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<document id="CE72A735E0C3FFA276302A1F436F5CDD" ID-DOI="10.26028/cybium/2017-412-009" ID-ISSN="2101-0315" ID-Zenodo-Dep="13484144" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1723042309133" checkinUser="felipe" docAuthor="Hoese, Douglass F., Allen, Gerald R. &amp; Hadiaty, Renny K." docDate="2017" docId="03BB87FBFFBCF80B2FCBFF6DED6CC6CE" docLanguage="en" docName="Cybium.41.2.179-193.pdf" docOrigin="Cybium 41 (2)" docSource="http://dx.doi.org/10.26028/cybium/2017-412-009" docStyle="DocumentStyle:73DAD84611431DC6606F8B63B2C6039C.1:Cybium.2013-2018.journal_article.type1" docStyleId="73DAD84611431DC6606F8B63B2C6039C" docStyleName="Cybium.2013-2018.journal_article.type1" docStyleVersion="1" docTitle="Glossogobius pumilus Hoese, Allen &amp; Hadiaty, 2017, new species" docType="treatment" docVersion="3" lastPageNumber="192" masterDocId="FF82FF83FFB6F8062C66FFB4EF47C07C" masterDocTitle="Description of three new species of dwarf Glossogobius from New Guinea and northern Australia" masterLastPageNumber="193" masterPageNumber="179" pageNumber="189" updateTime="1724937597013" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="33D1986E9A695030F77C0A30E557D678">Description of three new species of dwarf Glossogobius from New Guinea and northern Australia</mods:title>
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<mods:namePart id="E0148C59064B6648D1A48892150868C1">Hoese, Douglass F.</mods:namePart>
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<mods:namePart id="75C2F69099F38ED6089E9B9B15297963">Allen, Gerald R.</mods:namePart>
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<mods:namePart id="2E79E4AA63BBA81FA1040A0070B1AAED">Hadiaty, Renny K.</mods:namePart>
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<mods:title id="0671DDBFADF3FAE53B6C9B930D93C3FF">Cybium</mods:title>
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<paragraph id="8BAD36EDFFBCF80C2FCBFF6DEA68C08D" blockId="10.[941,1327,217,242]" box="[941,1327,217,242]" pageId="10" pageNumber="189">
<heading id="D0E58181FFBCF80C2FCBFF6DEA68C08D" bold="true" box="[941,1327,217,242]" centered="true" fontSize="9" level="2" pageId="10" pageNumber="189" reason="2">
<emphasis id="B966EAFFFFBCF80C2FCBFF6DEA68C08D" bold="true" box="[941,1327,217,242]" pageId="10" pageNumber="189">
<emphasis id="B966EAFFFFBCF80C2FCBFF6DEBE1C08D" bold="true" box="[941,1190,217,241]" italics="true" pageId="10" pageNumber="189">
<taxonomicName id="4C124D6EFFBCF80C2FCBFF6DEBE7C08D" authorityName="Hoese &amp; Allen &amp; Hadiaty" authorityYear="2017" box="[941,1184,217,241]" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="189" phylum="Chordata" rank="species" species="pumilus" status="sp. nov.">Glossogobius pumilus</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2555784FFBCF80C28CAFF6EEA68C08D" box="[1196,1327,217,242]" pageId="10" pageNumber="189" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8BAD36EDFFBCF80C2FFCFF4EEA06C16F" blockId="10.[922,1345,250,275]" box="[922,1345,250,275]" pageId="10" pageNumber="189">
(
<figureCitation id="13292A68FFBCF80C2FC7FF4EECB0C16F" box="[929,1015,250,275]" captionStart="Figure 1" captionStartId="1.[132,195,1250,1272]" captionTargetBox="[138,775,221,1227]" captionTargetId="figure-676@1.[132,775,215,1233]" captionTargetPageId="1" captionText="Figure 1. Sensory papillae in three dwarf species of Glossogobius, composite drawing, based largely on holotypes. A: G. gnomus; B: G. nanus; C: G. pumilus. Numbers indicate papilla lines after Akhito and Meguro (1975) and in descriptions here. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13484154" httpUri="https://zenodo.org/record/13484154/files/figure.png" pageId="10" pageNumber="189">Figs 1C</figureCitation>
,
<figureCitation id="13292A68FFBCF80C2864FF4FEB07C16E" box="[1026,1088,251,275]" captionStart="Figure 4" captionStartId="2.[1077,1142,1693,1715]" captionTargetBox="[134,1038,1499,1898]" captionTargetId="figure-687@2.[132,1039,1499,1901]" captionTargetPageId="2" captionText="Figure 4. Dark area indicating anterior extent of predorsal and prepelvic scales and head pores in various populations and species of dwarf Glossogobius. A: Glossogobius gnomus; B: G. nanus, Australia; C: G. nanus, Fly River; D: G. pumilus, Bensbach River; E: G. pumilus, Timika region. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13484166" httpUri="https://zenodo.org/record/13484166/files/figure.png" pageId="10" pageNumber="189">4D, E</figureCitation>
,
<figureCitation id="13292A68FFBCF80C282BFF4FEB1BC16F" box="[1101,1116,251,275]" captionStart="Figure 9" captionStartId="11.[132,195,422,444]" captionTargetBox="[548,1455,215,486]" captionTargetId="figure-435@11.[548,1455,215,486]" captionTargetPageId="11" captionText="Figure 9. Preserved holotype of Glos- sogobius pumilus, WAM P.27823-011, 32 mm SL male." figureDoi="http://doi.org/10.5281/zenodo.13484194" httpUri="https://zenodo.org/record/13484194/files/figure.png" pageId="10" pageNumber="189">9</figureCitation>
,
<figureCitation id="13292A68FFBCF80C2801FF4FEBC3C16F" box="[1127,1156,251,275]" captionStart="Figure 10" captionStartId="11.[132,199,653,675]" captionTargetBox="[560,1449,545,725]" captionTargetId="figure-433@11.[548,1455,525,741]" captionTargetPageId="11" captionText="Figure 10. Freshly collected para- type of Glossogobius pumilus, WAM P.27823-011, 33 mm SL male from Bensbach River. Photo G.Allen" figureDoi="http://doi.org/10.5281/zenodo.13484202" httpUri="https://zenodo.org/record/13484202/files/figure.png" pageId="10" pageNumber="189">10</figureCitation>
,
<figureCitation id="13292A68FFBCF80C28E8FF4FEBEEC16F" box="[1166,1193,251,275]" captionStart="Figure 11" captionStartId="11.[132,195,1171,1193]" captionTargetBox="[548,1455,779,1235]" captionTargetId="figure-434@11.[548,1455,779,1235]" captionTargetPageId="11" captionText="Figure 11. Live individual from Timi- ka area of Glossogobius pumilus. Photo G. Allen." figureDoi="http://doi.org/10.5281/zenodo.13484206" httpUri="https://zenodo.org/record/13484206/files/figure.png" pageId="10" pageNumber="189">11</figureCitation>
, 12; Tabs I-V)
</paragraph>
<paragraph id="8BAD36EDFFBCF80C2F34FE8FEAE8C12F" blockId="10.[813,1455,315,372]" box="[850,1455,315,339]" pageId="10" pageNumber="189">
<taxonomicName id="4C124D6EFFBCF80C2F34FE8FEB64C12F" box="[850,1059,315,339]" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="189" phylum="Chordata" rank="species" species="undefined-2">
<emphasis id="B966EAFFFFBCF80C2F34FE8FECA1C12F" box="[850,998,315,339]" italics="true" pageId="10" pageNumber="189">Glossogobius</emphasis>
sp. 2
</taxonomicName>
.
<bibRefCitation id="EF834B1CFFBCF80C2827FE8FEBB9C12F" author="ALLEN G. R. &amp; HORTLE K. G. &amp; RENYAAN S. J." box="[1089,1278,315,339]" pageId="10" pageNumber="189" refId="ref13686" refString="ALLEN G. R., HORTLE K. G. &amp; RENYAAN S. J., 2000. - Freshwater Fishes of the Timika Region, New Guinea. 175 p. Timika: Freeport Indonesia." type="book" year="2000">
Allen
<emphasis id="B966EAFFFFBCF80C28E2FE88EB85C12F" box="[1156,1218,315,339]" italics="true" pageId="10" pageNumber="189">et al.,</emphasis>
2000
</bibRefCitation>
(in part, Timika
</paragraph>
<paragraph id="8BAD36EDFFBCF80C2F4BFEE8EB5BC108" blockId="10.[813,1455,315,372]" box="[813,1052,348,372]" pageId="10" pageNumber="189">
Region, New
<collectingCountry id="F305767DFFBCF80C2FD9FEE8EB56C108" box="[959,1041,348,372]" name="Guinea-Bissau" pageId="10" pageNumber="189">Guinea</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C3086566FFBCF80C2F4BFE28EA00C322" pageId="10" pageNumber="189" type="materials_examined">
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<emphasis id="B966EAFFFFBCF80C2F4BFE28EB43C1C8" bold="true" box="[813,1028,412,436]" pageId="10" pageNumber="189">Material examined</emphasis>
</paragraph>
<paragraph id="8BAD36EDFFBCF80C2F34FE09EB51C26B" blockId="10.[813,1456,412,862]" pageId="10" pageNumber="189">
<materialsCitation id="3B7A3CB0FFBCF80C2F34FE09EB51C26B" collectingDate="1982-10-01" collectionCode="WAM" collectorName="G. Allen &amp; J. Paska." latitude="-8.883333" longLatPrecision="1300" longitude="141.2" pageId="10" pageNumber="189" specimenCount="1" typeStatus="holotype">
<emphasis id="B966EAFFFFBCF80C2F34FE09ECFAC1A9" box="[850,957,445,469]" italics="true" pageId="10" pageNumber="189">
<typeStatus id="54A9884FFFBCF80C2F34FE09ECFFC1A9" box="[850,952,445,469]" pageId="10" pageNumber="189" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED03AE28FFBCF80C2FB0FE0AEB5FC1AA" box="[982,1048,446,470]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="10" pageNumber="189" type="Museum">WAM</collectionCode>
P.27823-011,
<quantity id="4CEA9B08FFBCF80C28D5FE09EBB9C1A9" box="[1203,1278,445,469]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="10" pageNumber="189" unit="mm" value="32.0">32 mm</quantity>
SL male,
<quantity id="4CEA9B08FFBCF80C290DFE09EAE8C1A9" box="[1387,1455,445,469]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="189" unit="km" value="17.0">17 km</quantity>
upstream of Bensbach lodge,
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,
<geoCoordinate id="EE26502AFFBCF80C28DDFE6AEA66C18A" box="[1211,1313,478,502]" degrees="141" direction="east" minutes="12" orientation="longitude" pageId="10" pageNumber="189" precision="925" value="141.2">141°12E</geoCoordinate>
,
<date id="FFAC102DFFBCF80C2948FE6AEAECC18A" box="[1326,1451,478,502]" pageId="10" pageNumber="189" value="1982-10-01">
<collectingDate id="EFE8E9C5FFBCF80C2948FE6AEAECC18A" box="[1326,1451,478,502]" pageId="10" pageNumber="189" value="1982-10-01">1 Oct. 1982</collectingDate>
</date>
,
<collectorName id="26E7533BFFBCF80C2F4BFE4BECC1C26B" box="[813,902,511,535]" pageId="10" pageNumber="189">G. Allen</collectorName>
and
<collectorName id="26E7533BFFBCF80C2FDFFE4BEB51C26B" box="[953,1046,511,535]" pageId="10" pageNumber="189">J. Paska.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BAD36EDFFBCF80C2F34FD94EA00C322" blockId="10.[813,1456,412,862]" pageId="10" pageNumber="189">
<materialsCitation id="3B7A3CB0FFBCF80C2F34FD94EC89C224" country="Papua New Guinea" location="Papua New Guinea" pageId="10" pageNumber="189" specimenCount="1" typeStatus="paratype">
<emphasis id="B966EAFFFFBCF80C2F34FD94ECA7C24B" box="[850,992,544,567]" italics="true" pageId="10" pageNumber="189">
<typeStatus id="54A9884FFFBCF80C2F34FD94EC80C24B" box="[850,967,544,567]" pageId="10" pageNumber="189" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<emphasis id="B966EAFFFFBCF80C2F8EFD94EB8AC24B" bold="true" box="[1000,1229,543,568]" pageId="10" pageNumber="189">
<collectingCountry id="F305767DFFBCF80C2F8EFD94EB8AC24B" box="[1000,1229,543,568]" name="Papua New Guinea" pageId="10" pageNumber="189">Papua New Guinea</collectingCountry>
</emphasis>
: AMS I.27507-001, 31 (21-32) and
</materialsCitation>
<materialsCitation id="3B7A3CB0FFBCF80C2FB3FDF4ECD5C205" collectionCode="WAM" country="Papua New Guinea" pageId="10" pageNumber="189" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED03AE28FFBCF80C2FB3FDF4EB50C224" box="[981,1047,576,600]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="10" pageNumber="189" type="Museum">WAM</collectionCode>
P.27823-001, 109(13-35), taken with
<typeStatus id="54A9884FFFBCF80C2F4BFDD5ECD5C205" box="[813,914,609,633]" pageId="10" pageNumber="189" type="holotype">holotype</typeStatus>
</materialsCitation>
;
<materialsCitation id="3B7A3CB0FFBCF80C2FFAFDD5EB40C2C6" collectingDate="1982-09-29" collectionCode="WAM" collectorName="G. Allen &amp; J. Paska" country="Papua New Guinea" latitude="-8.85" location="Bensbach" longLatPrecision="1300" longitude="141.21666" pageId="10" pageNumber="189" specimenCount="1" stateProvince="Papua Barat" typeStatus="paratype">
<collectionCode id="ED03AE28FFBCF80C2FFAFDD5EC98C205" box="[924,991,609,633]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="10" pageNumber="189" type="Museum">WAM</collectionCode>
P.27817-006, 2(31-32),
<quantity id="4CEA9B08FFBCF80C2888FDD5EA62C205" box="[1262,1317,609,633]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="189" unit="km" value="5.0">5 km</quantity>
upstream of
<location id="8ECD6036FFBCF80C2F4BFD36ECD0C2E6" LSID="urn:lsid:plazi:treatment:03BB87FBFFBCF80B2FCBFF6DED6CC6CE:8ECD6036FFBCF80C2F4BFD36ECD0C2E6" box="[813,919,642,666]" country="Papua New Guinea" latitude="-8.85" longLatPrecision="1300" longitude="141.21666" name="Bensbach" pageId="10" pageNumber="189" stateProvince="Papua Barat">Bensbach</location>
lodge,
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,
<geoCoordinate id="EE26502AFFBCF80C2827FD36EBEEC2E5" box="[1089,1193,642,666]" degrees="141" direction="east" minutes="13" orientation="longitude" pageId="10" pageNumber="189" precision="925" value="141.21666">141°13E</geoCoordinate>
,
<date id="FFAC102DFFBCF80C28D1FD36EA0FC2E6" box="[1207,1352,642,666]" pageId="10" pageNumber="189" value="1982-09-29">
<collectingDate id="EFE8E9C5FFBCF80C28D1FD36EA0FC2E6" box="[1207,1352,642,666]" pageId="10" pageNumber="189" value="1982-09-29">29 Sep. 1982</collectingDate>
</date>
,
<collectorName id="26E7533BFFBCF80C2935FD36EAE8C2E6" box="[1363,1455,642,666]" pageId="10" pageNumber="189">G. Allen</collectorName>
and
<collectorName id="26E7533BFFBCF80C2F3FFD17ECF4C2C6" box="[857,947,674,698]" pageId="10" pageNumber="189">J. Paska</collectorName>
;
<collectingRegion id="49D6F80FFFBCF80C2FDBFD16EB40C2C6" box="[957,1031,674,698]" country="Indonesia" name="Papua Barat" pageId="10" pageNumber="189">
<emphasis id="B966EAFFFFBCF80C2FDBFD16EB40C2C6" bold="true" box="[957,1031,674,698]" pageId="10" pageNumber="189">Papua</emphasis>
</collectingRegion>
</materialsCitation>
<emphasis id="B966EAFFFFBCF80C2861FD16EB4BC2C6" bold="true" box="[1031,1036,674,698]" pageId="10" pageNumber="189">,</emphasis>
<materialsCitation id="3B7A3CB0FFBCF80C2874FD16ECC6C2A7" collectionCode="WAM" country="Indonesia" location="Indonesia" pageId="10" pageNumber="189" specimenCount="1" typeStatus="paratype">
<emphasis id="B966EAFFFFBCF80C2874FD16EBC6C2C6" bold="true" box="[1042,1153,674,698]" pageId="10" pageNumber="189">
<collectingCountry id="F305767DFFBCF80C2874FD16EBC6C2C6" box="[1042,1153,674,698]" name="Indonesia" pageId="10" pageNumber="189">Indonesia</collectingCountry>
</emphasis>
:
<collectionCode id="ED03AE28FFBCF80C28E8FD17EB88C2C7" box="[1166,1231,675,699]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="10" pageNumber="189" type="Museum">WAM</collectionCode>
P.31296-003, 11(25- 40) and
</materialsCitation>
<materialsCitation id="3B7A3CB0FFBCF80C2FEEFD70EB23C361" collectingDate="1997-04-22" collectionCode="MZB" collectorName="G. Allen &amp; Party" country="Indonesia" latitude="-4.6245003" location="Portsite" longLatPrecision="922" longitude="136.95" municipality="Timika" pageId="10" pageNumber="189" specimenCode="MZB 23337, 2" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB49E96FFBCF80C2FEEFD70EB55C2A7" box="[904,1042,707,731]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="189" type="Museum">MZB 23337</specimenCode>
,
<specimenCode id="DBB49E96FFBCF80C2878FD77EB6AC2A7" box="[1054,1069,707,731]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="189" type="Museum">2</specimenCode>
(29-32), south of
<collectingMunicipality id="6BC9AC97FFBCF80C2889FD77EA06C2A7" box="[1263,1345,707,731]" pageId="10" pageNumber="189">Timika</collectingMunicipality>
, E side of
<location id="8ECD6036FFBCF80C2F4BFD50EC39C280" LSID="urn:lsid:plazi:treatment:03BB87FBFFBCF80B2FCBFF6DED6CC6CE:8ECD6036FFBCF80C2F4BFD50EC39C280" box="[813,894,740,764]" country="Indonesia" latitude="-4.6245003" longLatPrecision="922" longitude="136.95" municipality="Timika" name="Portsite" pageId="10" pageNumber="189">Portsite</location>
road, approximate km 24,
<date id="FFAC102DFFBCF80C28FCFD50EA61C280" box="[1178,1318,740,764]" pageId="10" pageNumber="189" value="1997-04-22">
<collectingDate id="EFE8E9C5FFBCF80C28FCFD50EA61C280" box="[1178,1318,740,764]" pageId="10" pageNumber="189" value="1997-04-22">22 Apr. 1997</collectingDate>
</date>
,
<geoCoordinate id="EE26502AFFBCF80C2949FD50EAEEC280" box="[1327,1449,740,764]" degrees="04" direction="south" minutes="37.47" orientation="latitude" pageId="10" pageNumber="189" precision="9" value="-4.6245003">04°37.47S</geoCoordinate>
,
<geoCoordinate id="EE26502AFFBCF80C2F4BFCB0ECD4C360" box="[813,915,772,796]" degrees="136" direction="east" minutes="57" orientation="longitude" pageId="10" pageNumber="189" precision="925" value="136.95">136°57E</geoCoordinate>
,
<collectorName id="26E7533BFFBCF80C2FF9FCB1ECB0C360" box="[927,1015,772,797]" pageId="10" pageNumber="189">G. Allen</collectorName>
and
<collectorName id="26E7533BFFBCF80C284EFCB1EB23C361" box="[1064,1124,773,797]" pageId="10" pageNumber="189">Party</collectorName>
</materialsCitation>
;
<materialsCitation id="3B7A3CB0FFBCF80C280AFCB1EA03C322" collectingDate="2000-08-07" collectionCode="WAM" collectorName="Allen &amp; Hortle" country="Indonesia" latitude="-4.6833334" location="Timika" longLatPrecision="925" longitude="136.90517" pageId="10" pageNumber="189" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED03AE28FFBCF80C280AFCB1EBEAC361" box="[1132,1197,773,797]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="10" pageNumber="189" type="Museum">WAM</collectionCode>
P.31756-002, 22(15-29) and AMS I.47250-001, 6(21-27), south of
<location id="8ECD6036FFBCF80C2893FC91EA02C341" LSID="urn:lsid:plazi:treatment:03BB87FBFFBCF80B2FCBFF6DED6CC6CE:8ECD6036FFBCF80C2893FC91EA02C341" box="[1269,1349,805,829]" country="Indonesia" latitude="-4.6833334" longLatPrecision="925" longitude="136.90517" name="Timika" pageId="10" pageNumber="189">Timika</location>
,
<geoCoordinate id="EE26502AFFBCF80C2936FC91EAEFC341" box="[1360,1448,805,829]" degrees="04" direction="south" minutes="41" orientation="latitude" pageId="10" pageNumber="189" precision="925" value="-4.6833334">04°41S</geoCoordinate>
,
<geoCoordinate id="EE26502AFFBCF80C2F4BFCF2ECF3C321" box="[813,948,838,862]" degrees="136" direction="east" minutes="54.31" orientation="longitude" pageId="10" pageNumber="189" precision="9" value="136.90517">136°54.31E</geoCoordinate>
,
<date id="FFAC102DFFBCF80C2FA7FCF2EB01C322" box="[961,1094,838,862]" pageId="10" pageNumber="189" value="2000-08-07">
<collectingDate id="EFE8E9C5FFBCF80C2FA7FCF2EB01C322" box="[961,1094,838,862]" pageId="10" pageNumber="189" value="2000-08-07">7 Aug. 2000</collectingDate>
</date>
, G.
<collectorName id="26E7533BFFBCF80C2808FCF2EBEEC322" box="[1134,1193,838,862]" pageId="10" pageNumber="189">Allen</collectorName>
and K.
<collectorName id="26E7533BFFBCF80C289AFCF2EA03C322" box="[1276,1348,838,862]" pageId="10" pageNumber="189">Hortle</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3086566FFBCF80C2F4BFC32ECC6C5B6" pageId="10" pageNumber="189" type="diagnosis">
<paragraph id="8BAD36EDFFBCF80C2F4BFC32ECDCC3E2" blockId="10.[813,1456,902,1482]" box="[813,923,902,926]" pageId="10" pageNumber="189">
<emphasis id="B966EAFFFFBCF80C2F4BFC32ECDCC3E2" bold="true" box="[813,923,902,926]" pageId="10" pageNumber="189">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BAD36EDFFBCF80C2F34FC13ECC6C5B6" blockId="10.[813,1456,902,1482]" pageId="10" pageNumber="189">Mouth small, reaching to below a point between anterior margin of pupil and anterior margin of eye; mental fraenum a small rounded protrusion or triangular, without distinct lateral lobes; predorsal partly scaled, midline scaled forward in adult to above a point between middle of operculum and just behind posterior preopercular margin; cheek and operculum naked; pectoral-fin base and prepelvic area naked; pelvic disc thin and oval, longer than wide and reaching to anus in adults; first dorsal fin with black spot posteriorly surrounding area near base of sixth dorsal spine; second dorsal rays usually I,9; anal rays usually I,7; pectoral rays 13-16; longitudinal scale count 24-27; predorsal scale count 0-9; transverse scale count (TRB) usually 7.5; vertebrae usually 11+16 = 27, preoperculum without pores, no lateral canal tube above operculum; lateral canal pore between infraorbital pore present in adults and terminal lateral canal pore present.</paragraph>
</subSubSection>
<subSubSection id="C3086566FFBCF80B2F4BFA47EFA0C54E" lastPageId="13" lastPageNumber="192" pageId="10" pageNumber="189" type="description">
<paragraph id="8BAD36EDFFBCF80C2F4BFA47ECF6C677" blockId="10.[813,1456,1523,1908]" box="[813,945,1523,1547]" pageId="10" pageNumber="189">
<emphasis id="B966EAFFFFBCF80C2F4BFA47ECF6C677" bold="true" box="[813,945,1523,1547]" pageId="10" pageNumber="189">Description</emphasis>
</paragraph>
<paragraph id="8BAD36EDFFBCF80C2F34F9A0EBEEC70F" blockId="10.[813,1456,1523,1908]" pageId="10" pageNumber="189">
Counts based on
<specimenCount id="9D14FD64FFBCF80C287CF9A0EBFEC650" box="[1050,1209,1556,1580]" count="59" pageId="10" pageNumber="189" type="generic">59 specimens</specimenCount>
, measurements taken on
<specimenCount id="9D14FD64FFBCF80C2F29F981ECA4C631" box="[847,995,1589,1613]" count="24" pageId="10" pageNumber="189" type="generic">24 specimens</specimenCount>
<quantity id="4CEA9B08FFBCF80C2F8CF981EB1CC631" box="[1002,1115,1589,1613]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="4.1" metricValueMin="2.4" pageId="10" pageNumber="189" unit="mm" value="32.5" valueMax="41.0" valueMin="24.0">24-41 mm</quantity>
SL. First dorsal spines 6(40*); second dorsal rays I,8-10, usually I,9; anal rays I,6-7, usually I,7; pectoral rays 13-16, usually 14; longitudinal scale count 24(5), 25(11), 26(14). 27(5*), 28(1); predorsal scale count 0-9; transverse scale count (TRB) 7/5(28), 8.5(9*); gill rakers on outer face of first arch 1+0+5(1), 1+1+5(2), 1+1+6(3), 2+1+7(1) = 6-9; gill rakers on outer face of second arch 0+6(6), 0+7(1) = 6-7; segmented caudal rays 8/7(1), 9/8(29*); branched caudal rays 7/6(5), 7/7(8*), 8/7(19). Other meristics shown in tables I-V.
</paragraph>
<caption id="DF6D6665FFBDF80D2CE2FE12EE64C190" ID-DOI="http://doi.org/10.5281/zenodo.13484194" ID-Zenodo-Dep="13484194" httpUri="https://zenodo.org/record/13484194/files/figure.png" pageId="11" pageNumber="190" startId="11.[132,195,422,444]" targetBox="[548,1455,215,486]" targetPageId="11" targetType="figure">
<paragraph id="8BAD36EDFFBDF80D2CE2FE12EEB8C1A8" blockId="11.[132,511,422,492]" pageId="11" pageNumber="190">
Figure 9. Preserved holotype of
<taxonomicName id="4C124D6EFFBDF80D2DAFFE12EED7C1A8" authority=", WAM P." authorityName="WAM P." family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="190" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B966EAFFFFBDF80D2DAFFE12EE77C1AF" italics="true" pageId="11" pageNumber="190">Glossogobius pumilus</emphasis>
, WAM P.
</taxonomicName>
27823-011,
</paragraph>
<paragraph id="8BAD36EDFFBDF80D2CE2FE63EE64C190" blockId="11.[132,511,422,492]" box="[132,291,470,492]" pageId="11" pageNumber="190">32 mm SL male.</paragraph>
</caption>
<caption id="DF6D6665FFBDF80D2CE2FD39EEF3C297" ID-DOI="http://doi.org/10.5281/zenodo.13484202" ID-Zenodo-Dep="13484202" httpUri="https://zenodo.org/record/13484202/files/figure.png" pageId="11" pageNumber="190" startId="11.[132,199,653,675]" targetBox="[560,1449,545,725]" targetPageId="11" targetType="figure">
<paragraph id="8BAD36EDFFBDF80D2CE2FD39EEB9C2C6" blockId="11.[132,510,653,747]" pageId="11" pageNumber="190">
Figure 10. Freshly collected paratype of
<taxonomicName id="4C124D6EFFBDF80D2CB5FD11EEB9C2C6" box="[211,510,677,698]" pageId="11" pageNumber="190">
<emphasis id="B966EAFFFFBDF80D2CB5FD11EEF2C2C6" box="[211,437,677,698]" italics="true" pageId="11" pageNumber="190">Glossogobius pumilus</emphasis>
, WAM
</taxonomicName>
</paragraph>
<paragraph id="8BAD36EDFFBDF80D2CE2FD09EEB9C2AF" blockId="11.[132,510,653,747]" box="[132,510,701,723]" pageId="11" pageNumber="190">P.27823-011, 33 mm SL male from</paragraph>
<paragraph id="8BAD36EDFFBDF80D2CE2FD61EEF3C297" blockId="11.[132,510,653,747]" box="[132,436,725,747]" pageId="11" pageNumber="190">Bensbach River. Photo G. Allen</paragraph>
</caption>
<caption id="DF6D6665FFBDF80D2CE2FB27EF9BC4A5" ID-DOI="http://doi.org/10.5281/zenodo.13484206" ID-Zenodo-Dep="13484206" httpUri="https://zenodo.org/record/13484206/files/figure.png" pageId="11" pageNumber="190" startId="11.[132,195,1171,1193]" targetBox="[548,1455,779,1235]" targetPageId="11" targetType="figure">
<paragraph id="8BAD36EDFFBDF80D2CE2FB27EEB9C4BD" blockId="11.[132,510,1171,1241]" pageId="11" pageNumber="190">
Figure 11. Live individual from Timika area of
<taxonomicName id="4C124D6EFFBDF80D2C81FB1FEEFCC4BC" box="[231,443,1195,1216]" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="190" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B966EAFFFFBDF80D2C81FB1FEEFCC4BC" box="[231,443,1195,1216]" italics="true" pageId="11" pageNumber="190">Glossogobius pumilus</emphasis>
</taxonomicName>
. Photo
</paragraph>
<paragraph id="8BAD36EDFFBDF80D2CE2FB77EF9BC4A5" blockId="11.[132,510,1171,1241]" box="[132,220,1219,1241]" pageId="11" pageNumber="190">G. Allen.</paragraph>
</caption>
<paragraph id="8BAD36EDFFBDF80A2CCCFB4FEAE8C6AD" blockId="11.[132,776,1274,1907]" lastBlockId="12.[132,1456,217,1906]" lastPageId="12" lastPageNumber="191" pageId="11" pageNumber="190">
Head slightly depressed, length 29.2-33.5% SL; cheeks bulbous in adults over
<quantity id="4CEA9B08FFBDF80D2D1BFAAFEE8EC54E" box="[381,457,1307,1331]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="190" unit="mm" value="22.0">22 mm</quantity>
SL, slightly bulbous in juveniles, head width at posterior preopercular margin 16.4- 20.3% SL; depth at posterior preopercular margin 14.3-17.9% SL. Snout rounded in dorsal view; convex or concave in side view, with distinct notch formed from ascending process of premaxilla just before eye in some specimens; 7.1-11.5% SL. Eye slightly longer than snout in juveniles and subequal to snout in adults above
<quantity id="4CEA9B08FFBDF80D2DF6FA4FEE9BC66E" box="[400,476,1531,1555]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="190" unit="mm" value="20.0">20 mm</quantity>
SL, 6.3-8.5% SL in adults. Small bump between nostrils below anterior nostril present, but very low. Anterior nostril at end of short tube, slightly less than one nostril diameter above upper lip. Posterior nostril a pore without elevated rim, about 1.5-3 nostril diameters, from eye and 1.5-2 nostril diameters from anterior nostril. Preoperculum short, distance from end of eye to upper posterior preopercular margin slightly less than snout length. Postorbital moderate, subequal to distance from tip of snout to a point above end of pupil to just behind end of eye. Gill opening reaching to below a point just behind posterior preopercular margin. Jaws forming an angle of 37-47° with body axis; upper margin of upper jaw in line with or just below lower margin of eye, posterior end below anterior margin of eye in females and immature males and below anterior half of pupil in mature males, upper jaw length 12.2-15.6% SL in adult males and 10.4-12.2% SL in adult females. Teeth in upper jaw: outer row of teeth conical, slightly enlarged teeth extending full length of premaxilla; an inner row of smaller conical teeth; an innermost row of enlarged depressible conical teeth anteriorly on premaxilla. Teeth in lower jaw: teeth in outer row conical, slightly enlarged and close-set confined to anterior half of dentary; a second inner row of slightly smaller teeth extending the full length of the dentary becoming larger posteriorly; an innermost row of posteriorly directed, curved depressible teeth anteriorly on inner face of dentary. Tongue tip distinctly bilobed. Gill rakers on outer face of first arch triangular, very short, much less than filament length. Rakers on inner face of first arch and other arches shorter than rakers on outer face of first arch and denticulate. Body covered mostly with large ctenoid scales, cycloid before a line from pectoral fin insertion to below first dorsal fin; belly usually with a few cycloid scales before anus and a large naked patch behind lum (smallest specimen examined
<quantity id="4CEA9B08FFBAF80A28FBFF6DEBAFC08D" box="[1181,1256,217,241]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="191" unit="mm" value="19.0">19 mm</quantity>
SL); no short tube pelvic fin insertion and a few rows of cycloid scales extend- above operculum; no preopercular pores (
<figureCitation id="13292A68FFBAF80A2880FF4DEA6CC16D" box="[1254,1323,249,273]" captionStart="Figure 2" captionStartId="1.[813,876,640,662]" captionTargetBox="[841,1432,218,618]" captionTargetId="figure-677@1.[836,1432,215,622]" captionTargetPageId="1" captionText="Figure 2. Generalised drawing of head showing terminology used for sensory pore in dwarf species of Glossogobius. See Methods for explanation of abbreviations. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13484148" httpUri="https://zenodo.org/record/13484148/files/figure.png" pageId="12" pageNumber="191">Figs 2</figureCitation>
,
<figureCitation id="13292A68FFBAF80A2950FF4DEAC7C16D" box="[1334,1408,249,273]" captionStart="Figure 4" captionStartId="2.[1077,1142,1693,1715]" captionTargetBox="[134,1038,1499,1898]" captionTargetId="figure-687@2.[132,1039,1499,1901]" captionTargetPageId="2" captionText="Figure 4. Dark area indicating anterior extent of predorsal and prepelvic scales and head pores in various populations and species of dwarf Glossogobius. A: Glossogobius gnomus; B: G. nanus, Australia; C: G. nanus, Fly River; D: G. pumilus, Bensbach River; E: G. pumilus, Timika region. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13484166" httpUri="https://zenodo.org/record/13484166/files/figure.png" pageId="12" pageNumber="191">4D, 4E</figureCitation>
). ing forward to below pectoral fin insertion ventrally. First dorsal fin low and rounded in both sexes, lower than body
<emphasis id="B966EAFFFFBAF80A2F4BFE8DECC0C12D" box="[813,903,313,337]" italics="true" pageId="12" pageNumber="191">Papillae</emphasis>
depth at anal fin origin, fourth to sixth spines reach to about Line 1(before nasal pore) present and very short, not the same point just before to just before origin of second dor- reaching snout tip. Line 2 extending from behind nasal pore sal fin when depressed; first dorsal fin origin above a point arching upwards and backwards to opposite side of head. just behind pelvic fin insertion. Second dorsal fin slightly Line 5 (suborbital) short, extending from below front half of higher than first dorsal fin, but lower than body depth at anal eye to infraorbital pore. Line 6 (suborbital branch) absent. fin origin; posterior rays reach just over half way to upper Line 7 (LL cheek row) extending from below front of eye caudal fin base in mature females and to base of upper cau- and usually continuous with papillae behind jaws, extenddal fin in mature males. Anal fin slightly lower than second ing posteriorly to just beyond eye. Line 8 (LT cheek row = dorsal fin, posterior most rays very elongate in mature males, row
<emphasis id="B966EAFFFFBAF80A2F3BFDEDEC2DC20D" box="[861,874,601,625]" italics="true" pageId="12" pageNumber="191">b</emphasis>
) extending from jaws to end of preoperculum. Line 9 reaching to lower rays of caudal fin when depressed, reach- (LL cheek row) short extending from below front half miding over half way to caudal fin in mature females. Pectoral dle of eye arching upward then downward to just behind eye. fin with rounded margin, reaching to above a point between Line 10 (LL cheek row) extending from jaws to just beyond anus and behind anal fin origin, 23.3-31.2% SL. Pelvic fin end of eye and not reaching preopercular margin. Line 11 thin and elongate, reaching to or just beyond anus in males (LT row
<emphasis id="B966EAFFFFBAF80A2FECFD4DECD0C36D" box="[906,919,761,785]" italics="true" pageId="12" pageNumber="191">d</emphasis>
) extending to below posterior end of eye. Line 12 (21.2-28.5% SL) and to or just anterior to anus in females (Outer POP-mandibular) continuous from chin to near lower (19.7-23.9% SL), fifth ray with few long branches and about preopercular pore, usually with a gap of no papillae behind 8-12 terminal tips. posterior tip of jaws. Line 13 (Inner POP-mandibular) continuous from chin to lower preopercular pore. Line 20 (Oper-
<emphasis id="B966EAFFFFBAF80A2CE2FC2DEFB9C3CD" box="[132,254,921,945]" italics="true" pageId="12" pageNumber="191">Head pores</emphasis>
cular VT) composed of single row of papillae extending Posterior nasal pore just medial and slightly above poste- from just behind upper preopercular pore to lower margin of rior nostril; anterior interorbital pore above a point between operculum. Line 21 (Upper OT) separate from Line 20 slopanterior margin of pupil to half way to anterior margin of ing downward posteriorly, ending close to posterior opercueye; posterior interorbital pore above posterior margin of lar margin. Line 22 (Lower OT) almost horizontal and compupil; postorbital pore behind upper margin of eye; infraor- posted of 6-10 papillae reaching to near end of operculum. bital pore below postorbital and behind middle of eye; lat- Other papillae shown in figure 1C. eral canal pore above preoperculum present just before posterior preopercular margin; terminal lateral canal pore above
<emphasis id="B966EAFFFFBAF80A2F4BFB2DECEFC4CD" bold="true" box="[813,936,1177,1201]" pageId="12" pageNumber="191">Coloration</emphasis>
a point just behind posterior preopercular margin of opercu- Head and body brown. Head with scattered dark brown or black small spots and with larger mottling in adults; a broad dark stripe from anteroventral margin of eye to behind middle of upper lip, extending onto lips followed posteriorly by a broad white stripe extending from eye to over posterior third of lips, followed by a thin dark brown to black stripe from posterior margin of eye to behind posterior tips of jaws, becoming broader ventrally and often broken into two or three round spots; a small dark brown to black spot, smaller than pupil diameter almost in contact with posteroventral margin of eye; rest of cheek with large scattered dark brown to black mottling; chin and isthmus dark brown in mature males, chin dark brown with rest of ventral surface of head lighter in mature females; mental fraenum dark brown,
</paragraph>
<paragraph id="8BAD36EDFFBAF80B2FF8F96DEE1AC44D" blockId="12.[132,1456,217,1906]" lastBlockId="13.[132,775,217,1330]" lastPageId="13" lastPageNumber="192" pageId="12" pageNumber="191">
sometimes only at sides of fraenum; lips dark brown except white at posterior tip, bar from Figure 12. Map showing known locations of dwarf species of
<taxonomicName id="4C124D6EFFBAF80A2E95F8A1EC3FC756" authorityName="Gill" authorityYear="1860" box="[755,888,1813,1834]" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="191" phylum="Chordata" rank="genus">
<emphasis id="B966EAFFFFBAF80A2E95F8A1EC3FC756" box="[755,888,1813,1834]" italics="true" pageId="12" pageNumber="191">Glossogobius</emphasis>
</taxonomicName>
anteroventral margin of eye bars crossing lips, but from
<collectingCountry id="F305767DFFBAF80A2CDEF899EE56C73F" box="[184,273,1837,1859]" name="Australia" pageId="12" pageNumber="191">Australia</collectingCountry>
and New
<collectingCountry id="F305767DFFBAF80A2D15F899EEFBC73F" box="[371,444,1837,1859]" name="Guinea-Bissau" pageId="12" pageNumber="191">Guinea</collectingCountry>
, each symbol may represent multiple samples for
<taxonomicName id="4C124D6EFFBAF80A2CC0F8F1EFBCC726" box="[166,251,1861,1882]" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="191" phylum="Chordata" rank="species" species="nanus">
<emphasis id="B966EAFFFFBAF80A2CC0F8F1EFBCC726" box="[166,251,1861,1882]" italics="true" pageId="12" pageNumber="191">G. nanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C124D6EFFBAF80A2D4FF8F1EED6C726" box="[297,401,1861,1882]" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="191" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B966EAFFFFBAF80A2D4FF8F1EED6C726" box="[297,401,1861,1882]" italics="true" pageId="12" pageNumber="191">G. pumilus</emphasis>
</taxonomicName>
; solid circle:
<taxonomicName id="4C124D6EFFBAF80A2E74F8F1ED2FC726" box="[530,616,1861,1882]" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="191" phylum="Chordata" rank="species" species="nanus">
<emphasis id="B966EAFFFFBAF80A2E74F8F1ED2FC726" box="[530,616,1861,1882]" italics="true" pageId="12" pageNumber="191">G. nanus</emphasis>
</taxonomicName>
; square:
<taxonomicName id="4C124D6EFFBAF80A2ED9F8F1EC60C726" authorityName="Hoese &amp; Allen &amp; Hadiaty" authorityYear="2017" box="[703,807,1861,1882]" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="191" phylum="Chordata" rank="species" species="gnomus">
<emphasis id="B966EAFFFFBAF80A2ED9F8F1EC60C726" box="[703,807,1861,1882]" italics="true" pageId="12" pageNumber="191">G. gnomus</emphasis>
</taxonomicName>
; star:
<emphasis id="B966EAFFFFBAF80A2F07F8F1EC3EC726" box="[865,889,1861,1882]" italics="true" pageId="12" pageNumber="191">G.</emphasis>
lips often dark and band not distinct on lips. Body
<taxonomicName id="4C124D6EFFBAF80A2CE2F8E9EF97C70E" box="[132,208,1885,1906]" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="191" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B966EAFFFFBAF80A2CE2F8E9EF97C70E" box="[132,208,1885,1906]" italics="true" pageId="12" pageNumber="191">pumilus</emphasis>
</taxonomicName>
. brown, with an irregular shaped dark brown stripe dorsally on midside breaking up into irregularly shaped dark brown spots on caudal peduncle (
<figureCitation id="13292A68FFBBF80B2D80FF4DED73C16D" box="[486,564,249,273]" captionStart="Figure 11" captionStartId="11.[132,195,1171,1193]" captionTargetBox="[548,1455,779,1235]" captionTargetId="figure-434@11.[548,1455,779,1235]" captionTargetPageId="11" captionText="Figure 11. Live individual from Timi- ka area of Glossogobius pumilus. Photo G. Allen." figureDoi="http://doi.org/10.5281/zenodo.13484206" httpUri="https://zenodo.org/record/13484206/files/figure.png" pageId="13" pageNumber="192">Fig. 11</figureCitation>
). A series of 8 elongate spots on midside of body, with X-shaped extensions dorsally and ventrally, each covering 1-2 scales in a longitudinal direction along midside; first spot below anterior quarter of first dorsal fin; second below posterior quarter of first dorsal fin, sometimes connected to first spot, third spot below a point between first and second dorsal fin; fourth and fifth spots below second dorsal fin; sixth and seventh spot on caudal peduncle; eighth spot triangular at posterior end of caudal peduncle, without dorsal or ventral extensions and apex pointing anteriorly. First dorsal fin with a thin black stripe near base, connecting to an ocellated black spot surrounding middle of sixth dorsal spine, spot fainter in females and immature males. Second dorsal fin with black spots forming 2 rows ventrally, distal margin of fin dark brown. Pectoral fin base with a round to oval spot, smaller than pupil diameter dorsally and in some specimens a second smaller spot near ventral base, fin translucent without spots. Caudal fin with upper rays covered with brown spots forming wavy bands (usually 46) on dorsal two-thirds of fin, ventral third gray to dark brown; a large black median spot on dorsal midline of caudal peduncle followed by a white spot at base of caudal fin, followed by a small dark spot on caudal fin. Anal fin whitish to translucent in young and gray in adults of both sexes. Pelvic fins translucent in young, gray to dark brown in adults of both sexes.
</paragraph>
<paragraph id="8BAD36EDFFBBF80B2CCCFB8EEFA0C54E" blockId="13.[132,775,217,1330]" pageId="13" pageNumber="192">
In freshly collected individuals from Bensbach River (
<figureCitation id="13292A68FFBBF80B2CEAFBEEEF99C40E" box="[140,222,1114,1138]" captionStart="Figure 10" captionStartId="11.[132,199,653,675]" captionTargetBox="[560,1449,545,725]" captionTargetId="figure-433@11.[548,1455,525,741]" captionTargetPageId="11" captionText="Figure 10. Freshly collected para- type of Glossogobius pumilus, WAM P.27823-011, 33 mm SL male from Bensbach River. Photo G.Allen" figureDoi="http://doi.org/10.5281/zenodo.13484202" httpUri="https://zenodo.org/record/13484202/files/figure.png" pageId="13" pageNumber="192">Fig. 10</figureCitation>
), the overall coloration is similar to that described for preserved material, except as follows: distal tips of two dorsal and caudal fins orange and anal fin and ventral quarter of caudal fin gray with an orange tinge and the pectoral fins white. A live individual from the Timika region is similar in coloration, except that the pectoral fin is translucent (
<figureCitation id="13292A68FFBBF80B2CEAFAAEEF9EC54E" box="[140,217,1306,1330]" captionStart="Figure 11" captionStartId="11.[132,195,1171,1193]" captionTargetBox="[548,1455,779,1235]" captionTargetId="figure-434@11.[548,1455,779,1235]" captionTargetPageId="11" captionText="Figure 11. Live individual from Timi- ka area of Glossogobius pumilus. Photo G. Allen." figureDoi="http://doi.org/10.5281/zenodo.13484206" httpUri="https://zenodo.org/record/13484206/files/figure.png" pageId="13" pageNumber="192">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3086566FFBBF80B2CE2FAEEED6CC6CE" pageId="13" pageNumber="192" type="distribution">
<paragraph id="8BAD36EDFFBBF80B2CE2FAEEEE48C50E" blockId="13.[132,776,1370,1715]" box="[132,271,1370,1394]" pageId="13" pageNumber="192">
<emphasis id="B966EAFFFFBBF80B2CE2FAEEEE48C50E" bold="true" box="[132,271,1370,1394]" pageId="13" pageNumber="192">Distribution</emphasis>
</paragraph>
<paragraph id="8BAD36EDFFBBF80B2CCCFACEED6CC6CE" blockId="13.[132,776,1370,1715]" pageId="13" pageNumber="192">
<materialsCitation id="3B7A3CB0FFBBF80B2CCCFACEED60C6CE" country="Papua New Guinea" county="The" elevation="50" location="Timika Region" municipality="Bensbach River" pageId="13" pageNumber="192" specimenCount="1" stateProvince="Papua Barat">
<collectingCounty id="62CC4E61FFBBF80B2CCCFACEEF91C5EE" box="[170,214,1402,1426]" pageId="13" pageNumber="192">The</collectingCounty>
species is known only from three localities, the
<collectingMunicipality id="6BC9AC97FFBBF80B2CE2FA2EEE07C5CE" box="[132,320,1434,1458]" pageId="13" pageNumber="192">Bensbach River</collectingMunicipality>
in
<collectingCountry id="F305767DFFBBF80B2D0FFA2FED0BC5CE" box="[361,588,1434,1458]" name="Papua New Guinea" pageId="13" pageNumber="192">Papua New Guinea</collectingCountry>
and the
<location id="8ECD6036FFBBF80B2ED3FA2EEF93C5AE" LSID="urn:lsid:plazi:treatment:03BB87FBFFBCF80B2FCBFF6DED6CC6CE:8ECD6036FFBBF80B2ED3FA2EEF93C5AE" country="Papua New Guinea" county="The" municipality="Bensbach River" name="Timika Region" pageId="13" pageNumber="192" stateProvince="Papua Barat">Timika Region</location>
in
<collectingRegion id="49D6F80FFFBBF80B2C9EFA0FEE78C5AE" box="[248,319,1467,1490]" country="Indonesia" name="Papua Barat" pageId="13" pageNumber="192">Papua</collectingRegion>
,
<collectingCountry id="F305767DFFBBF80B2D2FFA0EEEFEC5AE" box="[329,441,1466,1490]" name="Indonesia" pageId="13" pageNumber="192">Indonesia</collectingCountry>
.
<location id="8ECD6036FFBBF80B2DA5FA0EEEA9C5AE" LSID="urn:lsid:plazi:treatment:03BB87FBFFBCF80B2FCBFF6DED6CC6CE:8ECD6036FFBBF80B2DA5FA0EEEA9C5AE" box="[451,494,1466,1490]" country="Papua New Guinea" county="The" municipality="Bensbach River" name="The" pageId="13" pageNumber="192" stateProvince="Papua Barat">The</location>
species appears common in lowland clear to turbid streams (less than
<quantity id="4CEA9B08FFBBF80B2E39FA6EEDD2C58E" box="[607,661,1498,1522]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="192" unit="m" value="50.0">
<elevation id="003FD1DEFFBBF80B2E39FA6EEDD2C58E" box="[607,661,1498,1522]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="192" unit="m" value="50.0">50 m</elevation>
</quantity>
elevation) and also occurs in peat swamp habitat in the
<location id="8ECD6036FFBBF80B2E01FA4EEC43C66E" LSID="urn:lsid:plazi:treatment:03BB87FBFFBCF80B2FCBFF6DED6CC6CE:8ECD6036FFBBF80B2E01FA4EEC43C66E" box="[615,772,1530,1554]" country="Papua New Guinea" county="The" municipality="Bensbach River" name="Timika region" pageId="13" pageNumber="192" stateProvince="Papua Barat">Timika region</location>
.
<location id="8ECD6036FFBBF80B2CE2F9AFEFD2C64E" LSID="urn:lsid:plazi:treatment:03BB87FBFFBCF80B2FCBFF6DED6CC6CE:8ECD6036FFBBF80B2CE2F9AFEFD2C64E" box="[132,149,1563,1586]" country="Papua New Guinea" county="The" municipality="Bensbach River" name="It" pageId="13" pageNumber="192" stateProvince="Papua Barat">It</location>
belongs to a relatively diverse fish species assemblage at the
<typeStatus id="54A9884FFFBBF80B2CCAF98FEF9CC62F" box="[172,219,1595,1619]" pageId="13" pageNumber="192">type</typeStatus>
locality, consisting of 16 species that also includes clupeids, plotosids (2 species), atherinids, melanotaeniids (2 species), ambassids, apogonids (2 species), terapontids, toxotids, eleotrids (3 species), and soleiids
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>