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<document id="BADFF4638D0A0DAA5F587BDD7D3510EB" ID-CLB-Dataset="22067" ID-DOI="10.5852/ejt.2017.337" ID-GBIF-Dataset="5a7f73c5-5646-446f-a72c-3ea18d8f634f" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832483" ID-ZooBank="2669CB69-5064-4B5D-AC25-856680E3B9AA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589830819450" checkinUser="carolina" docAuthor="Budha, Prem B., Naggs, Fred &amp; Backeljau, Thierry" docDate="2017" docId="03C6CF41FFE8FFE5FE48414BD606FC31" docLanguage="en" docName="ejt-2017-337.pdf.imf" docOrigin="European Journal of Taxonomy 337" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Diplommatina munipurensis Godwin-Austen 1892" docType="treatment" docVersion="6" lastPageNumber="15" masterDocId="FFFFB739FFE4FFEBFFAE4762D552FF83" masterDocTitle="The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species" masterLastPageNumber="30" masterPageNumber="1" pageNumber="13" updateTime="1698840088040" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AA946DF065564B8AFA3C02581E8E8782">The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species</mods:title>
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<mods:namePart id="0B22D10E66D7AFD15C3FD541E70BCC3C">Budha, Prem B.</mods:namePart>
<mods:affiliation id="C0B5C01CA45AE7294E0444878ADCC53A">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Central Department of Zoology, Tribhuvan University, Kirtipur, Kathmandu, Nepal. &amp; Corresponding author: prembudha @ yahoo. com</mods:affiliation>
<mods:nameIdentifier id="C03BECD5E2FF379E40B35DAD82FD6EEF" type="email">prembudha@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="C15CBDECF3C06E809984AA5DD89D1A0D">Naggs, Fred</mods:namePart>
<mods:affiliation id="8C036D09F8891927ABDF6CE3394A93B7">Natural History Museum, Cromwell Road, London, SW 7 5 BD, UK. &amp; Email: f. naggs @ nhm. ac. uk</mods:affiliation>
<mods:nameIdentifier id="73868457F9F0CD7136A030A12E9DE27D" type="email">f.naggs@nhm.ac.uk</mods:nameIdentifier>
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<mods:namePart id="0B1B2E5CF57F19B9F31BE9EAAF557AE6">Backeljau, Thierry</mods:namePart>
<mods:affiliation id="1E36DD8B9458D6F5A36AE2E50CAC737A">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B- 1000, Brussels, Belgium. &amp; Email: thierry. backeljau @ naturalsciences. be</mods:affiliation>
<mods:nameIdentifier id="C44447A8C1B8B5952C26856CE42140DA" type="email">thierry.backeljau@naturalsciences.be</mods:nameIdentifier>
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<mods:title id="F4015DF56CCA6B9527D63890308C99AF">European Journal of Taxonomy</mods:title>
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<mods:date id="80F4F6661F4E5E62B0D4F9843DDA0D7D">2017</mods:date>
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<mods:number id="CFC915FA43C6108E38AA4760416DC164">2017-07-21</mods:number>
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<heading id="D098C93BFFE8FFE7FE48414BD11CF9C0" box="[486,1102,1577,1604]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="4C6F05D4FFE8FFE7FE48414BD11CF9C0" ID-CoL="36M39" authority="Godwin-Austen, 1892" authorityName="Godwin-Austen" authorityYear="1892" box="[486,1102,1577,1604]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="munipurensis">
<emphasis id="B91BA245FFE8FFE7FE48414BD613F9C0" bold="true" box="[486,833,1577,1604]" italics="true" pageId="12" pageNumber="13">Diplommatina munipurensis</emphasis>
<bibRefCitation id="EFFE03A6FFE8FFE7FCE7414BD11CF9C0" author="Godwin-Austen H. H." box="[841,1102,1577,1603]" pageId="12" pageNumber="13" pagination="509 - 520" refId="ref13295" refString="Godwin-Austen H. H. 1892. On new species and varieties of the land molluscan genus Diplommatina from the Garo, Naga and Munipur Hill-ranges, Assam. Proceedings of the Zoological Society of London 35: 509 - 520." type="journal article" year="1892">Godwin-Austen, 1892</bibRefCitation>
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<paragraph id="8BD07E57FFE8FFE7FD7C412FD633F9E4" blockId="12.[486,1102,1577,1640]" box="[722,865,1613,1640]" pageId="12" pageNumber="13">
<figureCitation id="135462D2FFE8FFE7FD7C412FD662F9EB" box="[722,816,1613,1640]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[189,1398,265,1879]" captionTargetId="figure@13.[189,1398,261,1881]" captionTargetPageId="13" captionText="Fig. 5. Diplommatina Benson, 1849 of Nepal. A. D. miriensis Godwin-Austen, 1917 (CDZMTU036). B. D.munipurensis Godwin-Austen, 1892 (CDZMTU039).C. D.oviformis Fulton, 1901 (CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="12" pageNumber="13">Figs 5B</figureCitation>
,
<figureCitation id="135462D2FFE8FFE7FC93412FD633F9E4" box="[829,865,1613,1639]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="12" pageNumber="13">9H</figureCitation>
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<treatmentCitationGroup id="AB7F5979FFE8FFE7FF1341EDD608F929" box="[189,858,1679,1706]" pageId="12" pageNumber="13">
<treatmentCitation id="0ACE5846FFE8FFE7FF1341EDD604F929" author="Godwin-Austen H. H." box="[189,854,1679,1706]" page="518" pageId="12" pageNumber="13" year="1892">
<taxonomicName id="4C6F05D4FFE8FFE7FF1341EDD604F929" ID-CoL="36M39" authority="Godwin-Austen, 1892: 518" authorityName="Godwin-Austen" authorityPageNumber="518" authorityYear="1892" box="[189,854,1679,1706]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="munipurensis">
<emphasis id="B91BA245FFE8FFE7FF1341EDD759F92A" box="[189,523,1679,1705]" italics="true" pageId="12" pageNumber="13">Diplommatina munipurensis</emphasis>
<bibRefCitation id="EFFE03A6FFE8FFE7FDBC41EDD604F929" author="Godwin-Austen H. H." box="[530,854,1679,1706]" pageId="12" pageNumber="13" pagination="509 - 520" refId="ref13295" refString="Godwin-Austen H. H. 1892. On new species and varieties of the land molluscan genus Diplommatina from the Garo, Naga and Munipur Hill-ranges, Assam. Proceedings of the Zoological Society of London 35: 509 - 520." type="journal article" year="1892">Godwin-Austen, 1892: 518</bibRefCitation>
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.
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</paragraph>
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<treatmentCitationGroup id="AB7F5979FFE8FFE7FF1341B3D1B6F88C" pageId="12" pageNumber="13">
<taxonomicName id="4C6F05D4FFE8FFE7FF1341B3D759F968" ID-CoL="36M39" authorityName="Godwin-Austen" authorityYear="1892" box="[189,523,1745,1771]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="munipurensis">
<emphasis id="B91BA245FFE8FFE7FF1341B3D759F968" box="[189,523,1745,1771]" italics="true" pageId="12" pageNumber="13">Diplommatina munipurensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ACE5846FFE8FFE7FD8641B3D607F968" author="Godwin-Austen H. H." box="[552,853,1745,1771]" page="14" pageId="12" pageNumber="13" year="1897">
<bibRefCitation id="EFFE03A6FFE8FFE7FD8641B3D607F968" author="Godwin-Austen H. H." box="[552,853,1745,1771]" pageId="12" pageNumber="13" pagination="1 - 46" refId="ref13337" refString="Godwin-Austen H. H. 1897. Land and freshwater Mollusca of India including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malay Peninsula, Ceylon and other Islands of the Indian ocean. Conchologia Indica 2 (7): 1 - 46." type="journal article" year="1897">Godwin-Austen 1897: 14</bibRefCitation>
</treatmentCitation>
, pl. 66, fig. 6. —
<treatmentCitation id="0ACE5846FFE8FFE7FB8C41B3D1B1F968" author="Gude G. K." box="[1058,1251,1745,1771]" page="323" pageId="12" pageNumber="13" year="1921">
<bibRefCitation id="EFFE03A6FFE8FFE7FB8C41B3D1B1F968" author="Gude G. K." box="[1058,1251,1745,1771]" pageId="12" pageNumber="13" refId="ref13576" refString="Gude G. K. 1921. The fauna of British India. Mollusca-III (Land operculates: Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Today &amp; Tomorrow's Printers &amp; Publishers, New Delhi." type="book" year="1921">Gude 1921: 323</bibRefCitation>
</treatmentCitation>
.
<taxonomicName id="4C6F05D4FFE8FFE7FF134196D79DF88D" ID-CoL="36M39" authorityName="Godwin-Austen" authorityYear="1892" box="[189,719,1780,1807]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="munipurensis" subGenus="Diplommatina">
<emphasis id="B91BA245FFE8FFE7FF134196D434F88D" box="[189,358,1780,1806]" italics="true" pageId="12" pageNumber="13">Diplommatina</emphasis>
(
<emphasis id="B91BA245FFE8FFE7FED64196D773F88D" box="[376,545,1780,1806]" italics="true" pageId="12" pageNumber="13">Diplommatina</emphasis>
)
<emphasis id="B91BA245FFE8FFE7FD9C4197D79DF88D" box="[562,719,1781,1806]" italics="true" pageId="12" pageNumber="13">munipurensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ACE5846FFE8FFE7FD424196D6ECF88C" author="Kobelt W." box="[748,958,1780,1807]" page="438" pageId="12" pageNumber="13" year="1902">
<bibRefCitation id="EFFE03A6FFE8FFE7FD424196D6ECF88C" author="Kobelt W." box="[748,958,1780,1807]" pageId="12" pageNumber="13" refId="ref13702" refString="Kobelt W. 1902. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. Mollusca: Cyclophoridae, Lief. 16. R. Friedlander und Sohn Verlag, Berlin." type="book" year="1902">Kobelt 1902: 438</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0ACE5846FFE8FFE7FC434196D18CF88C" author="Budha P. B. &amp; Naggs F. &amp; Backeljau T." box="[1005,1246,1780,1807]" page="7" pageId="12" pageNumber="13" year="2015">
<bibRefCitation id="EFFE03A6FFE8FFE7FC434196D18CF88C" author="Budha P. B. &amp; Naggs F. &amp; Backeljau T." box="[1005,1246,1780,1807]" pageId="12" pageNumber="13" pagination="1 - 48" refId="ref13118" refString="Budha P. B., Naggs F. &amp; Backeljau T. 2015. Annotated checklist of the terrestrial gastropods of Nepal. ZooKeys 492: 1 - 48. https: // doi. org / 10.3897 / zookeys. 492.9175" type="journal article" year="2015">
Budha
<emphasis id="B91BA245FFE8FFE7FBEF4197D12EF88D" box="[1089,1148,1780,1806]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2015: 7
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFE8FFE5FF134054D423FD9D" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8BD07E57FFE8FFE7FF134054D464F8D3" blockId="12.[189,310,1846,1872]" box="[189,310,1846,1872]" pageId="12" pageNumber="13">
<heading id="D098C93BFFE8FFE7FF134054D464F8D3" bold="true" box="[189,310,1846,1872]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B91BA245FFE8FFE7FF134054D464F8D3" bold="true" box="[189,310,1846,1872]" pageId="12" pageNumber="13">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFE8FFE5FF134004D423FD9D" blockId="12.[189,1399,1893,2027]" lastBlockId="14.[189,1399,267,542]" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
Measurements (n = 10): SH
<quantity id="4C97D3B2FFE8FFE7FDAB4007D7C7F803" box="[517,661,1893,1920]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="12" pageNumber="13" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.5–2.6 mm</quantity>
, SW
<quantity id="4C97D3B2FFE8FFE7FD7F4004D632F803" box="[721,864,1894,1920]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="12" pageNumber="13" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3–1.4 mm</quantity>
, Wh 6.0–6.5. Dextral, well rounded, regularly increasing, penultimate and body whorls nearly equal in width, suture impressed. Constriction at the middle of the parietal wall (
<figureCitation id="135462D2FFE8FFE7FDAD40CFD733F844" box="[515,609,1965,1991]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="12" pageNumber="13">Fig. 9H</figureCitation>
). One parietalis, moderate (
<figureCitation id="135462D2FFE8FFE7FC0840CFD151F84B" box="[934,1027,1965,1992]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[189,1398,265,1879]" captionTargetId="figure@13.[189,1398,261,1881]" captionTargetPageId="13" captionText="Fig. 5. Diplommatina Benson, 1849 of Nepal. A. D. miriensis Godwin-Austen, 1917 (CDZMTU036). B. D.munipurensis Godwin-Austen, 1892 (CDZMTU039).C. D.oviformis Fulton, 1901 (CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="12" pageNumber="13">Fig. 5B</figureCitation>
<figureCitation id="135462D2FFE8FFE7FBAC40CFD143F84B" box="[1026,1041,1965,1992]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="12" pageNumber="13">3</figureCitation>
). One vertical palatalis, which runs along the constriction, but does not reach the lower half of the whorl. No horizontal palatalis (
<figureCitation id="135462D2FFEAFFE5FF684669D476FEA6" box="[198,292,267,293]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="14" pageNumber="15">Fig. 9H</figureCitation>
). Columellaris close to the umbilicus. Columellar tooth weak. Protoconch smooth, Wh approx 2.0, with small pits distributed all over, no radial ribs (
<figureCitation id="135462D2FFEAFFE5FCFB464DD6E4FEC9" box="[853,950,303,330]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[189,1398,265,1879]" captionTargetId="figure@13.[189,1398,261,1881]" captionTargetPageId="13" captionText="Fig. 5. Diplommatina Benson, 1849 of Nepal. A. D. miriensis Godwin-Austen, 1917 (CDZMTU036). B. D.munipurensis Godwin-Austen, 1892 (CDZMTU039).C. D.oviformis Fulton, 1901 (CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="14" pageNumber="15">Fig. 5B</figureCitation>
<figureCitation id="135462D2FFEAFFE5FC18464DD693FEC9" box="[950,961,303,330]" captionStart="Fig" captionStartId="2.[189,235,1999,2025]" captionTargetBox="[189,1399,1129,1958]" captionTargetId="figure@2.[187,1401,1126,1960]" captionTargetPageId="2" captionText="Fig.1. Map of the localities and areas surveyed for Diplommatina spp. in Nepal." figureDoi="http://doi.org/10.5281/zenodo.3832486" httpUri="https://zenodo.org/record/3832486/files/figure.png" pageId="14" pageNumber="15">1</figureCitation>
). Teleoconch with dense radial ribs; about 8–10 ribs/
<quantity id="4C97D3B2FFEAFFE5FE2E4630D488FEEE" box="[384,474,338,365]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="15" unit="mm" value="0.5">0.5 mm</quantity>
on the penultimate whorl; 10–14 ribs/
<quantity id="4C97D3B2FFEAFFE5FC084630D153FEEE" box="[934,1025,338,365]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="15" unit="mm" value="0.5">0.5 mm</quantity>
on the body whorl towards the aperture (
<figureCitation id="135462D2FFEAFFE5FE804614D4DBFE12" box="[302,393,374,401]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[189,1398,265,1879]" captionTargetId="figure@13.[189,1398,261,1881]" captionTargetPageId="13" captionText="Fig. 5. Diplommatina Benson, 1849 of Nepal. A. D. miriensis Godwin-Austen, 1917 (CDZMTU036). B. D.munipurensis Godwin-Austen, 1892 (CDZMTU039).C. D.oviformis Fulton, 1901 (CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="14" pageNumber="15">Fig. 5B</figureCitation>
<figureCitation id="135462D2FFEAFFE5FE264614D4CAFE12" box="[392,408,374,401]" captionStart="Fig" captionStartId="4.[189,232,1895,1921]" captionTargetBox="[189,1398,285,1853]" captionTargetId="figure@4.[189,1398,285,1853]" captionTargetPageId="4" captionText="Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two)." figureDoi="http://doi.org/10.5281/zenodo.3832488" httpUri="https://zenodo.org/record/3832488/files/figure.png" pageId="14" pageNumber="15">2</figureCitation>
, B
<figureCitation id="135462D2FFEAFFE5FE144614D499FE13" box="[442,459,374,400]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="14" pageNumber="15">4</figureCitation>
). Fine spiral striation present. Umbilicus closed. Aperture circular, parietal wall thin and straight. Peristome with two close lips; angular edge of the peristome straight; outer palatal lip rounded; basal lip along the columellar margin slightly curved; inner lip thin. Operculum paucispiral, corneous, thin, transparent, slightly concave, outer surface smooth, inner surface with raised edge at the central portion.
</paragraph>
</subSubSection>
<caption id="DF102EDFFFE9FFE6FF1340E8D617F868" ID-DOI="http://doi.org/10.5281/zenodo.3832494" ID-Zenodo-Dep="3832494" httpUri="https://zenodo.org/record/3832494/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1930,1956]" targetBox="[189,1398,265,1879]" targetPageId="13">
<paragraph id="8BD07E57FFE9FFE6FF1340E8D617F868" blockId="13.[189,1399,1930,2027]" pageId="13" pageNumber="14">
<emphasis id="B91BA245FFE9FFE6FF1340E8D459F827" bold="true" box="[189,267,1930,1956]" pageId="13" pageNumber="14">Fig. 5.</emphasis>
<taxonomicName id="4C6F05D4FFE9FFE6FEBA40E8D73BF827" authority="Benson, 1849" authorityName="Benson" authorityYear="1849" box="[276,617,1930,1957]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
<emphasis id="B91BA245FFE9FFE6FEBA40E8D4EFF827" box="[276,445,1930,1956]" italics="true" pageId="13" pageNumber="14">Diplommatina</emphasis>
<bibRefCitation id="EFFE03A6FFE9FFE6FE6840E9D73BF827" author="Benson W. H." box="[454,617,1930,1957]" pageId="13" pageNumber="14" pagination="193 - 195" refId="ref12623" refString="Benson W. H. 1849. Characters of Diplommatina, a new genus of terrestrial mollusks belonging to the family of Carychiadae, and of a second species contained in it; also of a new species of Carychium inhabiting the Western Himalaya. Annals and Magazine of Natural History, Series 2 4: 193 - 195. https: // doi. org / 10.1080 / 03745486009494813" type="journal article" year="1849">Benson, 1849</bibRefCitation>
</taxonomicName>
of Nepal.
<emphasis id="B91BA245FFE9FFE6FD4940E8D7AEF827" bold="true" box="[743,764,1930,1956]" pageId="13" pageNumber="14">A</emphasis>
.
<taxonomicName id="4C6F05D4FFE9FFE6FCA240E9D1FEF827" authority="Godwin-Austen, 1917" authorityName="Godwin-Austen" authorityYear="1917" box="[780,1196,1930,1956]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="miriensis">
<emphasis id="B91BA245FFE9FFE6FCA240E9D6CFF827" box="[780,925,1931,1956]" italics="true" pageId="13" pageNumber="14">D. miriensis</emphasis>
<bibRefCitation id="EFFE03A6FFE9FFE6FC0B40E8D1FEF827" author="Godwin-Austen H. H." box="[933,1196,1930,1956]" pageId="13" pageNumber="14" pagination="547 - 559" refId="ref13391" refString="Godwin-Austen H. H. 1917. Zoological results of the Abor Expedition, Mollusca 6. Records of the Indian Museum 8: 547 - 559." type="journal article" year="1917">Godwin-Austen, 1917</bibRefCitation>
</taxonomicName>
(CDZMTU036).
<emphasis id="B91BA245FFE9FFE6FF1340CCD583F84B" bold="true" box="[189,209,1966,1992]" pageId="13" pageNumber="14">B</emphasis>
.
<taxonomicName id="4C6F05D4FFE9FFE6FF7240CCD7CDF84B" authority="Godwin-Austen, 1892" authorityName="Godwin-Austen" authorityYear="1892" box="[220,671,1965,1992]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="munipurensis">
<emphasis id="B91BA245FFE9FFE6FF7240CCD4CBF844" box="[220,409,1966,1991]" italics="true" pageId="13" pageNumber="14">D.munipurensis</emphasis>
<bibRefCitation id="EFFE03A6FFE9FFE6FE3340CFD7CDF84B" author="Godwin-Austen H. H." box="[413,671,1965,1992]" pageId="13" pageNumber="14" pagination="509 - 520" refId="ref13295" refString="Godwin-Austen H. H. 1892. On new species and varieties of the land molluscan genus Diplommatina from the Garo, Naga and Munipur Hill-ranges, Assam. Proceedings of the Zoological Society of London 35: 509 - 520." type="journal article" year="1892">Godwin-Austen, 1892</bibRefCitation>
</taxonomicName>
(CDZMTU039).
<emphasis id="B91BA245FFE9FFE6FCC740CFD62CF844" bold="true" box="[873,894,1965,1991]" pageId="13" pageNumber="14">C</emphasis>
.
<taxonomicName id="4C6F05D4FFE9FFE6FC2440CCD1E2F84B" authority="Fulton, 1901" authorityName="Fulton" authorityYear="1901" box="[906,1200,1965,1992]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="oviformis">
<emphasis id="B91BA245FFE9FFE6FC2440CCD14BF84B" box="[906,1049,1966,1992]" italics="true" pageId="13" pageNumber="14">D.oviformis</emphasis>
<bibRefCitation id="EFFE03A6FFE9FFE6FBB340CFD1E2F84B" author="Fulton H." box="[1053,1200,1965,1992]" pageId="13" pageNumber="14" pagination="242 - 245" refId="ref13161" refString="Fulton H. 1901. Descriptions of some supposed new species of Diplommatina, Opisthostoma, and a new variety of Alycaeus from N. Borneo, Banguey Island and Darjeeling. Annals and Magazine of Natural History, Series 7 8: 242 - 245. https: // doi. org / 10.1080 / 03745480109442915" type="journal article" year="1901">Fulton, 1901</bibRefCitation>
</taxonomicName>
(CDZMTU042). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm.
</paragraph>
</caption>
<subSubSection id="C3752DDCFFEAFFE5FF134524D025FD79" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BD07E57FFEAFFE5FF134524D4F8FDE3" blockId="14.[189,426,582,608]" box="[189,426,582,608]" pageId="14" pageNumber="15">
<heading id="D098C93BFFEAFFE5FF134524D4F8FDE3" bold="true" box="[189,426,582,608]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B91BA245FFEAFFE5FF134524D4F8FDE3" bold="true" box="[189,426,582,608]" pageId="14" pageNumber="15">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFEAFFE5FF134517D025FD79" blockId="14.[189,1399,629,763]" pageId="14" pageNumber="15">
<collectingCountry id="F3783EC7FFEAFFE5FF134517D44CFD0C" box="[189,286,629,655]" name="Nepal" pageId="14" pageNumber="15">NEPAL</collectingCountry>
: 6 shells, Syabrubesi, above Lama Hotel, Langtang National Park,
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,
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,
<geoCoordinate id="EE5B1890FFEAFFE5FF1345FBD401FD30" box="[189,339,665,692]" degrees="85.455117" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="85.45512">85.455117 E</geoCoordinate>
,
<date id="FFD15897FFEAFFE5FECE45FBD4ADFD30" box="[352,511,665,691]" pageId="14" pageNumber="15" value="2008-05-13">13 May 2008</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFEAFFE5FD6345FBD61DFD30" box="[717,847,665,691]" pageId="14" pageNumber="15">CDZMTU</collectionCode>
039); 5 shells, Syabru, Langtang National Park,
<quantity id="4C97D3B2FFEAFFE5FF1345DED445FD54" box="[189,279,700,727]" metricMagnitude="3" metricUnit="m" metricValue="3.2769999999999997" pageId="14" pageNumber="15" unit="m" value="3277.0">3277 m</quantity>
,
<geoCoordinate id="EE5B1890FFEAFFE5FE8D45DED4ECFD54" box="[291,446,700,727]" degrees="28.105855" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="28.105856">28.105855 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFEAFFE5FE6745DED732FD54" box="[457,608,700,727]" degrees="85.326567" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="85.32657">85.326567 E</geoCoordinate>
,
<date id="FFD15897FFEAFFE5FDC245DED7A4FD55" box="[620,758,700,727]" pageId="14" pageNumber="15" value="2007-06-06">6 Jun. 2007</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFEAFFE5FC6C45DED116FD55" box="[962,1092,700,726]" pageId="14" pageNumber="15">CDZMTU</collectionCode>
040); 7 shells, Phulchowki Hill, Lalitpur District,
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,
<geoCoordinate id="EE5B1890FFEAFFE5FD864582D7E6FD79" box="[552,692,736,763]" degrees="27.57159" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="27.57159">27.57159 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFEAFFE5FD114582D607FD79" box="[703,853,736,763]" degrees="85.401647" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="85.40165">85.401647 E</geoCoordinate>
,
<date id="FFD15897FFEAFFE5FCCF4582D6A2FD79" box="[865,1008,736,762]" pageId="14" pageNumber="15" value="2007-05-04">4 May 2007</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFEAFFE5FB124582D012FD79" box="[1212,1344,736,762]" pageId="14" pageNumber="15">CDZMTU</collectionCode>
041).
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFEAFFE5FF134440D606FC31" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BD07E57FFEAFFE5FF134440D47DFCBF" blockId="14.[189,303,802,828]" box="[189,303,802,828]" pageId="14" pageNumber="15">
<heading id="D098C93BFFEAFFE5FF134440D47DFCBF" bold="true" box="[189,303,802,828]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B91BA245FFEAFFE5FF134440D47DFCBF" bold="true" box="[189,303,802,828]" pageId="14" pageNumber="15">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFEAFFE5FF134433D606FC31" blockId="14.[189,1399,849,946]" pageId="14" pageNumber="15">
<bibRefCitation id="EFFE03A6FFEAFFE5FF134433D49EFCE8" author="Godwin-Austen H. H." box="[189,460,849,875]" pageId="14" pageNumber="15" pagination="509 - 520" refId="ref13295" refString="Godwin-Austen H. H. 1892. On new species and varieties of the land molluscan genus Diplommatina from the Garo, Naga and Munipur Hill-ranges, Assam. Proceedings of the Zoological Society of London 35: 509 - 520." type="journal article" year="1892">Godwin-Austen (1892)</bibRefCitation>
gave the
<typeStatus id="54D4C0F5FFEAFFE5FD994433D738FCE8" box="[567,618,849,875]" pageId="14" pageNumber="15">type</typeStatus>
locality as “South of the Barak River, between the Mao villages and Munipur” and noted that “from the number of specimens found, this appears a very abundant species; I never got anything like it in the Khasi Hills to the west”.
</paragraph>
</subSubSection>
</treatment>
</document>
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