202 lines
20 KiB
XML
202 lines
20 KiB
XML
<document id="79B08F8ADEB83BA331E6086E5E675B1A" ID-DOI="10.5281/zenodo.195747" ID-GBIF-Dataset="c371f527-f7c6-456b-ab07-5bef541b10e6" ID-ISSN="1175-5326" ID-Zenodo-Dep="195747" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460232772886" checkinUser="plazi" docAuthor="Saether, Ole A., Andersen, Trond, Pinho, Luiz C. & Mendes, Humberto F." docDate="2010" docId="03BC87900E12FFCCFF44C012CD0C71DC" docLanguage="en" docName="zt02497p036.pdf" docOrigin="Zootaxa 2497" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Polypedilum" docType="treatment" docVersion="10" lastPageNumber="14" masterDocId="FF85FFE80E1EFFC1FFD3C738CC1E7200" masterDocTitle="The problems with Polypedilum Kieffer (Diptera: Chironomidae), with the description of Probolum subgen. n." masterLastPageNumber="36" masterPageNumber="1" pageNumber="13" updateTime="1698239121165" updateUser="plazi">
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<mods:mods id="3826775EE18DD6D3E26FAC2AB19643AE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="70535A2E888877147D5658D22707E27A">
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<mods:title id="5162BD0D32951822B43E7B9043AB49C0">The problems with Polypedilum Kieffer (Diptera: Chironomidae), with the description of Probolum subgen. n.</mods:title>
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</mods:titleInfo>
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<mods:name id="2ED795F433E2583AFD96A46DE651723A" type="personal">
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<mods:role id="3BCBD90789EFE0152E1C25BAC76C745E">
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<mods:roleTerm id="CA82C4152BC3CC7544BE07CE270DA0B8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="27AFB5930E2683E35BC3618C1588AA2B">Saether, Ole A.</mods:namePart>
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</mods:name>
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<mods:name id="FA00F4B1470A5585579202610A9EEB04" type="personal">
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<mods:role id="C7CE305E3699382CF8757EA56E322C34">
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<mods:roleTerm id="E60BA33D3F7A02F96A271BA34CC73C30">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9CC9BAF06358833D4916CCA7B166112D">Andersen, Trond</mods:namePart>
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</mods:name>
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<mods:name id="BEEED93F0BAFE57D5AA36ECE05A428C8" type="personal">
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<mods:role id="4BF745EA627201AFE2CFB58D67DB5558">
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<mods:roleTerm id="F3DCB04BE1083D18970FC6121EDC2F65">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7E7BAFFF27B8DE75AB4B540531ABF010">Pinho, Luiz C.</mods:namePart>
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</mods:name>
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<mods:name id="29148C41C284E3425B392E4BB1018F6A" type="personal">
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<mods:roleTerm id="49E1A6D5C192B1E473B42C45BE2686E7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="422C7F02A8A2267B2E3F01FC3C985EC6">Mendes, Humberto F.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="536DC55455D9E83DF2E1FC75FEAB089B">text</mods:typeOfResource>
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<mods:titleInfo id="B55F31B370F0CEA985128646B53A3B1B">
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<mods:title id="887B7E3E8964A8C9DDD32FC77B132BEB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="655C4FE0330A1A4850C3EBDEBEF85220">
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<mods:date id="DFFF534CCEF8874818439A5A447A268A">2010</mods:date>
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<mods:detail id="024B68526598D1928FDDDF6F54F092F7" type="volume">
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<mods:number id="75D685861E70FC482E2322333C7315C7">2497</mods:number>
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</mods:detail>
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<mods:extent id="A48080E920E2B2C84F1C270533A79513" unit="page">
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<mods:start id="3ABF185C355D26F7CDCB3BBADA4C3591">1</mods:start>
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<mods:end id="37A75D03418A5D4A0078F7DD1DF00FC8">36</mods:end>
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<mods:classification id="601434D1A57FA732A8C5E755C747AC52">journal article</mods:classification>
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<mods:identifier id="7B93FDE3930DBED3CDB51240EFEBC626" type="DOI">10.5281/zenodo.195747</mods:identifier>
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<mods:identifier id="3AD57F6AB39AB118433B38E2519CF03A" type="GBIF-Dataset">c371f527-f7c6-456b-ab07-5bef541b10e6</mods:identifier>
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<mods:identifier id="08D83C50D434859BD7F2C6E0B73D61F5" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5112EBCADAC15DF3BEFEF539D7225C33" type="Zenodo-Dep">195747</mods:identifier>
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<treatment id="03BC87900E12FFCCFF44C012CD0C71DC" ID-DOI="http://doi.org/10.5281/zenodo.5030728" ID-GBIF-Taxon="119387542" ID-Zenodo-Dep="5030728" LSID="urn:lsid:plazi:treatment:03BC87900E12FFCCFF44C012CD0C71DC" httpUri="http://treatment.plazi.org/id/03BC87900E12FFCCFF44C012CD0C71DC" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
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<subSubSection id="C30F650D0E12FFCDFF44C012CF337544" box="[151,813,1834,1860]" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="8BAA36860E12FFCDFF44C012CF337544" blockId="12.[151,813,1834,1860]" box="[151,813,1834,1860]" pageId="12" pageNumber="13">
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<heading id="D0E281EA0E12FFCDFF44C012CF337544" bold="true" box="[151,813,1834,1860]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
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<emphasis id="B961EA940E12FFCDFF44C012CF337544" bold="true" box="[151,813,1834,1860]" pageId="12" pageNumber="13">
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<taxonomicName id="4C154D050E12FFCDFF44C012CD307544" ID-CoL="63MXN" ID-ENA="54968" box="[151,302,1834,1860]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EA940E12FFCDFF44C012CD307544" bold="true" box="[151,302,1834,1860]" italics="true" pageId="12" pageNumber="13">Polypedilum</emphasis>
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</taxonomicName>
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subgenus
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<taxonomicName id="4C154D050E12FFCDFE63C012CF337544" authority="Oyewo et Saether" authorityName="Oyewo et Saether" box="[432,813,1834,1860]" class="Insecta" family="Chironomidae" genus="Uresipedilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EA940E12FFCDFE63C012CE4D7544" bold="true" box="[432,595,1834,1860]" italics="true" pageId="12" pageNumber="13">Uresipedilum</emphasis>
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Oyewo
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<emphasis id="B961EA940E12FFCDFD6BC012CED37544" bold="true" box="[696,717,1834,1860]" italics="true" pageId="12" pageNumber="13">et</emphasis>
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Saether
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F650D0E12FFCDFF44C04ACF4575C9" pageId="12" pageNumber="13" type="reference_group">
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<paragraph id="8BAA36860E12FFCDFF44C04ACF4575C9" blockId="12.[151,1008,1906,1993]" pageId="12" pageNumber="13">
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<treatmentCitationGroup id="AB0511A80E12FFCDFF44C04ACF4575C9" pageId="12" pageNumber="13">
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<taxonomicName id="4C154D050E12FFCDFF44C04ACD007589" ID-CoL="63MXN" box="[151,286,1906,1929]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EA940E12FFCDFF44C04ACD007589" box="[151,286,1906,1929]" italics="true" pageId="12" pageNumber="13">Polypedilum</emphasis>
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</taxonomicName>
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subgenus
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<taxonomicName id="4C154D050E12FFCDFE5DC04ACF597589" authority="Sasa et Okazawa, 1991: 54" authorityName="Sasa et Okazawa" authorityPageNumber="54" authorityYear="1991" box="[398,839,1906,1929]" class="Insecta" family="Chironomidae" genus="Uresipedilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EA940E12FFCDFE5DC04ACE007589" box="[398,542,1906,1929]" italics="true" pageId="12" pageNumber="13">Uresipedilum</emphasis>
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<treatmentCitation id="0AB410970E12FFCDFDF6C04ACF597589" author="Sasa" box="[549,839,1906,1929]" page="54" pageId="12" pageNumber="13" year="1991">
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<bibRefCitation id="EF844B770E12FFCDFDF6C04ACF3E7589" author="Sasa" box="[549,800,1906,1929]" pageId="12" pageNumber="31" refString="Sasa, M. & Okazawa, T. (1991) Studies on the chironomids of the Joganji River and other places. Part 1. Studies on the chironomids of the Joganji River, Toyama. Research Report from Toyama Prefectural Environmental Pollution Research Centre, 1991, 52 - 67." type="journal article" year="1991">
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Sasa
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<emphasis id="B961EA940E12FFCDFD8FC04ACE717589" box="[604,623,1906,1929]" italics="true" pageId="12" pageNumber="13">et</emphasis>
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Okazawa, 1991
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</bibRefCitation>
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: 54
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</treatmentCitation>
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</taxonomicName>
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,
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<emphasis id="B961EA940E12FFCDFC81C04ACD0075A9" italics="true" pageId="12" pageNumber="13">
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nomen nudum.
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<taxonomicName id="4C154D050E12FFCDFF44C0AACD0075A9" ID-CoL="63MXN" box="[151,286,1938,1961]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Polypedilum</taxonomicName>
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</emphasis>
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subgenus
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<taxonomicName id="4C154D050E12FFCDFE5DC0AACF5875A9" authority="Sasa et Kikuchi, 1995: 119" authorityName="Sasa et Kikuchi" authorityPageNumber="119" authorityYear="1995" box="[398,838,1938,1961]" class="Insecta" family="Chironomidae" genus="Uresipedilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EA940E12FFCDFE5DC0AACE0075A9" box="[398,542,1938,1961]" italics="true" pageId="12" pageNumber="13">Uresipedilum</emphasis>
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<treatmentCitation id="0AB410970E12FFCDFDF6C0AACF5875A9" author="Sasa" box="[549,838,1938,1961]" page="119" pageId="12" pageNumber="13" year="1995">
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<bibRefCitation id="EF844B770E12FFCDFDF6C0AACF0D75A9" author="Sasa" box="[549,787,1938,1961]" pageId="12" pageNumber="31" refString="Sasa, M. & Kikuchi, M. (1995) Chironomidae (Diptera) of Japan. University of Tokyo Press, Tokyo, 333 pp." type="book" year="1995">
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Sasa
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<emphasis id="B961EA940E12FFCDFD8FC0AACE7175A9" box="[604,623,1938,1961]" italics="true" pageId="12" pageNumber="13">et</emphasis>
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Kikuchi, 1995
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</bibRefCitation>
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: 119
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</treatmentCitation>
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</taxonomicName>
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,
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<emphasis id="B961EA940E12FFCDFC82C0AACD0075C9" italics="true" pageId="12" pageNumber="13">
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nomen nudum.
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<taxonomicName id="4C154D050E12FFCDFF44C08ACD0075C9" ID-CoL="63MXN" box="[151,286,1970,1993]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Polypedilum</taxonomicName>
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</emphasis>
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subgenus
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<taxonomicName id="4C154D050E12FFCDFE5DC08ACF4975C9" authority="Oyewo et Saether, 1998: 317" authorityName="Oyewo et Saether" authorityPageNumber="317" authorityYear="1998" box="[398,855,1970,1993]" class="Insecta" family="Chironomidae" genus="Uresipedilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EA940E12FFCDFE5DC08ACE0075C9" box="[398,542,1970,1993]" italics="true" pageId="12" pageNumber="13">Uresipedilum</emphasis>
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<treatmentCitation id="0AB410970E12FFCDFDF6C08ACF4975C9" author="Oyewo" box="[549,855,1970,1993]" page="317" pageId="12" pageNumber="13" year="1998">
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<bibRefCitation id="EF844B770E12FFCDFDF6C08ACF3C75C9" author="Oyewo" box="[549,802,1970,1993]" lastPageId="12" lastPageNumber="31" pageId="12" pageNumber="30" refString="Oyewo, E. A. & Saether, O. A. (1998) Revision of Afrotropical Polypedilum Kieffer subgen. Uresipedilum Sasa et Kikuchi, 1995 (Diptera: Chironomidae), with a review of the subgenus. Annales de Limnologie, 34, 315 - 362." type="journal article" year="1998">
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Oyewo
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<emphasis id="B961EA940E12FFCDFDABC08ACE9275C9" box="[632,652,1970,1993]" italics="true" pageId="12" pageNumber="13">et</emphasis>
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Saether, 1998
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</bibRefCitation>
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: 317
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F650D0E13FFCCFF44C7A0CD3572B2" box="[151,299,152,178]" pageId="13" pageNumber="14" type="materials_examined">
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<paragraph id="8BAA36860E13FFCCFF44C7A0CD3572B2" blockId="13.[151,723,152,244]" box="[151,299,152,178]" pageId="13" pageNumber="14">
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<typeStatus id="54AE88240E13FFCCFF44C7A0CCD172B2" box="[151,207,152,178]" pageId="13" pageNumber="14">Type</typeStatus>
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species
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F650D0E13FFCCFF44C782CFE073AC" pageId="13" pageNumber="14" type="reference_group">
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<paragraph id="8BAA36860E13FFCCFF44C782CE9972F4" blockId="13.[151,723,152,244]" pageId="13" pageNumber="14">
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<treatmentCitationGroup id="AB0511A80E13FFCCFF44C782CE9972F4" pageId="13" pageNumber="14">
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<taxonomicName id="4C154D050E13FFCCFF44C782CECD72D4" ID-CoL="8DG86" authority="Walker" authorityName="Walker" box="[151,723,186,212]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="convictum" subGenus="Uresipedilum">
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<emphasis id="B961EA940E13FFCCFF44C782CE7D72D3" box="[151,611,186,211]" italics="true" pageId="13" pageNumber="14">Polypedilum (Uresipedilum) convictum</emphasis>
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(Walker)
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</taxonomicName>
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<taxonomicName id="4C154D050E13FFCCFF14C7E5CE9D72F4" authority="Walker, 1856: 161" authorityName="Walker" authorityPageNumber="161" authorityYear="1856" box="[199,643,221,244]" class="Insecta" family="Chironomidae" genus="Chironomus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="convictus">
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<emphasis id="B961EA940E13FFCCFF14C7E5CDAB72F4" box="[199,437,221,244]" italics="true" pageId="13" pageNumber="14">Chironomus convictus</emphasis>
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<treatmentCitation id="0AB410970E13FFCCFE6FC7E5CE9D72F4" author="Walker" box="[444,643,221,244]" page="161" pageId="13" pageNumber="14" year="1856">
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<bibRefCitation id="EF844B770E13FFCCFE6FC7E5CE5372F4" author="Walker" box="[444,589,221,244]" pageId="13" pageNumber="32" refString="Walker, F. (1856) Insecta Britannica, Diptera. Volume 3. Reeve & Benham, London, xxiv + 352 pp." type="book" year="1856">Walker, 1856</bibRefCitation>
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: 161
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BAA36860E13FFCCFF44C61ACFE073AC" blockId="13.[151,1437,290,988]" pageId="13" pageNumber="14">
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<emphasis id="B961EA940E13FFCCFF44C61ACDB1733C" bold="true" box="[151,431,290,316]" pageId="13" pageNumber="14">Diagnostic characters.</emphasis>
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The male imagines of the subgenus are distinguished by having the basal portion of the superior volsella much longer than wide, with an apicomedian projection without setae arising from the inner margin of the base and directed medially and without prominent inner projection; wing membrane without markings or setae and fore tibial scale nearly always without spur.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F650D0E13FFCCFF17C680CE7470FC" pageId="13" pageNumber="14" type="description">
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<paragraph id="8BAA36860E13FFCCFF17C680CD977042" blockId="13.[151,1437,290,988]" pageId="13" pageNumber="14">Pupae of the subgenus lack dorsal setae on the anal lobe; have few branches in the thoracic horn; reduced cephalic tubercles and mostly single anal spur with few teeth, but often numerous fine lateral spinules. Conjunctive III/IV is often without spinules and pedes spurii A often absent. The anal lobe is nearly always without dorsal setae.</paragraph>
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<paragraph id="8BAA36860E13FFCCFF17C575CE7470FC" blockId="13.[151,1437,290,988]" pageId="13" pageNumber="14">Larvae of the subgenus are distinguished by having the four median teeth set off from the rest of the mentum and in contact with the anteriorly produced median ends of the ventromental plates and with well developed to at least indicated posterior lobes on the ventromental plates. Antennal segment 3 is very slightly to more distinctly shorter than segment 4. The mentum has the first lateral teeth much or only slightly lower than median and/or second lateral teeth.</paragraph>
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</subSubSection>
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<subSubSection id="C30F650D0E13FFCCFF15C43FCD0C71DC" pageId="13" pageNumber="14" type="discussion">
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<paragraph id="8BAA36860E13FFCCFF15C43FCDF37147" blockId="13.[151,1437,290,988]" pageId="13" pageNumber="14">
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<emphasis id="B961EA940E13FFCCFF15C43FCD217121" bold="true" box="[198,319,775,801]" pageId="13" pageNumber="14">Remarks.</emphasis>
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The pupae of
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<taxonomicName id="4C154D050E13FFCCFDDEC430CEB57121" box="[525,683,776,801]" class="Insecta" family="Chironomidae" genus="Uresipedilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EA940E13FFCCFDDEC430CEB57121" box="[525,683,776,801]" italics="true" pageId="13" pageNumber="14">Uresipedilum</emphasis>
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</taxonomicName>
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are not distinguishable from those of
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<emphasis id="B961EA940E13FFCCFB79C430C98B7121" box="[1194,1429,776,801]" italics="true" pageId="13" pageNumber="14">
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<taxonomicName id="4C154D050E13FFCCFB79C430C9207121" box="[1194,1342,776,801]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" sensu="stricto">Polypedilum</taxonomicName>
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<taxonomicNameLabel id="A25257EF0E13FFCCFA9DC430C98B7121" box="[1358,1429,776,801]" pageId="13" pageNumber="14" sensu="stricto">s. str.</taxonomicNameLabel>
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</emphasis>
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,
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<taxonomicName id="4C154D050E13FFCCFF44C415CD247146" box="[151,314,813,838]" class="Insecta" family="Chironomidae" genus="Pentapedilum" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EA940E13FFCCFF44C415CD247146" box="[151,314,813,838]" italics="true" pageId="13" pageNumber="14">Pentapedilum</emphasis>
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</taxonomicName>
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and
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<emphasis id="B961EA940E13FFCCFEA0C415CDF97146" box="[371,487,813,838]" italics="true" pageId="13" pageNumber="14">Probolum</emphasis>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BAA36860E13FFCCFF17C46ACD0C71DC" blockId="13.[151,1437,290,988]" pageId="13" pageNumber="14">
|
|
<taxonomicName id="4C154D050E13FFCCFF17C46ACD72716B" box="[196,364,850,875]" class="Insecta" family="Chironomidae" genus="Uresipedilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
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|
<emphasis id="B961EA940E13FFCCFF17C46ACD72716B" box="[196,364,850,875]" italics="true" pageId="13" pageNumber="14">Uresipedilum</emphasis>
|
|
</taxonomicName>
|
|
as previously defined is not monophyletic as it includes the new subgenus
|
|
<emphasis id="B961EA940E13FFCCFAF1C46AC982716B" box="[1314,1436,850,875]" italics="true" pageId="13" pageNumber="14">Probolum</emphasis>
|
|
described below. If the association of the immatures described by
|
|
<bibRefCitation id="EF844B770E13FFCCFC04C440C9437192" author="Grodhaus" box="[983,1373,888,914]" lastPageId="13" lastPageNumber="30" pageId="13" pageNumber="29" refString="Grodhaus, G. & Rotramel, G. L. (1980) Immature stages of Polypedilum pedatum excelsius (Diptera: Chironomidae) from seasonally flooded treeholes. Acta Universitatis Carolinae - Biologica, 1978, 69 - 76." type="journal article" year="1980">Grodhaus and Rotramel (1980)</bibRefCitation>
|
|
with
|
|
<taxonomicName id="4C154D050E13FFCCFF44C4A5CE6B71B7" authority="Townes" authorityName="Townes" box="[151,629,925,951]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="pedatum" subSpecies="excelsius">
|
|
<emphasis id="B961EA940E13FFCCFF44C4A5CE0F71B6" box="[151,529,925,950]" italics="true" pageId="13" pageNumber="14">Polypedilum pedatum excelsius</emphasis>
|
|
Townes
|
|
</taxonomicName>
|
|
is correct not all imagines of
|
|
<taxonomicName id="4C154D050E13FFCCFC0CC4A5C89E71B6" box="[991,1152,925,950]" class="Insecta" family="Chironomidae" genus="Uresipedilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B961EA940E13FFCCFC0CC4A5C89E71B6" box="[991,1152,925,950]" italics="true" pageId="13" pageNumber="14">Uresipedilum</emphasis>
|
|
</taxonomicName>
|
|
may be separable from
|
|
<emphasis id="B961EA940E13FFCCFF44C4FACD1571DB" box="[151,267,962,987]" italics="true" pageId="13" pageNumber="14">Probolum</emphasis>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |