<mods:roleTerm id="74F00D9319313851A4900423F9F05FE1">Author</mods:roleTerm>
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<mods:namePart id="DEC483A7115FA78E6B8D8D6061636DC4">Grinang, Jongkar</mods:namePart>
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<mods:name id="E7AB989F3879C97CC09AEA3168005943" type="personal">
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<mods:roleTerm id="DFB05462A103B4B734E545280DECA2BE">Author</mods:roleTerm>
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<mods:namePart id="A885827F8CA8AFBA9A275B5392E91F0B">Ng, Peter K. L.</mods:namePart>
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<mods:typeOfResource id="B0E81CD7B80F7F4D6B5F6EB291C345BB">text</mods:typeOfResource>
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<mods:title id="C3036F65724A9C49445F88A84CB70E98">Zootaxa</mods:title>
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<mods:part id="C077BAC898C5558F9D7CD940793B195B">
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<mods:date id="B40FC806FE96A4DC7E5C4DBB60FC99AD">2015</mods:date>
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<mods:detail id="523C9A83D130499DFEE19696A8569128" type="volume">
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<mods:number id="D1599418AFDACF409ABE6D91AFDB5545">3946</mods:number>
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<mods:detail id="AA13DFFD9095C2FB89CC62B75B2415C9" type="issue">
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<mods:number id="B15D5B1DD57D64B8ACE92BB67550FBD8">3</mods:number>
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<mods:classification id="117BB8BE2451FA9AC515208B02F03847">journal article</mods:classification>
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<mods:identifier id="FBF3F0F55F19AAEE449D632806B84FDB" type="DOI">10.11646/zootaxa.3946.3.2</mods:identifier>
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<mods:identifier id="FBE94759D2BC6EE08CBD3B11C90B26CC" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03DF87F3FFC26E73FF4FFE3FD03D2FF6" ID-DOI="http://doi.org/10.5281/zenodo.5695393" ID-GBIF-Taxon="119608783" ID-Zenodo-Dep="5695393" LSID="urn:lsid:plazi:treatment:03DF87F3FFC26E73FF4FFE3FD03D2FF6" httpUri="http://treatment.plazi.org/id/03DF87F3FFC26E73FF4FFE3FD03D2FF6" lastPageId="14" lastPageNumber="344" pageId="10" pageNumber="341">
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<subSubSection id="C36C656EFFC26E77FF4FFE3FD2D42FB9" box="[151,465,429,455]" pageId="10" pageNumber="341" type="nomenclature">
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<paragraph id="8BC936E5FFC26E77FF4FFE3FD2D42FB9" blockId="10.[151,468,429,488]" box="[151,465,429,455]" pageId="10" pageNumber="341">
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<heading id="D0818189FFC26E77FF4FFE3FD2D42FB9" bold="true" box="[151,465,429,455]" fontSize="11" level="1" pageId="10" pageNumber="341" reason="1">
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<emphasis id="B902EAF7FFC26E77FF4FFE3FD2D42FB9" bold="true" box="[151,465,429,455]" pageId="10" pageNumber="341">
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<taxonomicName id="4C764D66FFC26E77FF4FFE3FD28D2FB9" ID-CoL="55HXM" box="[151,392,429,455]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="mas" status="sp. nov.">
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<emphasis id="B902EAF7FFC26E77FF4FFE3FD28D2FB9" bold="true" box="[151,392,429,455]" italics="true" pageId="10" pageNumber="341">Terrathelphusa mas</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A231578CFFC26E77FE57FE3FD2D42FB9" box="[399,465,429,455]" pageId="10" pageNumber="341" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C656EFFC26E76FF4FFE5DD1212F69" lastPageId="11" lastPageNumber="342" pageId="10" pageNumber="341" type="materials_examined">
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<paragraph id="8BC936E5FFC26E76FF4FFE5DD1212F69" blockId="10.[151,468,429,488]" lastBlockId="11.[151,1437,151,961]" lastPageId="11" lastPageNumber="342" pageId="10" pageNumber="341">
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(
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<figureCitation id="134D2A60FFC26E77FF47FE5DD3EE2F96" box="[159,235,463,488]" captionStart="FIGURE 9" captionStartId="10.[151,250,1848,1870]" captionTargetBox="[338,1245,542,1820]" captionTargetId="figure@10.[338,1245,542,1820]" captionTargetPageId="10" captionText="FIGURE 9. Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244750/files/figure.png" pageId="10" pageNumber="341">Figs. 9</figureCitation>
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A–H, 10A–E,
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<date id="FFC81025FFC26E77FE56FE42D2B02F96" box="[398,437,464,488]" pageId="10" pageNumber="341" value="2011-01">11I</date>
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, J)
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<emphasis id="B902EAF7FFC36E76FF4FFF05D2852ECE" bold="true" box="[151,384,151,176]" pageId="11" pageNumber="342">Material examined</emphasis>
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.
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<typeStatus id="54CD8847FFC36E76FE4CFF05D1052ECE" box="[404,512,151,176]" pageId="11" pageNumber="342" type="holotype">Holotype</typeStatus>
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: male (35.5 ×
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<quantity id="4C8E9B00FFC36E76FD64FF05D02D2ECE" box="[700,808,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="11" pageNumber="342" unit="mm" value="26.4">26.4 mm</quantity>
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) (
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<collectionCode id="ED67AE20FFC36E76FC9DFF0AD0782ECE" box="[837,893,152,176]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="342">ZRC</collectionCode>
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2015.0005), open green golf course, Borneo Highlands Resort, Gunung Penrissen, Kuching,
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<collectingCountry id="F3617675FFC36E76FD6FFF2ED01F2EAB" box="[695,794,188,213]" name="Malaysia" pageId="11" pageNumber="342">Sarawak</collectingCountry>
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, 1O07’43.8”N 110O13’07.3”E,
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<quantity id="4C8E9B00FFC36E76FBB7FF2ED7B22EAA" box="[1135,1207,188,213]" metricMagnitude="2" metricUnit="m" metricValue="8.52" pageId="11" pageNumber="342" unit="m" value="852.0">852 m</quantity>
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asl, coll. J. Grinang
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<emphasis id="B902EAF7FFC36E76FF4FFF4CD3C82E88" box="[151,205,221,246]" italics="true" pageId="11" pageNumber="342">et al</emphasis>
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.,
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<date id="FFC81025FFC36E76FF3FFF4FD2CB2E88" box="[231,462,221,246]" pageId="11" pageNumber="342" value="2013-09-16">16 September 2013</date>
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.
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<typeStatus id="54CD8847FFC36E76FE06FF4CD1412E88" box="[478,580,222,246]" pageId="11" pageNumber="342" type="paratype">Paratype</typeStatus>
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:
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<specimenCount id="9D70FD6CFFC36E76FD8CFF4CD1C02E88" box="[596,709,221,246]" pageId="11" pageNumber="342" type="female">2 females</specimenCount>
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(larger 26.3 ×
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<quantity id="4C8E9B00FFC36E76FC58FF4CD0EF2E88" box="[896,1002,222,246]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="11" pageNumber="342" unit="mm" value="20.4">20.4 mm</quantity>
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) (SBC.C.00374–75);
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<specimenCount id="9D70FD6CFFC36E76FB32FF4CD6552E88" box="[1258,1360,221,246]" pageId="11" pageNumber="342" type="female">1 female</specimenCount>
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(
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<collectionCode id="ED67AE20FFC36E76FABCFF4CD6992E88" box="[1380,1436,222,246]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="342">ZRC</collectionCode>
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2014.0840); data same as
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<typeStatus id="54CD8847FFC36E76FE62FF6CD1252F69" box="[442,544,254,279]" pageId="11" pageNumber="342" type="holotype">holotype</typeStatus>
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||
.
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</paragraph>
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||
</subSubSection>
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<caption id="DF09666DFFC26E77FF4FF8AAD1AB29D4" httpUri="https://zenodo.org/record/244750/files/figure.png" pageId="10" pageNumber="341" targetBox="[338,1245,542,1820]" targetPageId="10">
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<paragraph id="8BC936E5FFC26E77FF4FF8AAD1AB29D4" blockId="10.[151,1436,1848,1962]" pageId="10" pageNumber="341">
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<emphasis id="B902EAF7FFC26E77FF4FF8AAD2142930" bold="true" box="[151,273,1848,1870]" pageId="10" pageNumber="341">FIGURE 9</emphasis>
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.
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<taxonomicName id="4C764D66FFC26E77FEFAF8AAD2F52930" box="[290,496,1848,1870]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="mas" status="sp. nov.">
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||
<emphasis id="B902EAF7FFC26E77FEFAF8AAD2F52930" box="[290,496,1848,1870]" italics="true" pageId="10" pageNumber="341">Terrathelphusa mas</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A231578CFFC26E77FE22F8AAD13D2930" box="[506,568,1848,1870]" pageId="10" pageNumber="341" rank="species">
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<emphasis id="B902EAF7FFC26E77FE22F8AAD13D2930" bold="true" box="[506,568,1848,1870]" pageId="10" pageNumber="341">n. sp.</emphasis>
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</taxonomicNameLabel>
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, holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela.
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||
</paragraph>
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||
</caption>
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||
<subSubSection id="C36C656EFFC36E76FF1FFEB2D0F32CBA" pageId="11" pageNumber="342" type="diagnosis">
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||
<paragraph id="8BC936E5FFC36E76FF1FFEB2D0B52CDE" blockId="11.[151,1437,151,961]" pageId="11" pageNumber="342">
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<emphasis id="B902EAF7FFC36E76FF1FFEB2D2442F47" bold="true" box="[199,321,288,313]" pageId="11" pageNumber="342">Diagnosis.</emphasis>
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Carapace broader than long, conspicuously swollen, surface convex, smooth; cervical grooves broad, deep; epigastric cristae high, separated by deep epigastric groove; H-groove deep, long almost confluent with cervical grooves; epibranchial tooth indistinct but can be separated from external orbital angle by notch, anterolateral, frontal regions appearing compressed, supraorbital margin almost parallel with frontal margin; epistome median lobe blunt to bifurcated (
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<figureCitation id="134D2A60FFC36E76FD5CFE22D1CC2FB6" box="[644,713,432,457]" captionStart="FIGURE 9" captionStartId="10.[151,250,1848,1870]" captionTargetBox="[338,1245,542,1820]" captionTargetId="figure@10.[338,1245,542,1820]" captionTargetPageId="10" captionText="FIGURE 9. Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244750/files/figure.png" pageId="11" pageNumber="342">Fig. 9</figureCitation>
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A–C). Male abdomen with elongated somite 6, lateral margins gently concave (
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<figureCitation id="134D2A60FFC36E76FE83FE46D2A12F92" box="[347,420,468,493]" captionStart="FIGURE 9" captionStartId="10.[151,250,1848,1870]" captionTargetBox="[338,1245,542,1820]" captionTargetId="figure@10.[338,1245,542,1820]" captionTargetPageId="10" captionText="FIGURE 9. Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244750/files/figure.png" pageId="11" pageNumber="342">Fig. 9</figureCitation>
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D). Third maxilliped ischium rectangular, about 2.1 times maximum width (
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<figureCitation id="134D2A60FFC36E76FAEBFE46D6792F92" box="[1331,1404,468,493]" captionStart="FIGURE 9" captionStartId="10.[151,250,1848,1870]" captionTargetBox="[338,1245,542,1820]" captionTargetId="figure@10.[338,1245,542,1820]" captionTargetPageId="10" captionText="FIGURE 9. Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244750/files/figure.png" pageId="11" pageNumber="342">Fig. 9</figureCitation>
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E). Ambulatory legs smooth, relatively slender, length about 4.4 times width (
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<figureCitation id="134D2A60FFC36E76FBD7FE6AD75D2C6E" box="[1039,1112,504,529]" captionStart="FIGURE 9" captionStartId="10.[151,250,1848,1870]" captionTargetBox="[338,1245,542,1820]" captionTargetId="figure@10.[338,1245,542,1820]" captionTargetPageId="10" captionText="FIGURE 9. Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244750/files/figure.png" pageId="11" pageNumber="342">Fig. 9</figureCitation>
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F). Cheliped carpus rugose, without granules (
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<figureCitation id="134D2A60FFC36E76FEBEFD8ED2AF2C4A" box="[358,426,540,565]" captionStart="FIGURE 9" captionStartId="10.[151,250,1848,1870]" captionTargetBox="[338,1245,542,1820]" captionTargetId="figure@10.[338,1245,542,1820]" captionTargetPageId="10" captionText="FIGURE 9. Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244750/files/figure.png" pageId="11" pageNumber="342">Fig. 9</figureCitation>
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G, H), inner angle with narrower, acutely triangular tooth; gap between fingers of major chela conspicuously wide (
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<figureCitation id="134D2A60FFC36E76FE0EFDD2D11A2C26" box="[470,543,576,601]" captionStart="FIGURE 9" captionStartId="10.[151,250,1848,1870]" captionTargetBox="[338,1245,542,1820]" captionTargetId="figure@10.[338,1245,542,1820]" captionTargetPageId="10" captionText="FIGURE 9. Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244750/files/figure.png" pageId="11" pageNumber="342">Fig. 9</figureCitation>
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||
A). G1 distinctly curving outwards; terminal segment cone-shaped, tapered, gently curving upwards to tip, about 0.3 times length of subterminal segment (
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<figureCitation id="134D2A60FFC36E76FC25FDF6D7482C02" box="[1021,1101,612,637]" captionStart="FIGURE 10" captionStartId="11.[151,250,1771,1793]" captionTargetBox="[188,1396,1011,1741]" captionTargetId="figure@11.[182,1406,1005,1753]" captionTargetPageId="11" captionText="FIGURE 10. Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244751/files/figure.png" pageId="11" pageNumber="342">Fig. 10</figureCitation>
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A, B, D, E). Distal segment of G2 relatively long, about 0.8 times length of basal segment (
|
||
<figureCitation id="134D2A60FFC36E76FCE4FD1AD08A2CDE" box="[828,911,648,673]" captionStart="FIGURE 10" captionStartId="11.[151,250,1771,1793]" captionTargetBox="[188,1396,1011,1741]" captionTargetId="figure@11.[182,1406,1005,1753]" captionTargetPageId="11" captionText="FIGURE 10. Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244751/files/figure.png" pageId="11" pageNumber="342">Fig. 10</figureCitation>
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C).
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</paragraph>
|
||
<paragraph id="8BC936E5FFC36E76FF1FFD3ED0F32CBA" blockId="11.[151,1437,151,961]" box="[199,1014,684,709]" pageId="11" pageNumber="342">
|
||
<emphasis id="B902EAF7FFC36E76FF1FFD3ED24A2CBB" bold="true" box="[199,335,684,709]" pageId="11" pageNumber="342">Life colour.</emphasis>
|
||
Body and appendages are bright golden yellow (
|
||
<figureCitation id="134D2A60FFC36E76FCACFD3ED0C12CBA" box="[884,964,684,709]" captionStart="FIGURE 11" captionStartId="12.[151,250,1856,1878]" captionTargetBox="[157,1429,195,1832]" captionTargetId="figure@12.[157,1430,195,1834]" captionTargetPageId="12" captionText="FIGURE 11. Life colour of five species of Terrathelphusa from Kuching Division, Sarawak. A, B, Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), shortly after collection, Gunung Temurang; C, D, Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), Gunung Penrissen; E, F, Terrathelphusa cerina n. sp., female (24.2 × 20.1 mm), captured-marked-released, Gunung Singai; G, H, Terrathelphusa kundong n. sp., female (30.8 × 23.0 mm) (SBC. C. 00364 - 65), Gunung Serambu; I, J, Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), Gunung Penrissen." httpUri="https://zenodo.org/record/244752/files/figure.png" pageId="11" pageNumber="342">
|
||
Fig.
|
||
<date id="FFC81025FFC36E76FC70FD3ED0C12CBA" box="[936,964,684,708]" pageId="11" pageNumber="342" value="2011-01">11</date>
|
||
</figureCitation>
|
||
I, J).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C656EFFC36E76FF1FFD42D2442D07" pageId="11" pageNumber="342" type="description">
|
||
<paragraph id="8BC936E5FFC36E76FF1FFD42D2682D4F" blockId="11.[151,1437,151,961]" pageId="11" pageNumber="342">
|
||
<emphasis id="B902EAF7FFC36E76FF1FFD42D23E2C97" bold="true" box="[199,315,720,745]" pageId="11" pageNumber="342">Variation.</emphasis>
|
||
The
|
||
<typeStatus id="54CD8847FFC36E76FEA4FD43D2ED2C97" box="[380,488,721,745]" pageId="11" pageNumber="342" type="paratype">paratypes</typeStatus>
|
||
agree well with the
|
||
<typeStatus id="54CD8847FFC36E76FD0CFD42D0332C97" box="[724,822,720,745]" pageId="11" pageNumber="342" type="holotype">holotype</typeStatus>
|
||
male except for variations in a few characters. In the larger specimens, the ambulatory legs are relatively longer and the cervical grooves are somewhat broader than in smaller specimens.
|
||
</paragraph>
|
||
<paragraph id="8BC936E5FFC36E76FF1FFCAED2442D07" blockId="11.[151,1437,151,961]" pageId="11" pageNumber="342">
|
||
<emphasis id="B902EAF7FFC36E76FF1FFCAED22F2D2B" bold="true" box="[199,298,828,853]" pageId="11" pageNumber="342">Habitat.</emphasis>
|
||
The species is sympatric with
|
||
<taxonomicName id="4C764D66FFC36E76FDA7FCAFD1E72D2B" box="[639,738,828,853]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="342" phylum="Arthropoda" rank="species" species="aglaia" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFC36E76FDA7FCAFD1E72D2B" box="[639,738,828,853]" italics="true" pageId="11" pageNumber="342">T. aglaia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFC36E76FD30FCAED0232D2B" box="[744,806,828,853]" pageId="11" pageNumber="342" rank="species">
|
||
<emphasis id="B902EAF7FFC36E76FD30FCAED0232D2B" bold="true" box="[744,806,828,853]" pageId="11" pageNumber="342">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, at high elevations on Gunung Penrissen, but it absent at low elevations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C656EFFC36E73FF1FFC16D03D2FF6" lastPageId="14" lastPageNumber="345" pageId="11" pageNumber="342" type="etymology">
|
||
<paragraph id="8BC936E5FFC36E76FF1FFC16D16E2DBF" blockId="11.[151,1437,151,961]" pageId="11" pageNumber="342">
|
||
<emphasis id="B902EAF7FFC36E76FF1FFC16D24B2DE3" bold="true" box="[199,334,900,925]" pageId="11" pageNumber="342">Etymology.</emphasis>
|
||
Derived from the Iban word
|
||
<taxonomicName id="4C764D66FFC36E76FD47FC17D1C92DE2" box="[671,716,901,924]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="342" phylum="Arthropoda" rank="species" species="mas">
|
||
<emphasis id="B902EAF7FFC36E76FD47FC17D1C92DE2" box="[671,716,901,924]" italics="true" pageId="11" pageNumber="342">mas</emphasis>
|
||
</taxonomicName>
|
||
for gold alluding to the golden appearance of the new species. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
<caption id="DF09666DFFC36E76FF4FF979D7862940" httpUri="https://zenodo.org/record/244751/files/figure.png" pageId="11" pageNumber="342" targetBox="[188,1396,1011,1741]" targetPageId="11">
|
||
<paragraph id="8BC936E5FFC36E76FF4FF979D7862940" blockId="11.[151,1436,1771,1854]" pageId="11" pageNumber="342">
|
||
<emphasis id="B902EAF7FFC36E76FF4FF979D219297F" bold="true" box="[151,284,1771,1793]" pageId="11" pageNumber="342">FIGURE 10</emphasis>
|
||
.
|
||
<taxonomicName id="4C764D66FFC36E76FEF5F979D2FF297F" box="[301,506,1771,1793]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="342" phylum="Arthropoda" rank="species" species="mas" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFC36E76FEF5F979D2FF297F" box="[301,506,1771,1793]" italics="true" pageId="11" pageNumber="342">Terrathelphusa mas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFC36E76FDDCF979D144297F" box="[516,577,1771,1793]" pageId="11" pageNumber="342" rank="species">
|
||
<emphasis id="B902EAF7FFC36E76FDDCF979D144297F" bold="true" box="[516,577,1771,1793]" pageId="11" pageNumber="342">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, ventral view of left G1; B, ventral view of distal part of left G1; C, dorsal view of left G2; D, dorsal view of G1; E, dorsal view of distal part of left G1. Scales: A, C, D = 1 mm; B, E = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF09666DFFC46E71FF4FF8D2D14B2991" httpUri="https://zenodo.org/record/244752/files/figure.png" pageId="12" pageNumber="343" targetBox="[157,1429,195,1832]" targetPageId="12">
|
||
<paragraph id="8BC936E5FFC46E71FF4FF8D2D14B2991" blockId="12.[151,1436,1856,2031]" pageId="12" pageNumber="343">
|
||
<emphasis id="B902EAF7FFC46E71FF4FF8D2D2252928" bold="true" box="[151,288,1856,1878]" pageId="12" pageNumber="343">FIGURE 11</emphasis>
|
||
. Life colour of five species of
|
||
<taxonomicName id="4C764D66FFC46E71FD48F8D2D02F2928" box="[656,810,1856,1878]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="343" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EAF7FFC46E71FD48F8D2D02F2928" box="[656,810,1856,1878]" italics="true" pageId="12" pageNumber="343">Terrathelphusa</emphasis>
|
||
</taxonomicName>
|
||
from Kuching Division, Sarawak. A, B,
|
||
<taxonomicName id="4C764D66FFC46E71FADAF8D2D2BE290B" authority="(Nobili, 1901)" authorityName="Nobili" authorityYear="1901" baseAuthorityName="Nobili" baseAuthorityYear="1901" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="343" phylum="Arthropoda" rank="species" species="kuchingensis">
|
||
<emphasis id="B902EAF7FFC46E71FADAF8D2D21F290B" italics="true" pageId="12" pageNumber="343">Terrathelphusa kuchingensis</emphasis>
|
||
(Nobili, 1901)
|
||
</taxonomicName>
|
||
, male (36.0 × 26.4 mm) (ZRC 2015.0001), shortly after collection, Gunung Temurang; C, D,
|
||
<taxonomicName id="4C764D66FFC46E71FF4FF8EFD27E29ED" box="[151,379,1917,1939]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="343" phylum="Arthropoda" rank="species" species="aglaia" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFC46E71FF4FF8EFD27E29ED" box="[151,379,1917,1939]" italics="true" pageId="12" pageNumber="343">Terrathelphusa aglaia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFC46E71FE5CF8EFD2C429ED" box="[388,449,1917,1939]" pageId="12" pageNumber="343" rank="species">
|
||
<emphasis id="B902EAF7FFC46E71FE5CF8EFD2C429ED" bold="true" box="[388,449,1917,1939]" pageId="12" pageNumber="343">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), Gunung Penrissen; E, F,
|
||
<taxonomicName id="4C764D66FFC46E71FADAF8EFD3DD29CC" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="343" phylum="Arthropoda" rank="species" species="cerina" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFC46E71FADAF8EFD3DD29CC" italics="true" pageId="12" pageNumber="343">Terrathelphusa cerina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFC46E71FF3AF80ED21A29CC" box="[226,287,1948,1970]" pageId="12" pageNumber="343" rank="species">
|
||
<emphasis id="B902EAF7FFC46E71FF3AF80ED21A29CC" bold="true" box="[226,287,1948,1970]" pageId="12" pageNumber="343">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female (24.2 × 20.1 mm), captured-marked-released, Gunung Singai; G, H,
|
||
<taxonomicName id="4C764D66FFC46E71FB8CF80ED64B29CC" box="[1108,1358,1948,1970]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="343" phylum="Arthropoda" rank="species" species="kundong" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFC46E71FB8CF80ED64B29CC" box="[1108,1358,1948,1970]" italics="true" pageId="12" pageNumber="343">Terrathelphusa kundong</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFC46E71FA81F80ED69329CC" box="[1369,1430,1948,1970]" pageId="12" pageNumber="343" rank="species">
|
||
<emphasis id="B902EAF7FFC46E71FA81F80ED69329CC" bold="true" box="[1369,1430,1948,1970]" pageId="12" pageNumber="343">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female (30.8 × 23.0 mm) (SBC.C.00364-65), Gunung Serambu; I, J,
|
||
<taxonomicName id="4C764D66FFC46E71FCBDF829D73529AF" box="[869,1072,1979,2001]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="343" phylum="Arthropoda" rank="species" species="mas" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFC46E71FCBDF829D73529AF" box="[869,1072,1979,2001]" italics="true" pageId="12" pageNumber="343">Terrathelphusa mas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFC46E71FBE0F829D77629AF" box="[1080,1139,1979,2001]" pageId="12" pageNumber="343" rank="species">
|
||
<emphasis id="B902EAF7FFC46E71FBE0F829D77629AF" bold="true" box="[1080,1139,1979,2001]" pageId="12" pageNumber="343">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), Gunung Penrissen.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF09666DFFC56E70FFA5FD84D7322C52" ID-Table-UUID="DF09666DFFC56E70FFA5FD84D7322C52" box="[125,1079,534,556]" httpUri="http://table.plazi.org/id/DF09666DFFC56E70FFA5FD84D7322C52" pageId="13" pageNumber="344" targetBox="[125,1879,736,1664]" targetIsTable="true" targetPageId="13">
|
||
<paragraph id="8BC936E5FFC56E70FFA5FD84D7322C52" blockId="13.[125,1875,534,704]" box="[125,1079,534,556]" pageId="13" pageNumber="344">
|
||
<emphasis id="B902EAF7FFC56E70FFA5FD84D3D72C52" bold="true" box="[125,210,534,556]" pageId="13" pageNumber="344">TABLE</emphasis>
|
||
1. Differences between eight
|
||
<taxonomicName id="4C764D66FFC56E70FE26FD84D19C2C52" box="[510,665,534,556]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="344" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EAF7FFC56E70FE26FD84D19C2C52" box="[510,665,534,556]" italics="true" pageId="13" pageNumber="344">Terrathelphusa</emphasis>
|
||
</taxonomicName>
|
||
species from Borneo [* from Ng (1997)]
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC936E5FFC56E70FFA5FDD0D4382CF2" blockId="13.[125,1875,534,704]" pageId="13" pageNumber="344">
|
||
Characters
|
||
<emphasis id="B902EAF7FFC56E70FEFBFDD6D4092C28" box="[291,1804,578,598]" italics="true" pageId="13" pageNumber="344">
|
||
<taxonomicName id="4C764D66FFC56E70FEFBFDD6D2702C28" box="[291,373,578,598]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="344" phylum="Arthropoda" rank="species" species="aglaia">T. aglaia</taxonomicName>
|
||
<taxonomicName id="4C764D66FFC56E70FE25FDD6D14A2C28" box="[509,591,580,598]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="344" phylum="Arthropoda" rank="species" species="cerina">T. cerina</taxonomicName>
|
||
<taxonomicName id="4C764D66FFC56E70FD10FDD6D0512C28" box="[712,852,578,598]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="344" phylum="Arthropoda" rank="species" species="kuchingensis">T. kuchingensis</taxonomicName>
|
||
<taxonomicName id="4C764D66FFC56E70FC4BFDD6D0FC2C28" box="[915,1017,578,598]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="344" phylum="Arthropoda" rank="species" species="kundong">T. kundong</taxonomicName>
|
||
<taxonomicName id="4C764D66FFC56E70FB9FFDD6D7DB2C28" box="[1095,1246,578,598]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="344" phylum="Arthropoda" rank="species" species="loxophthalma">T. loxophthalma</taxonomicName>
|
||
*
|
||
<taxonomicName id="4C764D66FFC56E70FAC3FDD6D6522C28" box="[1307,1367,580,598]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="344" phylum="Arthropoda" rank="species" species="mas">T. mas</taxonomicName>
|
||
<taxonomicName id="4C764D66FFC56E70FA2CFDD6D5372C28" box="[1524,1586,579,598]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="344" phylum="Arthropoda" rank="species" species="ovis">T. ovis</taxonomicName>
|
||
*
|
||
<taxonomicName id="4C764D66FFC56E70F966FDD6D4092C28" authority="Third Length" authorityName="Third Length" box="[1726,1804,578,598]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="344" phylum="Arthropoda" rank="species" species="telur">T. telur*</taxonomicName>
|
||
</emphasis>
|
||
Third Length of ischium Length of ischium Length of ischium Length of Length of ischium Length of ischium Length of ischium Length of maxilliped about 1.9 times about 2.0 times about 1.8 times ischium about about 1.8 times about 2.1 times about 1.6 times ischium about
|
||
</paragraph>
|
||
<paragraph id="8BC936E5FFC56E70FEFBFD00D4142CD8" blockId="13.[125,1875,534,704]" box="[291,1809,658,678]" pageId="13" pageNumber="344">maximum width maximum width maximum width 1.9 times maximum width maximum width maximum width 1.5 times</paragraph>
|
||
<paragraph id="8BC936E5FFC56E73FC4BFD39D03D2FF6" blockId="13.[125,1875,534,704]" lastBlockId="14.[151,1437,151,392]" lastPageId="14" lastPageNumber="345" pageId="13" pageNumber="344">
|
||
maximum width maximum width
|
||
<emphasis id="B902EAF7FFC66E73FF1FFF05D23E2ECE" bold="true" box="[199,315,151,176]" pageId="14" pageNumber="345">Remarks.</emphasis>
|
||
<taxonomicName id="4C764D66FFC66E73FE9CFF05D1262ECE" box="[324,547,151,176]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="345" phylum="Arthropoda" rank="species" species="mas" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFC66E73FE9CFF05D1262ECE" box="[324,547,151,176]" italics="true" pageId="14" pageNumber="345">Terrathelphusa mas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFC66E73FDF4FF05D1682ECE" box="[556,621,151,176]" pageId="14" pageNumber="345" rank="species">
|
||
<emphasis id="B902EAF7FFC66E73FDF4FF05D1682ECE" bold="true" box="[556,621,151,176]" pageId="14" pageNumber="345">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, can be distinguished from
|
||
<taxonomicName id="4C764D66FFC66E73FC76FF0BD7582ECE" box="[942,1117,151,176]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="345" phylum="Arthropoda" rank="species" species="kuchingensis">
|
||
<emphasis id="B902EAF7FFC66E73FC76FF0BD7582ECE" box="[942,1117,151,176]" italics="true" pageId="14" pageNumber="345">T. kuchingensis</emphasis>
|
||
</taxonomicName>
|
||
and the other species being described by a combination of characters: inner angle of carpus having a narrow and acutely triangular tooth (
|
||
<figureCitation id="134D2A60FFC66E73FAB7FF2ED3A02E86" captionStart="FIGURE 9" captionStartId="10.[151,250,1848,1870]" captionTargetBox="[338,1245,542,1820]" captionTargetId="figure@10.[338,1245,542,1820]" captionTargetPageId="10" captionText="FIGURE 9. Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244750/files/figure.png" pageId="14" pageNumber="345">Fig. 9</figureCitation>
|
||
A) (versus broad and acutely triangular tooth in
|
||
<taxonomicName id="4C764D66FFC66E73FD19FF73D0682E86" box="[705,877,223,248]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="345" phylum="Arthropoda" rank="species" species="kuchingensis">
|
||
<emphasis id="B902EAF7FFC66E73FD19FF73D0682E86" box="[705,877,223,248]" italics="true" pageId="14" pageNumber="345">T. kuchingensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134D2A60FFC66E73FCA4FF4DD0BF2E86" box="[892,954,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[302,1284,591,1881]" captionTargetId="figure@2.[302,1284,591,1881]" captionTargetPageId="2" captionText="FIGURE 1. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244742/files/figure.png" pageId="14" pageNumber="345">Fig. 1</figureCitation>
|
||
A),
|
||
<taxonomicName id="4C764D66FFC66E73FC32FF73D7482E86" box="[1002,1101,223,248]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="345" phylum="Arthropoda" rank="species" species="aglaia" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFC66E73FC32FF73D7482E86" box="[1002,1101,223,248]" italics="true" pageId="14" pageNumber="345">T. aglaia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFC66E73FB8CFF4DD7962E86" box="[1108,1171,223,248]" pageId="14" pageNumber="345" rank="species">
|
||
<emphasis id="B902EAF7FFC66E73FB8CFF4DD7962E86" bold="true" box="[1108,1171,223,248]" pageId="14" pageNumber="345">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="134D2A60FFC66E73FB7BFF4DD7E12E86" box="[1187,1252,223,248]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1924]" captionTargetBox="[356,1228,673,1884]" captionTargetId="figure@4.[356,1228,673,1884]" captionTargetPageId="4" captionText="FIGURE 3. Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244744/files/figure.png" pageId="14" pageNumber="345">Fig. 3</figureCitation>
|
||
A) and
|
||
<taxonomicName id="4C764D66FFC66E73FAE1FF73D69C2E86" box="[1337,1433,225,248]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="345" phylum="Arthropoda" rank="species" species="cerina" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFC66E73FAE1FF73D69C2E86" box="[1337,1433,225,248]" italics="true" pageId="14" pageNumber="345">T. cerina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFC66E73FF4FFE96D3DF2F63" box="[151,218,260,285]" pageId="14" pageNumber="345" rank="species">
|
||
<emphasis id="B902EAF7FFC66E73FF4FFE96D3DF2F63" bold="true" box="[151,218,260,285]" pageId="14" pageNumber="345">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="134D2A60FFC66E73FF35FE96D2362F63" box="[237,307,260,285]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[341,1248,375,1692]" captionTargetId="figure@6.[341,1248,375,1692]" captionTargetPageId="6" captionText="FIGURE 5. Terrathelphusa cerina n. sp., holotype male (24.7 × 19.3 mm) (ZRC 2015.0003), sandstone outcrop, Gunung Singai, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244746/files/figure.png" pageId="14" pageNumber="345">Fig. 5</figureCitation>
|
||
A), and broadly triangular tooth in
|
||
<taxonomicName id="4C764D66FFC66E73FD0AFE97D0572F63" box="[722,850,260,285]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="345" phylum="Arthropoda" rank="species" species="kundong" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFC66E73FD0AFE97D0572F63" box="[722,850,260,285]" italics="true" pageId="14" pageNumber="345">T. kundong</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFC66E73FC84FE96D09A2F63" box="[860,927,260,285]" pageId="14" pageNumber="345" rank="species">
|
||
<emphasis id="B902EAF7FFC66E73FC84FE96D09A2F63" bold="true" box="[860,927,260,285]" pageId="14" pageNumber="345">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="134D2A60FFC66E73FC6BFE96D0FF2F62" box="[947,1018,260,285]" captionStart="FIGURE 7" captionStartId="8.[151,250,1725,1747]" captionTargetBox="[206,1384,197,1688]" captionTargetId="figure@8.[206,1384,197,1689]" captionTargetPageId="8" captionText="FIGURE 7. Terrathelphusa kundong n. sp., holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244748/files/figure.png" pageId="14" pageNumber="345">Fig. 7</figureCitation>
|
||
A); and the G1 is distinctly curving outwards (
|
||
<figureCitation id="134D2A60FFC66E73FED5FEB5D25B2F3E" box="[269,350,295,320]" captionStart="FIGURE 10" captionStartId="11.[151,250,1771,1793]" captionTargetBox="[188,1396,1011,1741]" captionTargetId="figure@11.[182,1406,1005,1753]" captionTargetPageId="11" captionText="FIGURE 10. Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244751/files/figure.png" pageId="14" pageNumber="345">Fig. 10</figureCitation>
|
||
A, B, C, D) (versus gently curving outwards in others,
|
||
<figureCitation id="134D2A60FFC66E73FC10FEB5D7102F3E" box="[968,1045,295,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1052,1074]" captionTargetBox="[159,1427,198,1024]" captionTargetId="figure@3.[153,1434,193,1030]" captionTargetPageId="3" captionText="FIGURE 2. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244743/files/figure.png" pageId="14" pageNumber="345">Figs. 2</figureCitation>
|
||
A, B, D, E; 4A, B, D, E; 6A, B, D, E; 8A, B, D, E).
|
||
<taxonomicName id="4C764D66FFC66E73FE87FEDED10C2F1B" box="[351,521,332,357]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="345" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EAF7FFC66E73FE87FEDED10C2F1B" box="[351,521,332,357]" italics="true" pageId="14" pageNumber="345">Terrathelphusa</emphasis>
|
||
</taxonomicName>
|
||
species from the Kuching region differ from taxa in the northern region
|
||
<emphasis id="B902EAF7FFC66E73FA80FEDFD2342FF6" italics="true" pageId="14" pageNumber="345">
|
||
viz.
|
||
<taxonomicName id="4C764D66FFC66E73FA5FFEDFD2342FF6" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="345" phylum="Arthropoda" rank="species" species="loxophthalma">T. loxophthalma</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C764D66FFC66E73FEE7FEE3D2822FF6" box="[319,391,369,392]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="345" phylum="Arthropoda" rank="species" species="ovis">
|
||
<emphasis id="B902EAF7FFC66E73FEE7FEE3D2822FF6" box="[319,391,369,392]" italics="true" pageId="14" pageNumber="345">T. ovis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C764D66FFC66E73FE66FEE3D10B2FF6" box="[446,526,367,392]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="345" phylum="Arthropoda" rank="species" species="telur">
|
||
<emphasis id="B902EAF7FFC66E73FE66FEE3D10B2FF6" box="[446,526,367,392]" italics="true" pageId="14" pageNumber="345">T. telur</emphasis>
|
||
</taxonomicName>
|
||
as summarised in
|
||
<tableCitation id="C6F4035EFFC66E73FD06FEFDD0372FF6" box="[734,818,367,392]" captionStart="TABLE 1" captionStartId="13.[125,210,534,556]" captionTargetBox="[125,1879,736,1664]" captionTargetPageId="13" captionText="TABLE 1. Differences between eight Terrathelphusa species from Borneo [* from Ng (1997)]" httpUri="http://table.plazi.org/id/DF09666DFFC56E70FFA5FD84D7322C52" pageId="14" pageNumber="345" tableUuid="DF09666DFFC56E70FFA5FD84D7322C52">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |