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<document id="ADF83C36F89F064322116A3F761BC654" ID-CLB-Dataset="28423" ID-DOI="10.11646/zootaxa.4564.2.11" ID-GBIF-Dataset="c088902a-755d-4218-8dda-9820e2b1e333" ID-ISSN="1175-5326" ID-Zenodo-Dep="2588961" ID-ZooBank="7378731F-9FB5-407A-ADB9-5430C15917B6" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552193110972" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2019" docId="03CFA937556C1100FF1DE7F9FEA4FE48" docLanguage="en" docName="zootaxa.4564.2.11.pdf" docOrigin="Zootaxa 4564 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guajirolus ektrapeloglossa Flowers 1985" docType="treatment" docVersion="7" lastPageNumber="552" masterDocId="FFF6D14F55641115FF8AE263FFBCFFC0" masterDocTitle="Systematics of Guajirolus ektrapeloglossa Flowers 1985 (Ephemeroptera: Baetidae)" masterLastPageNumber="553" masterPageNumber="531" pageNumber="539" updateTime="1698700815134" updateUser="ExternalLinkService">
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<mods:title id="16BD68E2912C702EB77BC63C89BB7C95">Systematics of Guajirolus ektrapeloglossa Flowers 1985 (Ephemeroptera: Baetidae)</mods:title>
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<mods:namePart id="36CC22A11C19F629D0BC4A4C5E6CF025">Kluge, Nikita J.</mods:namePart>
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<treatment id="03CFA937556C1100FF1DE7F9FEA4FE48" ID-DOI="http://doi.org/10.5281/zenodo.5923793" ID-GBIF-Taxon="154366494" ID-Zenodo-Dep="5923793" LSID="urn:lsid:plazi:treatment:03CFA937556C1100FF1DE7F9FEA4FE48" httpUri="http://treatment.plazi.org/id/03CFA937556C1100FF1DE7F9FEA4FE48" lastPageId="21" lastPageNumber="552" pageId="8" pageNumber="539">
<subSubSection id="C37C4BAA556C111DFF1DE7F9FD34FA74" box="[151,648,1434,1460]" pageId="8" pageNumber="539" type="nomenclature">
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<heading id="D091AF4D556C111DFF1DE7F9FD34FA74" bold="true" box="[151,648,1434,1460]" fontSize="11" level="1" pageId="8" pageNumber="539" reason="1">
<taxonomicName id="4C6663A2556C111DFF1DE7F9FD34FA74" ID-CoL="3HHYL" authority="Flowers 1985" authorityName="Flowers" authorityYear="1985" box="[151,648,1434,1460]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C433556C111DFF1DE7F9FD34FA74" bold="true" box="[151,648,1434,1460]" pageId="8" pageNumber="539">
<emphasis id="B912C433556C111DFF1DE7F9FE67FA74" bold="true" box="[151,475,1434,1460]" italics="true" pageId="8" pageNumber="539">Guajirolus ektrapeloglossa</emphasis>
<bibRefCitation id="EFF765D0556C111DFE69E7F9FD34FA74" author="Flowers, R. W." box="[483,648,1434,1460]" pageId="8" pageNumber="539" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers 1985</bibRefCitation>
</emphasis>
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<subSubSection id="C37C4BAA556C111DFF1DE7DEFC93F9DB" pageId="8" pageNumber="539" type="description">
<paragraph id="8BD91821556C111DFF1DE7DEFEA5FA16" blockId="8.[151,648,1434,1494]" box="[151,281,1469,1494]" pageId="8" pageNumber="539">
(
<figureCitation id="135D04A4556C111DFF15E7DEFEACFA16" box="[159,272,1469,1494]" captionStart-0="FIGURES 1–3" captionStart-1="FIGURES 4–11" captionStart-10="FIGURES 59–69" captionStart-11="FIGURES 70–74" captionStart-12="FIGURES 75–80" captionStart-13="FIGURES 81–82" captionStart-14="FIGURES 83–88" captionStart-2="FIGURES 12–16" captionStart-3="FIGURES 17–21" captionStart-4="FIGURES 22–28" captionStart-5="FIGURES 29–35" captionStart-6="FIGURES 36–40" captionStart-7="FIGURES 41–43" captionStart-8="FIGURES 44–50" captionStart-9="FIGURES 51–58" captionStartId-0="2.[151,264,2010,2033]" captionStartId-1="3.[151,264,1708,1731]" captionStartId-10="16.[151,264,1942,1965]" captionStartId-11="17.[151,264,1969,1992]" captionStartId-12="18.[151,264,1754,1777]" captionStartId-13="19.[151,264,1894,1917]" captionStartId-14="21.[151,264,1913,1936]" captionStartId-2="5.[151,264,1907,1930]" captionStartId-3="6.[151,264,1909,1932]" captionStartId-4="7.[151,264,1417,1440]" captionStartId-5="8.[151,264,1275,1298]" captionStartId-6="10.[151,264,1934,1957]" captionStartId-7="13.[151,264,1877,1900]" captionStartId-8="14.[151,264,1937,1960]" captionStartId-9="15.[151,264,2007,2030]" captionTargetBox-0="[157,1428,650,1986]" captionTargetBox-1="[184,1405,451,1693]" captionTargetBox-10="[178,1421,193,1929]" captionTargetBox-11="[159,1427,223,1913]" captionTargetBox-12="[158,1428,193,1733]" captionTargetBox-13="[158,1429,547,1844]" captionTargetBox-14="[163,1423,465,1892]" captionTargetBox-2="[157,1430,193,1886]" captionTargetBox-3="[171,1428,199,1815]" captionTargetBox-4="[189,1417,267,1396]" captionTargetBox-5="[158,1428,195,1254]" captionTargetBox-6="[157,1429,326,1896]" captionTargetBox-7="[156,1431,355,1848]" captionTargetBox-8="[156,1430,258,1857]" captionTargetBox-9="[156,1430,243,1914]" captionTargetId-0="figure@2.[157,1429,628,1986]" captionTargetId-1="figure@3.[184,1405,440,1693]" captionTargetId-10="figure@16.[176,1421,193,1929]" captionTargetId-11="figure@17.[159,1427,223,1913]" captionTargetId-12="figure@18.[158,1428,193,1733]" captionTargetId-13="figure@19.[158,1429,547,1844]" captionTargetId-14="figure@21.[163,1423,465,1892]" captionTargetId-2="figure@5.[157,1430,193,1886]" captionTargetId-3="figure@6.[157,1429,193,1889]" captionTargetId-4="figure@7.[169,1417,267,1396]" captionTargetId-5="figure@8.[158,1428,195,1254]" captionTargetId-6="figure@10.[157,1429,326,1896]" captionTargetId-7="figure@13.[156,1431,337,1856]" captionTargetId-8="figure@14.[156,1430,258,1857]" captionTargetId-9="figure@15.[156,1430,243,1914]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-10="16" captionTargetPageId-11="17" captionTargetPageId-12="18" captionTargetPageId-13="19" captionTargetPageId-14="21" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="10" captionTargetPageId-7="13" captionTargetPageId-8="14" captionTargetPageId-9="15" captionText-0="FIGURES 1–3. Female larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón." captionText-1="FIGURES 4–11. Labrum of last instar larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 4, labrum; 5, exuviae of labrum; 6, longitudinal section of lateral part; 7–8, one and the same median emargination differently compressed; 9–11, fragments of exuviae of one labrum: 9, with focus on distal margin; 10, with focus on dorsal side; 11, detached ventral cuticle (right side at right)." captionText-10="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." captionText-11="FIGURES 70–74. Guajirolus ektrapeloglossa from Quebrada Barrigón. 70–71, male imagines; 72, subimaginal exuviae of left half of mesonotum; 73, subimaginal exuviae of left half of meso- and metapleuron; 74, subimago extracted from larval exuviae." captionText-12="FIGURES 75–80. Male genitals of Guajirolus ektrapeloglossa from Quebrada Barrigón. 75, genital of imago (ventral view; muscles and internal parts of penis shown by interrupted lines; right gonostylar and gonovectal muscles not shown); 76, subimaginal genitals developing under larval cuticle; 77–78 and 79–80, genitals of two specimens. Abbreviations: 1, 2, 3, segments of gonostylus; pgs, larval protogonostylus; usg, unistyliger." captionText-13="FIGURES 81–82. Female imagines of Guajirolus ektrapeloglossa from Quebrada Barrigón." captionText-14="FIGURES 83–88. Eggs of Guajirolus ektrapeloglossa. 83–85, from Quebrada Barrigón (Panama); 86–88, from Rio Sotorani (Peru)." captionText-2="FIGURES 12–16. Mandibles of Guajirolus ektrapeloglossa from Quebrada Barrigón. 12–13 right and left mandibles inside old cuticle before molt from penultimate to last larval instar; 14–15, biting edges of right and left mandibles; 16, lateral part of left mandible." captionText-3="FIGURES 17–21. Guajirolus ektrapeloglossa from Quebrada Barrigón. 17, maxilla (dorsal view); 18, apex of maxilla (ventral view); 19, distal margin of labrum (dorsal view); 20–21, biting denticles of left and right mandibles. Abbreviations: 1d, distal subsegment of first palpomere; 1p, proximal subsegment of first palpomere; 2, second palpomere; ds1, ds2, ds3, dentisetae; in, incisor; kd, kinetodontium; prs, prostheca." captionText-4="FIGURES 22–28. Guajirolus ektrapeloglossa from Quebrada Barrigón. 22–23, exuviae of maxillae; 24, maxilla; 25, exuviae of hypopharynx; 26–28, hypopharynx." captionText-5="FIGURES 29–35. Labium of Guajirolus ektrapeloglossa from Quebrada Barrigón. 29–31, labium (29, ventral view; 30–31, in translucent light, with focus on glossae and on palps); 32–33, one and the same labial palp differently compressed; 34–35, left and right labial palps of one and the same individual." captionText-6="FIGURES 36–40. Setation of labium of Guajirolus ektrapeloglossa from Quebrada Barrigón. 36, exuviae of glossae and paraglossa (focus on ventral side); 37, glossae (focus on dorsal side); 38–40, exuviae of palp and paraglossa (38, 40, focus on ventral side; 39, focus on dorsal side)." captionText-7="FIGURES 41–43. Larval thorax of Guajirolus ektrapeloglossa from Quebrada Barrigón. 41, exuviae of left half of pronotum and mesonotum; 42, posterior margin of mesonotum and protoptera; 43, exuviae of sterna and pleura (fore leg turned anteriorly, middle and hind legs detached). Abbreviations: abd1, first abdominal tergum; msth, mesothorax; mtth, metathorax; prth, prothorax." captionText-8="FIGURES 44–50. Larval legs of Guajirolus ektrapeloglossa from Quebrada Barrigón. 44–45, fore and middle legs of darkest specimen; 46–47, claws; 48, fragment of fore femur (focus on anterior side); 49, fragment of fore tibia (focus on anterior side); 50, fragment of hind tibia (focus on posterior side)." captionText-9="FIGURES 51–58. Larval coloration of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri-0="https://zenodo.org/record/2588963/files/figure.png" httpUri-1="https://zenodo.org/record/2588965/files/figure.png" httpUri-10="https://zenodo.org/record/2588983/files/figure.png" httpUri-11="https://zenodo.org/record/2588985/files/figure.png" httpUri-12="https://zenodo.org/record/2588987/files/figure.png" httpUri-13="https://zenodo.org/record/2588989/files/figure.png" httpUri-14="https://zenodo.org/record/2588991/files/figure.png" httpUri-2="https://zenodo.org/record/2588967/files/figure.png" httpUri-3="https://zenodo.org/record/2588969/files/figure.png" httpUri-4="https://zenodo.org/record/2588971/files/figure.png" httpUri-5="https://zenodo.org/record/2588973/files/figure.png" httpUri-6="https://zenodo.org/record/2588975/files/figure.png" httpUri-7="https://zenodo.org/record/2588977/files/figure.png" httpUri-8="https://zenodo.org/record/2588979/files/figure.png" httpUri-9="https://zenodo.org/record/2588981/files/figure.png" pageId="8" pageNumber="539">Figs 1–88</figureCitation>
)
</paragraph>
<paragraph id="8BD91821556C111DFF1DE466FC93F9DB" blockId="8.[151,1163,1541,1686]" box="[151,815,1541,1563]" pageId="8" pageNumber="539">
Genus 3 nr.
<taxonomicName id="4C6663A2556C111DFE9BE466FDD4F9DB" authority=": Roback 1966: 135" authorityName=": Roback" authorityPageNumber="135" authorityYear="1966" box="[273,616,1541,1563]" class="Insecta" family="Baetidae" genus="Pseudocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="genus">
<emphasis id="B912C433556C111DFE9BE466FE23F9DB" box="[273,415,1541,1563]" italics="true" pageId="8" pageNumber="539">Pseudocloeon</emphasis>
:
<bibRefCitation id="EFF765D0556C111DFE26E466FD8BF9DB" author="Roback, S. S." box="[428,567,1541,1563]" pageId="8" pageNumber="539" pagination="129 - 199" refId="ref9176" refString="Roback, S. S. (1966) The Catherwood Foundation Peruvia-Amazon expedition. VI-Ephemeroptera nymphs. Monographs of the Academy of Natural Sciences of Philadelphia, No. 14, 129 - 199." type="book chapter" year="1966">Roback 1966</bibRefCitation>
: 135
</taxonomicName>
, figs 58–63 (larva).
</paragraph>
</subSubSection>
<subSubSection id="C37C4BAA556C111DFF1DE447FB37F955" pageId="8" pageNumber="539" type="reference_group">
<paragraph id="8BD91821556C111DFF1DE447FCE0F9FA" blockId="8.[151,1163,1541,1686]" box="[151,860,1572,1594]" pageId="8" pageNumber="539">
<treatmentCitationGroup id="AB763F0F556C111DFF1DE447FCE0F9FA" box="[151,860,1572,1594]" pageId="8" pageNumber="539">
<taxonomicName id="4C6663A2556C111DFF1DE447FDDDF9FA" ID-CoL="3HHYL" authority="Flowers 1985: 29" authorityName="Flowers" authorityPageNumber="29" authorityYear="1985" box="[151,609,1572,1594]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C433556C111DFF1DE447FE16F9FA" box="[151,426,1572,1594]" italics="true" pageId="8" pageNumber="539">Guajirolus ektrapeloglossa</emphasis>
<treatmentCitation id="0AC73E30556C111DFE3BE447FDDDF9FA" author="Flowers, R. W." box="[433,609,1572,1594]" page="29" pageId="8" pageNumber="539" year="1985">
<bibRefCitation id="EFF765D0556C111DFE3BE447FD82F9FA" author="Flowers, R. W." box="[433,574,1572,1594]" pageId="8" pageNumber="539" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers 1985</bibRefCitation>
: 29
</treatmentCitation>
</taxonomicName>
(
<specimenCount id="9D60D3A8556C111DFDFAE447FD38F9FA" box="[624,644,1572,1594]" pageId="8" pageNumber="539" type="male">♂</specimenCount>
and
<specimenCount id="9D60D3A8556C111DFD3EE446FD7BF9FA" box="[692,711,1573,1594]" pageId="8" pageNumber="539" type="female">♀</specimenCount>
imago, larva).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD91821556C111DFF1DE421FCF6F998" blockId="8.[151,1163,1541,1686]" box="[151,842,1602,1624]" pageId="8" pageNumber="539">
<treatmentCitationGroup id="AB763F0F556C111DFF1DE421FCF6F998" box="[151,842,1602,1624]" pageId="8" pageNumber="539">
<taxonomicName id="4C6663A2556C111DFF1DE421FD08F998" ID-CoL="3HHYN" authority="Lugo-Ortiz &amp; McCafferty 1995: 58" authorityName="Lugo-Ortiz &amp; McCafferty" authorityPageNumber="58" authorityYear="1995" box="[151,692,1602,1624]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C433556C111DFF1DE421FEFBF998" box="[151,327,1602,1624]" italics="true" pageId="8" pageNumber="539">Guajirolus nanus</emphasis>
<treatmentCitation id="0AC73E30556C111DFEC7E421FD08F998" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[333,692,1602,1624]" page="58" pageId="8" pageNumber="539" year="1995">
<bibRefCitation id="EFF765D0556C111DFEC7E421FD2DF998" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[333,657,1602,1624]" pageId="8" pageNumber="539" pagination="68 - 70" refId="ref8926" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1995) Guajirolus nanus (Ephemeroptera: Baetidae), a new species from Costa Rica. Entomological News, 106 (2), 68 - 70." type="journal article" year="1995">Lugo-Ortiz &amp; McCafferty 1995</bibRefCitation>
: 58
</treatmentCitation>
</taxonomicName>
(larva)
<emphasis id="B912C433556C111DFC8EE420FCF6F998" bold="true" box="[772,842,1603,1624]" pageId="8" pageNumber="539">syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD91821556C111DFF1DE402FC8BF9B6" blockId="8.[151,1163,1541,1686]" box="[151,823,1633,1655]" pageId="8" pageNumber="539">
<treatmentCitationGroup id="AB763F0F556C111DFF1DE402FC8BF9B6" box="[151,823,1633,1655]" pageId="8" pageNumber="539">
<taxonomicName id="4C6663A2556C111DFF1DE402FDA5F9B7" ID-CoL="3HHYP" authority="Nieto 2003: 153" authorityName="Nieto" authorityPageNumber="153" authorityYear="2003" box="[151,537,1633,1655]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="species" species="queremba">
<emphasis id="B912C433556C111DFF1DE402FED2F9B7" box="[151,366,1633,1655]" italics="true" pageId="8" pageNumber="539">Guajirolus queremba</emphasis>
<treatmentCitation id="0AC73E30556C111DFEFEE402FDA5F9B7" author="Nieto, C." box="[372,537,1633,1655]" page="153" pageId="8" pageNumber="539" year="2003">
<bibRefCitation id="EFF765D0556C111DFEFEE402FE55F9B7" author="Nieto, C." box="[372,489,1633,1655]" pageId="8" pageNumber="539" pagination="153 - 158" refId="ref8969" refString="Nieto, C. (2003) A new species of Guajirolus (Ephemeroptera: Baetidae) from Argentina and description of a new genus from Bolivia. In: Gaino E. (Ed.), Research Update on Ephemeroptera &amp; Plecoptera (Proc. 2001 Int. Joint Meeting-X Int. Conf. on Ephemeroptera &amp; XIV Int. Symp. on Plecoptera, August 5 - 11, 2001 in Perugia). Univ. Perugia, Italy, 153 - 158." type="book chapter" year="2003">Nieto 2003</bibRefCitation>
: 153
</treatmentCitation>
</taxonomicName>
(
<specimenCount id="9D60D3A8556C111DFDA0E402FD87F9B7" box="[554,571,1633,1655]" pageId="8" pageNumber="539" type="female">♀</specimenCount>
subimago, larva)
<emphasis id="B912C433556C111DFD78E402FC8BF9B6" bold="true" box="[754,823,1633,1654]" pageId="8" pageNumber="539">syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD91821556C111DFF1DE4E3FB37F955" blockId="8.[151,1163,1541,1686]" box="[151,1163,1664,1686]" pageId="8" pageNumber="539">
<taxonomicName id="4C6663A2556C111DFF1DE4E3FC49F956" ID-CoL="4RBZ" authority="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth)" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[151,1013,1664,1686]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="genus">
<emphasis id="B912C433556C111DFF1DE4E3FEE0F956" box="[151,348,1664,1686]" italics="true" pageId="8" pageNumber="539">Guajirolus flowersi</emphasis>
Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth) 2005: 21
</taxonomicName>
(larva)
<emphasis id="B912C433556C111DFBCFE4E3FB37F955" bold="true" box="[1093,1163,1664,1685]" pageId="8" pageNumber="539">syn. n.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37C4BAA556C111CFF1DE4A1FD44FD45" lastPageId="9" lastPageNumber="540" pageId="8" pageNumber="539" type="materials_examined">
<paragraph id="8BD91821556C111DFF1DE4A1FD19F873" blockId="8.[151,1437,1730,2007]" pageId="8" pageNumber="539">
<emphasis id="B912C433556C111DFF1DE4A1FE38F91B" bold="true" box="[151,388,1730,1755]" pageId="8" pageNumber="539">Material examined.</emphasis>
<materialsCitation id="3B0E127C556C111DFE04E4A0FAFCF886" ID-GBIF-Occurrence="2012767228" collectingDate="2018-01-19" collectingDateMax="2018-01-22" collectingDateMin="2018-01-19" collectorName="N. Kluge &amp; L. Sheyko" country="Panama" county="La Esperanza" latitude="8.578611" location="Nuevo" longLatPrecision="21" longitude="-82.32333" municipality="Rio" pageId="8" pageNumber="539" specimenCount="10" specimenCount-female="5" specimenCount-male="5" stateProvince="Chiriqui">
<collectingCountry id="F37158B1556C111DFE04E4A0FDB4F91B" box="[398,520,1731,1755]" name="Panama" pageId="8" pageNumber="539">PANAMA</collectingCountry>
,
<collectingRegion id="49A2D6C3556C111DFD9CE4A1FD53F91B" box="[534,751,1730,1755]" country="Panama" name="Chiriqui" pageId="8" pageNumber="539">Provincia Chiriquí</collectingRegion>
:
<collectingCounty id="62B860AD556C111DFD76E4A0FC2BF91A" box="[764,919,1731,1754]" pageId="8" pageNumber="539">La Esperanza</collectingCounty>
(
<locationDeviation id="9A6B0103556C111DFC21E4A0FB1CF91B" box="[939,1184,1730,1755]" location="Gualaca" pageId="8" pageNumber="539">
<quantity id="4C9EB5C4556C111DFC21E4A0FC54F91B" box="[939,1000,1730,1755]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="539" unit="km" value="7.0">7 km</quantity>
NNW Gualaca
</locationDeviation>
),
<collectingMunicipality id="6BBD825B556C111DFB3DE4A1FB63F91B" box="[1207,1247,1730,1755]" pageId="8" pageNumber="539">Rio</collectingMunicipality>
<collectingRegion id="49A2D6C3556C111DFB63E4A1FAFAF91B" box="[1257,1350,1730,1755]" country="Panama" name="Chiriqui" pageId="8" pageNumber="539">Chiriquí</collectingRegion>
<location id="8EB94EFA556C111DFADBE4A0FA20F91A" LSID="urn:lsid:plazi:treatment:03CFA937556C1100FF1DE7F9FEA4FE48:8EB94EFA556C111DFADBE4A0FA20F91A" box="[1361,1436,1731,1754]" country="Panama" county="La Esperanza" latitude="8.578611" longLatPrecision="21" longitude="-82.32333" municipality="Rio" name="Nuevo" pageId="8" pageNumber="539" stateProvince="Chiriqui">Nuevo</location>
between dam and power station, downstream of footbridge,
<collectingDate id="EF9CC709556C111DFCCDE484FC09F93F" box="[839,949,1767,1791]" pageId="8" pageNumber="539" value="2018-01-19">
<date id="FFD83EE1556C111DFCCDE484FC09F93F" box="[839,949,1767,1791]" pageId="8" pageNumber="539" value="2018-01-19">19.I.2018</date>
</collectingDate>
, coll.
<collectorName id="26937DF7556C111DFC76E484FBD5F93E" box="[1020,1129,1766,1790]" pageId="8" pageNumber="539">N. Kluge</collectorName>
: 1 L
<specimenCount id="9D60D3A8556C111DFB29E484FB08F93E" box="[1187,1204,1767,1790]" pageId="8" pageNumber="539" type="female">♀</specimenCount>
;
<location id="8EB94EFA556C111DFB49E485FA21F93E" LSID="urn:lsid:plazi:treatment:03CFA937556C1100FF1DE7F9FEA4FE48:8EB94EFA556C111DFB49E485FA21F93E" box="[1219,1437,1766,1790]" country="Panama" county="La Esperanza" latitude="8.578611" longLatPrecision="21" longitude="-82.32333" municipality="Rio" name="Quebrada Barrigon" pageId="8" pageNumber="539" stateProvince="Chiriqui">Quebrada Barrigón</location>
just upstream of the road bridge
<locationDeviation id="9A6B0103556C111DFD8FE569FC28F8E3" box="[517,916,1802,1827]" firstLocation="La Esperanza" pageId="8" pageNumber="539" secondLocation="Gualaca">between La Esperanza and Gualaca</locationDeviation>
(
<geoCoordinate id="EE527EE6556C111DFC2EE568FBA7F8E3" box="[932,1051,1803,1827]" direction="north" orientation="latitude" pageId="8" pageNumber="539" precision="15" value="8.578611">8°34'43''N</geoCoordinate>
,
<geoCoordinate id="EE527EE6556C111DFBADE568FB0FF8E3" box="[1063,1203,1803,1827]" direction="west" orientation="longitude" pageId="8" pageNumber="539" precision="15" value="-82.32333">82°19'24''W</geoCoordinate>
),
<collectingDate id="EF9CC709556C111DFB42E568FAE3F8E3" box="[1224,1375,1803,1827]" pageId="8" pageNumber="539" value="2018-01-20" valueMax="2018-01-22" valueMin="2018-01-20">
<date id="FFD83EE1556C111DFB42E568FAE3F8E3" box="[1224,1375,1803,1827]" pageId="8" pageNumber="539" value="2018-01-20" valueMax="2018-01-22" valueMin="2018-01-20">20–22.I.2018</date>
</collectingDate>
, coll.
<collectorName id="26937DF7556C111DFF1DE54CFF43F887" box="[151,255,1838,1863]" pageId="8" pageNumber="539">N. Kluge</collectorName>
&amp;
<collectorName id="26937DF7556C111DFEAFE54CFE22F887" box="[293,414,1838,1863]" pageId="8" pageNumber="539">L. Sheyko</collectorName>
: 2 L-S-I
<specimenCount id="9D60D3A8556C111DFD88E54CFDA9F887" box="[514,533,1839,1863]" pageId="8" pageNumber="539" type="male">♂</specimenCount>
, 1 S-I
<specimenCount id="9D60D3A8556C111DFDD7E54CFDCDF887" box="[605,625,1839,1863]" pageId="8" pageNumber="539" type="male">♂</specimenCount>
, 2 L-S
<specimenCount id="9D60D3A8556C111DFD48E54CFD6AF887" box="[706,726,1839,1863]" pageId="8" pageNumber="539" type="male">♂</specimenCount>
, 1 L/S
<specimenCount id="9D60D3A8556C111DFCAFE54CFC85F887" box="[805,825,1839,1863]" pageId="8" pageNumber="539" type="male">♂</specimenCount>
, 19 L
<specimenCount id="9D60D3A8556C111DFCF5E54CFC28F886" box="[895,916,1839,1862]" pageId="8" pageNumber="539" type="male">♂</specimenCount>
, 8 L-S-I
<specimenCount id="9D60D3A8556C111DFC77E54CFBB7F887" box="[1021,1035,1839,1863]" pageId="8" pageNumber="539" type="female">♀</specimenCount>
, 1 L-S/I
<specimenCount id="9D60D3A8556C111DFBF8E54CFB3DF887" box="[1138,1153,1839,1863]" pageId="8" pageNumber="539" type="female">♀</specimenCount>
, 1 L-S
<specimenCount id="9D60D3A8556C111DFB5CE54CFB59F887" box="[1238,1253,1839,1863]" pageId="8" pageNumber="539" type="female">♀</specimenCount>
, 48 L
<specimenCount id="9D60D3A8556C111DFAA5E54CFAFCF886" box="[1327,1344,1839,1862]" pageId="8" pageNumber="539" type="female">♀</specimenCount>
</materialsCitation>
.
<materialsCitation id="3B0E127C556C111DFAC7E54CFD19F873" ID-GBIF-Occurrence="2012767227" collectingDate="2006-01-07" collectingDateMax="2013-08-21" collectingDateMin="2006-01-07" collectorName="N. Kluge &amp; L. Sheyko" country="Peru" county="Region Junun" location="Rio Sotorani" municipality="Provincia Chanchamayo" pageId="8" pageNumber="539" specimenCount="2" specimenCount-female="2" stateProvince="Loreto">
<collectingCountry id="F37158B1556C111DFAC7E54CFA25F886" box="[1357,1433,1839,1862]" name="Peru" pageId="8" pageNumber="539">PERU</collectingCountry>
:
<collectingCounty id="62B860AD556C111DFF1DE531FE80F8AA" box="[151,316,1874,1899]" pageId="8" pageNumber="539">Region Junun</collectingCounty>
,
<collectingMunicipality id="6BBD825B556C111DFEC7E531FDD6F8AB" box="[333,618,1874,1899]" pageId="8" pageNumber="539">Provincia Chanchamayo</collectingMunicipality>
,
<location id="8EB94EFA556C111DFDF6E531FCADF8AB" LSID="urn:lsid:plazi:treatment:03CFA937556C1100FF1DE7F9FEA4FE48:8EB94EFA556C111DFDF6E531FCADF8AB" box="[636,785,1874,1899]" country="Peru" county="Region Junun" municipality="Provincia Chanchamayo" name="Rio Sotorani" pageId="8" pageNumber="539" stateProvince="Loreto">Rio Sotorani</location>
(right tributary of
<location id="8EB94EFA556C111DFC75E531FB3DF8AA" LSID="urn:lsid:plazi:treatment:03CFA937556C1100FF1DE7F9FEA4FE48:8EB94EFA556C111DFC75E531FB3DF8AA" box="[1023,1153,1874,1899]" country="Peru" county="Region Junun" municipality="Provincia Chanchamayo" name="Rio Perene" pageId="8" pageNumber="539" stateProvince="Loreto">Rio Perene</location>
downstream
<location id="8EB94EFA556C111DFAADE531FA24F8AB" LSID="urn:lsid:plazi:treatment:03CFA937556C1100FF1DE7F9FEA4FE48:8EB94EFA556C111DFAADE531FA24F8AB" box="[1319,1432,1874,1899]" country="Peru" county="Region Junun" municipality="Provincia Chanchamayo" name="Pichanaki" pageId="8" pageNumber="539" stateProvince="Loreto">Pichanaki</location>
,
<collectingDate id="EF9CC709556C111DFF1DE515FF4AF84E" box="[151,246,1910,1934]" pageId="8" pageNumber="539" value="2006-01-07">
<date id="FFD83EE1556C111DFF1DE515FF4AF84E" box="[151,246,1910,1934]" pageId="8" pageNumber="539" value="2006-01-07">7.I.2006</date>
</collectingDate>
, coll.
<collectorName id="26937DF7556C111DFEB4E514FE17F84E" box="[318,427,1910,1934]" pageId="8" pageNumber="539">N. Kluge</collectorName>
: 1 L-S-I
<specimenCount id="9D60D3A8556C111DFD9EE514FD9EF84F" box="[532,546,1911,1935]" pageId="8" pageNumber="539" type="female">♀</specimenCount>
;
<collectingRegion id="49A2D6C3556C111DFDBAE515FD6BF84E" box="[560,727,1910,1934]" country="Peru" name="Loreto" pageId="8" pageNumber="539">Region Loreto</collectingRegion>
, Provincia
<collectingRegion id="49A2D6C3556C111DFCD2E515FC0EF84E" box="[856,946,1910,1934]" country="Peru" name="Ucayali" pageId="8" pageNumber="539">Ucayali</collectingRegion>
, Pampa Hermosa,
<collectingDate id="EF9CC709556C111DFB03E514FA85F84F" box="[1161,1337,1911,1935]" pageId="8" pageNumber="539" value="2013-08-09" valueMax="2013-08-21" valueMin="2013-08-09">
<date id="FFD83EE1556C111DFB03E514FA85F84F" box="[1161,1337,1911,1935]" pageId="8" pageNumber="539" value="2013-08-09" valueMax="2013-08-21" valueMin="2013-08-09">9–21.VIII.2013</date>
</collectingDate>
, coll.
<collectorName id="26937DF7556C111DFA0BE514FF60F873" pageId="8" pageNumber="539">N. Kluge</collectorName>
&amp;
<collectorName id="26937DF7556C111DFF75E5F8FECAF873" box="[255,374,1946,1971]" pageId="8" pageNumber="539">L. Sheyko</collectorName>
: exuviae of
<specimenCount id="9D60D3A8556C111DFE77E5F8FDE2F873" box="[509,606,1946,1971]" pageId="8" pageNumber="539" type="female">1 female</specimenCount>
larva.
</materialsCitation>
</paragraph>
<paragraph id="8BD91821556C111DFF4DE5DDFA20F816" blockId="8.[151,1437,1730,2007]" box="[199,1436,1982,2007]" pageId="8" pageNumber="539">
<emphasis id="B912C433556C111DFF4DE5DDFE0AF817" bold="true" box="[199,438,1982,2007]" pageId="8" pageNumber="539">
Descriptions.
<emphasis id="B912C433556C111DFEE1E5DCFE13F816" bold="true" box="[363,431,1983,2006]" italics="true" pageId="8" pageNumber="539">Larva</emphasis>
.
</emphasis>
CUTICULAR COLORATION. Head, pronotum, mesonotum and abdominal terga I–IX
</paragraph>
<paragraph id="8BD91821556D111CFF1DE2FBFE7FFDA0" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="540">
at most part brown, with or without variable unpaired contrasting colorless blanks (
<figureCitation id="135D04A4556D111CFBBFE2F4FBC7FF70" box="[1077,1147,151,176]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,2010,2033]" captionTargetBox="[157,1428,650,1986]" captionTargetId="figure@2.[157,1429,628,1986]" captionTargetPageId="2" captionText="FIGURES 1–3. Female larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri="https://zenodo.org/record/2588963/files/figure.png" pageId="9" pageNumber="540">Figs 1</figureCitation>
,
<figureCitation id="135D04A4556D111CFB03E2FBFB6DFF70" box="[1161,1233,152,176]" captionStart="FIGURES 41–43" captionStartId="13.[151,264,1877,1900]" captionTargetBox="[156,1431,355,1848]" captionTargetId="figure@13.[156,1431,337,1856]" captionTargetPageId="13" captionText="FIGURES 41–43. Larval thorax of Guajirolus ektrapeloglossa from Quebrada Barrigón. 41, exuviae of left half of pronotum and mesonotum; 42, posterior margin of mesonotum and protoptera; 43, exuviae of sterna and pleura (fore leg turned anteriorly, middle and hind legs detached). Abbreviations: abd1, first abdominal tergum; msth, mesothorax; mtth, metathorax; prth, prothorax." httpUri="https://zenodo.org/record/2588977/files/figure.png" pageId="9" pageNumber="540">41–42</figureCitation>
,
<figureCitation id="135D04A4556D111CFB56E2F4FA9AFF70" box="[1244,1318,151,176]" captionStart="FIGURES 51–58" captionStartId="15.[151,264,2007,2030]" captionTargetBox="[156,1430,243,1914]" captionTargetId="figure@15.[156,1430,243,1914]" captionTargetPageId="15" captionText="FIGURES 51–58. Larval coloration of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri="https://zenodo.org/record/2588981/files/figure.png" pageId="9" pageNumber="540">51–58</figureCitation>
). Thoracic pleura and sterna with brown sclerites and colorless membranes (
<figureCitation id="135D04A4556D111CFC04E2DFFC59FF14" box="[910,997,188,213]" captionStart="FIGURES 41–43" captionStartId="13.[151,264,1877,1900]" captionTargetBox="[156,1431,355,1848]" captionTargetId="figure@13.[156,1431,337,1856]" captionTargetPageId="13" captionText="FIGURES 41–43. Larval thorax of Guajirolus ektrapeloglossa from Quebrada Barrigón. 41, exuviae of left half of pronotum and mesonotum; 42, posterior margin of mesonotum and protoptera; 43, exuviae of sterna and pleura (fore leg turned anteriorly, middle and hind legs detached). Abbreviations: abd1, first abdominal tergum; msth, mesothorax; mtth, metathorax; prth, prothorax." httpUri="https://zenodo.org/record/2588977/files/figure.png" pageId="9" pageNumber="540">Fig. 43</figureCitation>
). Legs with colorless background and following brown markings: femur with two more or less expressed brown bands, among which distal band darker than proximal; tibia with more or less expressed brown band near base; tarsus with more or less expressed brown band at base (
<figureCitation id="135D04A4556D111CFECFE344FE6CFE80" box="[325,464,295,320]" captionStart="FIGURES 44–50" captionStartId="14.[151,264,1937,1960]" captionTargetBox="[156,1430,258,1857]" captionTargetId="figure@14.[156,1430,258,1857]" captionTargetPageId="14" captionText="FIGURES 44–50. Larval legs of Guajirolus ektrapeloglossa from Quebrada Barrigón. 44–45, fore and middle legs of darkest specimen; 46–47, claws; 48, fragment of fore femur (focus on anterior side); 49, fragment of fore tibia (focus on anterior side); 50, fragment of hind tibia (focus on posterior side)." httpUri="https://zenodo.org/record/2588979/files/figure.png" pageId="9" pageNumber="540">Figs 44–45</figureCitation>
); sometimes brown markings better expressed on fore legs, worse on hind legs. Abdominal terga I–IX with or without unpaired blanks, which either form integral longitudinal median line, or separated one from another; sometimes blanks on terga IV and/or VII wide, transverse and occupy most part of tergum (
<figureCitation id="135D04A4556D111CFF72E3F7FEF3FE6D" box="[248,335,404,429]" captionStart="FIGURES 51–58" captionStartId="15.[151,264,2007,2030]" captionTargetBox="[156,1430,243,1914]" captionTargetId="figure@15.[156,1430,243,1914]" captionTargetPageId="15" captionText="FIGURES 51–58. Larval coloration of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri="https://zenodo.org/record/2588981/files/figure.png" pageId="9" pageNumber="540">Fig. 58</figureCitation>
); sometimes blanks absent and terga I–IX entirely dark brown. Abdominal sterna I–IX brown, often lighter than terga, unicolor or with diffusive color pattern. Last abdominal segment (tergum X and paraprocts) light, often entirely colorless. Tergalii more or less colored with brown, lighter at periphery (
<figureCitation id="135D04A4556D111CFB14E3BFFB4EFE35" box="[1182,1266,476,501]" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="9" pageNumber="540">Fig. 67</figureCitation>
). Caudalii with brown and colorless portions at following succession: proximally light brown—gradually turn to dark brown—contrastingly changed to colorless—contrastingly changed to dark brown—gradually become lighter toward tip (
<figureCitation id="135D04A4556D111CFE92E024FEE3FDA0" box="[280,351,583,608]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,2010,2033]" captionTargetBox="[157,1428,650,1986]" captionTargetId="figure@2.[157,1429,628,1986]" captionTargetPageId="2" captionText="FIGURES 1–3. Female larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri="https://zenodo.org/record/2588963/files/figure.png" pageId="9" pageNumber="540">Figs 1</figureCitation>
,
<figureCitation id="135D04A4556D111CFEE7E024FE37FDA0" box="[365,395,583,608]" captionStart="FIGURES 51–58" captionStartId="15.[151,264,2007,2030]" captionTargetBox="[156,1430,243,1914]" captionTargetId="figure@15.[156,1430,243,1914]" captionTargetPageId="15" captionText="FIGURES 51–58. Larval coloration of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri="https://zenodo.org/record/2588981/files/figure.png" pageId="9" pageNumber="540">52</figureCitation>
,
<figureCitation id="135D04A4556D111CFE1DE024FE0BFDA0" box="[407,439,583,608]" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="9" pageNumber="540">59</figureCitation>
).
</paragraph>
<paragraph id="8BD91821556D111CFF4DE00FFD44FD45" blockId="9.[151,1437,151,2013]" box="[199,760,620,645]" pageId="9" pageNumber="540">HYPODERMAL COLORATION. Not expressed.</paragraph>
</subSubSection>
<subSubSection id="C37C4BAA556D1119FF4DE0F3FD56FE48" lastPageId="12" lastPageNumber="543" pageId="9" pageNumber="540" type="description">
<paragraph id="8BD91821556D111CFF4DE0F3FEADFD30" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="540">
STRUCTURE. Head and thorax somewhat compressed laterally and arched dorsally (
<figureCitation id="135D04A4556D111CFB0CE0ECFB50FD68" box="[1158,1260,655,680]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,2010,2033]" captionTargetBox="[157,1428,650,1986]" captionTargetId="figure@2.[157,1429,628,1986]" captionTargetPageId="2" captionText="FIGURES 1–3. Female larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri="https://zenodo.org/record/2588963/files/figure.png" pageId="9" pageNumber="540">Figs 1–3</figureCitation>
). Frontal suture sinuate (
<figureCitation id="135D04A4556D111CFF73E0D7FEFDFD0C" box="[249,321,692,717]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,2010,2033]" captionTargetBox="[157,1428,650,1986]" captionTargetId="figure@2.[157,1429,628,1986]" captionTargetPageId="2" captionText="FIGURES 1–3. Female larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri="https://zenodo.org/record/2588963/files/figure.png" pageId="9" pageNumber="540">Fig. 2</figureCitation>
); antennae bases brought together, frons between them keel-shaped and elevated above clypeus (
<figureCitation id="135D04A4556D111CFF15E0B4FEB8FD30" box="[159,260,727,752]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,2010,2033]" captionTargetBox="[157,1428,650,1986]" captionTargetId="figure@2.[157,1429,628,1986]" captionTargetPageId="2" captionText="FIGURES 1–3. Female larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri="https://zenodo.org/record/2588963/files/figure.png" pageId="9" pageNumber="540">Figs 1–3</figureCitation>
).
</paragraph>
<paragraph id="8BD91821556D111CFF4DE09FFAA2FC9C" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="540">
Labrum as characterized above [
<taxonomicName id="4C6663A2556D111CFDCAE09FFD02FCD5" authorityName="Flowers" authorityYear="1985" box="[576,702,764,789]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="553" phylum="Arthropoda" rank="genus">Guajirolus</taxonomicName>
/g1 (1)]; median emargination with 6 (i.e. 3 pairs) minute stout setae exposed on outer side (
<figureCitation id="135D04A4556D111CFE6EE17CFD85FCF8" box="[484,569,799,824]" captionStart="FIGURES 17–21" captionStartId="6.[151,264,1909,1932]" captionTargetBox="[171,1428,199,1815]" captionTargetId="figure@6.[157,1429,193,1889]" captionTargetPageId="6" captionText="FIGURES 17–21. Guajirolus ektrapeloglossa from Quebrada Barrigón. 17, maxilla (dorsal view); 18, apex of maxilla (ventral view); 19, distal margin of labrum (dorsal view); 20–21, biting denticles of left and right mandibles. Abbreviations: 1d, distal subsegment of first palpomere; 1p, proximal subsegment of first palpomere; 2, second palpomere; ds1, ds2, ds3, dentisetae; in, incisor; kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2588969/files/figure.png" pageId="9" pageNumber="540">Fig. 19</figureCitation>
); among setae-like projections, located on inner side and directed medially, projections of distal group in right half more stout and blunt than others and form regular row (
<figureCitation id="135D04A4556D111CFB35E127FAADFC9C" box="[1215,1297,836,861]" captionStart="FIGURES 4–11" captionStartId="3.[151,264,1708,1731]" captionTargetBox="[184,1405,451,1693]" captionTargetId="figure@3.[184,1405,440,1693]" captionTargetPageId="3" captionText="FIGURES 4–11. Labrum of last instar larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 4, labrum; 5, exuviae of labrum; 6, longitudinal section of lateral part; 7–8, one and the same median emargination differently compressed; 9–11, fragments of exuviae of one labrum: 9, with focus on distal margin; 10, with focus on dorsal side; 11, detached ventral cuticle (right side at right)." httpUri="https://zenodo.org/record/2588965/files/figure.png" pageId="9" pageNumber="540">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8BD91821556D111CFF4DE104FEFDFBD0" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="540">
Mandibles as characterized above [
<taxonomicName id="4C6663A2556D111CFDE1E104FD55FC40" authorityName="Flowers" authorityYear="1985" box="[619,745,871,896]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="553" phylum="Arthropoda" rank="genus">Guajirolus</taxonomicName>
/g1 (2), (3)], stout and convex; lateral surface pigmented, covered with minute protuberances and sparse fine setae (
<figureCitation id="135D04A4556D111CFCA7E1EFFC3EFC65" box="[813,898,908,933]" captionStart="FIGURES 12–16" captionStartId="5.[151,264,1907,1930]" captionTargetBox="[157,1430,193,1886]" captionTargetId="figure@5.[157,1430,193,1886]" captionTargetPageId="5" captionText="FIGURES 12–16. Mandibles of Guajirolus ektrapeloglossa from Quebrada Barrigón. 12–13 right and left mandibles inside old cuticle before molt from penultimate to last larval instar; 14–15, biting edges of right and left mandibles; 16, lateral part of left mandible." httpUri="https://zenodo.org/record/2588967/files/figure.png" pageId="9" pageNumber="540">Fig. 16</figureCitation>
); on both mandibles incisor reduced to 2 small denticles; kinetodontium (fused with incisor) on left mandible represented by 3 large and several small denticles, on right mandible by 4 large denticles; on both mandibles prostheca large and sharply widened apically (
<figureCitation id="135D04A4556D111CFAE0E1B7FF63FBD0" captionStart="FIGURES 12–16" captionStartId="5.[151,264,1907,1930]" captionTargetBox="[157,1430,193,1886]" captionTargetId="figure@5.[157,1430,193,1886]" captionTargetPageId="5" captionText="FIGURES 12–16. Mandibles of Guajirolus ektrapeloglossa from Quebrada Barrigón. 12–13 right and left mandibles inside old cuticle before molt from penultimate to last larval instar; 14–15, biting edges of right and left mandibles; 16, lateral part of left mandible." httpUri="https://zenodo.org/record/2588967/files/figure.png" pageId="9" pageNumber="540">Figs 12–13</figureCitation>
,
<figureCitation id="135D04A4556D111CFF61E19BFE89FBD0" box="[235,309,1016,1040]" captionStart="FIGURES 17–21" captionStartId="6.[151,264,1909,1932]" captionTargetBox="[171,1428,199,1815]" captionTargetId="figure@6.[157,1429,193,1889]" captionTargetPageId="6" captionText="FIGURES 17–21. Guajirolus ektrapeloglossa from Quebrada Barrigón. 17, maxilla (dorsal view); 18, apex of maxilla (ventral view); 19, distal margin of labrum (dorsal view); 20–21, biting denticles of left and right mandibles. Abbreviations: 1d, distal subsegment of first palpomere; 1p, proximal subsegment of first palpomere; 2, second palpomere; ds1, ds2, ds3, dentisetae; in, incisor; kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2588969/files/figure.png" pageId="9" pageNumber="540">20–21</figureCitation>
).
</paragraph>
<paragraph id="8BD91821556D111CFF4DE67FFE0CFB60" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="540">
Maxilla as characterized above [
<taxonomicName id="4C6663A2556D111CFDB8E67FFD0CFBF5" authorityName="Flowers" authorityYear="1985" box="[562,688,1052,1077]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="553" phylum="Arthropoda" rank="genus">Guajirolus</taxonomicName>
/g1 (5)]; galea-lacinia narrowed toward apex, proximally on lateral side with protuberance located more dorsally than base of palp (
<figureCitation id="135D04A4556D111CFCF6E65CFC69FB98" box="[892,981,1087,1112]" captionStart="FIGURES 17–21" captionStartId="6.[151,264,1909,1932]" captionTargetBox="[171,1428,199,1815]" captionTargetId="figure@6.[157,1429,193,1889]" captionTargetPageId="6" captionText="FIGURES 17–21. Guajirolus ektrapeloglossa from Quebrada Barrigón. 17, maxilla (dorsal view); 18, apex of maxilla (ventral view); 19, distal margin of labrum (dorsal view); 20–21, biting denticles of left and right mandibles. Abbreviations: 1d, distal subsegment of first palpomere; 1p, proximal subsegment of first palpomere; 2, second palpomere; ds1, ds2, ds3, dentisetae; in, incisor; kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2588969/files/figure.png" pageId="9" pageNumber="540">Figs 17</figureCitation>
,
<figureCitation id="135D04A4556D111CFC69E623FB92FB98" box="[995,1070,1088,1112]" captionStart="FIGURES 22–28" captionStartId="7.[151,264,1417,1440]" captionTargetBox="[189,1417,267,1396]" captionTargetId="figure@7.[169,1417,267,1396]" captionTargetPageId="7" captionText="FIGURES 22–28. Guajirolus ektrapeloglossa from Quebrada Barrigón. 22–23, exuviae of maxillae; 24, maxilla; 25, exuviae of hypopharynx; 26–28, hypopharynx." httpUri="https://zenodo.org/record/2588971/files/figure.png" pageId="9" pageNumber="540">22–24</figureCitation>
); inner-dorsal row of setae with distal end turned laterally (
<figureCitation id="135D04A4556D111CFE4CE607FDA7FBBC" box="[454,539,1124,1149]" captionStart="FIGURES 17–21" captionStartId="6.[151,264,1909,1932]" captionTargetBox="[171,1428,199,1815]" captionTargetId="figure@6.[157,1429,193,1889]" captionTargetPageId="6" captionText="FIGURES 17–21. Guajirolus ektrapeloglossa from Quebrada Barrigón. 17, maxilla (dorsal view); 18, apex of maxilla (ventral view); 19, distal margin of labrum (dorsal view); 20–21, biting denticles of left and right mandibles. Abbreviations: 1d, distal subsegment of first palpomere; 1p, proximal subsegment of first palpomere; 2, second palpomere; ds1, ds2, ds3, dentisetae; in, incisor; kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2588969/files/figure.png" pageId="9" pageNumber="540">Fig. 18</figureCitation>
); apical bunch of long setae consists of 8 setae arranged in 2 rows and one seta laterad of them (
<figureCitation id="135D04A4556D111CFEDAE6E4FE1FFB60" box="[336,419,1159,1184]" captionStart="FIGURES 17–21" captionStartId="6.[151,264,1909,1932]" captionTargetBox="[171,1428,199,1815]" captionTargetId="figure@6.[157,1429,193,1889]" captionTargetPageId="6" captionText="FIGURES 17–21. Guajirolus ektrapeloglossa from Quebrada Barrigón. 17, maxilla (dorsal view); 18, apex of maxilla (ventral view); 19, distal margin of labrum (dorsal view); 20–21, biting denticles of left and right mandibles. Abbreviations: 1d, distal subsegment of first palpomere; 1p, proximal subsegment of first palpomere; 2, second palpomere; ds1, ds2, ds3, dentisetae; in, incisor; kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2588969/files/figure.png" pageId="9" pageNumber="540">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="8BD91821556D111CFF4DE6CFFB51FB28" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="540">
Maxillary palp as characterized above [
<taxonomicName id="4C6663A2556D111CFD16E6CFFCA6FB05" authorityName="Flowers" authorityYear="1985" box="[668,794,1196,1221]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="553" phylum="Arthropoda" rank="genus">Guajirolus</taxonomicName>
/g1 (6)]; both subsegments of 1st palpomere long and movably articulated one with another; 2nd palpomere with proximal part swollen (
<figureCitation id="135D04A4556D111CFBBEE6ACFB37FB28" box="[1076,1163,1231,1256]" captionStart="FIGURES 17–21" captionStartId="6.[151,264,1909,1932]" captionTargetBox="[171,1428,199,1815]" captionTargetId="figure@6.[157,1429,193,1889]" captionTargetPageId="6" captionText="FIGURES 17–21. Guajirolus ektrapeloglossa from Quebrada Barrigón. 17, maxilla (dorsal view); 18, apex of maxilla (ventral view); 19, distal margin of labrum (dorsal view); 20–21, biting denticles of left and right mandibles. Abbreviations: 1d, distal subsegment of first palpomere; 1p, proximal subsegment of first palpomere; 2, second palpomere; ds1, ds2, ds3, dentisetae; in, incisor; kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2588969/files/figure.png" pageId="9" pageNumber="540">Figs 17</figureCitation>
,
<figureCitation id="135D04A4556D111CFB1CE6B3FB5EFB28" box="[1174,1250,1232,1256]" captionStart="FIGURES 22–28" captionStartId="7.[151,264,1417,1440]" captionTargetBox="[189,1417,267,1396]" captionTargetId="figure@7.[169,1417,267,1396]" captionTargetPageId="7" captionText="FIGURES 22–28. Guajirolus ektrapeloglossa from Quebrada Barrigón. 22–23, exuviae of maxillae; 24, maxilla; 25, exuviae of hypopharynx; 26–28, hypopharynx." httpUri="https://zenodo.org/record/2588971/files/figure.png" pageId="9" pageNumber="540">22–24</figureCitation>
).
</paragraph>
<paragraph id="8BD91821556D111CFF4DE697FC73F9C8" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="540">
Labium with glossae and paraglossae equally wide, paraglossae slightly longer than glossae; base of glossa very slightly overlap base of paraglossa in ventral view; base of paraglossa somewhat projected laterally (
<figureCitation id="135D04A4556D111CFACAE774FA2BFAF0" box="[1344,1431,1303,1328]" captionStart="FIGURES 29–35" captionStartId="8.[151,264,1275,1298]" captionTargetBox="[158,1428,195,1254]" captionTargetId="figure@8.[158,1428,195,1254]" captionTargetPageId="8" captionText="FIGURES 29–35. Labium of Guajirolus ektrapeloglossa from Quebrada Barrigón. 29–31, labium (29, ventral view; 30–31, in translucent light, with focus on glossae and on palps); 32–33, one and the same labial palp differently compressed; 34–35, left and right labial palps of one and the same individual." httpUri="https://zenodo.org/record/2588973/files/figure.png" pageId="9" pageNumber="540">Figs 30</figureCitation>
,
<figureCitation id="135D04A4556D111CFF1DE75FFF0BFA94" box="[151,183,1340,1364]" captionStart="FIGURES 36–40" captionStartId="10.[151,264,1934,1957]" captionTargetBox="[157,1429,326,1896]" captionTargetId="figure@10.[157,1429,326,1896]" captionTargetPageId="10" captionText="FIGURES 36–40. Setation of labium of Guajirolus ektrapeloglossa from Quebrada Barrigón. 36, exuviae of glossae and paraglossa (focus on ventral side); 37, glossae (focus on dorsal side); 38–40, exuviae of palp and paraglossa (38, 40, focus on ventral side; 39, focus on dorsal side)." httpUri="https://zenodo.org/record/2588975/files/figure.png" pageId="9" pageNumber="540">37</figureCitation>
). Ventral side of glossa with numerous irregularly situated long hair-like setae (
<figureCitation id="135D04A4556D111CFBC9E75FFB24FA95" box="[1091,1176,1340,1365]" captionStart="FIGURES 36–40" captionStartId="10.[151,264,1934,1957]" captionTargetBox="[157,1429,326,1896]" captionTargetId="figure@10.[157,1429,326,1896]" captionTargetPageId="10" captionText="FIGURES 36–40. Setation of labium of Guajirolus ektrapeloglossa from Quebrada Barrigón. 36, exuviae of glossae and paraglossa (focus on ventral side); 37, glossae (focus on dorsal side); 38–40, exuviae of palp and paraglossa (38, 40, focus on ventral side; 39, focus on dorsal side)." httpUri="https://zenodo.org/record/2588975/files/figure.png" pageId="9" pageNumber="540">Fig. 36</figureCitation>
). Dorsal side of glossa with especially stout long setae forming regular transverse row in distal part of glossa and shorter irregular transverse row more distally; middle part of dorsal side with small group of irregularly situated long hair-like setae (
<figureCitation id="135D04A4556D111CFF15E7C4FF48FA00" box="[159,244,1447,1472]" captionStart="FIGURES 36–40" captionStartId="10.[151,264,1934,1957]" captionTargetBox="[157,1429,326,1896]" captionTargetId="figure@10.[157,1429,326,1896]" captionTargetPageId="10" captionText="FIGURES 36–40. Setation of labium of Guajirolus ektrapeloglossa from Quebrada Barrigón. 36, exuviae of glossae and paraglossa (focus on ventral side); 37, glossae (focus on dorsal side); 38–40, exuviae of palp and paraglossa (38, 40, focus on ventral side; 39, focus on dorsal side)." httpUri="https://zenodo.org/record/2588975/files/figure.png" pageId="9" pageNumber="540">Fig. 37</figureCitation>
). Row of long setae along median side of glossa (usual for
<taxonomicName id="4C6663A2556D111CFC1EE7C4FC44FA00" baseAuthorityName="Kluge" baseAuthorityYear="2004" box="[916,1016,1447,1472]" class="Insecta" family="Baetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="540" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
) either regular (
<figureCitation id="135D04A4556D111CFB24E7C4FABEFA00" box="[1198,1282,1447,1472]" captionStart="FIGURES 36–40" captionStartId="10.[151,264,1934,1957]" captionTargetBox="[157,1429,326,1896]" captionTargetId="figure@10.[157,1429,326,1896]" captionTargetPageId="10" captionText="FIGURES 36–40. Setation of labium of Guajirolus ektrapeloglossa from Quebrada Barrigón. 36, exuviae of glossae and paraglossa (focus on ventral side); 37, glossae (focus on dorsal side); 38–40, exuviae of palp and paraglossa (38, 40, focus on ventral side; 39, focus on dorsal side)." httpUri="https://zenodo.org/record/2588975/files/figure.png" pageId="9" pageNumber="540">Fig. 37</figureCitation>
), or irregular. Paraglossa without setal rows on median and lateral sides; distal margin with numerous long setae (
<figureCitation id="135D04A4556D111CFA85E7AFFADFFA24" box="[1295,1379,1484,1509]" captionStart="FIGURES 36–40" captionStartId="10.[151,264,1934,1957]" captionTargetBox="[157,1429,326,1896]" captionTargetId="figure@10.[157,1429,326,1896]" captionTargetPageId="10" captionText="FIGURES 36–40. Setation of labium of Guajirolus ektrapeloglossa from Quebrada Barrigón. 36, exuviae of glossae and paraglossa (focus on ventral side); 37, glossae (focus on dorsal side); 38–40, exuviae of palp and paraglossa (38, 40, focus on ventral side; 39, focus on dorsal side)." httpUri="https://zenodo.org/record/2588975/files/figure.png" pageId="9" pageNumber="540">Fig. 40</figureCitation>
) and with 2 somewhat thicker long setae on dorsal side close to apex (
<figureCitation id="135D04A4556D111CFCE4E78CFC7EF9C8" box="[878,962,1519,1544]" captionStart="FIGURES 36–40" captionStartId="10.[151,264,1934,1957]" captionTargetBox="[157,1429,326,1896]" captionTargetId="figure@10.[157,1429,326,1896]" captionTargetPageId="10" captionText="FIGURES 36–40. Setation of labium of Guajirolus ektrapeloglossa from Quebrada Barrigón. 36, exuviae of glossae and paraglossa (focus on ventral side); 37, glossae (focus on dorsal side); 38–40, exuviae of palp and paraglossa (38, 40, focus on ventral side; 39, focus on dorsal side)." httpUri="https://zenodo.org/record/2588975/files/figure.png" pageId="9" pageNumber="540">Fig. 39</figureCitation>
).
</paragraph>
<paragraph id="8BD91821556D111CFF4DE477FC5FF9B4" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="540">
Labial palp as characterized above [
<taxonomicName id="4C6663A2556D111CFDDDE477FD69F9ED" authorityName="Flowers" authorityYear="1985" box="[599,725,1556,1581]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="553" phylum="Arthropoda" rank="genus">Guajirolus</taxonomicName>
/g1 (7)]; projection of its 2nd segment very thick and soft, sacklike. 2nd and 3rd segments with irregularly situated stout pointed setae (
<figureCitation id="135D04A4556D111CFC4DE454FBA0F990" box="[967,1052,1591,1616]" captionStart="FIGURES 36–40" captionStartId="10.[151,264,1934,1957]" captionTargetBox="[157,1429,326,1896]" captionTargetId="figure@10.[157,1429,326,1896]" captionTargetPageId="10" captionText="FIGURES 36–40. Setation of labium of Guajirolus ektrapeloglossa from Quebrada Barrigón. 36, exuviae of glossae and paraglossa (focus on ventral side); 37, glossae (focus on dorsal side); 38–40, exuviae of palp and paraglossa (38, 40, focus on ventral side; 39, focus on dorsal side)." httpUri="https://zenodo.org/record/2588975/files/figure.png" pageId="9" pageNumber="540">Fig. 39</figureCitation>
); ventral side of projection of 2nd segment and of apex of 3rd segment with dense minute fine setae (
<figureCitation id="135D04A4556D111CFC08E43FFC6AF9B4" box="[898,982,1628,1653]" captionStart="FIGURES 36–40" captionStartId="10.[151,264,1934,1957]" captionTargetBox="[157,1429,326,1896]" captionTargetId="figure@10.[157,1429,326,1896]" captionTargetPageId="10" captionText="FIGURES 36–40. Setation of labium of Guajirolus ektrapeloglossa from Quebrada Barrigón. 36, exuviae of glossae and paraglossa (focus on ventral side); 37, glossae (focus on dorsal side); 38–40, exuviae of palp and paraglossa (38, 40, focus on ventral side; 39, focus on dorsal side)." httpUri="https://zenodo.org/record/2588975/files/figure.png" pageId="9" pageNumber="540">Fig. 38</figureCitation>
).
</paragraph>
<paragraph id="8BD91821556D111CFF4DE4E2FBB3F88D" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="540">
Mesonotum with anterolateral corners projected anteriorly (
<figureCitation id="135D04A4556D111CFCEDE41CFC07F958" box="[871,955,1663,1688]" captionStart="FIGURES 41–43" captionStartId="13.[151,264,1877,1900]" captionTargetBox="[156,1431,355,1848]" captionTargetId="figure@13.[156,1431,337,1856]" captionTargetPageId="13" captionText="FIGURES 41–43. Larval thorax of Guajirolus ektrapeloglossa from Quebrada Barrigón. 41, exuviae of left half of pronotum and mesonotum; 42, posterior margin of mesonotum and protoptera; 43, exuviae of sterna and pleura (fore leg turned anteriorly, middle and hind legs detached). Abbreviations: abd1, first abdominal tergum; msth, mesothorax; mtth, metathorax; prth, prothorax." httpUri="https://zenodo.org/record/2588977/files/figure.png" pageId="9" pageNumber="540">Fig. 41</figureCitation>
). Fore protoptera brought together, but not fused with posterior margin of mesonotum (
<figureCitation id="135D04A4556D111CFD2DE4C7FD42F97C" box="[679,766,1700,1725]" captionStart="FIGURES 41–43" captionStartId="13.[151,264,1877,1900]" captionTargetBox="[156,1431,355,1848]" captionTargetId="figure@13.[156,1431,337,1856]" captionTargetPageId="13" captionText="FIGURES 41–43. Larval thorax of Guajirolus ektrapeloglossa from Quebrada Barrigón. 41, exuviae of left half of pronotum and mesonotum; 42, posterior margin of mesonotum and protoptera; 43, exuviae of sterna and pleura (fore leg turned anteriorly, middle and hind legs detached). Abbreviations: abd1, first abdominal tergum; msth, mesothorax; mtth, metathorax; prth, prothorax." httpUri="https://zenodo.org/record/2588977/files/figure.png" pageId="9" pageNumber="540">Fig. 42</figureCitation>
) (in contrast to
<taxonomicName id="4C6663A2556D111CFC4DE4C7FBC9F97C" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[967,1141,1700,1724]" class="Insecta" family="Baetidae" genus="Pseudopannota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="B912C433556D111CFC4DE4C7FBC9F97C" box="[967,1141,1700,1724]" italics="true" pageId="9" pageNumber="540">Pseudopannota</emphasis>
</taxonomicName>
). Lateral margin of fore protopteron with small incision corresponding to costal brace (
<figureCitation id="135D04A4556D111CFCF5E4A4FC64F920" box="[895,984,1735,1760]" captionStart="FIGURES 41–43" captionStartId="13.[151,264,1877,1900]" captionTargetBox="[156,1431,355,1848]" captionTargetId="figure@13.[156,1431,337,1856]" captionTargetPageId="13" captionText="FIGURES 41–43. Larval thorax of Guajirolus ektrapeloglossa from Quebrada Barrigón. 41, exuviae of left half of pronotum and mesonotum; 42, posterior margin of mesonotum and protoptera; 43, exuviae of sterna and pleura (fore leg turned anteriorly, middle and hind legs detached). Abbreviations: abd1, first abdominal tergum; msth, mesothorax; mtth, metathorax; prth, prothorax." httpUri="https://zenodo.org/record/2588977/files/figure.png" pageId="9" pageNumber="540">Fig. 41</figureCitation>
). Metanotum without vestiges of hind protoptera (
<figureCitation id="135D04A4556D111CFEA8E48FFEC3F8C4" box="[290,383,1772,1797]" captionStart="FIGURES 41–43" captionStartId="13.[151,264,1877,1900]" captionTargetBox="[156,1431,355,1848]" captionTargetId="figure@13.[156,1431,337,1856]" captionTargetPageId="13" captionText="FIGURES 41–43. Larval thorax of Guajirolus ektrapeloglossa from Quebrada Barrigón. 41, exuviae of left half of pronotum and mesonotum; 42, posterior margin of mesonotum and protoptera; 43, exuviae of sterna and pleura (fore leg turned anteriorly, middle and hind legs detached). Abbreviations: abd1, first abdominal tergum; msth, mesothorax; mtth, metathorax; prth, prothorax." httpUri="https://zenodo.org/record/2588977/files/figure.png" pageId="9" pageNumber="540">Fig. 43</figureCitation>
). Thoracic sterna partly sclerotized and pigmented: prosternum with unpaired sclerite, mesosternum and metasternum each with pair of sclerites of peculiar shape (
<figureCitation id="135D04A4556D111CFB8AE56CFBEAF8E8" box="[1024,1110,1807,1832]" captionStart="FIGURES 41–43" captionStartId="13.[151,264,1877,1900]" captionTargetBox="[156,1431,355,1848]" captionTargetId="figure@13.[156,1431,337,1856]" captionTargetPageId="13" captionText="FIGURES 41–43. Larval thorax of Guajirolus ektrapeloglossa from Quebrada Barrigón. 41, exuviae of left half of pronotum and mesonotum; 42, posterior margin of mesonotum and protoptera; 43, exuviae of sterna and pleura (fore leg turned anteriorly, middle and hind legs detached). Abbreviations: abd1, first abdominal tergum; msth, mesothorax; mtth, metathorax; prth, prothorax." httpUri="https://zenodo.org/record/2588977/files/figure.png" pageId="9" pageNumber="540">Fig. 43</figureCitation>
). Sclerites of thoracic pleura and sterna partly with scales in operculate sockets (similar to that of abdomen).
</paragraph>
<paragraph id="8BD91821556D111FFF4DE53AFBB1FF38" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1436,151,248]" lastPageId="10" lastPageNumber="541" pageId="9" pageNumber="540">
Legs with cuticle thick and denticulate (
<figureCitation id="135D04A4556D111CFD07E534FCAEF8B0" box="[653,786,1879,1904]" captionStart="FIGURES 44–50" captionStartId="14.[151,264,1937,1960]" captionTargetBox="[156,1430,258,1857]" captionTargetId="figure@14.[156,1430,258,1857]" captionTargetPageId="14" captionText="FIGURES 44–50. Larval legs of Guajirolus ektrapeloglossa from Quebrada Barrigón. 44–45, fore and middle legs of darkest specimen; 46–47, claws; 48, fragment of fore femur (focus on anterior side); 49, fragment of fore tibia (focus on anterior side); 50, fragment of hind tibia (focus on posterior side)." httpUri="https://zenodo.org/record/2588979/files/figure.png" pageId="9" pageNumber="540">Figs 48–50</figureCitation>
). Femora as characterized above [
<taxonomicName id="4C6663A2556D111CFB1AE534FAB2F8B0" authorityName="Flowers" authorityYear="1985" box="[1168,1294,1879,1904]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="553" phylum="Arthropoda" rank="genus">Guajirolus</taxonomicName>
/g1 (8)]; fore femur, besides stout setae on outer side, with such setae also on anterior surface—in distal part and along outer side (
<figureCitation id="135D04A4556D111CFF15E5C3FE99F878" box="[159,293,1952,1977]" captionStart="FIGURES 44–50" captionStartId="14.[151,264,1937,1960]" captionTargetBox="[156,1430,258,1857]" captionTargetId="figure@14.[156,1430,258,1857]" captionTargetPageId="14" captionText="FIGURES 44–50. Larval legs of Guajirolus ektrapeloglossa from Quebrada Barrigón. 44–45, fore and middle legs of darkest specimen; 46–47, claws; 48, fragment of fore femur (focus on anterior side); 49, fragment of fore tibia (focus on anterior side); 50, fragment of hind tibia (focus on posterior side)." httpUri="https://zenodo.org/record/2588979/files/figure.png" pageId="9" pageNumber="540">Figs 44, 48</figureCitation>
). Tibiae of all legs with patella-tibial suture short, so that on inner side patella occupies only 1/3 of tibia (
<figureCitation id="135D04A4556D111CFF52E5A7FEE7F81D" box="[216,347,1988,2013]" captionStart="FIGURES 44–50" captionStartId="14.[151,264,1937,1960]" captionTargetBox="[156,1430,258,1857]" captionTargetId="figure@14.[156,1430,258,1857]" captionTargetPageId="14" captionText="FIGURES 44–50. Larval legs of Guajirolus ektrapeloglossa from Quebrada Barrigón. 44–45, fore and middle legs of darkest specimen; 46–47, claws; 48, fragment of fore femur (focus on anterior side); 49, fragment of fore tibia (focus on anterior side); 50, fragment of hind tibia (focus on posterior side)." httpUri="https://zenodo.org/record/2588979/files/figure.png" pageId="9" pageNumber="540">Figs 44–45</figureCitation>
). Tibia of fore leg with sparse small stout pointed setae on anterior side and few small stout setae on inner side (
<figureCitation id="135D04A4556E111FFEB1E2F4FE2CFF70" box="[315,400,151,176]" captionStart="FIGURES 44–50" captionStartId="14.[151,264,1937,1960]" captionTargetBox="[156,1430,258,1857]" captionTargetId="figure@14.[156,1430,258,1857]" captionTargetPageId="14" captionText="FIGURES 44–50. Larval legs of Guajirolus ektrapeloglossa from Quebrada Barrigón. 44–45, fore and middle legs of darkest specimen; 46–47, claws; 48, fragment of fore femur (focus on anterior side); 49, fragment of fore tibia (focus on anterior side); 50, fragment of hind tibia (focus on posterior side)." httpUri="https://zenodo.org/record/2588979/files/figure.png" pageId="10" pageNumber="541">Fig. 49</figureCitation>
;
<bibRefCitation id="EFF765D0556E111FFE17E2F4FDC1FF70" author="Thomas, A. &amp; Dominique, Y." box="[413,637,151,176]" pageId="10" pageNumber="541" pagination="21 - 29" refId="ref9271" refString="Thomas, A., Dominique, Y. &amp; Orth, K. (2005) Les Ephemeres de la Guyane Francaise. 12. Description de Guajirolus flowersi n. sp. [Ephemeroptera, Baetidae]. Ephemera, (2004), 6 (1), 21 - 29." type="journal article" year="2005">Thomas &amp; al. 2005</bibRefCitation>
: fig. 9). Tibiae of middle and hind legs without stout setae on anterior side, with such setae on posterior and inner sides (
<figureCitation id="135D04A4556E111FFD51E2DFFC8DFF15" box="[731,817,188,213]" captionStart="FIGURES 44–50" captionStartId="14.[151,264,1937,1960]" captionTargetBox="[156,1430,258,1857]" captionTargetId="figure@14.[156,1430,258,1857]" captionTargetPageId="14" captionText="FIGURES 44–50. Larval legs of Guajirolus ektrapeloglossa from Quebrada Barrigón. 44–45, fore and middle legs of darkest specimen; 46–47, claws; 48, fragment of fore femur (focus on anterior side); 49, fragment of fore tibia (focus on anterior side); 50, fragment of hind tibia (focus on posterior side)." httpUri="https://zenodo.org/record/2588979/files/figure.png" pageId="10" pageNumber="541">Fig. 50</figureCitation>
). Tarsus of all legs with row of stout pointed setae on inner side only. Claws as characterized above [
<taxonomicName id="4C6663A2556E111FFD2BE2BCFCA3FF38" authorityName="Flowers" authorityYear="1985" box="[673,799,223,248]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="553" phylum="Arthropoda" rank="genus">Guajirolus</taxonomicName>
/g1 (9)] (
<figureCitation id="135D04A4556E111FFCF4E2BCFBBDFF38" box="[894,1025,223,248]" captionStart="FIGURES 44–50" captionStartId="14.[151,264,1937,1960]" captionTargetBox="[156,1430,258,1857]" captionTargetId="figure@14.[156,1430,258,1857]" captionTargetPageId="14" captionText="FIGURES 44–50. Larval legs of Guajirolus ektrapeloglossa from Quebrada Barrigón. 44–45, fore and middle legs of darkest specimen; 46–47, claws; 48, fragment of fore femur (focus on anterior side); 49, fragment of fore tibia (focus on anterior side); 50, fragment of hind tibia (focus on posterior side)." httpUri="https://zenodo.org/record/2588979/files/figure.png" pageId="10" pageNumber="541">Figs 46–47</figureCitation>
).
</paragraph>
<caption id="DF1948A9556E111FFF1DE5EDFDA5F822" httpUri="https://zenodo.org/record/2588975/files/figure.png" pageId="10" pageNumber="541" startId="10.[151,264,1934,1957]" targetBox="[157,1429,326,1896]" targetPageId="10">
<paragraph id="8BD91821556E111FFF1DE5EDFDA5F822" blockId="10.[151,1436,1934,2018]" pageId="10" pageNumber="541">
<emphasis id="B912C433556E111FFF1DE5EDFEEBF864" bold="true" box="[151,343,1934,1957]" pageId="10" pageNumber="541">FIGURES 36–40.</emphasis>
Setation of labium of
<taxonomicName id="4C6663A2556E111FFDC7E5ECFCD8F865" authorityName="Flowers" authorityYear="1985" box="[589,868,1935,1957]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="541" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C433556E111FFDC7E5ECFCD8F865" box="[589,868,1935,1957]" italics="true" pageId="10" pageNumber="541">Guajirolus ektrapeloglossa</emphasis>
</taxonomicName>
from Quebrada Barrigón. 36, exuviae of glossae and paraglossa (focus on ventral side); 37, glossae (focus on dorsal side); 38–40, exuviae of palp and paraglossa (38, 40, focus on ventral side; 39, focus on dorsal side).
</paragraph>
</caption>
<paragraph id="8BD91821556F111EFF4DE2F4FD78FE48" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="542">
Abdominal terga and sterna with dense scales in operculate sockets (
<figureCitation id="135D04A4556F111EFC4DE2F4FBF6FF70" box="[967,1098,151,176]" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="11" pageNumber="542">Figs 60–61</figureCitation>
). Posterior margin of tergum I smooth, without denticles. Posterior margins of terga II–X with row of triangular denticles; denticles of more anterior terga shorter and blunt (
<figureCitation id="135D04A4556F111EFD88E2BCFDEAFF38" box="[514,598,223,248]" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="11" pageNumber="542">Fig. 62</figureCitation>
), denticles of more posterior terga longer and pointed (
<figureCitation id="135D04A4556F111EFB48E2BCFAA9FF38" box="[1218,1301,223,248]" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="11" pageNumber="542">Fig. 63</figureCitation>
); on middle of posterior margin of tergum IX (behind submedian pair of setae) denticles smaller. Posterior margins of sterna I–VII smooth, without denticles; sternum VIII with blunt denticles on median part of posterior margin (
<figureCitation id="135D04A4556F111EFAB3E344FA33FE80" box="[1337,1423,295,320]" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="11" pageNumber="542">Fig. 64</figureCitation>
); sternum IX with pointed denticles on posterior margin (
<figureCitation id="135D04A4556F111EFCAEE32FFCC2FEA5" box="[804,894,332,357]" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="11" pageNumber="542">Figs 65</figureCitation>
,
<figureCitation id="135D04A4556F111EFC04E32FFC12FEA5" box="[910,942,332,357]" captionStart="FIGURES 75–80" captionStartId="18.[151,264,1754,1777]" captionTargetBox="[158,1428,193,1733]" captionTargetId="figure@18.[158,1428,193,1733]" captionTargetPageId="18" captionText="FIGURES 75–80. Male genitals of Guajirolus ektrapeloglossa from Quebrada Barrigón. 75, genital of imago (ventral view; muscles and internal parts of penis shown by interrupted lines; right gonostylar and gonovectal muscles not shown); 76, subimaginal genitals developing under larval cuticle; 77–78 and 79–80, genitals of two specimens. Abbreviations: 1, 2, 3, segments of gonostylus; pgs, larval protogonostylus; usg, unistyliger." httpUri="https://zenodo.org/record/2588987/files/figure.png" pageId="11" pageNumber="542">76</figureCitation>
). Paraprocts with 10–20 pointed denticles, increasing in size toward caudal margin (
<figureCitation id="135D04A4556F111EFDE9E30CFD0BFE48" box="[611,695,367,392]" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="11" pageNumber="542">Fig. 69</figureCitation>
).
</paragraph>
<paragraph id="8BD91821556F111EFF4DE3F7FA86FE35" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="542">
Tergalii I absent (
<figureCitation id="135D04A4556F111EFE19E3F7FE56FE6C" box="[403,490,404,429]" captionStart="FIGURES 41–43" captionStartId="13.[151,264,1877,1900]" captionTargetBox="[156,1431,355,1848]" captionTargetId="figure@13.[156,1431,337,1856]" captionTargetPageId="13" captionText="FIGURES 41–43. Larval thorax of Guajirolus ektrapeloglossa from Quebrada Barrigón. 41, exuviae of left half of pronotum and mesonotum; 42, posterior margin of mesonotum and protoptera; 43, exuviae of sterna and pleura (fore leg turned anteriorly, middle and hind legs detached). Abbreviations: abd1, first abdominal tergum; msth, mesothorax; mtth, metathorax; prth, prothorax." httpUri="https://zenodo.org/record/2588977/files/figure.png" pageId="11" pageNumber="542">Fig. 43</figureCitation>
). Tergalii II–V subequal, tergalii VI–VII gradually diminished (
<figureCitation id="135D04A4556F111EFB43E3F7FAF2FE6D" box="[1225,1358,404,429]" captionStart="FIGURES 51–58" captionStartId="15.[151,264,2007,2030]" captionTargetBox="[156,1430,243,1914]" captionTargetId="figure@15.[156,1430,243,1914]" captionTargetPageId="15" captionText="FIGURES 51–58. Larval coloration of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri="https://zenodo.org/record/2588981/files/figure.png" pageId="11" pageNumber="542">Figs 51–58</figureCitation>
). Each tergalius oval; costal and anal ribs fused apically, so that entire margin armed by costa; in distal half margin with pointed seta-bearing denticles (
<figureCitation id="135D04A4556F111EFE7EE3BFFDCBFE35" box="[500,631,476,501]" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="11" pageNumber="542">Figs 67–68</figureCitation>
); dorsal surface with scales in operculate sockets (
<figureCitation id="135D04A4556F111EFB21E3BFFA90FE35" box="[1195,1324,476,501]" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="11" pageNumber="542">Figs 66, 68</figureCitation>
).
</paragraph>
<paragraph id="8BD91821556F111EFF4DE063FE44FD67" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="542">
Paracercus as thick as cerci and somewhat shorter (
<figureCitation id="135D04A4556F111EFCADE39CFCCFFDD8" box="[807,883,511,536]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,2010,2033]" captionTargetBox="[157,1428,650,1986]" captionTargetId="figure@2.[157,1429,628,1986]" captionTargetPageId="2" captionText="FIGURES 1–3. Female larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri="https://zenodo.org/record/2588963/files/figure.png" pageId="11" pageNumber="542">Figs 1</figureCitation>
,
<figureCitation id="135D04A4556F111EFC0FE39CFC19FDD8" box="[901,933,511,536]" captionStart="FIGURES 51–58" captionStartId="15.[151,264,2007,2030]" captionTargetBox="[156,1430,243,1914]" captionTargetId="figure@15.[156,1430,243,1914]" captionTargetPageId="15" captionText="FIGURES 51–58. Larval coloration of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri="https://zenodo.org/record/2588981/files/figure.png" pageId="11" pageNumber="542">52</figureCitation>
). Cerci and paracercus with dense primary swimming setae at most their length and with slender distal portion lacking swimming setae. In proximal and distal parts of cercus segments cylindrical, in middle part slightly oblique (
<figureCitation id="135D04A4556F111EFC29E024FC44FDA0" box="[931,1016,583,608]" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="11" pageNumber="542">Fig. 59</figureCitation>
); posterior margins of segments with regular rows of flat denticles, nearly equally large on dorsal, ventral and lateral sides of cercus (
<figureCitation id="135D04A4556F111EFB46E00FFAA3FD45" box="[1228,1311,620,645]" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="11" pageNumber="542">Fig. 59</figureCitation>
), on dorsal and ventral sides of paracercus.
</paragraph>
<paragraph id="8BD91821556F111EFF4DE0D7FE4AFD2F" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="542">RESPIRATORY MOVENENTS. Larva unable for rhythmical respiratory movements by tergalii; cannot live for long time in stagnant water.</paragraph>
<paragraph id="8BD91821556F111EFF4DE098FCC8FC40" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="542">
<emphasis id="B912C433556F111EFF4DE098FEFFFCD5" bold="true" box="[199,323,763,789]" pageId="11" pageNumber="542">
<emphasis id="B912C433556F111EFF4DE098FE80FCD4" bold="true" box="[199,316,763,788]" italics="true" pageId="11" pageNumber="542">Subimago</emphasis>
.
</emphasis>
CUTICULAR COLORATION. Head and pronotum entirely colorless. Prealar bridge contrastingly dark brown (
<figureCitation id="135D04A4556F111EFEBCE17CFE2EFCF8" box="[310,402,799,824]" captionStart="FIGURES 70–74" captionStartId="17.[151,264,1969,1992]" captionTargetBox="[159,1427,223,1913]" captionTargetId="figure@17.[159,1427,223,1913]" captionTargetPageId="17" captionText="FIGURES 70–74. Guajirolus ektrapeloglossa from Quebrada Barrigón. 70–71, male imagines; 72, subimaginal exuviae of left half of mesonotum; 73, subimaginal exuviae of left half of meso- and metapleuron; 74, subimago extracted from larval exuviae." httpUri="https://zenodo.org/record/2588985/files/figure.png" pageId="11" pageNumber="542">Fig. 72</figureCitation>
). Mesonotum partly colorless, with certain areas shed with light brownish (
<figureCitation id="135D04A4556F111EFABEE17CFA2CFCF8" box="[1332,1424,799,824]" captionStart="FIGURES 70–74" captionStartId="17.[151,264,1969,1992]" captionTargetBox="[159,1427,223,1913]" captionTargetId="figure@17.[159,1427,223,1913]" captionTargetPageId="17" captionText="FIGURES 70–74. Guajirolus ektrapeloglossa from Quebrada Barrigón. 70–71, male imagines; 72, subimaginal exuviae of left half of mesonotum; 73, subimaginal exuviae of left half of meso- and metapleuron; 74, subimago extracted from larval exuviae." httpUri="https://zenodo.org/record/2588985/files/figure.png" pageId="11" pageNumber="542">Fig. 72</figureCitation>
). Pterothoracic pleura and sterna at most part colorless, with certain sclerites contrastingly brown (
<figureCitation id="135D04A4556F111EFB73E127FAECFC9C" box="[1273,1360,836,861]" captionStart="FIGURES 70–74" captionStartId="17.[151,264,1969,1992]" captionTargetBox="[159,1427,223,1913]" captionTargetId="figure@17.[159,1427,223,1913]" captionTargetPageId="17" captionText="FIGURES 70–74. Guajirolus ektrapeloglossa from Quebrada Barrigón. 70–71, male imagines; 72, subimaginal exuviae of left half of mesonotum; 73, subimaginal exuviae of left half of meso- and metapleuron; 74, subimago extracted from larval exuviae." httpUri="https://zenodo.org/record/2588985/files/figure.png" pageId="11" pageNumber="542">Fig. 73</figureCitation>
). Legs entirely colorless. Abdomen, genitals and cerci entirely colorless.
</paragraph>
<paragraph id="8BD91821556F111EFF4DE1EFFC0CFBD0" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="542">
HYPODERMAL COLORATION. Just after emergence from larva, most part of body lack hypodermal coloration, being pale ocher; certain areas of thorax brown; abdomen with intersegmental membranes of terga brown, terga with light brownish pigmentation forming paired diffusive maculae (
<figureCitation id="135D04A4556F111EFBC6E1B7FB1FFC2C" box="[1100,1187,980,1005]" captionStart="FIGURES 70–74" captionStartId="17.[151,264,1969,1992]" captionTargetBox="[159,1427,223,1913]" captionTargetId="figure@17.[159,1427,223,1913]" captionTargetPageId="17" captionText="FIGURES 70–74. Guajirolus ektrapeloglossa from Quebrada Barrigón. 70–71, male imagines; 72, subimaginal exuviae of left half of mesonotum; 73, subimaginal exuviae of left half of meso- and metapleuron; 74, subimago extracted from larval exuviae." httpUri="https://zenodo.org/record/2588985/files/figure.png" pageId="11" pageNumber="542">Fig. 74</figureCitation>
). During subimaginal development, thorax and abdominal terga get brown color as in imago.
</paragraph>
<paragraph id="8BD91821556F111EFF4DE67EFBD1FB97" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="542">TEXTURE. On all legs of male and female last tarsomere covered by pointed microlepides, previous tarsomeres covered mainly by blunt microlepides, with pointed microlepides near apex.</paragraph>
<paragraph id="8BD91821556F111EFF4DE607FC12FA5D" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="542">
<emphasis id="B912C433556F111EFF4DE607FEEBFBBC" bold="true" box="[199,343,1123,1148]" italics="true" pageId="11" pageNumber="542">Imago, male</emphasis>
(
<figureCitation id="135D04A4556F111EFEECE607FE55FBBC" box="[358,489,1124,1149]" captionStart="FIGURES 70–74" captionStartId="17.[151,264,1969,1992]" captionTargetBox="[159,1427,223,1913]" captionTargetId="figure@17.[159,1427,223,1913]" captionTargetPageId="17" captionText="FIGURES 70–74. Guajirolus ektrapeloglossa from Quebrada Barrigón. 70–71, male imagines; 72, subimaginal exuviae of left half of mesonotum; 73, subimaginal exuviae of left half of meso- and metapleuron; 74, subimago extracted from larval exuviae." httpUri="https://zenodo.org/record/2588985/files/figure.png" pageId="11" pageNumber="542">Figs 70–71</figureCitation>
). Head ocher. Turbinate eyes with stalks relatively short and sharply widened toward facetted surface; facetted surfaces very wide, elliptical, greatly divergent anteriorly; stalk and facetted surface brown, border between them narrowly outlined by dark brown. Thorax brown with ocher, equally dark dorsally and ventrally. Fore wing colorless, with veins pale yellowish, only bases of costa and radius brown. Hind wing absent. All legs unicolor pale yellowish. On middle and hind legs tarsus relatively short, 0.3 of tibia length. Length of leg segments (mm): femur, tibia and tarsomeres of fore leg 0.87: 1.70: 0.06:050: 0.30: 0.20: 0.13; of middle leg 0.78: 1.05: 0.10: 0.06: 0.03: 0.13; of hind leg 0.82: 1.05: 0.10: 0.06: 0.03: 0.13. Apical spines either present on two tarsomeres, or on one tarsomere, or absent. All abdominal terga nearly unicolor brown; sterna ocher, shed with gray anteriorly and laterally. Genitals brown. Cerci unicolor pale yellowish.
</paragraph>
<paragraph id="8BD91821556F111EFF4DE7C4FBA1F958" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="542">
<emphasis id="B912C433556F111EFF4DE7C4FD21FA00" bold="true" box="[199,669,1447,1472]" italics="true" pageId="11" pageNumber="542">Male genital structure and development</emphasis>
(
<figureCitation id="135D04A4556F111EFD24E7C4FC88FA00" box="[686,820,1447,1472]" captionStart="FIGURES 75–80" captionStartId="18.[151,264,1754,1777]" captionTargetBox="[158,1428,193,1733]" captionTargetId="figure@18.[158,1428,193,1733]" captionTargetPageId="18" captionText="FIGURES 75–80. Male genitals of Guajirolus ektrapeloglossa from Quebrada Barrigón. 75, genital of imago (ventral view; muscles and internal parts of penis shown by interrupted lines; right gonostylar and gonovectal muscles not shown); 76, subimaginal genitals developing under larval cuticle; 77–78 and 79–80, genitals of two specimens. Abbreviations: 1, 2, 3, segments of gonostylus; pgs, larval protogonostylus; usg, unistyliger." httpUri="https://zenodo.org/record/2588987/files/figure.png" pageId="11" pageNumber="542">Figs 75–80</figureCitation>
). Sterno-styligeral muscle present, but weak; it arises from base of abdominal sternum IX and terminates near its middle, far not reaching bases of unistyligers (
<figureCitation id="135D04A4556F111EFAB1E7AFFA2CFA25" box="[1339,1424,1484,1509]" captionStart="FIGURES 75–80" captionStartId="18.[151,264,1754,1777]" captionTargetBox="[158,1428,193,1733]" captionTargetId="figure@18.[158,1428,193,1733]" captionTargetPageId="18" captionText="FIGURES 75–80. Male genitals of Guajirolus ektrapeloglossa from Quebrada Barrigón. 75, genital of imago (ventral view; muscles and internal parts of penis shown by interrupted lines; right gonostylar and gonovectal muscles not shown); 76, subimaginal genitals developing under larval cuticle; 77–78 and 79–80, genitals of two specimens. Abbreviations: 1, 2, 3, segments of gonostylus; pgs, larval protogonostylus; usg, unistyliger." httpUri="https://zenodo.org/record/2588987/files/figure.png" pageId="11" pageNumber="542">Fig. 75</figureCitation>
). Gonovectes curved S-sharply; penial bridge with small prominent slerotized median projection. Unistyligers long, narrowed at midlength and widened apically. 1st segment of gonostylus with projected right angle on inner side apically; 2nd segment widened apically; 3rd segment small, with thin stem and thickened apically. In mature larva ready to molt to subimago, gonostyli folded by the «
<taxonomicName id="4C6663A2556F111EFD6CE43FFCE2F9B4" authorityName="Flowers" authorityYear="1985" box="[742,862,1628,1652]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="B912C433556F111EFD6CE43FFCE2F9B4" box="[742,862,1628,1652]" italics="true" pageId="11" pageNumber="542">Guajirolus</emphasis>
</taxonomicName>
-
<typeStatus id="54DDA683556F111EFCECE43EFC27F9B5" box="[870,923,1629,1653]" pageId="11" pageNumber="542">type</typeStatus>
», i.e. with 3rd segment attached caudally and retaining under larval protogonostylus, but turned by its apex medially (
<figureCitation id="135D04A4556F111EFC36E41CFBADF958" box="[956,1041,1663,1688]" captionStart="FIGURES 75–80" captionStartId="18.[151,264,1754,1777]" captionTargetBox="[158,1428,193,1733]" captionTargetId="figure@18.[158,1428,193,1733]" captionTargetPageId="18" captionText="FIGURES 75–80. Male genitals of Guajirolus ektrapeloglossa from Quebrada Barrigón. 75, genital of imago (ventral view; muscles and internal parts of penis shown by interrupted lines; right gonostylar and gonovectal muscles not shown); 76, subimaginal genitals developing under larval cuticle; 77–78 and 79–80, genitals of two specimens. Abbreviations: 1, 2, 3, segments of gonostylus; pgs, larval protogonostylus; usg, unistyliger." httpUri="https://zenodo.org/record/2588987/files/figure.png" pageId="11" pageNumber="542">Fig. 76</figureCitation>
).
</paragraph>
<paragraph id="8BD91821556F111EFF4DE4C7FB5FF97C" blockId="11.[151,1437,151,2013]" box="[199,1251,1699,1725]" pageId="11" pageNumber="542">
<emphasis id="B912C433556F111EFF4DE4C7FEC8F97D" bold="true" box="[199,372,1699,1725]" pageId="11" pageNumber="542">
<emphasis id="B912C433556F111EFF4DE4C7FED1F97C" bold="true" box="[199,365,1699,1724]" italics="true" pageId="11" pageNumber="542">Imago, female</emphasis>
.
</emphasis>
Coloration similar to that of male. Eyes small, widely separated (
<figureCitation id="135D04A4556F111EFBD9E4C7FB6AF97C" box="[1107,1238,1700,1725]" captionStart="FIGURES 81–82" captionStartId="19.[151,264,1894,1917]" captionTargetBox="[158,1429,547,1844]" captionTargetId="figure@19.[158,1429,547,1844]" captionTargetPageId="19" captionText="FIGURES 81–82. Female imagines of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri="https://zenodo.org/record/2588989/files/figure.png" pageId="11" pageNumber="542">Figs 81–82</figureCitation>
).
</paragraph>
<paragraph id="8BD91821556F111EFF4DE4ABFE1DF8C5" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="542">
<emphasis id="B912C433556F111EFF4DE4ABFEBDF91F" bold="true" box="[199,257,1736,1759]" italics="true" pageId="11" pageNumber="542">Eggs</emphasis>
(
<figureCitation id="135D04A4556F111EFE9BE4A4FE29F920" box="[273,405,1735,1760]" captionStart="FIGURES 83–88" captionStartId="21.[151,264,1913,1936]" captionTargetBox="[163,1423,465,1892]" captionTargetId="figure@21.[163,1423,465,1892]" captionTargetPageId="21" captionText="FIGURES 83–88. Eggs of Guajirolus ektrapeloglossa. 83–85, from Quebrada Barrigón (Panama); 86–88, from Rio Sotorani (Peru)." httpUri="https://zenodo.org/record/2588991/files/figure.png" pageId="11" pageNumber="542">Figs 83–88</figureCitation>
). Oval. Chorion partly smooth or irregularly crumpled, partly with fine net-like relief with cells small and shallow.
</paragraph>
<paragraph id="8BD91821556F111EFF4DE56CFCF0F8E8" blockId="11.[151,1437,151,2013]" box="[199,844,1807,1832]" pageId="11" pageNumber="542">
<emphasis id="B912C433556F111EFF4DE56CFEF0F8E8" bold="true" box="[199,332,1807,1832]" pageId="11" pageNumber="542">
<emphasis id="B912C433556F111EFF4DE56CFEF9F8E8" bold="true" box="[199,325,1807,1832]" italics="true" pageId="11" pageNumber="542">Dimension</emphasis>
.
</emphasis>
Fore wing length (and body length)
<quantity id="4C9EB5C4556F111EFD6DE56CFCFBF8E8" box="[743,839,1807,1832]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="11" pageNumber="542" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
.
</paragraph>
<paragraph id="8BD91821556F111EFF4DE557FAA0F81D" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="542">
<emphasis id="B912C433556F111EFF4DE557FE6CF88D" bold="true" box="[199,464,1844,1869]" pageId="11" pageNumber="542">Habitat and sex ratio.</emphasis>
Most part of specimens used in this study (
<specimenCount id="9D60D3A8556F111EFC4EE557FB92F88D" box="[964,1070,1844,1869]" pageId="11" pageNumber="542" type="larva">75 larvae</specimenCount>
and winged insects reared from them) were collected in Quebrada Barrigón, on a portion of its bottom formed by an integral flat rock.
<bibRefCitation id="EFF765D0556F111EFAC8E534FF5DF855" author="Flowers, R. W." pageId="11" pageNumber="542" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers (1985)</bibRefCitation>
also reported that he found larvae of
<taxonomicName id="4C6663A2556F111EFD09E51FFCEEF854" authorityName="Flowers" authorityYear="1985" box="[643,850,1916,1940]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="542" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C433556F111EFD09E51FFCEEF854" box="[643,850,1916,1940]" italics="true" pageId="11" pageNumber="542">G. ektrapeloglossa</emphasis>
</taxonomicName>
on rocky substrates. Among all specimens collected in Barrigón, 2/3 are females and only 1/3 are males. Three other specimens examined by me (one larva from Chiriquí Nuevo and two specimens from
<collectingCountry id="F37158B1556F111EFDECE5A7FD27F81C" box="[614,667,1988,2012]" name="Peru" pageId="11" pageNumber="542">Peru</collectingCountry>
) are females. Probably, females dominate in this species.
</paragraph>
<paragraph id="8BD9182155681119FF4DE2F4FD56FE48" blockId="12.[151,1437,151,1293]" pageId="12" pageNumber="543">
<emphasis id="B912C43355681119FF4DE2F4FE26FF70" bold="true" box="[199,410,151,176]" pageId="12" pageNumber="543">Stage association.</emphasis>
<bibRefCitation id="EFF765D055681119FE29E2F4FDEDFF70" author="Flowers, R. W." box="[419,593,151,176]" pageId="12" pageNumber="543" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers (1985)</bibRefCitation>
wrote that his association of male and female imagines with the nymphs was done «on the basis of color patterns», but did not specify, which features of coloration allowed him to do this. The brown hypodermal coloration of imaginal thorax and abdomen appears only in course of subimaginal development, being absent in larva and in just-molted subimago (
<figureCitation id="135D04A455681119FCF7E367FC6DFEDC" box="[893,977,260,285]" captionStart="FIGURES 70–74" captionStartId="17.[151,264,1969,1992]" captionTargetBox="[159,1427,223,1913]" captionTargetId="figure@17.[159,1427,223,1913]" captionTargetPageId="17" captionText="FIGURES 70–74. Guajirolus ektrapeloglossa from Quebrada Barrigón. 70–71, male imagines; 72, subimaginal exuviae of left half of mesonotum; 73, subimaginal exuviae of left half of meso- and metapleuron; 74, subimago extracted from larval exuviae." httpUri="https://zenodo.org/record/2588985/files/figure.png" pageId="12" pageNumber="543">Fig. 74</figureCitation>
), so it does not allow to associate larvae and imagines; legs and cerci have no hypodermal coloration at any stage of development. Cuticular coloration, being characteristic for larvae, is not retained in subimago and imago. Rearing imagines from larvae confirms correctness of the stage association done by Flowers.
</paragraph>
</subSubSection>
<subSubSection id="C37C4BAA55681119FF4DE3F7FCC0FE6D" box="[199,892,404,429]" pageId="12" pageNumber="543" type="distribution">
<paragraph id="8BD9182155681119FF4DE3F7FCC0FE6D" blockId="12.[151,1437,151,1293]" box="[199,892,404,429]" pageId="12" pageNumber="543">
<emphasis id="B912C43355681119FF4DE3F7FEDCFE6D" bold="true" box="[199,352,404,429]" pageId="12" pageNumber="543">Distribution.</emphasis>
Central and South America (see «Discussion»).
</paragraph>
</subSubSection>
<subSubSection id="C37C4BAA55681100FF4DE3D4FEA4FE48" lastPageId="21" lastPageNumber="552" pageId="12" pageNumber="543" type="discussion">
<paragraph id="8BD9182155681119FF4DE3D4FDB3FACC" blockId="12.[151,1437,151,1293]" pageId="12" pageNumber="543">
<emphasis id="B912C43355681119FF4DE3D4FEF0FE10" bold="true" box="[199,332,439,464]" pageId="12" pageNumber="543">Discussion.</emphasis>
The examined specimens from
<collectingCountry id="F37158B155681119FD4DE3DBFD47FE0F" box="[711,763,440,463]" name="Peru" pageId="12" pageNumber="543">Peru</collectingCountry>
and
<collectingCountry id="F37158B155681119FCB3E3DBFC2EFE0F" box="[825,914,440,463]" name="Panama" pageId="12" pageNumber="543">Panama</collectingCountry>
have the same structure of larva, subimago, female imago and eggs (compare
<figureCitation id="135D04A455681119FDAAE3BFFC9AFE35" box="[544,806,476,501]" captionStart="FIGURES 83–88" captionStartId="21.[151,264,1913,1936]" captionTargetBox="[163,1423,465,1892]" captionTargetId="figure@21.[163,1423,465,1892]" captionTargetPageId="21" captionText="FIGURES 83–88. Eggs of Guajirolus ektrapeloglossa. 83–85, from Quebrada Barrigón (Panama); 86–88, from Rio Sotorani (Peru)." httpUri="https://zenodo.org/record/2588991/files/figure.png" pageId="12" pageNumber="543">Figs 83–85 and 86–88</figureCitation>
), that allows to conclude that they belong to the same species, which is widely distributed in South and Central America. According to current literature, 6 species are reported in the genus
<taxonomicName id="4C6663A255681119FE07E047FDB9FDFC" authorityName="Flowers" authorityYear="1985" box="[397,517,548,572]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="B912C43355681119FE07E047FDB9FDFC" box="[397,517,548,572]" italics="true" pageId="12" pageNumber="543">Guajirolus</emphasis>
</taxonomicName>
: the species
<taxonomicName id="4C6663A255681119FD12E047FCD5FDFC" authorityName="Flowers" authorityYear="1985" box="[664,873,548,572]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355681119FD12E047FCD5FDFC" box="[664,873,548,572]" italics="true" pageId="12" pageNumber="543">G. ektrapeloglossa</emphasis>
</taxonomicName>
was originally described by
<bibRefCitation id="EFF765D055681119FB3AE047FAE1FDFD" author="Flowers, R. W." box="[1200,1373,548,573]" pageId="12" pageNumber="543" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers (1985)</bibRefCitation>
from
<collectingCountry id="F37158B155681119FF1DE02BFF4CFD9F" box="[151,240,584,607]" name="Panama" pageId="12" pageNumber="543">Panama</collectingCountry>
and
<collectingCountry id="F37158B155681119FEAFE024FE2FFDA0" box="[293,403,583,608]" name="Colombia" pageId="12" pageNumber="543">Colombia</collectingCountry>
as distinct from the unnamed species from
<collectingCountry id="F37158B155681119FCF3E02BFC11FD9F" box="[889,941,584,607]" name="Peru" pageId="12" pageNumber="543">Peru</collectingCountry>
reported by
<bibRefCitation id="EFF765D055681119FBB0E024FB5FFDA0" author="Roback, S. S." box="[1082,1251,583,608]" pageId="12" pageNumber="543" pagination="129 - 199" refId="ref9176" refString="Roback, S. S. (1966) The Catherwood Foundation Peruvia-Amazon expedition. VI-Ephemeroptera nymphs. Monographs of the Academy of Natural Sciences of Philadelphia, No. 14, 129 - 199." type="book chapter" year="1966">Roback (1966)</bibRefCitation>
; later, four other species were described—
<taxonomicName id="4C6663A255681119FE3EE00FFDA9FD44" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[436,533,620,644]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355681119FE3EE00FFDA9FD44" box="[436,533,620,644]" italics="true" pageId="12" pageNumber="543">G. nanus</emphasis>
</taxonomicName>
from
<collectingCountry id="F37158B155681119FDD0E00FFD6BFD45" box="[602,727,620,645]" name="Costa Rica" pageId="12" pageNumber="543">Costa Rica</collectingCountry>
,
<taxonomicName id="4C6663A255681119FD68E00FFCCCFD44" authorityName="Nieto" authorityYear="2003" box="[738,880,620,644]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="queremba">
<emphasis id="B912C43355681119FD68E00FFCCCFD44" box="[738,880,620,644]" italics="true" pageId="12" pageNumber="543">G. queremba</emphasis>
</taxonomicName>
from
<collectingCountry id="F37158B155681119FC3FE00FFB9BFD45" box="[949,1063,620,645]" name="Argentina" pageId="12" pageNumber="543">Argentina</collectingCountry>
,
<taxonomicName id="4C6663A255681119FBB8E00FFB17FD44" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[1074,1195,620,644]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355681119FBB8E00FFB17FD44" box="[1074,1195,620,644]" italics="true" pageId="12" pageNumber="543">G. flowersi</emphasis>
</taxonomicName>
from French
<collectingCountry id="F37158B155681119FACFE00FFA20FD44" box="[1349,1436,620,644]" name="Guyana" pageId="12" pageNumber="543">Guyana</collectingCountry>
and
<taxonomicName id="4C6663A255681119FF41E0F3FEFBFD68" authorityName="Salles" authorityYear="2007" box="[203,327,656,680]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="rondoni">
<emphasis id="B912C43355681119FF41E0F3FEFBFD68" box="[203,327,656,680]" italics="true" pageId="12" pageNumber="543">G. rondoni</emphasis>
</taxonomicName>
from
<collectingCountry id="F37158B155681119FE1FE0ECFE65FD68" box="[405,473,655,680]" name="Brazil" pageId="12" pageNumber="543">Brazil</collectingCountry>
(
<bibRefCitation id="EFF765D055681119FE67E0ECFCD8FD68" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[493,868,655,680]" pageId="12" pageNumber="543" pagination="68 - 70" refId="ref8926" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1995) Guajirolus nanus (Ephemeroptera: Baetidae), a new species from Costa Rica. Entomological News, 106 (2), 68 - 70." type="journal article" year="1995">Lugo-Ortiz &amp; McCafferty 1995</bibRefCitation>
,
<bibRefCitation id="EFF765D055681119FCFEE0ECFC46FD68" author="Nieto, C." box="[884,1018,655,680]" pageId="12" pageNumber="543" pagination="153 - 158" refId="ref8969" refString="Nieto, C. (2003) A new species of Guajirolus (Ephemeroptera: Baetidae) from Argentina and description of a new genus from Bolivia. In: Gaino E. (Ed.), Research Update on Ephemeroptera &amp; Plecoptera (Proc. 2001 Int. Joint Meeting-X Int. Conf. on Ephemeroptera &amp; XIV Int. Symp. on Plecoptera, August 5 - 11, 2001 in Perugia). Univ. Perugia, Italy, 153 - 158." type="book chapter" year="2003">Nieto 2003</bibRefCitation>
,
<bibRefCitation id="EFF765D055681119FB80E0ECFB48FD68" author="Thomas, A. &amp; Dominique, Y." box="[1034,1268,655,680]" pageId="12" pageNumber="543" pagination="21 - 29" refId="ref9271" refString="Thomas, A., Dominique, Y. &amp; Orth, K. (2005) Les Ephemeres de la Guyane Francaise. 12. Description de Guajirolus flowersi n. sp. [Ephemeroptera, Baetidae]. Ephemera, (2004), 6 (1), 21 - 29." type="journal article" year="2005">
Thomas
<emphasis id="B912C43355681119FBFAE0F2FB1BFD68" box="[1136,1191,656,680]" italics="true" pageId="12" pageNumber="543">et al</emphasis>
. 2005
</bibRefCitation>
,
<bibRefCitation id="EFF765D055681119FA8FE0ECFA2DFD68" author="Salles, F. F." box="[1285,1425,655,680]" pageId="12" pageNumber="543" pagination="404 - 409" refId="ref9212" refString="Salles, F. F. (2007) The presence of Chane Nieto and Guajirolus Flowers (Ephemeroptera, Baetidae) in Brazil with the description of a new species. Revista Brasileira de Entomologia, 51 (4), 404 - 409. https: // doi. org / 10.1590 / S 0085 - 56262007000400002" type="journal article" year="2007">Salles 2007</bibRefCitation>
). Diagnoses of all these species were based on larval characters (presumably determined imagines were described only for
<taxonomicName id="4C6663A255681119FF70E0BBFE77FD30" authorityName="Flowers" authorityYear="1985" box="[250,459,728,752]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355681119FF70E0BBFE77FD30" box="[250,459,728,752]" italics="true" pageId="12" pageNumber="543">G. ektrapeloglossa</emphasis>
</taxonomicName>
, and subimago—only for
<taxonomicName id="4C6663A255681119FD7DE0BBFC39FD30" authorityName="Nieto" authorityYear="2003" box="[759,901,728,752]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="queremba">
<emphasis id="B912C43355681119FD7DE0BBFC39FD30" box="[759,901,728,752]" italics="true" pageId="12" pageNumber="543">G. queremba</emphasis>
</taxonomicName>
). Each author, who described a new species of
<taxonomicName id="4C6663A255681119FF1DE09FFEB3FCD4" authorityName="Flowers" authorityYear="1985" box="[151,271,764,788]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="B912C43355681119FF1DE09FFEB3FCD4" box="[151,271,764,788]" italics="true" pageId="12" pageNumber="543">Guajirolus</emphasis>
</taxonomicName>
, compared its characters with the previously published descriptions and figures, but nobody compared specimens of different species.
<bibRefCitation id="EFF765D055681119FD9EE17CFD7BFCF8" author="Flowers, R. W." box="[532,711,799,824]" pageId="12" pageNumber="543" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers (1985)</bibRefCitation>
regarded
<taxonomicName id="4C6663A255681119FCCCE143FBA1FCF8" authorityName="Flowers" authorityYear="1985" box="[838,1053,800,824]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355681119FCCCE143FBA1FCF8" box="[838,1053,800,824]" italics="true" pageId="12" pageNumber="543">G. ektrapeloglossa</emphasis>
</taxonomicName>
as different from the unnamed Roback's species because it has 6 pairs of tergalii, instead of 7 reported by Roback [see (J) below].
<bibRefCitation id="EFF765D055681119FA8AE127FECCFC40" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." pageId="12" pageNumber="543" pagination="68 - 70" refId="ref8926" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1995) Guajirolus nanus (Ephemeroptera: Baetidae), a new species from Costa Rica. Entomological News, 106 (2), 68 - 70." type="journal article" year="1995">Lugo-Ortiz &amp; McCafferty (1995)</bibRefCitation>
regarded
<taxonomicName id="4C6663A255681119FE61E10BFDEEFC40" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[491,594,872,896]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355681119FE61E10BFDEEFC40" box="[491,594,872,896]" italics="true" pageId="12" pageNumber="543">G. nanus</emphasis>
</taxonomicName>
as different from
<taxonomicName id="4C6663A255681119FCBBE10BFBBBFC40" authorityName="Flowers" authorityYear="1985" box="[817,1031,872,896]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355681119FCBBE10BFBBBFC40" box="[817,1031,872,896]" italics="true" pageId="12" pageNumber="543">G. ektrapeloglossa</emphasis>
</taxonomicName>
based on presence of spicules on mandibles, shape of median process on hypopharynx, presence of proximal brown band on femur and number of spines on paraprocts; however, the original description of
<taxonomicName id="4C6663A255681119FCB7E1D3FBADFC08" authorityName="Flowers" authorityYear="1985" box="[829,1041,944,968]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355681119FCB7E1D3FBADFC08" box="[829,1041,944,968]" italics="true" pageId="12" pageNumber="543">G. ektrapeloglossa</emphasis>
</taxonomicName>
does not contain clear statements about presence or absence of these characters [see (B), (D), (F) and (H) below].
<bibRefCitation id="EFF765D055681119FBABE1B7FB0DFC2C" author="Nieto, C." box="[1057,1201,980,1005]" pageId="12" pageNumber="543" pagination="153 - 158" refId="ref8969" refString="Nieto, C. (2003) A new species of Guajirolus (Ephemeroptera: Baetidae) from Argentina and description of a new genus from Bolivia. In: Gaino E. (Ed.), Research Update on Ephemeroptera &amp; Plecoptera (Proc. 2001 Int. Joint Meeting-X Int. Conf. on Ephemeroptera &amp; XIV Int. Symp. on Plecoptera, August 5 - 11, 2001 in Perugia). Univ. Perugia, Italy, 153 - 158." type="book chapter" year="2003">Nieto (2003)</bibRefCitation>
regarded
<taxonomicName id="4C6663A255681119FAA8E1B7FA27FC2C" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[1314,1435,980,1004]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355681119FAA8E1B7FA27FC2C" box="[1314,1435,980,1004]" italics="true" pageId="12" pageNumber="543">G. flowersi</emphasis>
</taxonomicName>
as different from
<taxonomicName id="4C6663A255681119FEE9E19BFD8AFBD0" authorityName="Flowers" authorityYear="1985" box="[355,566,1016,1040]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355681119FEE9E19BFD8AFBD0" box="[355,566,1016,1040]" italics="true" pageId="12" pageNumber="543">G. ektrapeloglossa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6663A255681119FDF9E19BFD64FBD0" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[627,728,1016,1040]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355681119FDF9E19BFD64FBD0" box="[627,728,1016,1040]" italics="true" pageId="12" pageNumber="543">G. nanus</emphasis>
</taxonomicName>
based on combination of the same characters.
<bibRefCitation id="EFF765D055681119FB76E194FF5DFBF5" author="Thomas, A. &amp; Dominique, Y." pageId="12" pageNumber="543" pagination="21 - 29" refId="ref9271" refString="Thomas, A., Dominique, Y. &amp; Orth, K. (2005) Les Ephemeres de la Guyane Francaise. 12. Description de Guajirolus flowersi n. sp. [Ephemeroptera, Baetidae]. Ephemera, (2004), 6 (1), 21 - 29." type="journal article" year="2005">
Thomas
<emphasis id="B912C43355681119FAEBE19AFA29FBD0" box="[1377,1429,1016,1040]" italics="true" pageId="12" pageNumber="543">et al</emphasis>
. (2005)
</bibRefCitation>
regarded
<taxonomicName id="4C6663A255681119FED3E67FFE6AFBF4" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[345,470,1052,1076]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355681119FED3E67FFE6AFBF4" box="[345,470,1052,1076]" italics="true" pageId="12" pageNumber="543">G. flowersi</emphasis>
</taxonomicName>
as different from previously described species based on shape of distal margin of labrum [see (A)], maxillary palp interpreted as 4-segmented [see (C)] and combination of characters used by previous authors.
<bibRefCitation id="EFF765D055681119FEE0E607FDB8FBBC" author="Salles, F. F." box="[362,516,1124,1149]" pageId="12" pageNumber="543" pagination="404 - 409" refId="ref9212" refString="Salles, F. F. (2007) The presence of Chane Nieto and Guajirolus Flowers (Ephemeroptera, Baetidae) in Brazil with the description of a new species. Revista Brasileira de Entomologia, 51 (4), 404 - 409. https: // doi. org / 10.1590 / S 0085 - 56262007000400002" type="journal article" year="2007">Salles (2007)</bibRefCitation>
regarded
<taxonomicName id="4C6663A255681119FDF5E607FD46FBBC" authorityName="Salles" authorityYear="2007" box="[639,762,1124,1148]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="rondoni">
<emphasis id="B912C43355681119FDF5E607FD46FBBC" box="[639,762,1124,1148]" italics="true" pageId="12" pageNumber="543">G. rondoni</emphasis>
</taxonomicName>
as different from previously described species based on combination of the same characters plus narrower projection of 2nd segment of labial palp [see (E)]. Only the last character is species-specific; all other characters reported as species-specific in
<taxonomicName id="4C6663A255681119FB86E6CFFB38FB04" authorityName="Flowers" authorityYear="1985" box="[1036,1156,1196,1220]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="B912C43355681119FB86E6CFFB38FB04" box="[1036,1156,1196,1220]" italics="true" pageId="12" pageNumber="543">Guajirolus</emphasis>
</taxonomicName>
, either vary individually, or are caused by different interpretations made by different authors. The following characters were used to distinguish species of
<taxonomicName id="4C6663A255681119FE05E697FDBBFACC" authorityName="Flowers" authorityYear="1985" box="[399,519,1268,1292]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="B912C43355681119FE05E697FDBBFACC" box="[399,519,1268,1292]" italics="true" pageId="12" pageNumber="543">Guajirolus</emphasis>
</taxonomicName>
:
</paragraph>
<paragraph id="8BD9182155681119FF1DE75FFA99F958" blockId="12.[151,1437,1339,2013]" pageId="12" pageNumber="543">
(A)
<emphasis id="B912C43355681119FF4DE758FDF7FA95" bold="true" box="[199,587,1339,1365]" pageId="12" pageNumber="543">
<emphasis id="B912C43355681119FF4DE758FDFFFA94" bold="true" box="[199,579,1339,1364]" italics="true" pageId="12" pageNumber="543">Shape of distal margin of labrum</emphasis>
.
</emphasis>
<bibRefCitation id="EFF765D055681119FDDBE75FFC87FA95" author="Thomas, A. &amp; Dominique, Y." box="[593,827,1340,1365]" pageId="12" pageNumber="543" pagination="21 - 29" refId="ref9271" refString="Thomas, A., Dominique, Y. &amp; Orth, K. (2005) Les Ephemeres de la Guyane Francaise. 12. Description de Guajirolus flowersi n. sp. [Ephemeroptera, Baetidae]. Ephemera, (2004), 6 (1), 21 - 29." type="journal article" year="2005">
Thomas
<emphasis id="B912C43355681119FD38E75EFD5DFA94" box="[690,737,1340,1364]" italics="true" pageId="12" pageNumber="543">et al</emphasis>
. (2005)
</bibRefCitation>
stated that the new species
<taxonomicName id="4C6663A255681119FBE5E75FFB5BFA94" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[1135,1255,1340,1364]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355681119FBE5E75FFB5BFA94" box="[1135,1255,1340,1364]" italics="true" pageId="12" pageNumber="543">G. flowersi</emphasis>
</taxonomicName>
differs from the earlier described species of
<taxonomicName id="4C6663A255681119FD87E703FD39FAB8" authorityName="Flowers" authorityYear="1985" box="[525,645,1376,1400]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="B912C43355681119FD87E703FD39FAB8" box="[525,645,1376,1400]" italics="true" pageId="12" pageNumber="543">Guajirolus</emphasis>
</taxonomicName>
by the median («sagittal») emargination of labrum, whose width is subequal to width of lateral processes, while in
<taxonomicName id="4C6663A255681119FD64E7E7FC7CFA5C" authorityName="Flowers" authorityYear="1985" box="[750,960,1412,1436]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355681119FD64E7E7FC7CFA5C" box="[750,960,1412,1436]" italics="true" pageId="12" pageNumber="543">G. ektrapeloglossa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6663A255681119FC5BE7E7FB89FA5C" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[977,1077,1412,1436]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355681119FC5BE7E7FB89FA5C" box="[977,1077,1412,1436]" italics="true" pageId="12" pageNumber="543">G. nanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6663A255681119FBFAE7E7FABCFA5C" authorityName="Nieto" authorityYear="2003" box="[1136,1280,1412,1436]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="queremba">
<emphasis id="B912C43355681119FBFAE7E7FABCFA5C" box="[1136,1280,1412,1436]" italics="true" pageId="12" pageNumber="543">G. queremba</emphasis>
</taxonomicName>
width of this emargination is less than width of lateral processes.
<bibRefCitation id="EFF765D055681119FC81E7C4FC23FA00" author="Salles, F. F." box="[779,927,1447,1472]" pageId="12" pageNumber="543" pagination="404 - 409" refId="ref9212" refString="Salles, F. F. (2007) The presence of Chane Nieto and Guajirolus Flowers (Ephemeroptera, Baetidae) in Brazil with the description of a new species. Revista Brasileira de Entomologia, 51 (4), 404 - 409. https: // doi. org / 10.1590 / S 0085 - 56262007000400002" type="journal article" year="2007">Salles (2007)</bibRefCitation>
accepted this character in the key, to separate
<taxonomicName id="4C6663A255681119FF4DE7AFFEFDFA24" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[199,321,1484,1508]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355681119FF4DE7AFFEFDFA24" box="[199,321,1484,1508]" italics="true" pageId="12" pageNumber="543">G. flowersi</emphasis>
</taxonomicName>
and the new species
<taxonomicName id="4C6663A255681119FDBDE7AFFD12FA24" authorityName="Salles" authorityYear="2007" box="[567,686,1484,1508]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="rondoni">
<emphasis id="B912C43355681119FDBDE7AFFD12FA24" box="[567,686,1484,1508]" italics="true" pageId="12" pageNumber="543">G. rondoni</emphasis>
</taxonomicName>
from three other species; he also added that in
<taxonomicName id="4C6663A255681119FB5AE7AFFAF6FA24" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[1232,1354,1484,1508]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355681119FB5AE7AFFAF6FA24" box="[1232,1354,1484,1508]" italics="true" pageId="12" pageNumber="543">G. flowersi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6663A255681119FA0EE7AFFEA3F9C8" authorityName="Salles" authorityYear="2007" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="rondoni">
<emphasis id="B912C43355681119FA0EE7AFFEA3F9C8" italics="true" pageId="12" pageNumber="543">G. rondoni</emphasis>
</taxonomicName>
emargination of labrum has stout setae, in contrast to the three other species. Actually labrum of
<taxonomicName id="4C6663A255681119FF4DE477FE83F9EC" authorityName="Flowers" authorityYear="1985" box="[199,319,1556,1580]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="B912C43355681119FF4DE477FE83F9EC" box="[199,319,1556,1580]" italics="true" pageId="12" pageNumber="543">Guajirolus</emphasis>
</taxonomicName>
has unusual structure [see
<taxonomicName id="4C6663A255681119FDE5E477FD57F9ED" authorityName="Flowers" authorityYear="1985" box="[623,747,1556,1581]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="553" phylum="Arthropoda" rank="genus">Guajirolus</taxonomicName>
/g1 (1)], being soft and lacking sclerotization on distal margin (
<figureCitation id="135D04A455681119FF45E454FEAAF990" box="[207,278,1591,1616]" captionStart="FIGURES 4–11" captionStartId="3.[151,264,1708,1731]" captionTargetBox="[184,1405,451,1693]" captionTargetId="figure@3.[184,1405,440,1693]" captionTargetPageId="3" captionText="FIGURES 4–11. Labrum of last instar larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 4, labrum; 5, exuviae of labrum; 6, longitudinal section of lateral part; 7–8, one and the same median emargination differently compressed; 9–11, fragments of exuviae of one labrum: 9, with focus on distal margin; 10, with focus on dorsal side; 11, detached ventral cuticle (right side at right)." httpUri="https://zenodo.org/record/2588965/files/figure.png" pageId="12" pageNumber="543">Fig. 6</figureCitation>
). Because of this, shape of its distal margin depends on fixation (
<figureCitation id="135D04A455681119FB9CE454FB06F990" box="[1046,1210,1591,1616]" captionStart="FIGURES 4–11" captionStartId="3.[151,264,1708,1731]" captionTargetBox="[184,1405,451,1693]" captionTargetId="figure@3.[184,1405,440,1693]" captionTargetPageId="3" captionText="FIGURES 4–11. Labrum of last instar larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 4, labrum; 5, exuviae of labrum; 6, longitudinal section of lateral part; 7–8, one and the same median emargination differently compressed; 9–11, fragments of exuviae of one labrum: 9, with focus on distal margin; 10, with focus on dorsal side; 11, detached ventral cuticle (right side at right)." httpUri="https://zenodo.org/record/2588965/files/figure.png" pageId="12" pageNumber="543">Figs 4–5, 7–8</figureCitation>
). Small stout setae, reported by Salles, are present in all specimens; if the emargination is spread and looks wide, these setae are well visible, but if the emargination is compressed and looks narrow, these setae can be overlooked.
</paragraph>
<paragraph id="8BD9182155681118FF1DE4C7FDD2FEDC" blockId="12.[151,1437,1339,2013]" lastBlockId="13.[199,1436,151,285]" lastPageId="13" lastPageNumber="544" pageId="12" pageNumber="543">
(B)
<emphasis id="B912C43355681119FF4DE4C0FE6CF97D" bold="true" box="[199,464,1699,1725]" pageId="12" pageNumber="543">
<emphasis id="B912C43355681119FF4DE4C0FE75F97C" bold="true" box="[199,457,1699,1724]" italics="true" pageId="12" pageNumber="543">Spicules on mandibles</emphasis>
.
</emphasis>
<bibRefCitation id="EFF765D055681119FE5DE4C7FCF1F97D" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[471,845,1700,1725]" pageId="12" pageNumber="543" pagination="68 - 70" refId="ref8926" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1995) Guajirolus nanus (Ephemeroptera: Baetidae), a new species from Costa Rica. Entomological News, 106 (2), 68 - 70." type="journal article" year="1995">Lugo-Ortiz &amp; McCafferty (1995)</bibRefCitation>
stated that the new species
<taxonomicName id="4C6663A255681119FB0EE4C7FB5AF97C" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[1156,1254,1700,1724]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355681119FB0EE4C7FB5AF97C" box="[1156,1254,1700,1724]" italics="true" pageId="12" pageNumber="543">G. nanus</emphasis>
</taxonomicName>
differs from the earlier described
<taxonomicName id="4C6663A255681119FE07E4ABFDDCF920" authorityName="Flowers" authorityYear="1985" box="[397,608,1736,1760]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355681119FE07E4ABFDDCF920" box="[397,608,1736,1760]" italics="true" pageId="12" pageNumber="543">G. ektrapeloglossa</emphasis>
</taxonomicName>
by presence of minute spicules on lateral margins of mandibles. These authors did not examine specimens of
<taxonomicName id="4C6663A255681119FD1EE48FFCD5F8C4" authorityName="Flowers" authorityYear="1985" box="[660,873,1772,1796]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355681119FD1EE48FFCD5F8C4" box="[660,873,1772,1796]" italics="true" pageId="12" pageNumber="543">G. ektrapeloglossa</emphasis>
</taxonomicName>
and regarded that «those spicules appear to be absent» based only on the figures in the original description. However,
<bibRefCitation id="EFF765D055681119FB8DE56CFB0AF8E8" author="Flowers, R. W." box="[1031,1206,1807,1832]" pageId="12" pageNumber="543" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers (1985)</bibRefCitation>
did not pay special attention to presence or absence of these spicules; on his figures of mandibles, he showed only fine setae (
<bibRefCitation id="EFF765D055681119FF44E534FE65F8B0" author="Flowers, R. W." box="[206,473,1879,1904]" pageId="12" pageNumber="543" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers 1985: figs 6–7</bibRefCitation>
), which were ignored and not figured by
<bibRefCitation id="EFF765D055681119FC3AE534FA24F8B0" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[944,1432,1879,1904]" pageId="12" pageNumber="543" pagination="68 - 70" refId="ref8926" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1995) Guajirolus nanus (Ephemeroptera: Baetidae), a new species from Costa Rica. Entomological News, 106 (2), 68 - 70." type="journal article" year="1995">Lugo-Ortiz &amp; McCafferty (1995: figs 2–3)</bibRefCitation>
.
<bibRefCitation id="EFF765D055681119FF4DE51FFEE6F854" author="Nieto, C." box="[199,346,1916,1941]" pageId="12" pageNumber="543" pagination="153 - 158" refId="ref8969" refString="Nieto, C. (2003) A new species of Guajirolus (Ephemeroptera: Baetidae) from Argentina and description of a new genus from Bolivia. In: Gaino E. (Ed.), Research Update on Ephemeroptera &amp; Plecoptera (Proc. 2001 Int. Joint Meeting-X Int. Conf. on Ephemeroptera &amp; XIV Int. Symp. on Plecoptera, August 5 - 11, 2001 in Perugia). Univ. Perugia, Italy, 153 - 158." type="book chapter" year="2003">Nieto (2003)</bibRefCitation>
did not report presence or absence of spicules neither in the description of a new species
<taxonomicName id="4C6663A255681119FA0EE51FFE8AF878" authorityName="Nieto" authorityYear="2003" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="queremba">
<emphasis id="B912C43355681119FA0EE51FFE8AF878" italics="true" pageId="12" pageNumber="543">G. queremba</emphasis>
</taxonomicName>
, nor in the «Discussion», but used this character in the key to separate
<taxonomicName id="4C6663A255681119FBD4E5C2FB50F878" authorityName="Nieto" authorityYear="2003" box="[1118,1260,1952,1976]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="queremba">
<emphasis id="B912C43355681119FBD4E5C2FB50F878" box="[1118,1260,1952,1976]" italics="true" pageId="12" pageNumber="543">G. queremba</emphasis>
</taxonomicName>
from
<taxonomicName id="4C6663A255681119FAB9E5C2FA29F878" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[1331,1429,1953,1976]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="543" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355681119FAB9E5C2FA29F878" box="[1331,1429,1953,1976]" italics="true" pageId="12" pageNumber="543">G. nanus</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFF765D055681119FF4DE5A7FE0CF81D" author="Thomas, A. &amp; Dominique, Y." box="[199,432,1988,2013]" pageId="12" pageNumber="543" pagination="21 - 29" refId="ref9271" refString="Thomas, A., Dominique, Y. &amp; Orth, K. (2005) Les Ephemeres de la Guyane Francaise. 12. Description de Guajirolus flowersi n. sp. [Ephemeroptera, Baetidae]. Ephemera, (2004), 6 (1), 21 - 29." type="journal article" year="2005">
Thomas
<emphasis id="B912C43355681119FEADE5A6FEEBF81C" box="[295,343,1988,2012]" italics="true" pageId="12" pageNumber="543">et al</emphasis>
. (2005)
</bibRefCitation>
repeated the thesis and antithesis from the paper by
<bibRefCitation id="EFF765D055681119FC7FE5A7FB38F81C" author="Nieto, C." box="[1013,1156,1988,2013]" pageId="12" pageNumber="543" pagination="153 - 158" refId="ref8969" refString="Nieto, C. (2003) A new species of Guajirolus (Ephemeroptera: Baetidae) from Argentina and description of a new genus from Bolivia. In: Gaino E. (Ed.), Research Update on Ephemeroptera &amp; Plecoptera (Proc. 2001 Int. Joint Meeting-X Int. Conf. on Ephemeroptera &amp; XIV Int. Symp. on Plecoptera, August 5 - 11, 2001 in Perugia). Univ. Perugia, Italy, 153 - 158." type="book chapter" year="2003">Nieto (2003)</bibRefCitation>
and reported presence of spicules for the new species
<taxonomicName id="4C6663A255691118FD9EE2FBFD2CFF70" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[532,656,152,176]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="544" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355691118FD9EE2FBFD2CFF70" box="[532,656,152,176]" italics="true" pageId="13" pageNumber="544">G. flowersi</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFF765D055691118FD2BE2F4FC85FF70" author="Salles, F. F." box="[673,825,151,176]" pageId="13" pageNumber="544" pagination="404 - 409" refId="ref9212" refString="Salles, F. F. (2007) The presence of Chane Nieto and Guajirolus Flowers (Ephemeroptera, Baetidae) in Brazil with the description of a new species. Revista Brasileira de Entomologia, 51 (4), 404 - 409. https: // doi. org / 10.1590 / S 0085 - 56262007000400002" type="journal article" year="2007">Salles (2007)</bibRefCitation>
reported presence of spicules for the new species
<taxonomicName id="4C6663A255691118FA0EE2FBFEA3FF14" authorityName="Salles" authorityYear="2007" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="544" phylum="Arthropoda" rank="species" species="rondoni">
<emphasis id="B912C43355691118FA0EE2FBFEA3FF14" italics="true" pageId="13" pageNumber="544">G. rondoni</emphasis>
</taxonomicName>
. Actually these spicules are present in
<taxonomicName id="4C6663A255691118FC84E2DFFC5AFF14" authorityName="Flowers" authorityYear="1985" box="[782,998,188,212]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="544" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355691118FC84E2DFFC5AFF14" box="[782,998,188,212]" italics="true" pageId="13" pageNumber="544">G. ektrapeloglossa</emphasis>
</taxonomicName>
(
<figureCitation id="135D04A455691118FC74E2DFFBE6FF15" box="[1022,1114,188,213]" captionStart="FIGURES 12–16" captionStartId="5.[151,264,1907,1930]" captionTargetBox="[157,1430,193,1886]" captionTargetId="figure@5.[157,1430,193,1886]" captionTargetPageId="5" captionText="FIGURES 12–16. Mandibles of Guajirolus ektrapeloglossa from Quebrada Barrigón. 12–13 right and left mandibles inside old cuticle before molt from penultimate to last larval instar; 14–15, biting edges of right and left mandibles; 16, lateral part of left mandible." httpUri="https://zenodo.org/record/2588967/files/figure.png" pageId="13" pageNumber="544">Fig. 16</figureCitation>
). Similar spicules, besides mandibles, are present on mesonotum, on certain areas of thoracic pleura and on all segments of legs (
<figureCitation id="135D04A455691118FAE0E2BCFEACFEDD" captionStart="FIGURES 44–50" captionStartId="14.[151,264,1937,1960]" captionTargetBox="[156,1430,258,1857]" captionTargetId="figure@14.[156,1430,258,1857]" captionTargetPageId="14" captionText="FIGURES 44–50. Larval legs of Guajirolus ektrapeloglossa from Quebrada Barrigón. 44–45, fore and middle legs of darkest specimen; 46–47, claws; 48, fragment of fore femur (focus on anterior side); 49, fragment of fore tibia (focus on anterior side); 50, fragment of hind tibia (focus on posterior side)." httpUri="https://zenodo.org/record/2588979/files/figure.png" pageId="13" pageNumber="544">Figs 48–50</figureCitation>
;
<bibRefCitation id="EFF765D055691118FE96E367FE44FEDD" author="Thomas, A. &amp; Dominique, Y." box="[284,504,260,285]" pageId="13" pageNumber="544" pagination="21 - 29" refId="ref9271" refString="Thomas, A., Dominique, Y. &amp; Orth, K. (2005) Les Ephemeres de la Guyane Francaise. 12. Description de Guajirolus flowersi n. sp. [Ephemeroptera, Baetidae]. Ephemera, (2004), 6 (1), 21 - 29." type="journal article" year="2005">
Thomas
<emphasis id="B912C43355691118FEF7E366FE12FEDC" box="[381,430,260,284]" italics="true" pageId="13" pageNumber="544">et al</emphasis>
. 2005
</bibRefCitation>
: figs 8–9).
</paragraph>
<caption id="DF1948A955691118FF1DE536FEBDF808" httpUri="https://zenodo.org/record/2588977/files/figure.png" pageId="13" pageNumber="544" startId="13.[151,264,1877,1900]" targetBox="[156,1431,355,1848]" targetPageId="13">
<paragraph id="8BD9182155691118FF1DE536FEBDF808" blockId="13.[151,1436,1877,1992]" pageId="13" pageNumber="544">
<emphasis id="B912C43355691118FF1DE536FEE9F8AB" bold="true" box="[151,341,1877,1900]" pageId="13" pageNumber="544">FIGURES 41–43.</emphasis>
Larval thorax of
<taxonomicName id="4C6663A255691118FD82E536FCA1F8AB" authorityName="Flowers" authorityYear="1985" box="[520,797,1877,1899]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="544" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355691118FD82E536FCA1F8AB" box="[520,797,1877,1899]" italics="true" pageId="13" pageNumber="544">Guajirolus ektrapeloglossa</emphasis>
</taxonomicName>
from Quebrada Barrigón. 41, exuviae of left half of pronotum and mesonotum; 42, posterior margin of mesonotum and protoptera; 43, exuviae of sterna and pleura (fore leg turned anteriorly, middle and hind legs detached). Abbreviations: abd1, first abdominal tergum; msth, mesothorax; mtth, metathorax; prth, prothorax.
</paragraph>
</caption>
<caption id="DF1948A9556A111BFF1DE5F2FD22F825" httpUri="https://zenodo.org/record/2588979/files/figure.png" pageId="14" pageNumber="545" startId="14.[151,264,1937,1960]" targetBox="[156,1430,258,1857]" targetPageId="14">
<paragraph id="8BD91821556A111BFF1DE5F2FD22F825" blockId="14.[151,1436,1937,2021]" pageId="14" pageNumber="545">
<emphasis id="B912C433556A111BFF1DE5F2FEE8F867" bold="true" box="[151,340,1937,1960]" pageId="14" pageNumber="545">FIGURES 44–50.</emphasis>
Larval legs of
<taxonomicName id="4C6663A2556A111BFE65E5F1FCBFF868" authorityName="Flowers" authorityYear="1985" box="[495,771,1938,1960]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="545" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C433556A111BFE65E5F1FCBFF868" box="[495,771,1938,1960]" italics="true" pageId="14" pageNumber="545">Guajirolus ektrapeloglossa</emphasis>
</taxonomicName>
from Quebrada Barrigón. 44–45, fore and middle legs of darkest specimen; 46–47, claws; 48, fragment of fore femur (focus on anterior side); 49, fragment of fore tibia (focus on anterior side); 50, fragment of hind tibia (focus on posterior side).
</paragraph>
</caption>
<caption id="DF1948A9556B111AFF1DE5B4FBF9F82E" box="[151,1093,2007,2030]" httpUri="https://zenodo.org/record/2588981/files/figure.png" pageId="15" pageNumber="546" startId="15.[151,264,2007,2030]" targetBox="[156,1430,243,1914]" targetPageId="15">
<paragraph id="8BD91821556B111AFF1DE5B4FBF9F82E" blockId="15.[151,1093,2007,2030]" box="[151,1093,2007,2030]" pageId="15" pageNumber="546">
<emphasis id="B912C433556B111AFF1DE5B4FEEFF82D" bold="true" box="[151,339,2007,2030]" pageId="15" pageNumber="546">FIGURES 51–58.</emphasis>
Larval coloration of
<taxonomicName id="4C6663A2556B111AFDA0E5BBFC81F82E" authorityName="Flowers" authorityYear="1985" box="[554,829,2008,2030]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C433556B111AFDA0E5BBFC81F82E" box="[554,829,2008,2030]" italics="true" pageId="15" pageNumber="546">Guajirolus ektrapeloglossa</emphasis>
</taxonomicName>
from Quebrada Barrigón.
</paragraph>
</caption>
<caption id="DF1948A955741105FF1DE5F5FB22F82A" httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="16" pageNumber="547" startId="16.[151,264,1942,1965]" targetBox="[178,1421,193,1929]" targetPageId="16">
<paragraph id="8BD9182155741105FF1DE5F5FB22F82A" blockId="16.[151,1436,1942,2026]" pageId="16" pageNumber="547">
<emphasis id="B912C43355741105FF1DE5F5FEE8F86C" bold="true" box="[151,340,1942,1965]" pageId="16" pageNumber="547">FIGURES 59–69.</emphasis>
Larvae of
<taxonomicName id="4C6663A255741105FE4EE5F5FD64F86C" authorityName="Flowers" authorityYear="1985" box="[452,728,1942,1964]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="547" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355741105FE4EE5F5FD64F86C" box="[452,728,1942,1964]" italics="true" pageId="16" pageNumber="547">Guajirolus ektrapeloglossa</emphasis>
</taxonomicName>
from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts.
</paragraph>
</caption>
<caption id="DF1948A955751104FF1DE5D2FA21F826" httpUri="https://zenodo.org/record/2588985/files/figure.png" pageId="17" pageNumber="548" startId="17.[151,264,1969,1992]" targetBox="[159,1427,223,1913]" targetPageId="17">
<paragraph id="8BD9182155751104FF1DE5D2FA21F826" blockId="17.[151,1437,1969,2022]" pageId="17" pageNumber="548">
<emphasis id="B912C43355751104FF1DE5D2FEEFF806" bold="true" box="[151,339,1969,1992]" pageId="17" pageNumber="548">FIGURES 70–74.</emphasis>
<taxonomicName id="4C6663A255751104FED3E5D2FDD1F807" authorityName="Flowers" authorityYear="1985" box="[345,621,1969,1991]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="548" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355751104FED3E5D2FDD1F807" box="[345,621,1969,1991]" italics="true" pageId="17" pageNumber="548">Guajirolus ektrapeloglossa</emphasis>
</taxonomicName>
from Quebrada Barrigón. 70–71, male imagines; 72, subimaginal exuviae of left half of mesonotum; 73, subimaginal exuviae of left half of meso- and metapleuron; 74, subimago extracted from larval exuviae.
</paragraph>
</caption>
<caption id="DF1948A955761107FF1DE4B9FCE8F88C" httpUri="https://zenodo.org/record/2588987/files/figure.png" pageId="18" pageNumber="549" startId="18.[151,264,1754,1777]" targetBox="[158,1428,193,1733]" targetPageId="18">
<paragraph id="8BD9182155761107FF1DE4B9FCE8F88C" blockId="18.[151,1436,1754,1868]" pageId="18" pageNumber="549">
<emphasis id="B912C43355761107FF1DE4B9FEE9F92F" bold="true" box="[151,341,1754,1777]" pageId="18" pageNumber="549">FIGURES 75–80.</emphasis>
Male genitals of
<taxonomicName id="4C6663A255761107FD86E4B9FC9DF930" authorityName="Flowers" authorityYear="1985" box="[524,801,1754,1776]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="549" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355761107FD86E4B9FC9DF930" box="[524,801,1754,1776]" italics="true" pageId="18" pageNumber="549">Guajirolus ektrapeloglossa</emphasis>
</taxonomicName>
from Quebrada Barrigón. 75, genital of imago (ventral view; muscles and internal parts of penis shown by interrupted lines; right gonostylar and gonovectal muscles not shown); 76, subimaginal genitals developing under larval cuticle; 77–78 and 79–80, genitals of two specimens. Abbreviations: 1, 2, 3, segments of gonostylus; pgs, larval protogonostylus; usg, unistyliger.
</paragraph>
</caption>
<paragraph id="8BD9182155761107FF1DE517FA20F815" blockId="18.[151,1436,1907,2005]" pageId="18" pageNumber="549">
(C)
<emphasis id="B912C43355761107FF4DE510FD2EF84D" bold="true" box="[199,658,1907,1933]" pageId="18" pageNumber="549">
<emphasis id="B912C43355761107FF4DE510FD37F84C" bold="true" box="[199,651,1907,1932]" italics="true" pageId="18" pageNumber="549">Number of segments of maxillary palp</emphasis>
.
</emphasis>
<bibRefCitation id="EFF765D055761107FD11E517FC30F84D" author="Thomas, A. &amp; Dominique, Y." box="[667,908,1908,1933]" pageId="18" pageNumber="549" pagination="21 - 29" refId="ref9271" refString="Thomas, A., Dominique, Y. &amp; Orth, K. (2005) Les Ephemeres de la Guyane Francaise. 12. Description de Guajirolus flowersi n. sp. [Ephemeroptera, Baetidae]. Ephemera, (2004), 6 (1), 21 - 29." type="journal article" year="2005">
Thomas
<emphasis id="B912C43355761107FD74E516FC8EF84C" box="[766,818,1908,1932]" italics="true" pageId="18" pageNumber="549">et al</emphasis>
. (2005)
</bibRefCitation>
stated that the new species
<taxonomicName id="4C6663A255761107FB45E517FAF6F84C" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[1231,1354,1908,1932]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="549" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355761107FB45E517FAF6F84C" box="[1231,1354,1908,1932]" italics="true" pageId="18" pageNumber="549">G. flowersi</emphasis>
</taxonomicName>
differs from the earlier described species of
<taxonomicName id="4C6663A255761107FDEBE5FBFD65F870" authorityName="Flowers" authorityYear="1985" box="[609,729,1944,1968]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="549" phylum="Arthropoda" rank="genus">
<emphasis id="B912C43355761107FDEBE5FBFD65F870" box="[609,729,1944,1968]" italics="true" pageId="18" pageNumber="549">Guajirolus</emphasis>
</taxonomicName>
by 4-segmented maxillary palp, while in
<taxonomicName id="4C6663A255761107FB2DE5FBFACAF870" authorityName="Flowers" authorityYear="1985" box="[1191,1398,1944,1968]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="549" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355761107FB2DE5FBFACAF870" box="[1191,1398,1944,1968]" italics="true" pageId="18" pageNumber="549">G. ektrapeloglossa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6663A255761107FA0EE5FBFEB6F814" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="549" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355761107FA0EE5FBFEB6F814" italics="true" pageId="18" pageNumber="549">G. nanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6663A255761107FECBE5DFFE73F814" authorityName="Nieto" authorityYear="2003" box="[321,463,1980,2004]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="549" phylum="Arthropoda" rank="species" species="queremba">
<emphasis id="B912C43355761107FECBE5DFFE73F814" box="[321,463,1980,2004]" italics="true" pageId="18" pageNumber="549">G. queremba</emphasis>
</taxonomicName>
maxillary palp is 3-segmented.
<bibRefCitation id="EFF765D055761107FCB3E5DFFC72F814" author="Salles, F. F." box="[825,974,1980,2005]" pageId="18" pageNumber="549" pagination="404 - 409" refId="ref9212" refString="Salles, F. F. (2007) The presence of Chane Nieto and Guajirolus Flowers (Ephemeroptera, Baetidae) in Brazil with the description of a new species. Revista Brasileira de Entomologia, 51 (4), 404 - 409. https: // doi. org / 10.1590 / S 0085 - 56262007000400002" type="journal article" year="2007">Salles (2007)</bibRefCitation>
reported 3-segmented maxillary palp for
</paragraph>
<paragraph id="8BD9182155771106FF4DE2F4FEAEFE10" blockId="19.[199,1436,151,464]" pageId="19" pageNumber="550">
the new species
<taxonomicName id="4C6663A255771106FEF4E2FBFE4FFF70" authorityName="Salles" authorityYear="2007" box="[382,499,152,176]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="550" phylum="Arthropoda" rank="species" species="rondoni">
<emphasis id="B912C43355771106FEF4E2FBFE4FFF70" box="[382,499,152,176]" italics="true" pageId="19" pageNumber="550">G. rondoni</emphasis>
</taxonomicName>
and used this character to separate
<taxonomicName id="4C6663A255771106FC08E2FBFC45FF70" authorityName="Salles" authorityYear="2007" box="[898,1017,152,176]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="550" phylum="Arthropoda" rank="species" species="rondoni">
<emphasis id="B912C43355771106FC08E2FBFC45FF70" box="[898,1017,152,176]" italics="true" pageId="19" pageNumber="550">G. rondoni</emphasis>
</taxonomicName>
from
<taxonomicName id="4C6663A255771106FBB7E2FBFB0AFF70" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[1085,1206,152,176]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="550" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355771106FBB7E2FBFB0AFF70" box="[1085,1206,152,176]" italics="true" pageId="19" pageNumber="550">G. flowersi</emphasis>
</taxonomicName>
. Actually, maxillary palp has 2 primary segments, the 1st of which is subdivided into two secondary subsegments with one muscle passing through both of them [see
<taxonomicName id="4C6663A255771106FDE2E2BCFD58FF38" authorityName="Flowers" authorityYear="1985" box="[616,740,223,248]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="553" phylum="Arthropoda" rank="genus">Guajirolus</taxonomicName>
/g1 (6)]. The distal segment (2nd primary segment, lacking musculature) has a proximal swelling separated distally by an oblique concavity (
<figureCitation id="135D04A455771106FB03E367FB58FEDC" box="[1161,1252,260,285]" captionStart="FIGURES 17–21" captionStartId="6.[151,264,1909,1932]" captionTargetBox="[171,1428,199,1815]" captionTargetId="figure@6.[157,1429,193,1889]" captionTargetPageId="6" captionText="FIGURES 17–21. Guajirolus ektrapeloglossa from Quebrada Barrigón. 17, maxilla (dorsal view); 18, apex of maxilla (ventral view); 19, distal margin of labrum (dorsal view); 20–21, biting denticles of left and right mandibles. Abbreviations: 1d, distal subsegment of first palpomere; 1p, proximal subsegment of first palpomere; 2, second palpomere; ds1, ds2, ds3, dentisetae; in, incisor; kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2588969/files/figure.png" pageId="19" pageNumber="550">Figs 17</figureCitation>
,
<figureCitation id="135D04A455771106FB7EE367FA83FEDC" box="[1268,1343,260,284]" captionStart="FIGURES 22–28" captionStartId="7.[151,264,1417,1440]" captionTargetBox="[189,1417,267,1396]" captionTargetId="figure@7.[169,1417,267,1396]" captionTargetPageId="7" captionText="FIGURES 22–28. Guajirolus ektrapeloglossa from Quebrada Barrigón. 22–23, exuviae of maxillae; 24, maxilla; 25, exuviae of hypopharynx; 26–28, hypopharynx." httpUri="https://zenodo.org/record/2588971/files/figure.png" pageId="19" pageNumber="550">22–24</figureCitation>
); if this concavity is sharp, it can be interpreted as a joining between 3rd and 4th secondary segments; if it is shallow, it can be overlooked. The distal segment is soft, with very fine cuticle, so condition of this concavity depends on fixation. In the original description of
<taxonomicName id="4C6663A255771106FD1EE313FCD5FE48" authorityName="Flowers" authorityYear="1985" box="[660,873,368,392]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="550" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355771106FD1EE313FCD5FE48" box="[660,873,368,392]" italics="true" pageId="19" pageNumber="550">G. ektrapeloglossa</emphasis>
</taxonomicName>
, the 2nd primary segment of maxillary palp is correctly shown with obtuse-angled bent between the proximal swelling and the remainder part (
<bibRefCitation id="EFF765D055771106FB70E3F7FEB9FE10" author="Flowers, R. W." pageId="19" pageNumber="550" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers 1985: fig. 9</bibRefCitation>
).
</paragraph>
<caption id="DF1948A955771106FF1DE505FBFFF8BD" box="[151,1091,1894,1917]" httpUri="https://zenodo.org/record/2588989/files/figure.png" pageId="19" pageNumber="550" startId="19.[151,264,1894,1917]" targetBox="[158,1429,547,1844]" targetPageId="19">
<paragraph id="8BD9182155771106FF1DE505FBFFF8BD" blockId="19.[151,1091,1894,1917]" box="[151,1091,1894,1917]" pageId="19" pageNumber="550">
<emphasis id="B912C43355771106FF1DE505FEEFF8BC" bold="true" box="[151,339,1894,1917]" pageId="19" pageNumber="550">FIGURES 81–82.</emphasis>
Female imagines of
<taxonomicName id="4C6663A255771106FDADE504FC80F8BD" authorityName="Flowers" authorityYear="1985" box="[551,828,1895,1917]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="550" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355771106FDADE504FC80F8BD" box="[551,828,1895,1917]" italics="true" pageId="19" pageNumber="550">Guajirolus ektrapeloglossa</emphasis>
</taxonomicName>
from Quebrada Barrigón.
</paragraph>
</caption>
<paragraph id="8BD9182155771106FF1DE5C6FA20F821" blockId="19.[151,1436,1956,2017]" pageId="19" pageNumber="550">
(D)
<emphasis id="B912C43355771106FF4DE5C7FDDFF87D" bold="true" box="[199,611,1956,1981]" pageId="19" pageNumber="550">
<emphasis id="B912C43355771106FF4DE5C7FDE0F87D" bold="true" box="[199,604,1956,1981]" italics="true" pageId="19" pageNumber="550">Median projection of hypopharynx</emphasis>
.
</emphasis>
<bibRefCitation id="EFF765D055771106FDE0E5C7FCABF87D" author="Flowers, R. W." box="[618,791,1956,1981]" pageId="19" pageNumber="550" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers (1985)</bibRefCitation>
did not give wordily characteristic of hypopharynx of
<taxonomicName id="4C6663A255771106FA0EE5C6FEC4F821" authorityName="Flowers" authorityYear="1985" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="550" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355771106FA0EE5C6FEC4F821" italics="true" pageId="19" pageNumber="550">G. ektrapeloglossa</emphasis>
</taxonomicName>
, but only gave its drawing, on which it is terminated by three projections, with the medial
</paragraph>
<paragraph id="8BD9182155701101FF4DE2F4FA20FE6D" blockId="20.[151,1437,151,2013]" pageId="20" pageNumber="551">
projection pointed (
<bibRefCitation id="EFF765D055701101FE20E2F4FD20FF70" author="Flowers, R. W." box="[426,668,151,176]" pageId="20" pageNumber="551" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers 1985: fig. 8</bibRefCitation>
).
<bibRefCitation id="EFF765D055701101FD3FE2F4FB8AFF70" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[693,1078,151,176]" pageId="20" pageNumber="551" pagination="68 - 70" refId="ref8926" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1995) Guajirolus nanus (Ephemeroptera: Baetidae), a new species from Costa Rica. Entomological News, 106 (2), 68 - 70." type="journal article" year="1995">Lugo-Ortiz &amp; McCafferty (1995)</bibRefCitation>
stated that the new species
<taxonomicName id="4C6663A255701101FA0EE2FBFEB6FF14" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355701101FA0EE2FBFEB6FF14" italics="true" pageId="20" pageNumber="551">G. nanus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C6663A255701101FE2EE2DFFDC9FF14" authorityName="Flowers" authorityYear="1985" box="[420,629,188,212]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355701101FE2EE2DFFDC9FF14" box="[420,629,188,212]" italics="true" pageId="20" pageNumber="551">G. ektrapeloglossa</emphasis>
</taxonomicName>
by round medial projection of hypopharynx.
<bibRefCitation id="EFF765D055701101FB09E2DFFAA9FF14" author="Nieto, C." box="[1155,1301,188,213]" pageId="20" pageNumber="551" pagination="153 - 158" refId="ref8969" refString="Nieto, C. (2003) A new species of Guajirolus (Ephemeroptera: Baetidae) from Argentina and description of a new genus from Bolivia. In: Gaino E. (Ed.), Research Update on Ephemeroptera &amp; Plecoptera (Proc. 2001 Int. Joint Meeting-X Int. Conf. on Ephemeroptera &amp; XIV Int. Symp. on Plecoptera, August 5 - 11, 2001 in Perugia). Univ. Perugia, Italy, 153 - 158." type="book chapter" year="2003">Nieto (2003)</bibRefCitation>
used round medial projection to separate
<taxonomicName id="4C6663A255701101FD9DE283FDC6FF38" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[535,634,224,248]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355701101FD9DE283FDC6FF38" box="[535,634,224,248]" italics="true" pageId="20" pageNumber="551">G. nanus</emphasis>
</taxonomicName>
and the new species
<taxonomicName id="4C6663A255701101FCE4E283FC41FF38" authorityName="Nieto" authorityYear="2003" box="[878,1021,224,248]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="queremba">
<emphasis id="B912C43355701101FCE4E283FC41FF38" box="[878,1021,224,248]" italics="true" pageId="20" pageNumber="551">G. queremba</emphasis>
</taxonomicName>
from
<taxonomicName id="4C6663A255701101FBCEE283FAA9FF38" authorityName="Flowers" authorityYear="1985" box="[1092,1301,224,248]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355701101FBCEE283FAA9FF38" box="[1092,1301,224,248]" italics="true" pageId="20" pageNumber="551">G. ektrapeloglossa</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFF765D055701101FAAFE2BCFE8BFEDD" author="Thomas, A. &amp; Dominique, Y." pageId="20" pageNumber="551" pagination="21 - 29" refId="ref9271" refString="Thomas, A., Dominique, Y. &amp; Orth, K. (2005) Les Ephemeres de la Guyane Francaise. 12. Description de Guajirolus flowersi n. sp. [Ephemeroptera, Baetidae]. Ephemera, (2004), 6 (1), 21 - 29." type="journal article" year="2005">
Thomas
<emphasis id="B912C43355701101FA02E282FF61FEDC" italics="true" pageId="20" pageNumber="551">et al</emphasis>
. (2005)
</bibRefCitation>
stated that the new species
<taxonomicName id="4C6663A255701101FDF6E367FD4BFEDC" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[636,759,260,284]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355701101FDF6E367FD4BFEDC" box="[636,759,260,284]" italics="true" pageId="20" pageNumber="551">G. flowersi</emphasis>
</taxonomicName>
has medial projection reduced and flattened, in contrast to pointed in
<taxonomicName id="4C6663A255701101FEB7E34BFDB0FE80" authorityName="Flowers" authorityYear="1985" box="[317,524,296,320]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355701101FEB7E34BFDB0FE80" box="[317,524,296,320]" italics="true" pageId="20" pageNumber="551">G. ektrapeloglossa</emphasis>
</taxonomicName>
and rounded in
<taxonomicName id="4C6663A255701101FD4AE34BFC9DFE80" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[704,801,296,320]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355701101FD4AE34BFC9DFE80" box="[704,801,296,320]" italics="true" pageId="20" pageNumber="551">G. nanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6663A255701101FCDFE34BFC5EFE80" authorityName="Nieto" authorityYear="2003" box="[853,994,296,320]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="queremba">
<emphasis id="B912C43355701101FCDFE34BFC5EFE80" box="[853,994,296,320]" italics="true" pageId="20" pageNumber="551">G. queremba</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFF765D055701101FC7AE344FB38FE80" author="Salles, F. F." box="[1008,1156,295,320]" pageId="20" pageNumber="551" pagination="404 - 409" refId="ref9212" refString="Salles, F. F. (2007) The presence of Chane Nieto and Guajirolus Flowers (Ephemeroptera, Baetidae) in Brazil with the description of a new species. Revista Brasileira de Entomologia, 51 (4), 404 - 409. https: // doi. org / 10.1590 / S 0085 - 56262007000400002" type="journal article" year="2007">Salles (2007)</bibRefCitation>
reported rounded medial projection for the new species
<taxonomicName id="4C6663A255701101FD97E32FFD2EFEA4" authorityName="Salles" authorityYear="2007" box="[541,658,332,356]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="rondoni">
<emphasis id="B912C43355701101FD97E32FFD2EFEA4" box="[541,658,332,356]" italics="true" pageId="20" pageNumber="551">G. rondoni</emphasis>
</taxonomicName>
. Actually, the area between paired apical projections of hypopharynx is soft and has no constant shape; depending on condition of specimen, it can be widened and flattened (
<figureCitation id="135D04A455701101FAE5E30CFF5AFE6C" captionStart="FIGURES 22–28" captionStartId="7.[151,264,1417,1440]" captionTargetBox="[189,1417,267,1396]" captionTargetId="figure@7.[169,1417,267,1396]" captionTargetPageId="7" captionText="FIGURES 22–28. Guajirolus ektrapeloglossa from Quebrada Barrigón. 22–23, exuviae of maxillae; 24, maxilla; 25, exuviae of hypopharynx; 26–28, hypopharynx." httpUri="https://zenodo.org/record/2588971/files/figure.png" pageId="20" pageNumber="551">Fig. 28</figureCitation>
), or narrowed forming a median projection, which can be either rounded (
<figureCitation id="135D04A455701101FBA9E3F7FBCBFE6C" box="[1059,1143,404,429]" captionStart="FIGURES 22–28" captionStartId="7.[151,264,1417,1440]" captionTargetBox="[189,1417,267,1396]" captionTargetId="figure@7.[169,1417,267,1396]" captionTargetPageId="7" captionText="FIGURES 22–28. Guajirolus ektrapeloglossa from Quebrada Barrigón. 22–23, exuviae of maxillae; 24, maxilla; 25, exuviae of hypopharynx; 26–28, hypopharynx." httpUri="https://zenodo.org/record/2588971/files/figure.png" pageId="20" pageNumber="551">Fig. 27</figureCitation>
), or pointed (
<figureCitation id="135D04A455701101FA86E3F7FA2CFE6D" box="[1292,1424,404,429]" captionStart="FIGURES 22–28" captionStartId="7.[151,264,1417,1440]" captionTargetBox="[189,1417,267,1396]" captionTargetId="figure@7.[169,1417,267,1396]" captionTargetPageId="7" captionText="FIGURES 22–28. Guajirolus ektrapeloglossa from Quebrada Barrigón. 22–23, exuviae of maxillae; 24, maxilla; 25, exuviae of hypopharynx; 26–28, hypopharynx." httpUri="https://zenodo.org/record/2588971/files/figure.png" pageId="20" pageNumber="551">Figs 25–26</figureCitation>
).
</paragraph>
<paragraph id="8BD9182155701101FF1DE3DBFD35FCF7" blockId="20.[151,1437,151,2013]" pageId="20" pageNumber="551">
(E)
<emphasis id="B912C43355701101FF4DE3D4FD20FE10" bold="true" box="[199,668,439,464]" pageId="20" pageNumber="551">
<emphasis id="B912C43355701101FF4DE3D4FD2AFE10" bold="true" box="[199,662,439,464]" italics="true" pageId="20" pageNumber="551">Projection of 2nd segment of labial palp</emphasis>
.
</emphasis>
Labial palp has unusually massive projection directed medially [see
<taxonomicName id="4C6663A255701101FF4DE3BFFEFFFE35" authorityName="Flowers" authorityYear="1985" box="[199,323,476,501]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="553" phylum="Arthropoda" rank="genus">Guajirolus</taxonomicName>
/g1 (7)].
<bibRefCitation id="EFF765D055701101FE29E3BFFD2DFE35" author="Thomas, A. &amp; Dominique, Y." box="[419,657,476,501]" pageId="20" pageNumber="551" pagination="21 - 29" refId="ref9271" refString="Thomas, A., Dominique, Y. &amp; Orth, K. (2005) Les Ephemeres de la Guyane Francaise. 12. Description de Guajirolus flowersi n. sp. [Ephemeroptera, Baetidae]. Ephemera, (2004), 6 (1), 21 - 29." type="journal article" year="2005">
Thomas
<emphasis id="B912C43355701101FD8FE3BEFD8AFE34" box="[517,566,476,500]" italics="true" pageId="20" pageNumber="551">et al</emphasis>
. (2005)
</bibRefCitation>
stated that in the new species
<taxonomicName id="4C6663A255701101FC64E3BFFBD4FE34" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[1006,1128,476,500]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355701101FC64E3BFFBD4FE34" box="[1006,1128,476,500]" italics="true" pageId="20" pageNumber="551">G. flowersi</emphasis>
</taxonomicName>
this projection is narrower than in the earlier described
<taxonomicName id="4C6663A255701101FD87E063FD63FDD8" authorityName="Flowers" authorityYear="1985" box="[525,735,512,536]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355701101FD87E063FD63FDD8" box="[525,735,512,536]" italics="true" pageId="20" pageNumber="551">G. ektrapeloglossa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6663A255701101FD64E063FCEEFDD8" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[750,850,512,536]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355701101FD64E063FCEEFDD8" box="[750,850,512,536]" italics="true" pageId="20" pageNumber="551">G. nanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6663A255701101FC06E063FBA7FDD8" authorityName="Nieto" authorityYear="2003" box="[908,1051,512,536]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="queremba">
<emphasis id="B912C43355701101FC06E063FBA7FDD8" box="[908,1051,512,536]" italics="true" pageId="20" pageNumber="551">G. queremba</emphasis>
</taxonomicName>
. On their drawing (
<bibRefCitation id="EFF765D055701101FB74E39CFEBFFDFD" author="Thomas, A. &amp; Dominique, Y." pageId="20" pageNumber="551" pagination="21 - 29" refId="ref9271" refString="Thomas, A., Dominique, Y. &amp; Orth, K. (2005) Les Ephemeres de la Guyane Francaise. 12. Description de Guajirolus flowersi n. sp. [Ephemeroptera, Baetidae]. Ephemera, (2004), 6 (1), 21 - 29." type="journal article" year="2005">
Thomas
<emphasis id="B912C43355701101FAE8E062FA29FDD8" box="[1378,1429,512,536]" italics="true" pageId="20" pageNumber="551">et al</emphasis>
. 2005
</bibRefCitation>
: fig. 6) projection of left palp has apex as broad as base, and projection of right palp has apex narrower than base.
<bibRefCitation id="EFF765D055701101FEB4E024FE6FFDA0" author="Salles, F. F." box="[318,467,583,608]" pageId="20" pageNumber="551" pagination="404 - 409" refId="ref9212" refString="Salles, F. F. (2007) The presence of Chane Nieto and Guajirolus Flowers (Ephemeroptera, Baetidae) in Brazil with the description of a new species. Revista Brasileira de Entomologia, 51 (4), 404 - 409. https: // doi. org / 10.1590 / S 0085 - 56262007000400002" type="journal article" year="2007">Salles (2007)</bibRefCitation>
stated that
<taxonomicName id="4C6663A255701101FDDEE02BFD71FDA0" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[596,717,584,608]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355701101FDDEE02BFD71FDA0" box="[596,717,584,608]" italics="true" pageId="20" pageNumber="551">G. flowersi</emphasis>
</taxonomicName>
and the new species
<taxonomicName id="4C6663A255701101FC31E02BFB8DFDA0" authorityName="Salles" authorityYear="2007" box="[955,1073,584,608]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="rondoni">
<emphasis id="B912C43355701101FC31E02BFB8DFDA0" box="[955,1073,584,608]" italics="true" pageId="20" pageNumber="551">G. rondoni</emphasis>
</taxonomicName>
differ from others by projection of the 2nd segment of labial palp with apex narrower than base. The 2nd segment of labial palp is soft and sack-like, so its shape has individual variability (
<figureCitation id="135D04A455701101FC81E0ECFC2CFD68" box="[779,912,655,680]" captionStart="FIGURES 29–35" captionStartId="8.[151,264,1275,1298]" captionTargetBox="[158,1428,195,1254]" captionTargetId="figure@8.[158,1428,195,1254]" captionTargetPageId="8" captionText="FIGURES 29–35. Labium of Guajirolus ektrapeloglossa from Quebrada Barrigón. 29–31, labium (29, ventral view; 30–31, in translucent light, with focus on glossae and on palps); 32–33, one and the same labial palp differently compressed; 34–35, left and right labial palps of one and the same individual." httpUri="https://zenodo.org/record/2588973/files/figure.png" pageId="20" pageNumber="551">Figs 34–35</figureCitation>
) and depends upon flattening on slide (
<figureCitation id="135D04A455701101FAE0E0ECFEAEFD0C" captionStart="FIGURES 29–35" captionStartId="8.[151,264,1275,1298]" captionTargetBox="[158,1428,195,1254]" captionTargetId="figure@8.[158,1428,195,1254]" captionTargetPageId="8" captionText="FIGURES 29–35. Labium of Guajirolus ektrapeloglossa from Quebrada Barrigón. 29–31, labium (29, ventral view; 30–31, in translucent light, with focus on glossae and on palps); 32–33, one and the same labial palp differently compressed; 34–35, left and right labial palps of one and the same individual." httpUri="https://zenodo.org/record/2588973/files/figure.png" pageId="20" pageNumber="551">Figs 32–33</figureCitation>
). According to the figure of
<taxonomicName id="4C6663A255701101FDD8E0D7FD77FD0C" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[594,715,692,716]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355701101FDD8E0D7FD77FD0C" box="[594,715,692,716]" italics="true" pageId="20" pageNumber="551">G. flowersi</emphasis>
</taxonomicName>
, it has no significant difference from
<taxonomicName id="4C6663A255701101FBFCE0D7FAFAFD0C" authorityName="Flowers" authorityYear="1985" box="[1142,1350,692,716]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355701101FBFCE0D7FAFAFD0C" box="[1142,1350,692,716]" italics="true" pageId="20" pageNumber="551">G. ektrapeloglossa</emphasis>
</taxonomicName>
, but on the drawing of
<taxonomicName id="4C6663A255701101FEF7E0BBFE44FD30" authorityName="Salles" authorityYear="2007" box="[381,504,728,752]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="rondoni">
<emphasis id="B912C43355701101FEF7E0BBFE44FD30" box="[381,504,728,752]" italics="true" pageId="20" pageNumber="551">G. rondoni</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF765D055701101FD81E0B4FD2BFD30" author="Salles, F. F." box="[523,663,727,752]" pageId="20" pageNumber="551" pagination="404 - 409" refId="ref9212" refString="Salles, F. F. (2007) The presence of Chane Nieto and Guajirolus Flowers (Ephemeroptera, Baetidae) in Brazil with the description of a new species. Revista Brasileira de Entomologia, 51 (4), 404 - 409. https: // doi. org / 10.1590 / S 0085 - 56262007000400002" type="journal article" year="2007">Salles 2007</bibRefCitation>
: fig. 8;
<bibRefCitation id="EFF765D055701101FD72E0B4FC6CFD30" author="Falcao, J. N. &amp; Salles, F. F. &amp; Hamada, N." box="[760,976,727,752]" pageId="20" pageNumber="551" pagination="516 - 548" refId="ref8527" refString="Falcao, J. N., Salles, F. F. &amp; Hamada, N. (2011) Baetidae (Insecta, Ephemeroptera) ocorrentes em Roraima, Brasil: novos registros e chaves para generos e especies no estagio ninfal. Revista Brasileira de Entomologia, 55, 516 - 548. https: // doi. org / 10.1590 / S 0085 - 56262011005000048" type="journal article" year="2011">
Falcão
<emphasis id="B912C43355701101FCC7E0BAFC3FFD30" box="[845,899,728,752]" italics="true" pageId="20" pageNumber="551">et al</emphasis>
. 2011
</bibRefCitation>
: fig. 111) projection of 2nd segment is narrower than in examined specimens from
<collectingCountry id="F37158B155701101FD37E09EFCAAFCD4" box="[701,790,765,788]" name="Panama" pageId="20" pageNumber="551">Panama</collectingCountry>
and
<collectingCountry id="F37158B155701101FCC5E09EFC3AFCD4" box="[847,902,765,788]" name="Peru" pageId="20" pageNumber="551">Peru</collectingCountry>
; this is the single known character in which
<taxonomicName id="4C6663A255701101FA0EE09FFEA3FCF8" authorityName="Salles" authorityYear="2007" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="rondoni">
<emphasis id="B912C43355701101FA0EE09FFEA3FCF8" italics="true" pageId="20" pageNumber="551">G. rondoni</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C6663A255701101FE38E143FD3EFCF8" authorityName="Flowers" authorityYear="1985" box="[434,642,800,824]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355701101FE38E143FD3EFCF8" box="[434,642,800,824]" italics="true" pageId="20" pageNumber="551">G. ektrapeloglossa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD9182155701101FF1DE127FD24FBF4" blockId="20.[151,1437,151,2013]" pageId="20" pageNumber="551">
(F)
<emphasis id="B912C43355701101FF4DE127FD9BFC9D" bold="true" box="[199,551,835,861]" pageId="20" pageNumber="551">
<emphasis id="B912C43355701101FF4DE127FD9DFC9C" bold="true" box="[199,545,835,860]" italics="true" pageId="20" pageNumber="551">Brown bands on larval femur</emphasis>
.
</emphasis>
<bibRefCitation id="EFF765D055701101FDBAE127FD62FC9D" author="Flowers, R. W." box="[560,734,836,861]" pageId="20" pageNumber="551" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers (1985)</bibRefCitation>
reported and figured one brown band on larval femur. Lugo-
<bibRefCitation id="EFF765D055701101FF4DE104FE4BFC40" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[199,503,871,896]" pageId="20" pageNumber="551" pagination="68 - 70" refId="ref8926" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1995) Guajirolus nanus (Ephemeroptera: Baetidae), a new species from Costa Rica. Entomological News, 106 (2), 68 - 70." type="journal article" year="1995">Ortiz &amp; McCafferty (1995)</bibRefCitation>
stated that the new species
<taxonomicName id="4C6663A255701101FCA7E10BFC33FC40" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[813,911,872,896]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355701101FCA7E10BFC33FC40" box="[813,911,872,896]" italics="true" pageId="20" pageNumber="551">G. nanus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C6663A255701101FBA8E10BFB4FFC40" authorityName="Flowers" authorityYear="1985" box="[1058,1267,872,896]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355701101FBA8E10BFB4FFC40" box="[1058,1267,872,896]" italics="true" pageId="20" pageNumber="551">G. ektrapeloglossa</emphasis>
</taxonomicName>
by presence of two brown bands on femora.
<bibRefCitation id="EFF765D055701101FD9BE1EFFD1EFC64" author="Nieto, C." box="[529,674,908,933]" pageId="20" pageNumber="551" pagination="153 - 158" refId="ref8969" refString="Nieto, C. (2003) A new species of Guajirolus (Ephemeroptera: Baetidae) from Argentina and description of a new genus from Bolivia. In: Gaino E. (Ed.), Research Update on Ephemeroptera &amp; Plecoptera (Proc. 2001 Int. Joint Meeting-X Int. Conf. on Ephemeroptera &amp; XIV Int. Symp. on Plecoptera, August 5 - 11, 2001 in Perugia). Univ. Perugia, Italy, 153 - 158." type="book chapter" year="2003">Nieto (2003)</bibRefCitation>
and
<bibRefCitation id="EFF765D055701101FD53E1EFFC79FC65" author="Thomas, A. &amp; Dominique, Y." box="[729,965,908,933]" pageId="20" pageNumber="551" pagination="21 - 29" refId="ref9271" refString="Thomas, A., Dominique, Y. &amp; Orth, K. (2005) Les Ephemeres de la Guyane Francaise. 12. Description de Guajirolus flowersi n. sp. [Ephemeroptera, Baetidae]. Ephemera, (2004), 6 (1), 21 - 29." type="journal article" year="2005">
Thomas
<emphasis id="B912C43355701101FCB1E1EEFCD0FC64" box="[827,876,908,932]" italics="true" pageId="20" pageNumber="551">et al</emphasis>
. (2005)
</bibRefCitation>
reported one brown band on larval femur of
<taxonomicName id="4C6663A255701101FF6FE1D3FECEFC08" authorityName="Nieto" authorityYear="2003" box="[229,370,944,968]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="queremba">
<emphasis id="B912C43355701101FF6FE1D3FECEFC08" box="[229,370,944,968]" italics="true" pageId="20" pageNumber="551">G. queremba</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6663A255701101FE2DE1D3FDA3FC08" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[423,543,944,968]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355701101FE2DE1D3FDA3FC08" box="[423,543,944,968]" italics="true" pageId="20" pageNumber="551">G. flowersi</emphasis>
</taxonomicName>
, and used this character to separate these species from
<taxonomicName id="4C6663A255701101FB0CE1D3FB54FC08" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[1158,1256,944,968]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355701101FB0CE1D3FB54FC08" box="[1158,1256,944,968]" italics="true" pageId="20" pageNumber="551">G. nanus</emphasis>
</taxonomicName>
. Actually larval femur has more intensively colored subapical brown band and less intensive proximal brown coloration, which can be either interpreted as a second band, or not (
<figureCitation id="135D04A455701101FCA7E194FC0AFBD0" box="[813,950,1015,1040]" captionStart="FIGURES 44–50" captionStartId="14.[151,264,1937,1960]" captionTargetBox="[156,1430,258,1857]" captionTargetId="figure@14.[156,1430,258,1857]" captionTargetPageId="14" captionText="FIGURES 44–50. Larval legs of Guajirolus ektrapeloglossa from Quebrada Barrigón. 44–45, fore and middle legs of darkest specimen; 46–47, claws; 48, fragment of fore femur (focus on anterior side); 49, fragment of fore tibia (focus on anterior side); 50, fragment of hind tibia (focus on posterior side)." httpUri="https://zenodo.org/record/2588979/files/figure.png" pageId="20" pageNumber="551">Figs 44–45</figureCitation>
); if cuticular coloration of legs is poorly expressed, only the distal band is present.
</paragraph>
<paragraph id="8BD9182155701101FF1DE623FDB9FAF0" blockId="20.[151,1437,151,2013]" pageId="20" pageNumber="551">
(G)
<emphasis id="B912C43355701101FF4DE65CFCDEFB98" bold="true" box="[199,866,1087,1112]" pageId="20" pageNumber="551">
<emphasis id="B912C43355701101FF4DE65CFCE6FB97" bold="true" box="[199,858,1087,1112]" italics="true" pageId="20" pageNumber="551">Light medial longitudinal strip on abdominal terga I–IX</emphasis>
.
</emphasis>
<bibRefCitation id="EFF765D055701101FCE0E65CFBA4FB98" author="Flowers, R. W." box="[874,1048,1087,1112]" pageId="20" pageNumber="551" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers (1985)</bibRefCitation>
figured integral light stripe on all abdominal terga I–IX (
<bibRefCitation id="EFF765D055701101FE59E607FD7BFBBC" author="Flowers, R. W." box="[467,711,1124,1149]" pageId="20" pageNumber="551" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers 1985: fig. 4</bibRefCitation>
), but in the text wrote: «Abdomen brown; terga with median yellow bands which sometimes do not reach posterior margin».
<bibRefCitation id="EFF765D055701101FC31E6E4FB0EFB60" author="Thomas, A. &amp; Dominique, Y." box="[955,1202,1159,1184]" pageId="20" pageNumber="551" pagination="21 - 29" refId="ref9271" refString="Thomas, A., Dominique, Y. &amp; Orth, K. (2005) Les Ephemeres de la Guyane Francaise. 12. Description de Guajirolus flowersi n. sp. [Ephemeroptera, Baetidae]. Ephemera, (2004), 6 (1), 21 - 29." type="journal article" year="2005">
Thomas
<emphasis id="B912C43355701101FBAAE6EAFBE9FB60" box="[1056,1109,1160,1184]" italics="true" pageId="20" pageNumber="551">et al</emphasis>
. (2005)
</bibRefCitation>
stated that the new species
<taxonomicName id="4C6663A255701101FE94E6CFFE2AFB04" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[286,406,1196,1220]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355701101FE94E6CFFE2AFB04" box="[286,406,1196,1220]" italics="true" pageId="20" pageNumber="551">G. flowersi</emphasis>
</taxonomicName>
differs from other species of
<taxonomicName id="4C6663A255701101FD54E6CFFCEAFB04" authorityName="Flowers" authorityYear="1985" box="[734,854,1196,1220]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="B912C43355701101FD54E6CFFCEAFB04" box="[734,854,1196,1220]" italics="true" pageId="20" pageNumber="551">Guajirolus</emphasis>
</taxonomicName>
by absence of this band, and described light median blanks on certain terga (without illustration). Actually, unpaired colorless blanks on dark cuticle of abdominal terga greatly vary individually, sometimes forming integral longitudinal stripe, sometimes not (
<figureCitation id="135D04A455701101FA98E697FA2BFACD" box="[1298,1431,1268,1293]" captionStart="FIGURES 51–58" captionStartId="15.[151,264,2007,2030]" captionTargetBox="[156,1430,243,1914]" captionTargetId="figure@15.[156,1430,243,1914]" captionTargetPageId="15" captionText="FIGURES 51–58. Larval coloration of Guajirolus ektrapeloglossa from Quebrada Barrigón." httpUri="https://zenodo.org/record/2588981/files/figure.png" pageId="20" pageNumber="551">Figs 51–58</figureCitation>
;
<bibRefCitation id="EFF765D055701101FF4DE774FE2EFAF0" author="Falcao, J. N. &amp; Salles, F. F. &amp; Hamada, N." box="[199,402,1303,1328]" pageId="20" pageNumber="551" pagination="516 - 548" refId="ref8527" refString="Falcao, J. N., Salles, F. F. &amp; Hamada, N. (2011) Baetidae (Insecta, Ephemeroptera) ocorrentes em Roraima, Brasil: novos registros e chaves para generos e especies no estagio ninfal. Revista Brasileira de Entomologia, 55, 516 - 548. https: // doi. org / 10.1590 / S 0085 - 56262011005000048" type="journal article" year="2011">
Falcão
<emphasis id="B912C43355701101FE92E77AFEF6FAF0" box="[280,330,1304,1328]" italics="true" pageId="20" pageNumber="551">et al</emphasis>
. 2011
</bibRefCitation>
: fig. 105).
</paragraph>
<paragraph id="8BD9182155701101FF1DE75FFE46F920" blockId="20.[151,1437,151,2013]" pageId="20" pageNumber="551">
(H)
<emphasis id="B912C43355701101FF4DE758FE5EFA95" bold="true" box="[199,482,1339,1365]" pageId="20" pageNumber="551">
<emphasis id="B912C43355701101FF4DE758FE67FA93" bold="true" box="[199,475,1339,1364]" italics="true" pageId="20" pageNumber="551">Denticles on paraprocts</emphasis>
.
</emphasis>
<bibRefCitation id="EFF765D055701101FE63E75FFD2AFA95" author="Flowers, R. W." box="[489,662,1340,1365]" pageId="20" pageNumber="551" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers (1985)</bibRefCitation>
did not give wordily description of paraproct of
<taxonomicName id="4C6663A255701101FB4EE75FFA29FA94" authorityName="Flowers" authorityYear="1985" box="[1220,1429,1340,1364]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355701101FB4EE75FFA29FA94" box="[1220,1429,1340,1364]" italics="true" pageId="20" pageNumber="551">G. ektrapeloglossa</emphasis>
</taxonomicName>
, but gave its drawing (
<bibRefCitation id="EFF765D055701101FE47E73CFD6EFAB8" author="Flowers, R. W." box="[461,722,1375,1400]" pageId="20" pageNumber="551" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers 1985: fig. 17</bibRefCitation>
).
<bibRefCitation id="EFF765D055701101FD66E73CFBCCFAB8" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[748,1136,1375,1400]" pageId="20" pageNumber="551" pagination="68 - 70" refId="ref8926" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1995) Guajirolus nanus (Ephemeroptera: Baetidae), a new species from Costa Rica. Entomological News, 106 (2), 68 - 70." type="journal article" year="1995">Lugo-Ortiz &amp; McCafferty (1995)</bibRefCitation>
interpreted this figure as «paraprocts with unorganized spination» and stated that the new species
<taxonomicName id="4C6663A255701101FB8AE7E7FBDFFA5C" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[1024,1123,1412,1436]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355701101FB8AE7E7FBDFFA5C" box="[1024,1123,1412,1436]" italics="true" pageId="20" pageNumber="551">G. nanus</emphasis>
</taxonomicName>
differs by «paraprocts with organized spination». They did not explain, what means «organized» and «unorganized» spination.
<bibRefCitation id="EFF765D055701101FAD4E7C4FEADFA24" author="Nieto, C." pageId="20" pageNumber="551" pagination="153 - 158" refId="ref8969" refString="Nieto, C. (2003) A new species of Guajirolus (Ephemeroptera: Baetidae) from Argentina and description of a new genus from Bolivia. In: Gaino E. (Ed.), Research Update on Ephemeroptera &amp; Plecoptera (Proc. 2001 Int. Joint Meeting-X Int. Conf. on Ephemeroptera &amp; XIV Int. Symp. on Plecoptera, August 5 - 11, 2001 in Perugia). Univ. Perugia, Italy, 153 - 158." type="book chapter" year="2003">Nieto (2003)</bibRefCitation>
separated
<taxonomicName id="4C6663A255701101FE1EE7AFFE47FA24" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[404,507,1484,1508]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355701101FE1EE7AFFE47FA24" box="[404,507,1484,1508]" italics="true" pageId="20" pageNumber="551">G. nanus</emphasis>
</taxonomicName>
and the new species
<taxonomicName id="4C6663A255701101FC8EE7AFFC2BFA24" authorityName="Nieto" authorityYear="2003" box="[772,919,1484,1508]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="queremba">
<emphasis id="B912C43355701101FC8EE7AFFC2BFA24" box="[772,919,1484,1508]" italics="true" pageId="20" pageNumber="551">G. queremba</emphasis>
</taxonomicName>
from
<taxonomicName id="4C6663A255701101FC6DE7AFFB00FA24" authorityName="Flowers" authorityYear="1985" box="[999,1212,1484,1508]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355701101FC6DE7AFFB00FA24" box="[999,1212,1484,1508]" italics="true" pageId="20" pageNumber="551">G. ektrapeloglossa</emphasis>
</taxonomicName>
by the «organized spination».
<bibRefCitation id="EFF765D055701101FEC0E78CFD8BF9C8" author="Thomas, A. &amp; Dominique, Y." box="[330,567,1519,1544]" pageId="20" pageNumber="551" pagination="21 - 29" refId="ref9271" refString="Thomas, A., Dominique, Y. &amp; Orth, K. (2005) Les Ephemeres de la Guyane Francaise. 12. Description de Guajirolus flowersi n. sp. [Ephemeroptera, Baetidae]. Ephemera, (2004), 6 (1), 21 - 29." type="journal article" year="2005">
Thomas
<emphasis id="B912C43355701101FE26E792FE62F9C8" box="[428,478,1520,1544]" italics="true" pageId="20" pageNumber="551">et al</emphasis>
. (2005)
</bibRefCitation>
reported for the new species
<taxonomicName id="4C6663A255701101FC0FE793FC42F9C8" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[901,1022,1520,1544]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355701101FC0FE793FC42F9C8" box="[901,1022,1520,1544]" italics="true" pageId="20" pageNumber="551">G. flowersi</emphasis>
</taxonomicName>
5–6 denticles on paraproct and used this character to separate this species from all others, for which they report from 11 to 22 denticles based on descriptions and/or figures.
<bibRefCitation id="EFF765D055701101FD89E454FD24F990" author="Salles, F. F." box="[515,664,1591,1616]" pageId="20" pageNumber="551" pagination="404 - 409" refId="ref9212" refString="Salles, F. F. (2007) The presence of Chane Nieto and Guajirolus Flowers (Ephemeroptera, Baetidae) in Brazil with the description of a new species. Revista Brasileira de Entomologia, 51 (4), 404 - 409. https: // doi. org / 10.1590 / S 0085 - 56262007000400002" type="journal article" year="2007">Salles (2007)</bibRefCitation>
stated that in the new species
<taxonomicName id="4C6663A255701101FC7EE45BFBD6F990" authorityName="Salles" authorityYear="2007" box="[1012,1130,1592,1616]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="rondoni">
<emphasis id="B912C43355701101FC7EE45BFBD6F990" box="[1012,1130,1592,1616]" italics="true" pageId="20" pageNumber="551">G. rondoni</emphasis>
</taxonomicName>
paraproct has «organized» spination with about 15 spines, and used these characters («organized» and «unorganized» spination and number of spines) in the species key. Actually paraprocts of
<taxonomicName id="4C6663A255701101FC2BE4E3FBCAF958" authorityName="Flowers" authorityYear="1985" box="[929,1142,1664,1688]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355701101FC2BE4E3FBCAF958" box="[929,1142,1664,1688]" italics="true" pageId="20" pageNumber="551">G. ektrapeloglossa</emphasis>
</taxonomicName>
have normally about 15 denticles, varying approximately from 10 to 20, among which apical denticles are larger, and proximal gradually smaller (
<figureCitation id="135D04A455701101FE13E4A4FE52F920" box="[409,494,1735,1760]" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="20" pageNumber="551">Fig. 69</figureCitation>
).
</paragraph>
<paragraph id="8BD9182155701100FF1DE48FFE4AFF38" blockId="20.[151,1437,151,2013]" lastBlockId="21.[199,1436,151,248]" lastPageId="21" lastPageNumber="552" pageId="20" pageNumber="551">
(J)
<emphasis id="B912C43355701101FF4DE488FE11F8C5" bold="true" box="[199,429,1771,1797]" pageId="20" pageNumber="551">
<emphasis id="B912C43355701101FF4DE488FE1BF8C4" bold="true" box="[199,423,1771,1796]" italics="true" pageId="20" pageNumber="551">Number of tergalii</emphasis>
.
</emphasis>
<bibRefCitation id="EFF765D055701101FE32E48FFDD8F8C5" author="Roback, S. S." box="[440,612,1772,1797]" pageId="20" pageNumber="551" pagination="129 - 199" refId="ref9176" refString="Roback, S. S. (1966) The Catherwood Foundation Peruvia-Amazon expedition. VI-Ephemeroptera nymphs. Monographs of the Academy of Natural Sciences of Philadelphia, No. 14, 129 - 199." type="book chapter" year="1966">Roback (1966)</bibRefCitation>
described larva of
<taxonomicName id="4C6663A255701101FCC6E48FFC78F8C4" authorityName="Flowers" authorityYear="1985" box="[844,964,1772,1796]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="B912C43355701101FCC6E48FFC78F8C4" box="[844,964,1772,1796]" italics="true" pageId="20" pageNumber="551">Guajirolus</emphasis>
</taxonomicName>
from the river Tulumayo in
<collectingCountry id="F37158B155701101FA97E48EFAECF8C4" box="[1309,1360,1773,1796]" name="Peru" pageId="20" pageNumber="551">Peru</collectingCountry>
under arbitrary name «Genus 3 nr.
<taxonomicName id="4C6663A255701101FD8FE573FD1EF8E8" authorityName=": Roback" authorityYear="1966" box="[517,674,1808,1832]" class="Insecta" family="Baetidae" genus="Pseudocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="B912C43355701101FD8FE573FD1EF8E8" box="[517,674,1808,1832]" italics="true" pageId="20" pageNumber="551">Pseudocloeon</emphasis>
</taxonomicName>
» and reported presence of all 7 pairs of tergalii as «darker gills,
<figureCitation id="135D04A455701101FAFAE56CFF5AF88D" captionStart="FIGURES 59–69" captionStartId="16.[151,264,1942,1965]" captionTargetBox="[178,1421,193,1929]" captionTargetId="figure@16.[176,1421,193,1929]" captionTargetPageId="16" captionText="FIGURES 59–69. Larvae of Guajirolus ektrapeloglossa from Quebrada Barrigón. 59, middle part of cercus; 60–61, scales on abdominal terga; 62, posterior margin of tergum II; 63, posterior margin of tergum VIII; 64, posterior margin of sternum VIII; 56, posterior margin of sternum IX of female; 66, tergalii VI–VII; 67–68, tergalius III; 59, paraprocts." httpUri="https://zenodo.org/record/2588983/files/figure.png" pageId="20" pageNumber="551">Fig. 65</figureCitation>
, ovate, single on 1–7» (
<bibRefCitation id="EFF765D055701101FE7BE557FD79F88D" author="Roback, S. S." box="[497,709,1844,1869]" pageId="20" pageNumber="551" pagination="129 - 199" refId="ref9176" refString="Roback, S. S. (1966) The Catherwood Foundation Peruvia-Amazon expedition. VI-Ephemeroptera nymphs. Monographs of the Academy of Natural Sciences of Philadelphia, No. 14, 129 - 199." type="book chapter" year="1966">Roback 1966: 136</bibRefCitation>
).
<bibRefCitation id="EFF765D055701101FD52E557FC38F88D" author="Flowers, R. W." box="[728,900,1844,1869]" pageId="20" pageNumber="551" pagination="27 - 31" refId="ref8596" refString="Flowers, R. W. (1985) Guajirolus, a new genus of Neotropical Baetidae (Ephemeroptera). Studies on the Neotropical Fauna and Environment, 20 (1), 27 - 31. https: // doi. org / 10.1080 / 01650528509360667" type="journal article" year="1985">Flowers (1985)</bibRefCitation>
wrote: «The Peruvian nymphs are not included in
<taxonomicName id="4C6663A255701101FF6EE53BFE0FF8B0" authorityName="Flowers" authorityYear="1985" box="[228,435,1880,1904]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355701101FF6EE53BFE0FF8B0" box="[228,435,1880,1904]" italics="true" pageId="20" pageNumber="551">G. ektrapeloglossa</emphasis>
</taxonomicName>
because they have gills on abdominal segment 1 (
<bibRefCitation id="EFF765D055701101FC6FE534FB3CF8B0" author="Roback, S. S." box="[997,1152,1879,1904]" pageId="20" pageNumber="551" pagination="129 - 199" refId="ref9176" refString="Roback, S. S. (1966) The Catherwood Foundation Peruvia-Amazon expedition. VI-Ephemeroptera nymphs. Monographs of the Academy of Natural Sciences of Philadelphia, No. 14, 129 - 199." type="book chapter" year="1966">Roback 1966</bibRefCitation>
)...». In the generic key to South American mayflies (
<bibRefCitation id="EFF765D055701101FE76E51FFCB4F854" author="Dominguez, E. &amp; Hubbard, M. D. &amp; Peters, W. L." box="[508,776,1916,1940]" pageId="20" pageNumber="551" pagination="1 - 32" refId="ref8421" refString="Dominguez, E., Hubbard, M. D. &amp; Peters, W. L. (1992) Clave para ninfas y adultos de las familias y generos de Ephemeroptera (Insecta) sudamericanos. Biologia Acuatica, 16, 1 - 32, figs. a-c, 1 - 39." type="journal article" year="1992">
Domínguez
<emphasis id="B912C43355701101FD00E51EFD02F854" box="[650,702,1916,1940]" italics="true" pageId="20" pageNumber="551">et al</emphasis>
. 1992
</bibRefCitation>
) number of tergalii in
<taxonomicName id="4C6663A255701101FB99E51FFB37F854" authorityName="Flowers" authorityYear="1985" box="[1043,1163,1916,1940]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="B912C43355701101FB99E51FFB37F854" box="[1043,1163,1916,1940]" italics="true" pageId="20" pageNumber="551">Guajirolus</emphasis>
</taxonomicName>
is reported as variable: «Branquias en los segmentos abdominales 1 ó 2–7». In the monograph on South American mayflies (
<bibRefCitation id="EFF765D055701101FF44E5A7FE69F81D" author="Dominguez, E. &amp; Molineri, C. &amp; Pescador, M. L. &amp; Hubbard, M. &amp; Nieto, C." box="[206,469,1988,2013]" pageId="20" pageNumber="551" refId="ref8476" refString="Dominguez, E., Molineri, C., Pescador, M. L., Hubbard, M. &amp; Nieto, C. (2006) Ephemeroptera of South America. Aquatic Biodiversity in Latin America, vol. 2. Pensoft, Sofia-Moscow, 646 pp." type="book" year="2006">
Domínguez
<emphasis id="B912C43355701101FED0E5A6FE30F81C" box="[346,396,1988,2012]" italics="true" pageId="20" pageNumber="551">et al</emphasis>
. 2006
</bibRefCitation>
) invariably 6 pairs of tergalii are reported for
<taxonomicName id="4C6663A255701101FC56E5A7FBE8F81C" authorityName="Flowers" authorityYear="1985" box="[988,1108,1988,2012]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="B912C43355701101FC56E5A7FBE8F81C" box="[988,1108,1988,2012]" italics="true" pageId="20" pageNumber="551">Guajirolus</emphasis>
</taxonomicName>
: «abdominal gills present on segments II–VII»; in this book, the Roback's species is ignored, and the genus
<taxonomicName id="4C6663A255711100FBB6E2FBFB08FF70" authorityName="Flowers" authorityYear="1985" box="[1084,1204,152,176]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="552" phylum="Arthropoda" rank="genus">
<emphasis id="B912C43355711100FBB6E2FBFB08FF70" box="[1084,1204,152,176]" italics="true" pageId="21" pageNumber="552">Guajirolus</emphasis>
</taxonomicName>
is not reported from
<collectingCountry id="F37158B155711100FF4DE2DEFF42FF14" box="[199,254,189,212]" name="Peru" pageId="21" pageNumber="552">Peru</collectingCountry>
. Two larvae, collected by me in
<collectingCountry id="F37158B155711100FD0BE2DEFD0BFF14" box="[641,695,189,212]" name="Peru" pageId="21" pageNumber="552">Peru</collectingCountry>
, have 6 pairs of tergalii. Probably, the Roback's sentence about tergalii «1–7» is a mistake.
</paragraph>
<paragraph id="8BD9182155711100FF4DE344FEA4FE48" blockId="21.[151,1436,295,392]" pageId="21" pageNumber="552">
Thus, the existent descriptions of
<taxonomicName id="4C6663A255711100FDCBE34BFD1FFE80" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1995" box="[577,675,296,320]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B912C43355711100FDCBE34BFD1FFE80" box="[577,675,296,320]" italics="true" pageId="21" pageNumber="552">G. nanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6663A255711100FD3AE34BFC82FE80" authorityName="Nieto" authorityYear="2003" box="[688,830,296,320]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="queremba">
<emphasis id="B912C43355711100FD3AE34BFC82FE80" box="[688,830,296,320]" italics="true" pageId="21" pageNumber="552">G. queremba</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6663A255711100FCFEE34BFC51FE80" authorityName="Thomas &amp; Dominique (in Thomas, Dominique &amp; Orth" authorityYear="2005" box="[884,1005,296,320]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B912C43355711100FCFEE34BFC51FE80" box="[884,1005,296,320]" italics="true" pageId="21" pageNumber="552">G. flowersi</emphasis>
</taxonomicName>
do not allow to distinguish them from
<taxonomicName id="4C6663A255711100FF1DE32FFED5FEA4" authorityName="Flowers" authorityYear="1985" box="[151,361,332,356]" class="Insecta" family="Baetidae" genus="Guajirolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="552" phylum="Arthropoda" rank="species" species="ektrapeloglossa">
<emphasis id="B912C43355711100FF1DE32FFED5FEA4" box="[151,361,332,356]" italics="true" pageId="21" pageNumber="552">G. ektrapeloglossa</emphasis>
</taxonomicName>
; these names should be regarded as synonyms, if new species-specific characters will not be discovered.
</paragraph>
</subSubSection>
</treatment>
</document>
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