369 lines
57 KiB
XML
369 lines
57 KiB
XML
<document id="0538E53210A5061A633132B4020A38D4" ID-CLB-Dataset="22067" ID-DOI="10.5852/ejt.2017.337" ID-GBIF-Dataset="5a7f73c5-5646-446f-a72c-3ea18d8f634f" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832483" ID-ZooBank="2669CB69-5064-4B5D-AC25-856680E3B9AA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589830819450" checkinUser="carolina" docAuthor="Budha, Prem B., Naggs, Fred & Backeljau, Thierry" docDate="2017" docId="03C6CF41FFF2FFFCFF1342F7D025F9D2" docLanguage="en" docName="ejt-2017-337.pdf.imf" docOrigin="European Journal of Taxonomy 337" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Diplommatina Benson 1849" docType="key" docVersion="6" lastPageNumber="24" masterDocId="FFFFB739FFE4FFEBFFAE4762D552FF83" masterDocTitle="The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species" masterLastPageNumber="30" masterPageNumber="1" pageNumber="23" updateTime="1698840088040" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="BF702F37A441DDA1A7D9AE58FDFC41CB">
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<mods:title id="36E1D18AAA44476D0ED87ECD91506510">The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species</mods:title>
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<mods:name id="57FF9711116BB8C4071D06013D760B10" type="personal">
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<mods:roleTerm id="1C39C6EBFBD6BB2602819233E9180E30">Author</mods:roleTerm>
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<mods:namePart id="3D5C1FCAB9A053FEC700B241F8F4FB51">Budha, Prem B.</mods:namePart>
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<mods:affiliation id="694771403AE2333268059C986640EAC6">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. & Central Department of Zoology, Tribhuvan University, Kirtipur, Kathmandu, Nepal. & Corresponding author: prembudha @ yahoo. com</mods:affiliation>
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<mods:nameIdentifier id="FADFDC15F82C1B50C00602578A037450" type="email">prembudha@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="192679FAEA7165187AE3E672A4B6327E" type="personal">
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<mods:roleTerm id="CC76E57595F9BC11ADACC9D4AEA51A78">Author</mods:roleTerm>
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<mods:namePart id="D8D2B829669885CC53AAFB67622F97BE">Naggs, Fred</mods:namePart>
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<mods:affiliation id="7B72D0AB9FFB237362DE523A66278CAF">Natural History Museum, Cromwell Road, London, SW 7 5 BD, UK. & Email: f. naggs @ nhm. ac. uk</mods:affiliation>
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<mods:nameIdentifier id="B7355332D7B35F328731141424504B9A" type="email">f.naggs@nhm.ac.uk</mods:nameIdentifier>
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<mods:roleTerm id="A3DBEE1AC76C8603A6B29663F63DD092">Author</mods:roleTerm>
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<mods:namePart id="51FD49686EE04D14985F9F692F328D93">Backeljau, Thierry</mods:namePart>
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<mods:affiliation id="A5A222AD50795A9FED709903B6AA4F59">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. & Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B- 1000, Brussels, Belgium. & Email: thierry. backeljau @ naturalsciences. be</mods:affiliation>
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<mods:nameIdentifier id="A1ADEC441D97D97F55860C011B184AAF" type="email">thierry.backeljau@naturalsciences.be</mods:nameIdentifier>
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<mods:typeOfResource id="98319BF9D466CAF7A5265352FE7F857C">text</mods:typeOfResource>
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<mods:title id="DE12CDE281C6032225E9287B88CC7B40">European Journal of Taxonomy</mods:title>
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<mods:part id="99DC5F5125D2F67364B967DF7B65C4FA">
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<mods:date id="A32B2BF46DFADC642578D5AD789451EC">2017</mods:date>
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<mods:number id="C94DCCF727EDA605AEC905A1AF26A9C8">2017-07-21</mods:number>
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<mods:number id="AF329FF376002EA1AA71E29C1B7ECFB7">337</mods:number>
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<treatment id="03C6CF41FFF2FFFCFF1342F7D025F9D2" ID-DOI="http://doi.org/10.5281/zenodo.3850662" ID-GBIF-Taxon="164167119" ID-Zenodo-Dep="3850662" LSID="urn:lsid:plazi:treatment:03C6CF41FFF2FFFCFF1342F7D025F9D2" httpUri="http://treatment.plazi.org/id/03C6CF41FFF2FFFCFF1342F7D025F9D2" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<subSubSection id="C3752DDCFFF2FFFDFF1342F7D795FA2D" box="[189,711,1428,1455]" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph id="8BD07E57FFF2FFFDFF1342F7D795FA2D" blockId="22.[189,711,1428,1455]" box="[189,711,1428,1455]" pageId="22" pageNumber="23">
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<heading id="D098C93BFFF2FFFDFF1342F7D795FA2D" bold="true" box="[189,711,1428,1455]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
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<emphasis id="B91BA245FFF2FFFDFF1342F7D795FA2D" bold="true" box="[189,711,1428,1455]" pageId="22" pageNumber="23">
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Key to the Nepalese
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<taxonomicName id="4C6F05D4FFF2FFFDFE1542F6D73BFA2C" ID-CoL="7NVSX" authorityName="Benson" authorityYear="1849" box="[443,617,1428,1455]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
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<emphasis id="B91BA245FFF2FFFDFE1542F6D73BFA2C" bold="true" box="[443,617,1428,1455]" italics="true" pageId="22" pageNumber="23">Diplommatina</emphasis>
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</taxonomicName>
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species
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3752DDCFFF2FFFDFF1342A6D076F9C8" pageId="22" pageNumber="23" type="discussion">
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<paragraph id="8BD07E57FFF2FFFDFF1342A6D076F9C8" blockId="22.[189,1399,1476,1611]" pageId="22" pageNumber="23">
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A key to 15 species of
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<taxonomicName id="4C6F05D4FFF2FFFDFE4F42A6D7D8FA5D" authorityName="Benson" authorityYear="1849" box="[481,650,1476,1502]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
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<emphasis id="B91BA245FFF2FFFDFE4F42A6D7D8FA5D" box="[481,650,1476,1502]" italics="true" pageId="22" pageNumber="23">Diplommatina</emphasis>
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</taxonomicName>
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is provided. Two species of
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<taxonomicName id="4C6F05D4FFF2FFFDFC5C42A6D1C9FA5D" authorityName="Benson" authorityYear="1849" box="[1010,1179,1476,1502]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
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<emphasis id="B91BA245FFF2FFFDFC5C42A6D1C9FA5D" box="[1010,1179,1476,1502]" italics="true" pageId="22" pageNumber="23">Diplommatina</emphasis>
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</taxonomicName>
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(
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<taxonomicName id="4C6F05D4FFF2FFFDFB1F42A7D012FA5D" authorityName="Beddome" authorityYear="1875" box="[1201,1344,1477,1502]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="canarica">
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<emphasis id="B91BA245FFF2FFFDFB1F42A7D012FA5D" box="[1201,1344,1477,1502]" italics="true" pageId="22" pageNumber="23">D. canarica</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C6F05D4FFF2FFFDFF13428BD42DF981" authorityName="Benson" authorityYear="1857" box="[189,383,1512,1538]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="pachycheilus">
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<emphasis id="B91BA245FFF2FFFDFF13428BD42DF981" box="[189,383,1512,1538]" italics="true" pageId="22" pageNumber="23">D. pachycheilus</emphasis>
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</taxonomicName>
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) reported by
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<bibRefCitation id="EFFE03A6FFF2FFFDFD82428BD6FAF980" author="Kuznetsov A. G. & Schileyko A. A." box="[556,936,1512,1539]" pageId="22" pageNumber="23" pagination="133 - 140" refId="ref13735" refString="Kuznetsov A. G. & Schileyko A. A. 1997. New data on Enidae (Gastropoda, Pulmonata) of Nepal. Ruthenica 7: 133 - 140." type="journal article" year="1997">Kuznetsov & Schileyko (1997)</bibRefCitation>
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were excluded from the key because neither species were found in the present surveys and no Nepalese material was available to examine internal lamellae. Moreover,
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<taxonomicName id="4C6F05D4FFF2FFFDFDBF4153D7CFF9C9" authorityName="Beddome" authorityYear="1875" box="[529,669,1585,1610]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="canarica">
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<emphasis id="B91BA245FFF2FFFDFDBF4153D7CFF9C9" box="[529,669,1585,1610]" italics="true" pageId="22" pageNumber="23">D. canarica</emphasis>
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</taxonomicName>
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is endemic in the Western Ghats (
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<bibRefCitation id="EFFE03A6FFF2FFFDFB814153D046F9C8" author="Raheem D. C. & Taylor H. & Ablett J. & Preece R. C. & Aravind N. A. & Naggs F." box="[1071,1300,1585,1611]" pageId="22" pageNumber="23" pagination="1 - 294" refId="ref14199" refString="Raheem D. C., Taylor H., Ablett J., Preece R. C., Aravind N. A. & Naggs F. 2014. A systematic revision of the land snails of the Western Ghats of India. Tropical Natural History, Supplement 4: 1 - 294." type="journal article" year="2014">
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Raheem
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<emphasis id="B91BA245FFF2FFFDFB394153D183F9C8" box="[1175,1233,1585,1611]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
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2014
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</bibRefCitation>
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).
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3752DDCFFF2FFFCFF134111D025F9D2" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="key">
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<key id="FFFE2966FFF2FFFCFF134111D025F9D2" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<keyStep id="309B66F2FFF2FFFDFF134111D024F979" pageId="22" pageNumber="23">
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<paragraph id="8BD07E57FFF2FFFDFF134111D025F90E" blockId="22.[189,1399,1651,1786]" box="[189,1399,1651,1678]" pageId="22" pageNumber="23">
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<keyLead id="309EDD62FFF2FFFDFF134111D025F90E" box="[189,1399,1651,1678]" pageId="22" pageNumber="23">1. Spiral striation present ...................................................................................................................... 2</keyLead>
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</paragraph>
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<paragraph id="8BD07E57FFF2FFFDFF6F41FAD024F979" blockId="22.[189,1399,1651,1786]" pageId="22" pageNumber="23">
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<keyLead id="309EDD62FFF2FFFDFF6F41FAD024F979" pageId="22" pageNumber="23">
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– Spiral striation absent, constriction at the left side of the parietal edge of the aperture (
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<figureCitation id="135462D2FFF2FFFDFAA541F5D03AF932" box="[1291,1384,1687,1713]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[190,1398,279,1890]" captionTargetId="figure@20.[190,1398,276,1893]" captionTargetPageId="20" captionText="Fig. 8. A. Diplommatina silvicola Godwin-Austen, 1886 (CDZMTU050). B. D. sperata W.T. Blanford, 1862 (CDZMTU062). C. D. syabrubesiensis sp. nov., holotype (CDZMTU051). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832500" httpUri="https://zenodo.org/record/3832500/files/figure.png" pageId="22" pageNumber="23">Fig. 8E</figureCitation>
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), presence of two parietalis (Prt1 and Prt2), two vertical palatalis (Vpt1 and Vpt2) and two horizontal palatalis (Hpt1 and Hpt2) ...........................................................................
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<taxonomicName id="4C6F05D4FFF2FFFDFBE44182D049F979" ID-CoL="6CXRM" authority="Budha & Naggs & Backeljau, 2017" authorityName="Budha & Naggs & Backeljau" authorityYear="2017" box="[1098,1307,1759,1786]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="godawariensis" status="sp. nov.">
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<emphasis id="B91BA245FFF2FFFDFBE44182D049F979" bold="true" box="[1098,1307,1759,1786]" italics="true" pageId="22" pageNumber="23">D. godawariensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2281F3EFFF2FFFDFA8C4182D024F979" box="[1314,1398,1760,1786]" pageId="22" pageNumber="23" rank="species">sp. nov</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="309B66F2FFF2FFFDFF134040D025F8E2" pageId="22" pageNumber="23">
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<paragraph id="8BD07E57FFF2FFFDFF134040D025F8BF" blockId="22.[189,1399,1826,1889]" box="[189,1399,1826,1852]" pageId="22" pageNumber="23">
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<keyLead id="309EDD62FFF2FFFDFF134040D025F8BF" box="[189,1399,1826,1852]" pageId="22" pageNumber="23">2. Spiral striation fairly visible ............................................................................................................. 3</keyLead>
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</paragraph>
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<paragraph id="8BD07E57FFF2FFFDFF6F4025D025F8E2" blockId="22.[189,1399,1826,1889]" box="[193,1399,1862,1889]" pageId="22" pageNumber="23">
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<keyLead id="309EDD62FFF2FFFDFF6F4025D025F8E2" box="[193,1399,1862,1889]" pageId="22" pageNumber="23">– Spiral striation distinctly visible ....................................................................................................... 5</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="309B66F2FFF2FFFDFF1340EAD024F868" pageId="22" pageNumber="23">
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<paragraph id="8BD07E57FFF2FFFDFF1340EAD025F821" blockId="22.[189,1399,1928,2027]" box="[189,1399,1928,1955]" pageId="22" pageNumber="23">
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<keyLead id="309EDD62FFF2FFFDFF1340EAD025F821" box="[189,1399,1928,1955]" pageId="22" pageNumber="23">3. Dorsal fold present ............................................................................................................................ 4</keyLead>
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</paragraph>
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<paragraph id="8BD07E57FFF2FFFDFF6F40CFD024F868" blockId="22.[189,1399,1928,2027]" pageId="22" pageNumber="23">
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<keyLead id="309EDD62FFF2FFFDFF6F40CFD024F868" pageId="22" pageNumber="23">
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– Dorsal fold absent, constriction at AeP (
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<figureCitation id="135462D2FFF2FFFDFC8340CED6F0F844" box="[813,930,1964,1991]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="22" pageNumber="23">Fig. 9M</figureCitation>
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), parietal wall slightly tilted left (
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<figureCitation id="135462D2FFF2FFFDFF5D40B3D41CF868" box="[243,334,2001,2027]" captionStart="Fig" captionStartId="17.[189,232,1930,1956]" captionTargetBox="[189,1397,264,1875]" captionTargetId="figure@17.[189,1397,261,1877]" captionTargetPageId="17" captionText="Fig. 7. Diplommatina Benson, 1849 of Nepal. A. D. regularis Fulton, 1901 (CDZMTU045). B. D. salgharica sp. nov., holotype (CDZMTU046). C. D. shivapuriensis sp. nov., holotype (CDZMTU048). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832498" httpUri="https://zenodo.org/record/3832498/files/figure.png" pageId="22" pageNumber="23">Fig. 7C</figureCitation>
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) .....................................................................................................
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<taxonomicName id="4C6F05D4FFF2FFFDFBE640B3D048F868" ID-CoL="36M3V" authority="Budha & Naggs & Backeljau, 2017" authorityName="Budha & Naggs & Backeljau" authorityYear="2017" box="[1096,1306,2000,2027]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="shivapuriensis" status="sp. nov.">
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<emphasis id="B91BA245FFF2FFFDFBE640B3D048F868" bold="true" box="[1096,1306,2000,2027]" italics="true" pageId="22" pageNumber="23">D. shivapuriensis</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="A2281F3EFFF2FFFDFA8C40B3D024F868" box="[1314,1398,2001,2027]" pageId="22" pageNumber="23" rank="species">sp. nov</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309B66F2FFF3FFFCFF134669D024FECA" pageId="23" pageNumber="24">
|
||
<paragraph id="8BD07E57FFF3FFFCFF134669D024FEA5" blockId="23.[189,1398,267,329]" box="[189,1398,267,294]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF134669D024FEA5" box="[189,1398,267,294]" pageId="23" pageNumber="24">
|
||
4. Dorsal fold distinct (
|
||
<figureCitation id="135462D2FFF3FFFCFE734669D764FEA6" box="[477,566,267,293]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="23" pageNumber="24">Fig. 3C</figureCitation>
|
||
<figureCitation id="135462D2FFF3FFFCFD984669D714FEA6" box="[566,582,267,293]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="23" pageNumber="24">4</figureCitation>
|
||
) ..............................................................................
|
||
<taxonomicName id="4C6F05D4FFF3FFFCFB3B466ED049FEA5" ID-CoL="36M29" authority="Budha & Naggs & Backeljau, 2017" authorityName="Budha & Naggs & Backeljau" authorityYear="2017" box="[1173,1307,267,294]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="fistulata" status="sp. nov.">
|
||
<emphasis id="B91BA245FFF3FFFCFB3B466ED049FEA5" bold="true" box="[1173,1307,267,294]" italics="true" pageId="23" pageNumber="24">D. fistulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2281F3EFFF3FFFCFA8C466ED024FEA5" box="[1314,1398,268,294]" pageId="23" pageNumber="24" rank="species">sp. nov</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD07E57FFF3FFFCFF6F464DD024FECA" blockId="23.[189,1398,267,329]" box="[193,1398,302,329]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF6F464DD024FECA" box="[193,1398,302,329]" pageId="23" pageNumber="24">
|
||
– Dorsal fold fairly distinct (
|
||
<figureCitation id="135462D2FFF3FFFCFD8A464CD72FFECA" box="[548,637,302,329]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[190,1398,279,1890]" captionTargetId="figure@20.[190,1398,276,1893]" captionTargetPageId="20" captionText="Fig. 8. A. Diplommatina silvicola Godwin-Austen, 1886 (CDZMTU050). B. D. sperata W.T. Blanford, 1862 (CDZMTU062). C. D. syabrubesiensis sp. nov., holotype (CDZMTU051). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832500" httpUri="https://zenodo.org/record/3832500/files/figure.png" pageId="23" pageNumber="24">Fig. 8C</figureCitation>
|
||
<figureCitation id="135462D2FFF3FFFCFDD3464DD7DFFECA" box="[637,653,303,329]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="23" pageNumber="24">4</figureCitation>
|
||
) .......................................................
|
||
<taxonomicName id="4C6F05D4FFF3FFFCFB97464DD048FECA" ID-CoL="36M46" authority="Budha & Naggs & Backeljau, 2017" authorityName="Budha & Naggs & Backeljau" authorityYear="2017" box="[1081,1306,302,329]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="syabrubesiensis" status="sp. nov.">
|
||
<emphasis id="B91BA245FFF3FFFCFB97464DD048FECA" bold="true" box="[1081,1306,302,329]" italics="true" pageId="23" pageNumber="24">D. syabrubesiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2281F3EFFF3FFFCFA8C464DD024FECA" box="[1314,1398,303,329]" pageId="23" pageNumber="24" rank="species">sp. nov</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309B66F2FFF3FFFCFF134612D024FE2E" pageId="23" pageNumber="24">
|
||
<paragraph id="8BD07E57FFF3FFFCFF134612D025FE09" blockId="23.[189,1399,368,430]" box="[189,1399,368,395]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF134612D025FE09" box="[189,1399,368,395]" pageId="23" pageNumber="24">
|
||
5. Radial ribs widely spaced, 5–10 ribs/
|
||
<quantity id="4C97D3B2FFF3FFFCFD304612D7A5FE09" box="[670,759,368,395]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="24" unit="mm" value="0.5">0.5 mm</quantity>
|
||
................................................................................... 6
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD07E57FFF3FFFCFF6F46F6D024FE2E" blockId="23.[189,1399,368,430]" box="[193,1398,403,430]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF6F46F6D024FE2E" box="[193,1398,403,430]" pageId="23" pageNumber="24">
|
||
– Radial ribs narrowly spaced, 10–20 ribs/
|
||
<quantity id="4C97D3B2FFF3FFFCFD6946F1D672FE2D" box="[711,800,403,430]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="24" unit="mm" value="0.5">0.5 mm</quantity>
|
||
........................................................................... 10
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309B66F2FFF3FFFCFF1346B6D025FD91" pageId="23" pageNumber="24">
|
||
<paragraph id="8BD07E57FFF3FFFCFF1346B6D025FE6C" blockId="23.[189,1399,468,530]" box="[189,1399,468,495]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF1346B6D025FE6C" box="[189,1399,468,495]" pageId="23" pageNumber="24">6. Constriction at the middle of the parietal wall of the peristome ....................................................... 7</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD07E57FFF3FFFCFF6F469AD025FD91" blockId="23.[189,1399,468,530]" box="[193,1399,504,530]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF6F469AD025FD91" box="[193,1399,504,530]" pageId="23" pageNumber="24">– Constriction not at the middle of the parietal wall of the peristome ................................................ 8</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309B66F2FFF3FFFCFF134558D025FD1A" pageId="23" pageNumber="24">
|
||
<paragraph id="8BD07E57FFF3FFFCFF134558D024FDF4" blockId="23.[189,1399,569,666]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF134558D024FDF4" pageId="23" pageNumber="24">
|
||
7. Two short parietalis (Prt1 and Prt2) (
|
||
<figureCitation id="135462D2FFF3FFFCFD37455BD7A7FDD0" box="[665,757,569,595]" captionStart="Fig" captionStartId="4.[189,232,1895,1921]" captionTargetBox="[189,1398,285,1853]" captionTargetId="figure@4.[189,1398,285,1853]" captionTargetPageId="4" captionText="Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two)." figureDoi="http://doi.org/10.5281/zenodo.3832488" httpUri="https://zenodo.org/record/3832488/files/figure.png" pageId="23" pageNumber="24">Fig. 2B</figureCitation>
|
||
), columellaris moderate (
|
||
<figureCitation id="135462D2FFF3FFFCFB8E455BD12FFDD0" box="[1056,1149,569,595]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="23" pageNumber="24">Fig. 4A</figureCitation>
|
||
<figureCitation id="135462D2FFF3FFFCFBD3455BD1D8FDD0" box="[1149,1162,569,595]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="23" pageNumber="24">3</figureCitation>
|
||
) ............................. ...........................................................................................................
|
||
<taxonomicName id="4C6F05D4FFF3FFFCFBAA453FD024FDF4" ID-CoL="36M2D" authority="(L. Pfeiffer, 1846)" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1846" box="[1028,1398,604,631]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="folliculus">
|
||
<emphasis id="B91BA245FFF3FFFCFBAA453FD1C8FDF4" bold="true" box="[1028,1178,604,631]" italics="true" pageId="23" pageNumber="24">D. folliculus</emphasis>
|
||
(L.
|
||
<bibRefCitation id="EFFE03A6FFF3FFFCFB65453ED03CFDF4" author="Pfeiffer L." box="[1227,1390,604,631]" pageId="23" pageNumber="24" refId="ref14017" refString="Pfeiffer L. 1846. Symbolae ad Historiam Heliceorum. Fischer Verlag, Kassel, Germany. https: // doi. org / 10.5962 / bhl. title. 11903" type="book" year="1846">Pfeiffer, 1846</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD07E57FFF3FFFCFF6F45E2D025FD1A" blockId="23.[189,1399,569,666]" box="[193,1399,639,666]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF6F45E2D025FD1A" box="[193,1399,639,666]" pageId="23" pageNumber="24">
|
||
– One long parietalis, columellaris strong (
|
||
<figureCitation id="135462D2FFF3FFFCFD6445E2D674FD19" box="[714,806,640,666]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[190,1398,279,1890]" captionTargetId="figure@20.[190,1398,276,1893]" captionTargetPageId="20" captionText="Fig. 8. A. Diplommatina silvicola Godwin-Austen, 1886 (CDZMTU050). B. D. sperata W.T. Blanford, 1862 (CDZMTU062). C. D. syabrubesiensis sp. nov., holotype (CDZMTU051). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832500" httpUri="https://zenodo.org/record/3832500/files/figure.png" pageId="23" pageNumber="24">Fig. 8A</figureCitation>
|
||
<figureCitation id="135462D2FFF3FFFCFC8845E2D666FD19" box="[806,820,640,666]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="23" pageNumber="24">3</figureCitation>
|
||
) ....................
|
||
<taxonomicName id="4C6F05D4FFF3FFFCFC4D45E2D025FD1A" ID-CoL="36M3W" authority="Godwin-Austen, 1886" authorityName="Godwin-Austen" authorityYear="1886" box="[995,1399,639,666]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="silvicola">
|
||
<emphasis id="B91BA245FFF3FFFCFC4D45E2D138FD19" bold="true" box="[995,1130,639,666]" italics="true" pageId="23" pageNumber="24">D. silvicola</emphasis>
|
||
<bibRefCitation id="EFFE03A6FFF3FFFCFBDC45E2D025FD1A" author="Godwin-Austen H. H." box="[1138,1399,639,666]" pageId="23" pageNumber="24" pagination="165 - 185" refId="ref13270" refString="Godwin-Austen H. H. 1886. Land and freshwater Mollusca of India. Conchologia Indica 1 (5): 165 - 185." type="journal article" year="1886">Godwin-Austen, 1886</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309B66F2FFF3FFFCFF1345A3D025FCA1" pageId="23" pageNumber="24">
|
||
<paragraph id="8BD07E57FFF3FFFCFF1345A3D024FD7C" blockId="23.[189,1399,705,802]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF1345A3D024FD7C" pageId="23" pageNumber="24">
|
||
8. Constriction in the left side of PeP, two short horizontal palatalis (Hpt1 and Hpt2) (
|
||
<figureCitation id="135462D2FFF3FFFCFF5D4586D418FD7D" box="[243,330,740,766]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="23" pageNumber="24">Fig. 9F</figureCitation>
|
||
) ..................................................................................................
|
||
<taxonomicName id="4C6F05D4FFF3FFFCFB804587D048FD7C" ID-CoL="36M35" authority="Budha & Naggs & Backeljau, 2017" authorityName="Budha & Naggs & Backeljau" authorityYear="2017" box="[1070,1306,740,767]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="maipokhariensis" status="sp. nov.">
|
||
<emphasis id="B91BA245FFF3FFFCFB804587D048FD7C" bold="true" box="[1070,1306,740,767]" italics="true" pageId="23" pageNumber="24">D. maipokhariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2281F3EFFF3FFFCFA8C4587D024FD7C" box="[1314,1398,741,767]" pageId="23" pageNumber="24" rank="species">sp. nov</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD07E57FFF3FFFCFF6F446AD025FCA1" blockId="23.[189,1399,705,802]" box="[193,1399,775,802]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF6F446AD025FCA1" box="[193,1399,775,802]" pageId="23" pageNumber="24">– Constriction in the right side of PeP ................................................................................................. 9</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309B66F2FFF3FFFCFF13442BD024FC29" pageId="23" pageNumber="24">
|
||
<paragraph id="8BD07E57FFF3FFFCFF13442BD025FCE0" blockId="23.[189,1399,841,938]" box="[189,1399,841,867]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF13442BD025FCE0" box="[189,1399,841,867]" pageId="23" pageNumber="24">
|
||
9. Apical whorls slender (
|
||
<figureCitation id="135462D2FFF3FFFCFE52442BD705FCE0" box="[508,599,841,867]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="23" pageNumber="24">Fig. 3B</figureCitation>
|
||
), one parietalis .............................
|
||
<taxonomicName id="4C6F05D4FFF3FFFCFC5E442BD025FCE0" ID-CoL="36M28" authority="Godwin-Austen, 1886" authorityName="Godwin-Austen" authorityYear="1886" box="[1008,1399,841,867]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="exserta">
|
||
<emphasis id="B91BA245FFF3FFFCFC5E442BD138FCE0" bold="true" box="[1008,1130,841,867]" italics="true" pageId="23" pageNumber="24">D. exserta</emphasis>
|
||
<bibRefCitation id="EFFE03A6FFF3FFFCFBDC442BD025FCE0" author="Godwin-Austen H. H." box="[1138,1399,841,867]" pageId="23" pageNumber="24" pagination="165 - 185" refId="ref13270" refString="Godwin-Austen H. H. 1886. Land and freshwater Mollusca of India. Conchologia Indica 1 (5): 165 - 185." type="journal article" year="1886">Godwin-Austen, 1886</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD07E57FFF3FFFCFF6F440FD024FC29" blockId="23.[189,1399,841,938]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF6F440FD024FC29" pageId="23" pageNumber="24">
|
||
– Apical whorls not slender (
|
||
<figureCitation id="135462D2FFF3FFFCFD87440ED7D7FC05" box="[553,645,876,902]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="23" pageNumber="24">Fig. 4B</figureCitation>
|
||
), two parietalis (Prt1 and Prt2) (
|
||
<figureCitation id="135462D2FFF3FFFCFC51440ED10BFC05" box="[1023,1113,876,902]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="23" pageNumber="24">Fig. 9B</figureCitation>
|
||
) ................................... ...........................................................................................................................
|
||
<taxonomicName id="4C6F05D4FFF3FFFCFBD944F2D048FC29" ID-CoL="36M3T" authority="Budha & Naggs & Backeljau, 2017" authorityName="Budha & Naggs & Backeljau" authorityYear="2017" box="[1143,1306,911,938]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="salgharica" status="sp. nov.">
|
||
<emphasis id="B91BA245FFF3FFFCFBD944F2D048FC29" bold="true" box="[1143,1306,911,938]" italics="true" pageId="23" pageNumber="24">D. salgharica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2281F3EFFF3FFFCFA8C44F2D024FC29" box="[1314,1398,912,938]" pageId="23" pageNumber="24" rank="species">sp. nov</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309B66F2FFF3FFFCFF1344B3D024FB8D" pageId="23" pageNumber="24">
|
||
<paragraph id="8BD07E57FFF3FFFCFF1344B3D027FC68" blockId="23.[189,1398,977,1038]" box="[189,1397,977,1003]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF1344B3D027FC68" box="[189,1397,977,1003]" pageId="23" pageNumber="24">10. Constriction near or at the center of PeP .........................................................................................11</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD07E57FFF3FFFCFF6F4496D024FB8D" blockId="23.[189,1398,977,1038]" box="[193,1398,1012,1038]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF6F4496D024FB8D" box="[193,1398,1012,1038]" pageId="23" pageNumber="24">– Constriction not at the center of PeP ............................................................................................... 14</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309B66F2FFF3FFFCFF134357D024FBF0" pageId="23" pageNumber="24">
|
||
<paragraph id="8BD07E57FFF3FFFCFF134357D024FBCC" blockId="23.[189,1398,1077,1139]" box="[189,1398,1077,1104]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF134357D024FBCC" box="[189,1398,1077,1104]" pageId="23" pageNumber="24">11. Horizontal palatalis present ............................................................................................................ 12</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD07E57FFF3FFFCFF6F433BD024FBF0" blockId="23.[189,1398,1077,1139]" box="[193,1398,1112,1139]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF6F433BD024FBF0" box="[193,1398,1112,1139]" pageId="23" pageNumber="24">– Horizontal palatalis absent .............................................................................................................. 13</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309B66F2FFF3FFFCFF1343F8D025FB78" pageId="23" pageNumber="24">
|
||
<paragraph id="8BD07E57FFF3FFFCFF1343F8D024FB36" blockId="23.[189,1399,1178,1275]" box="[189,1398,1178,1205]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF1343F8D024FB36" box="[189,1398,1178,1205]" pageId="23" pageNumber="24">
|
||
12. AeP straight (
|
||
<figureCitation id="135462D2FFF3FFFCFE2543F8D4BBFB37" box="[395,489,1178,1204]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="23" pageNumber="24">Fig. 3A</figureCitation>
|
||
), parietalis thin and long (
|
||
<figureCitation id="135462D2FFF3FFFCFCB243F8D6D5FB37" box="[796,903,1178,1204]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="23" pageNumber="24">Fig. 3A3</figureCitation>
|
||
) ................................
|
||
<taxonomicName id="4C6F05D4FFF3FFFCFB2D43F8D049FB36" ID-CoL="36LZD" authority="Budha & Naggs & Backeljau, 2017" authorityName="Budha & Naggs & Backeljau" authorityYear="2017" box="[1155,1307,1178,1205]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="abiesiana" status="sp. nov.">
|
||
<emphasis id="B91BA245FFF3FFFCFB2D43F8D049FB36" bold="true" box="[1155,1307,1178,1205]" italics="true" pageId="23" pageNumber="24">D. abiesiana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2281F3EFFF3FFFCFA8C43F9D024FB36" box="[1314,1398,1179,1205]" pageId="23" pageNumber="24" rank="species">sp. nov</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD07E57FFF3FFFCFF6F43DCD025FB78" blockId="23.[189,1399,1178,1275]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF6F43DCD025FB78" pageId="23" pageNumber="24">
|
||
– AeP curved (
|
||
<figureCitation id="135462D2FFF3FFFCFE2D43DFD751FB54" box="[387,515,1213,1240]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[189,1398,265,1879]" captionTargetId="figure@13.[189,1398,261,1881]" captionTargetPageId="13" captionText="Fig. 5. Diplommatina Benson, 1849 of Nepal. A. D. miriensis Godwin-Austen, 1917 (CDZMTU036). B. D.munipurensis Godwin-Austen, 1892 (CDZMTU039).C. D.oviformis Fulton, 1901 (CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="23" pageNumber="24">Fig. 5A, A</figureCitation>
|
||
<figureCitation id="135462D2FFF3FFFCFDAC43DFD740FB54" box="[514,530,1213,1239]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="23" pageNumber="24">4</figureCitation>
|
||
), parietalis very weak, nearly invisible ...................................................... .....................................................................................................
|
||
<taxonomicName id="4C6F05D4FFF3FFFCFC764383D025FB78" ID-CoL="36M37" authority="Godwin-Austen, 1917" authorityName="Godwin-Austen" authorityYear="1917" box="[984,1399,1248,1275]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="miriensis">
|
||
<emphasis id="B91BA245FFF3FFFCFC764383D139FB79" bold="true" box="[984,1131,1248,1275]" italics="true" pageId="23" pageNumber="24">D. miriensis</emphasis>
|
||
<bibRefCitation id="EFFE03A6FFF3FFFCFBDC4382D025FB78" author="Godwin-Austen H. H." box="[1138,1399,1248,1275]" pageId="23" pageNumber="24" pagination="547 - 559" refId="ref13391" refString="Godwin-Austen H. H. 1917. Zoological results of the Abor Expedition, Mollusca 6. Records of the Indian Museum 8: 547 - 559." type="journal article" year="1917">Godwin-Austen, 1917</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309B66F2FFF3FFFCFF134240D025FA25" pageId="23" pageNumber="24">
|
||
<paragraph id="8BD07E57FFF3FFFCFF134240D025FADC" blockId="23.[189,1399,1314,1446]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF134240D025FADC" pageId="23" pageNumber="24">
|
||
13. Penultimate whorl slightly larger than body whorl, columellar tooth distinct (
|
||
<figureCitation id="135462D2FFF3FFFCFF5D4227D41FFADC" box="[243,333,1349,1375]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[190,1398,279,1890]" captionTargetId="figure@20.[190,1398,276,1893]" captionTargetPageId="20" captionText="Fig. 8. A. Diplommatina silvicola Godwin-Austen, 1886 (CDZMTU050). B. D. sperata W.T. Blanford, 1862 (CDZMTU062). C. D. syabrubesiensis sp. nov., holotype (CDZMTU051). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832500" httpUri="https://zenodo.org/record/3832500/files/figure.png" pageId="23" pageNumber="24">Fig. 8B</figureCitation>
|
||
) ............................................................................................
|
||
<taxonomicName id="4C6F05D4FFF3FFFCFBAD4227D025FADC" ID-CoL="36M43" authority="W. T. Blanford, 1862" authorityName="W. T. Blanford" authorityYear="1862" box="[1027,1399,1349,1376]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="sperata">
|
||
<emphasis id="B91BA245FFF3FFFCFBAD4227D12DFADC" bold="true" box="[1027,1151,1349,1375]" italics="true" pageId="23" pageNumber="24">D. sperata</emphasis>
|
||
W.T.
|
||
<bibRefCitation id="EFFE03A6FFF3FFFCFB6B4227D025FADC" author="Blanford W. T." box="[1221,1399,1349,1375]" pageId="23" pageNumber="24" pagination="135 - 145" refId="ref12986" refString="Blanford W. T. 1862. Contribution to Indian Malacology, No. III. Descriptions of new operculated land shells from Pegu, Arakan and the Khasi Hills. Journal of the Asiatic Society of Bengal 31: 135 - 145." type="journal article" year="1862">Blanford, 1862</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD07E57FFF3FFFCFF6F420BD025FA25" blockId="23.[189,1399,1314,1446]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF6F420BD025FA25" pageId="23" pageNumber="24">
|
||
– Penultimate whorl and body whorl nearly equal, columellar tooth not distinct (
|
||
<figureCitation id="135462D2FFF3FFFCFF5D42E9D41EFA25" box="[243,332,1419,1446]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[189,1398,265,1879]" captionTargetId="figure@13.[189,1398,261,1881]" captionTargetPageId="13" captionText="Fig. 5. Diplommatina Benson, 1849 of Nepal. A. D. miriensis Godwin-Austen, 1917 (CDZMTU036). B. D.munipurensis Godwin-Austen, 1892 (CDZMTU039).C. D.oviformis Fulton, 1901 (CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="23" pageNumber="24">Fig. 5B</figureCitation>
|
||
) ..............................................................................
|
||
<taxonomicName id="4C6F05D4FFF3FFFCFC0E42EED025FA25" ID-CoL="36M39" authority="Godwin-Austen, 1892" authorityName="Godwin-Austen" authorityYear="1892" box="[928,1399,1419,1446]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="munipurensis">
|
||
<emphasis id="B91BA245FFF3FFFCFC0E42EED138FA26" bold="true" box="[928,1130,1419,1446]" italics="true" pageId="23" pageNumber="24">D. munipurensis</emphasis>
|
||
<bibRefCitation id="EFFE03A6FFF3FFFCFBDC42E9D025FA25" author="Godwin-Austen H. H." box="[1138,1399,1419,1446]" pageId="23" pageNumber="24" pagination="509 - 520" refId="ref13295" refString="Godwin-Austen H. H. 1892. On new species and varieties of the land molluscan genus Diplommatina from the Garo, Naga and Munipur Hill-ranges, Assam. Proceedings of the Zoological Society of London 35: 509 - 520." type="journal article" year="1892">Godwin-Austen, 1892</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309B66F2FFF3FFFCFF1342AFD025F9D2" pageId="23" pageNumber="24">
|
||
<paragraph id="8BD07E57FFF3FFFCFF1342AFD025F989" blockId="23.[189,1399,1485,1617]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF1342AFD025F989" pageId="23" pageNumber="24">
|
||
14. Constriction at the left side from the center of the PeP (
|
||
<figureCitation id="135462D2FFF3FFFCFCF442AFD6FEFA64" box="[858,940,1485,1511]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="23" pageNumber="24">Fig. 9J</figureCitation>
|
||
), AeP curved (
|
||
<figureCitation id="135462D2FFF3FFFCFBFC42AFD1FFFA64" box="[1106,1197,1485,1511]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="23" pageNumber="24">Fig.A4</figureCitation>
|
||
), short horizontal palatalis (
|
||
<figureCitation id="135462D2FFF3FFFCFEF24292D4FCF989" box="[348,430,1520,1546]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="23" pageNumber="24">Fig. 9J</figureCitation>
|
||
) ........................................................................................
|
||
<taxonomicName id="4C6F05D4FFF3FFFCFBE64293D025F989" ID-CoL="36M3R" authority="Fulton, 1901" authorityName="Fulton" authorityYear="1901" box="[1096,1399,1520,1547]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="regularis">
|
||
<emphasis id="B91BA245FFF3FFFCFBE64293D18BF988" bold="true" box="[1096,1241,1520,1547]" italics="true" pageId="23" pageNumber="24">D. regularis</emphasis>
|
||
<bibRefCitation id="EFFE03A6FFF3FFFCFB4E4292D025F989" author="Fulton H." box="[1248,1399,1520,1546]" pageId="23" pageNumber="24" pagination="242 - 245" refId="ref13161" refString="Fulton H. 1901. Descriptions of some supposed new species of Diplommatina, Opisthostoma, and a new variety of Alycaeus from N. Borneo, Banguey Island and Darjeeling. Annals and Magazine of Natural History, Series 7 8: 242 - 245. https: // doi. org / 10.1080 / 03745480109442915" type="journal article" year="1901">Fulton, 1901</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD07E57FFF3FFFCFF6F4176D025F9D2" blockId="23.[189,1399,1485,1617]" pageId="23" pageNumber="24">
|
||
<keyLead id="309EDD62FFF3FFFCFF6F4176D025F9D2" pageId="23" pageNumber="24">
|
||
– Constriction far ahead from the aperture, very long horizontal palatalis and two vertical palatalis (Vpt1 and Vpt2) (
|
||
<figureCitation id="135462D2FFF3FFFCFE124154D75EF9D2" box="[444,524,1590,1617]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="23" pageNumber="24">Fig. 9I</figureCitation>
|
||
) ...........................................................................
|
||
<taxonomicName id="4C6F05D4FFF3FFFCFBED4155D025F9D2" ID-CoL="36M3G" authority="Fulton, 1901" authorityName="Fulton" authorityYear="1901" box="[1091,1399,1590,1617]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="oviformis">
|
||
<emphasis id="B91BA245FFF3FFFCFBED4155D18BF9D3" bold="true" box="[1091,1241,1590,1617]" italics="true" pageId="23" pageNumber="24">D. oviformis</emphasis>
|
||
<bibRefCitation id="EFFE03A6FFF3FFFCFB4E4154D025F9D2" author="Fulton H." box="[1248,1399,1590,1617]" pageId="23" pageNumber="24" pagination="242 - 245" refId="ref13161" refString="Fulton H. 1901. Descriptions of some supposed new species of Diplommatina, Opisthostoma, and a new variety of Alycaeus from N. Borneo, Banguey Island and Darjeeling. Annals and Magazine of Natural History, Series 7 8: 242 - 245. https: // doi. org / 10.1080 / 03745480109442915" type="journal article" year="1901">Fulton, 1901</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |