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<document id="DEE479B3D179BDD3F930EA7FB20ED15C" ID-DOI="10.11646/zootaxa.4033.3.3" ID-GBIF-Dataset="18e7e755-fb48-49b9-939b-b54073efb595" ID-ISSN="1175-5326" ID-Zenodo-Dep="238956" ID-ZooBank="5A0985CE-3B17-4BD0-866F-7763F1E2120B" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461297466117" checkinUser="plazi" docAuthor="Stroiński, Adam &amp; Szwedo, Jacek" docDate="2015" docId="03BC87C138280217E9E4F885FC616A42" docLanguage="en" docName="zt04033p379.pdf" docOrigin="Zootaxa 4033 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tupala Stroiński &amp; Szwedo, 2015, gen. nov." docType="treatment" docVersion="9" lastPageNumber="371" masterDocId="FF85FFB93829021FE973FFCBFF9B6B5F" masterDocTitle="Why so scarce? Dictyopharidae from Madagascar (Hemiptera: Fulgoromorpha)" masterLastPageNumber="379" masterPageNumber="363" pageNumber="364" updateTime="1698616727771" updateUser="plazi">
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<mods:title id="939B0DAE5FE92AA05A545EB0CF9A4121">Why so scarce? Dictyopharidae from Madagascar (Hemiptera: Fulgoromorpha)</mods:title>
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<mods:namePart id="8399AAF776B21E71A1E4F5ADA70A1F5B">Stroiński, Adam</mods:namePart>
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<treatment id="03BC87C138280217E9E4F885FC616A42" ID-DOI="http://doi.org/10.5281/zenodo.6097611" ID-GBIF-Taxon="119642901" ID-Zenodo-Dep="6097611" LSID="urn:lsid:plazi:treatment:03BC87C138280217E9E4F885FC616A42" httpUri="http://treatment.plazi.org/id/03BC87C138280217E9E4F885FC616A42" lastPageId="8" lastPageNumber="371" pageId="1" pageNumber="364">
<subSubSection id="C30F655C3828021EE9E4F885FED96CD5" pageId="1" pageNumber="364" type="nomenclature">
<paragraph id="8BAA36D73828021EE9E4F885FEFA6C37" blockId="1.[151,353,1870,1930]" box="[151,353,1870,1896]" pageId="1" pageNumber="364">
<heading id="D0E281BB3828021EE9E4F885FEFA6C37" bold="true" box="[151,353,1870,1896]" fontSize="11" level="1" pageId="1" pageNumber="364" reason="1">
<emphasis id="B961EAC53828021EE9E4F885FEFA6C37" bold="true" box="[151,353,1870,1896]" pageId="1" pageNumber="364">
<taxonomicName id="4C154D543828021EE9E4F885FF776C37" ID-CoL="83QY" box="[151,236,1870,1896]" class="Insecta" family="Dictyopharidae" genus="Tupala" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B961EAC53828021EE9E4F885FF776C37" bold="true" box="[151,236,1870,1896]" italics="true" pageId="1" pageNumber="364">Tupala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257BE3828021EE980F885FEFA6C37" box="[243,353,1870,1896]" pageId="1" pageNumber="364" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA36D73828021EE9E4F8BAFED96CD5" blockId="1.[151,353,1870,1930]" box="[151,322,1905,1930]" pageId="1" pageNumber="364">
(
<figureCitation id="132E2A523828021EE9ECF8BAFE896CD5" box="[159,274,1905,1930]" captionStart-0="FIGURES 1 7" captionStart-1="FIGURES 8 13" captionStart-2="FIGURES 14 17" captionStart-3="FIGURES 18 23" captionStart-4="FIGURES 24 29" captionStart-5="FIGURES 30 33" captionStart-6="FIGURES 40 43" captionStart-7="FIGURES 44 50" captionStartId-0="2.[151,264,1868,1891]" captionStartId-1="3.[151,264,1686,1709]" captionStartId-2="4.[151,264,1657,1680]" captionStartId-3="5.[151,264,1659,1682]" captionStartId-4="6.[151,264,1688,1711]" captionStartId-5="7.[151,264,1300,1323]" captionStartId-6="9.[151,264,1405,1428]" captionStartId-7="10.[151,264,1877,1900]" captionTargetBox-0="[164,1416,200,1834]" captionTargetBox-1="[151,1436,193,1652]" captionTargetBox-2="[151,1436,661,1630]" captionTargetBox-3="[151,1436,179,1640]" captionTargetBox-4="[151,1436,193,1653]" captionTargetBox-5="[151,1436,303,1274]" captionTargetBox-6="[169,1412,437,1376]" captionTargetBox-7="[172,1431,448,1851]" captionTargetId-0="figure@2.[151,1436,195,1845]" captionTargetId-1="figure@3.[151,1436,193,1666]" captionTargetId-2="figure@4.[151,1436,661,1636]" captionTargetId-3="figure@5.[151,1436,179,1647]" captionTargetId-4="figure@6.[151,1436,193,1667]" captionTargetId-5="figure@7.[151,1436,303,1279]" captionTargetId-6="figure@9.[169,1413,437,1388]" captionTargetId-7="figure@10.[151,1436,445,1856]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="9" captionTargetPageId-7="10" captionText-0="FIGURES 1 7. Tupala occulta gen. et sp. nov., holotype, male. (1) habitus, lateral view; (2) same, dorsal view; (3) right tegmen, dorsal view; (4 7) anterior part of body: (4) right side, lateral view, (5) left side, lateral view, (6) dorsal view, (7) frontal view." captionText-1="FIGURES 8 13. Tupala occulta gen. et sp. nov., holotype, male. (8) habitus, lateral view; (2) same, dorsal view; (10 12) anterior part of body: (10) dorsal view, (11) dorso-lateral view, (12) frontal view; (13) upper part of head capsule." captionText-2="FIGURES 14 17. Tupala occulta gen. et sp. nov., holotype, male. (14 15) vertex: (14) dorsal view, (15) dorsolateral view; (16 17) pronotum and mesonotum: (16) dorsal view; (17) dorsolateral view." captionText-3="FIGURES 18 23. Tupala occulta gen. et sp. nov., holotype, male. (18) anterior part of body, lateral view; (19 23) antenna and antennal plate organs." captionText-4="FIGURES 24 29. Tupala occulta gen. et sp. nov., holotype, male, tegmen. (24) dorsal view; (25) dorsolateral view; (26) lateral view; (27) apical part, dorsal view; (28) apical part, lateral view; (29) basal part, lateral view." captionText-5="FIGURES 30 33. Tupala occulta gen. et sp. nov., holotype, male, hind leg. (30) hind leg, ventral view; (31) apex of tibia and tarsomere; (32 33) tibia." captionText-6="FIGURES 40 43. Tupala occulta gen. et sp. nov., holotype, male. (40) abdomen, dorsal view; (41) male terminalia, ventral view; (42) hook-line process on gonostylus, ventral view, (43) apex of aedeagus, dorsal view." captionText-7="FIGURES 44 50. Tupala occulta gen. et sp. nov., holotype, male. (44 45) abdomen: (44) dorsal view, (45) ventral view; (46) gonostyles, lateral view; (47 50) aedeagus: (47) dorsal view, (48) lateral view, (49) ventrolateral view, (50) ventral view." httpUri-0="https://zenodo.org/record/238957/files/figure.png" httpUri-1="https://zenodo.org/record/238958/files/figure.png" httpUri-2="https://zenodo.org/record/238959/files/figure.png" httpUri-3="https://zenodo.org/record/238960/files/figure.png" httpUri-4="https://zenodo.org/record/238961/files/figure.png" httpUri-5="https://zenodo.org/record/238962/files/figure.png" httpUri-6="https://zenodo.org/record/238963/files/figure.png" httpUri-7="https://zenodo.org/record/238964/files/figure.png" pageId="1" pageNumber="364">Figs 150</figureCitation>
,
<figureCitation id="132E2A523828021EE86EF8BAFEA16CD5" box="[285,314,1905,1930]" captionStart="FIGURE 55" captionStartId="12.[151,250,1703,1725]" captionText="FIGURE 55. Dictyopharini of Madagascar, distribution map." pageId="1" pageNumber="364">55</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F655C3828021EE9E4F872FC996C8D" box="[151,770,1977,2002]" pageId="1" pageNumber="364" type="materials_examined">
<paragraph id="8BAA36D73828021EE9E4F872FC996C8D" blockId="1.[151,770,1977,2002]" box="[151,770,1977,2002]" pageId="1" pageNumber="364">
<emphasis id="B961EAC53828021EE9E4F872FEA96C8D" bold="true" box="[151,306,1977,2002]" pageId="1" pageNumber="364">
<typeStatus id="54AE88753828021EE9E4F872FF496C8D" box="[151,210,1977,2002]" pageId="1" pageNumber="364">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C154D543828021EE84AF872FE7B6C8D" box="[313,480,1977,2002]" class="Insecta" family="Dictyopharidae" genus="Tupala" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="species" species="occulta" status="sp. nov.">
<emphasis id="B961EAC53828021EE84AF872FE7B6C8D" box="[313,480,1977,2002]" italics="true" pageId="1" pageNumber="364">Tupala occulta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257BE3828021EE894F871FDDB6C8E" box="[487,576,1978,2001]" pageId="1" pageNumber="364" rank="species">sp. nov.</taxonomicNameLabel>
, here designated.
</paragraph>
</subSubSection>
<caption id="DF6A665F382B021DE9E4F887FE8C6CFF" httpUri="https://zenodo.org/record/238957/files/figure.png" pageId="2" pageNumber="365" targetBox="[164,1416,200,1834]" targetPageId="2">
<paragraph id="8BAA36D7382B021DE9E4F887FE8C6CFF" blockId="2.[151,1436,1868,1952]" pageId="2" pageNumber="365">
<emphasis id="B961EAC5382B021DE9E4F887FEAD6C3D" bold="true" box="[151,310,1868,1891]" pageId="2" pageNumber="365">FIGURES 17</emphasis>
.
<taxonomicName id="4C154D54382B021DE835F886FE7A6C3C" box="[326,481,1869,1891]" class="Insecta" family="Dictyopharidae" genus="Tupala" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="species" species="occulta">
<emphasis id="B961EAC5382B021DE835F886FE7A6C3C" box="[326,481,1869,1891]" italics="true" pageId="2" pageNumber="365">Tupala occulta</emphasis>
</taxonomicName>
<emphasis id="B961EAC5382B021DE899F886FD146C3D" bold="true" box="[490,655,1869,1890]" pageId="2" pageNumber="365">gen. et sp. nov.</emphasis>
, holotype, male. (1) habitus, lateral view; (2) same, dorsal view; (3) right tegmen, dorsal view; (47) anterior part of body: (4) right side, lateral view, (5) left side, lateral view, (6) dorsal view, (7) frontal view.
</paragraph>
</caption>
<caption id="DF6A665F382A021CE9E4F95DFA886D93" httpUri="https://zenodo.org/record/238958/files/figure.png" pageId="3" pageNumber="366" targetBox="[151,1436,193,1652]" targetPageId="3">
<paragraph id="8BAA36D7382A021CE9E4F95DFA886D93" blockId="3.[151,1435,1686,1740]" pageId="3" pageNumber="366">
<emphasis id="B961EAC5382A021CE9E4F95DFED86DF3" bold="true" box="[151,323,1686,1709]" pageId="3" pageNumber="366">FIGURES 813</emphasis>
.
<taxonomicName id="4C154D54382A021CE821F95CFE776DF2" box="[338,492,1687,1709]" class="Insecta" family="Dictyopharidae" genus="Tupala" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="366" phylum="Arthropoda" rank="species" species="occulta">
<emphasis id="B961EAC5382A021CE821F95CFE776DF2" box="[338,492,1687,1709]" italics="true" pageId="3" pageNumber="366">Tupala occulta</emphasis>
</taxonomicName>
<emphasis id="B961EAC5382A021CE886F95CFD016DF3" bold="true" box="[501,666,1687,1708]" pageId="3" pageNumber="366">gen. et sp. nov.</emphasis>
, holotype, male. (8) habitus, lateral view; (2) same, dorsal view; (1012) anterior part of body: (10) dorsal view, (11) dorso-lateral view, (12) frontal view; (13) upper part of head capsule.
</paragraph>
</caption>
<subSubSection id="C30F655C382A021CE9B4F933FDEC6CFE" pageId="3" pageNumber="366" type="etymology">
<paragraph id="8BAA36D7382A021CE9B4F933FDEC6CFE" blockId="3.[151,1436,1784,2025]" pageId="3" pageNumber="366">
<emphasis id="B961EAC5382A021CE9B4F933FED56C4E" bold="true" box="[199,334,1784,1809]" pageId="3" pageNumber="366">Etymology.</emphasis>
Generic name is an anagram of dictyopharid generic name
<taxonomicName id="4C154D54382A021CEA99F933FBAE6C4E" box="[1002,1077,1784,1809]" class="Insecta" family="Dictyopharidae" genus="Putala" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="366" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAC5382A021CEA99F933FBAE6C4E" box="[1002,1077,1784,1809]" italics="true" pageId="3" pageNumber="366">Putala</emphasis>
</taxonomicName>
. Gender: feminine.
<emphasis id="B961EAC5382A021CE9B4F8D7FEDA6C6A" bold="true" box="[199,321,1820,1845]" pageId="3" pageNumber="366">Diagnosis.</emphasis>
Differs from
<taxonomicName id="4C154D54382A021CE89DF8D7FDCF6C6A" box="[494,596,1820,1845]" class="Insecta" family="Dictyopharidae" genus="Zaputala" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="366" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAC5382A021CE89DF8D7FDCF6C6A" box="[494,596,1820,1845]" italics="true" pageId="3" pageNumber="366">Zaputala</emphasis>
</taxonomicName>
(
<figureCitation id="132E2A52382A021CEB1BF8D7FD746C6A" box="[616,751,1820,1845]" captionStart="FIGURES 51 54" captionStartId="11.[151,264,1752,1775]" captionTargetBox="[199,1411,202,1720]" captionTargetId="figure@11.[162,1424,193,1732]" captionTargetPageId="11" captionText="FIGURES 51 54. Zaputala bourgoini Emeljanov, 2008, holotype, female. (51) habitus, dorsal view; (52) same, ventral view; same, lateral view; (54) labels. Photo: Laurent Fauvre (MNHN, Paris)." httpUri="https://zenodo.org/record/238965/files/figure.png" pageId="3" pageNumber="366">Figs 5154</figureCitation>
) by: presence of longitudinal carina of vertex (absent in
<taxonomicName id="4C154D54382A021CE9E4F88BFF676C06" box="[151,252,1856,1881]" class="Insecta" family="Dictyopharidae" genus="Zaputala" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="366" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAC5382A021CE9E4F88BFF676C06" box="[151,252,1856,1881]" italics="true" pageId="3" pageNumber="366">Zaputala</emphasis>
</taxonomicName>
); tricarinate pronotum (single carina in
<taxonomicName id="4C154D54382A021CEB96F88BFCD16C06" box="[741,842,1856,1881]" class="Insecta" family="Dictyopharidae" genus="Zaputala" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="366" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAC5382A021CEB96F88BFCD16C06" box="[741,842,1856,1881]" italics="true" pageId="3" pageNumber="366">Zaputala</emphasis>
</taxonomicName>
); frons with median carina (absent in
<taxonomicName id="4C154D54382A021CEC56F88BFA116C06" box="[1317,1418,1856,1881]" class="Insecta" family="Dictyopharidae" genus="Zaputala" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="366" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAC5382A021CEC56F88BFA116C06" box="[1317,1418,1856,1881]" italics="true" pageId="3" pageNumber="366">Zaputala</emphasis>
</taxonomicName>
); different organization of head capsule extension. Lateral portions of pronotum with obsolete granulation (lateral portions of pronotum smooth in
<taxonomicName id="4C154D54382A021CEB71F843FDFC6CFE" box="[514,615,1928,1953]" class="Insecta" family="Dictyopharidae" genus="Zaputala" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="366" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAC5382A021CEB71F843FDFC6CFE" box="[514,615,1928,1953]" italics="true" pageId="3" pageNumber="366">Zaputala</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C30F655C382A0217E9B4F867FE0B6BA7" lastPageId="8" lastPageNumber="371" pageId="3" pageNumber="366" type="description">
<paragraph id="8BAA36D7382A021BE9B4F867FAC76B8A" blockId="3.[151,1436,1784,2025]" lastBlockId="4.[151,1436,151,608]" lastPageId="4" lastPageNumber="367" pageId="3" pageNumber="366">
<emphasis id="B961EAC5382A021CE9B4F867FEC26C9A" bold="true" box="[199,345,1964,1989]" pageId="3" pageNumber="366">Description.</emphasis>
Head with compound eyes distinctly narrower than pronotum. Vertex longer than wide at base, tapering apicad; anterior margin arcuate, lateral margins with shallow incision at about half of their length, posterior margin shallowly convex, indistinct; lateral margins carinate, anterior margin thickened, callose; disc of vertex flat, with faint longitudinal carina. Posterior margin of vertex at posterior ⅓ of compound eyes length.
</paragraph>
<paragraph id="8BAA36D7382D021BE9B4FF2BFE016900" blockId="4.[151,1436,151,608]" pageId="4" pageNumber="367">
Frons longer than wide at frontoclypeal suture, with lateral margins carinate, merely sigmoid, diverging towards frontoclypeal suture, with breaking point above the level of antennae base, then more distinctly diverging, lateral carinae reaching to half of frons length, in upper portion, forming obtuse extension, convex in upper portion towards margin with vertex, concave in lower portion towards disc of frons, median carina reaching frontoclypeal suture; disc of frons concave laterad, with median convexity at lower ⅓ of its length. Frontoclypeal suture straight, not distinct. Postclypeus with distinct lateral carinae and median carina, disc of clypeus convex, anteclypeus distinguishable, convex, with lateral and median carinae. Compound eyes with small posteroventral callus. Lateral ocelli present. Antennal fovea deep, emarginated; scapus short, pedicel bulbiform, with sensory plates in upper portion distributed in apical half, and sensory plates of lower hemisphere distributed in basal and apical parts; trichoid sensilla
<typeStatus id="54AE8875382D021BE825FDEEFE1D6962" box="[342,390,549,573]" pageId="4" pageNumber="367">type</typeStatus>
I present. Rostrum with apical segment almost as long as subapical one, apex of rostrum reaching to metacoxae.
</paragraph>
<caption id="DF6A665F382D021BE9E4F9B2FC066DF1" httpUri="https://zenodo.org/record/238959/files/figure.png" pageId="4" pageNumber="367" targetBox="[151,1436,661,1630]" targetPageId="4">
<paragraph id="8BAA36D7382D021BE9E4F9B2FC066DF1" blockId="4.[151,1436,1657,1710]" pageId="4" pageNumber="367">
<emphasis id="B961EAC5382D021BE9E4F9B2FED46DD0" bold="true" box="[151,335,1657,1680]" pageId="4" pageNumber="367">FIGURES 1417</emphasis>
.
<taxonomicName id="4C154D54382D021BE82EF9B1FE6D6DCF" box="[349,502,1658,1680]" class="Insecta" family="Dictyopharidae" genus="Tupala" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="occulta">
<emphasis id="B961EAC5382D021BE82EF9B1FE6D6DCF" box="[349,502,1658,1680]" italics="true" pageId="4" pageNumber="367">Tupala occulta</emphasis>
</taxonomicName>
<emphasis id="B961EAC5382D021BE88DF9B1FD3B6DD0" bold="true" box="[510,672,1658,1679]" pageId="4" pageNumber="367">gen. et sp. nov.</emphasis>
, holotype, male. (1415) vertex: (14) dorsal view, (15) dorsolateral view; (1617) pronotum and mesonotum: (16) dorsal view; (17) dorsolateral view.
</paragraph>
</caption>
<paragraph id="8BAA36D7382D021BE9B4F91CFCD46C21" blockId="4.[151,1437,1749,2027]" pageId="4" pageNumber="367">Pronotum heptagonal, with disc slightly elevated, delimited by straight anterior margin, incomplete anterolateral carinae, strongly diverging posteriad, not reaching to posterior margin, with distinct median carina, posterior margin of pronotum widely triangularly incised, with small but distinct triangular incision at terminus of median carina; lateral portions of pronotum descending lateroposteriad, with oblique postocular carina, reaching posterior margin, and lateral carina reaching posterior margin.</paragraph>
<paragraph id="8BAA36D7382D021BE9B4F841FDBA6CFC" blockId="4.[151,1437,1749,2027]" box="[199,545,1930,1955]" pageId="4" pageNumber="367">Tegulae large, without carinae.</paragraph>
<paragraph id="8BAA36D7382D021BE9B4F864FBF46CB4" blockId="4.[151,1437,1749,2027]" pageId="4" pageNumber="367">Mesonotum wider than long in mid line, with distinct median carina, reaching to level of scutellum and lateral carinae reaching posterior margin, delimiting flattened disc, lateral margins descending.</paragraph>
<caption id="DF6A665F382C021AE9E4F9B0FEEE6DEE" httpUri="https://zenodo.org/record/238960/files/figure.png" pageId="5" pageNumber="368" targetBox="[151,1436,179,1640]" targetPageId="5">
<paragraph id="8BAA36D7382C021AE9E4F9B0FEEE6DEE" blockId="5.[151,1435,1659,1713]" pageId="5" pageNumber="368">
<emphasis id="B961EAC5382C021AE9E4F9B0FED66DCE" bold="true" box="[151,333,1659,1682]" pageId="5" pageNumber="368">FIGURES 1823</emphasis>
.
<taxonomicName id="4C154D54382C021AE82AF9B7FE6B6DCD" box="[345,496,1660,1682]" class="Insecta" family="Dictyopharidae" genus="Tupala" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="occulta">
<emphasis id="B961EAC5382C021AE82AF9B7FE6B6DCD" box="[345,496,1660,1682]" italics="true" pageId="5" pageNumber="368">Tupala occulta</emphasis>
</taxonomicName>
<emphasis id="B961EAC5382C021AE886F9B7FD0A6DCE" bold="true" box="[501,657,1660,1681]" pageId="5" pageNumber="368">gen. et sp. nov.</emphasis>
, holotype, male. (18) anterior part of body, lateral view; (1923) antenna and antennal plate organs.
</paragraph>
</caption>
<paragraph id="8BAA36D7382C0218E9B4F912FB2E6BA4" blockId="5.[151,1437,1752,2038]" lastBlockId="7.[151,1436,151,251]" lastPageId="7" lastPageNumber="370" pageId="5" pageNumber="368">
Tegmen membranous, macropterous, with venation distinct; apex of clavus exceeding ⅔ of tegmen length. Costal margin arcuate, veins of costal complex fused, anteroapical angle wide, apex rounded, apical point at level of middle of tegmen width, posteroapical angle wide, tornus long. Basal cell narrow, elongate, stems ScP+R and M leaving basal cell with a short common stalk, about half as long as basal cell; stem ScP+R long, slightly curved, subparallel to costal margin, forked posteriad of stems M and CuA forkings, merely basad of level of claval apex, branch ScP+RA reaching margin with three terminals, terminal ScP+RA1 reaching margin apicad of claval apex; branch
<collectingCountry id="F3027647382C021AE999F87FFE976C94" box="[234,268,1972,1995]" name="Poland" pageId="5" pageNumber="368">RP</collectingCountry>
forked apicad of RA3 terminus at margin, reaching margin with two terminals. Stem M merely sigmoid, forked slightly basad of claval apex, basad of stem ScP+R forking, branch M1+2 forked slightly apicad of branch M3+4 forking, then M1 forked again on membrane, forking of branch M3+4 slightly apicad of claval apex, then branches M3 and M4 forked again on membrane, then stem M reaching margin with 7 terminals. Stem CuA straight, long, forked apicad of claval veins junction, branch CuA1 forked subapically on membrane, branch CuA2 single. Claval veins Pcu and A1 fused before half of claval suture (CuP) length. Veinlet
<emphasis id="B961EAC5382F0219ED45F8A4FB246CD9" box="[1078,1215,1903,1926]" italics="true" pageId="6" pageNumber="369">pccacp-scra</emphasis>
delimiting anterior margin of stigmal area placed slightly apicad of transverse veinlet
<emphasis id="B961EAC5382F0219EAD2F85EFC4C6CF2" box="[929,983,1941,1965]" italics="true" pageId="6" pageNumber="369">1r-m</emphasis>
, apicad of claval apex, callose at costal margin. Nodal veinlet
<emphasis id="B961EAC5382F0219E8EAF872FE546C8E" box="[409,463,1977,2001]" italics="true" pageId="6" pageNumber="369">1r-m</emphasis>
at level claval apex; nodal veinlet
<emphasis id="B961EAC5382F0219EA17F872FC316C8E" box="[868,938,1977,2001]" italics="true" pageId="6" pageNumber="369">1m-cu</emphasis>
slightly oblique, basad of
<emphasis id="B961EAC5382F0219EDAEF872FA8F6C8E" box="[1245,1300,1977,2001]" italics="true" pageId="6" pageNumber="369">1r-m</emphasis>
; veinlet
<emphasis id="B961EAC5382F0219EC09F871FA066C8E" box="[1402,1437,1978,2001]" italics="true" pageId="6" pageNumber="369">icu</emphasis>
oblique, reaching margin of tornus apicad of claval apex. Costal cell without transverse veinlets. Membrane with net of transverse veinlets delimiting polygonal cells. Stigmal area wide, dissected by ScP+RA1 and RA2 terminals. Cell C1 pentagonal, slightly longer than wide; cell C3 elongately pentagonal, longer than wide, about twice as long as C1; cell C5 lanceolate, tapering posteriad, more than twice as long as cell C3.
</paragraph>
<caption id="DF6A665F382F0219E9E4F953FB106D92" httpUri="https://zenodo.org/record/238961/files/figure.png" pageId="6" pageNumber="369" targetBox="[151,1436,193,1653]" targetPageId="6">
<paragraph id="8BAA36D7382F0219E9E4F953FB106D92" blockId="6.[151,1435,1688,1741]" pageId="6" pageNumber="369">
<emphasis id="B961EAC5382F0219E9E4F953FECB6DF2" bold="true" box="[151,336,1688,1711]" pageId="6" pageNumber="369">FIGURES 2429</emphasis>
.
<taxonomicName id="4C154D54382F0219E813F953FE606DF1" box="[352,507,1688,1710]" class="Insecta" family="Dictyopharidae" genus="Tupala" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="369" phylum="Arthropoda" rank="species" species="occulta">
<emphasis id="B961EAC5382F0219E813F953FE606DF1" box="[352,507,1688,1710]" italics="true" pageId="6" pageNumber="369">Tupala occulta</emphasis>
</taxonomicName>
<emphasis id="B961EAC5382F0219EB77F952FD316DF1" bold="true" box="[516,682,1689,1710]" pageId="6" pageNumber="369">gen. et sp. nov.</emphasis>
, holotype, male, tegmen. (24) dorsal view; (25) dorsolateral view; (26) lateral view; (27) apical part, dorsal view; (28) apical part, lateral view; (29) basal part, lateral view.
</paragraph>
</caption>
<caption id="DF6A665F382E0218E9E4FADFFE086E15" httpUri="https://zenodo.org/record/238962/files/figure.png" pageId="7" pageNumber="370" targetBox="[151,1436,303,1274]" targetPageId="7">
<paragraph id="8BAA36D7382E0218E9E4FADFFE086E15" blockId="7.[151,1436,1300,1354]" pageId="7" pageNumber="370">
<emphasis id="B961EAC5382E0218E9E4FADFFED66E75" bold="true" box="[151,333,1300,1323]" pageId="7" pageNumber="370">FIGURES 3033</emphasis>
.
<taxonomicName id="4C154D54382E0218E829FADEFE696E74" box="[346,498,1301,1323]" class="Insecta" family="Dictyopharidae" genus="Tupala" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="occulta">
<emphasis id="B961EAC5382E0218E829FADEFE696E74" box="[346,498,1301,1323]" italics="true" pageId="7" pageNumber="370">Tupala occulta</emphasis>
</taxonomicName>
<emphasis id="B961EAC5382E0218E88BFADDFD0E6E74" bold="true" box="[504,661,1302,1323]" pageId="7" pageNumber="370">gen. et sp. nov.</emphasis>
, holotype, male, hind leg. (30) hind leg, ventral view; (31) apex of tibia and tarsomere; (3233) tibia.
</paragraph>
</caption>
<paragraph id="8BAA36D7382E0218E9B4FAB9FF4E6DD9" blockId="7.[151,1437,1393,2030]" pageId="7" pageNumber="370">Legs long, slender, tibiae not flattened. Profemur longer than mesofemur. Metatibia longer than metafemur, with four lateral spines and 8 apical teeth arrange in row. Metabasitarsomere about as long as cumulative length of mid and apical metatarsomeres; Basimetatarsomere with row of apical teeth; subconical lateral ones and row of 22 dorsoventrally flattened and elongated teeth with subapical needle-shaped platellae, laterally compressed, with dorsal fuller in median portion. Midmetatarsomere with row of apical teethe, subconical lateral ones, and row of 18 dorsoventrally flattened and elongated teeth with subapical needle-shaped platellae, laterally compressed, with dorsal fuller in median portion. Apical metatarsomere about as long as preceding one; tarsal claws distinct, arolium wide.</paragraph>
<paragraph id="8BAA36D7382E0218E9B4F95AFAF66C01" blockId="7.[151,1437,1393,2030]" pageId="7" pageNumber="370">Male terminalia: pygofer laterally compressed, distinctly higher than long in lateral view, posterior margin smooth, with posterodorsal portion declining posteriad. Anal tube distinctly shorter than gonostyles, tubular, with short, triangular, posteroventral projections; anus placed apically. Gonostyles symmetrical, not fused; base narrow, expanded towards middle, than mildly tapering towards apex; dorsobasal margin straight, dorsoapical margin straight, apical portion bluntly rounded, ventral margin widely arcuate; upper margin with dorsal process in the middle, acutely tapering basad, hook-like process near sub-middle in lateral view, closer to dorsobasal margin.</paragraph>
<paragraph id="8BAA36D7382E0217E9B4F8A2FD986B8A" blockId="7.[151,1437,1393,2030]" lastBlockId="8.[151,1436,151,285]" lastPageId="8" lastPageNumber="371" pageId="7" pageNumber="370">Phallic complex almost symmetrical, with periandrium composed of two dorsolateral membranous sacs, with ventroapical sclerotized hemicircular lobes and sclerotized, double spine like processes present in laterobasal portion, dorsal spine distinctly bigger than ventral one; apical part of dorsal periandrium with two elongate, tapered, sclerotized processes. Aedeagus membranous sac-like, distinctly longer than periandrium, with double row of longitudinal pine-like processes, increasing in size from aedeagus base and posterodorsal portion covered with spiniferous microsculpture.</paragraph>
<paragraph id="8BAA36D738210217E9B4FF14FE0B6BA7" blockId="8.[151,1436,151,285]" box="[199,400,223,248]" pageId="8" pageNumber="371">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C30F655C38210217E9B4FECFFC616A42" box="[199,1018,260,285]" pageId="8" pageNumber="371" type="distribution">
<paragraph id="8BAA36D738210217E9B4FECFFC616A42" blockId="8.[151,1436,151,285]" box="[199,1018,260,285]" pageId="8" pageNumber="371">
<emphasis id="B961EAC538210217E9B4FECFFEFB6A42" bold="true" box="[199,352,260,285]" pageId="8" pageNumber="371">Distribution.</emphasis>
<collectingCountry id="F302764738210217E814FECFFE6B6A42" box="[359,496,260,285]" name="Madagascar" pageId="8" pageNumber="371">Madagascar</collectingCountry>
, Antsiranana Province, Sava Region (
<figureCitation id="132E2A5238210217EAEAFECFFC766A42" box="[921,1005,260,285]" captionStart="FIGURE 55" captionStartId="12.[151,250,1703,1725]" captionText="FIGURE 55. Dictyopharini of Madagascar, distribution map." pageId="8" pageNumber="371">Fig. 55</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>