treatments-xml/data/03/BB/87/03BB87F4FFB8FF9DE1FF22EFB810F926.xml
2024-06-21 12:22:17 +02:00

167 lines
24 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="6A0F23670B93B6928A1AFEB789287D34" ID="10.11646/zootaxa.4258.3.1SLASH10916" ID-CLB-Dataset="33146" ID-DOI="10.11646/zootaxa.4258.3.1" ID-GBIF-Dataset="97942778-cef4-4cf6-b4e4-c077e633daa8" ID-ISSN="1175-5326" ID-Zenodo-Dep="569729" ID-ZooBank="DE89E162-B255-474C-A824-20DFA551DBBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493363817835" checkinUser="plazi" docAuthor="Penz, Carla M., Casagrande, Mirna M., Devries, Phil &amp; Simonsen, Thomas J." docDate="2017" docId="03BB87F4FFB8FF9DE1FF22EFB810F926" docLanguage="en" docName="zootaxa.4258.3.1SLASH10916.pdf" docOrigin="Zootaxa 4258 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bia caelestis Penz &amp; DeVries, NEW SP." docType="treatment" docVersion="7" lastPageNumber="221" masterDocId="FF82FF8CFFACFF89E1682278BA7DFFFC" masterDocTitle="Documenting diversity in the Amazonian butterfly genus Bia (Lepidoptera, Nymphalidae)" masterLastPageNumber="237" masterPageNumber="201" pageNumber="221" updateTime="1698439134094" updateUser="ExternalLinkService">
<mods:mods id="4C199FA394F197D501B7641AF98A16F9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="819EE35AF440334BA692D184F475865F">
<mods:title id="45D9CE672D98620BE3C632C1E6A08B73">Documenting diversity in the Amazonian butterfly genus Bia (Lepidoptera, Nymphalidae)</mods:title>
</mods:titleInfo>
<mods:name id="DCD33E2A9A71B0CEEAD91D5977D68AFD" type="personal">
<mods:role id="6BD15408A3C9E6F1AB589282F7F24C19">
<mods:roleTerm id="7D6CC3F26005AE1F9070C39CA4C375C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="19F72AD3799328798647CBAD3C175DC1">Penz, Carla M.</mods:namePart>
</mods:name>
<mods:name id="0136DBE2A1866461713A737FCF36AFCE" type="personal">
<mods:role id="7E91AB2F37BBA636B1426155577AD0B3">
<mods:roleTerm id="8533FADFC04271AD841888511DE5A6DB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3DC9B7FD3409158618EC4704A1DB9EAC">Casagrande, Mirna M.</mods:namePart>
</mods:name>
<mods:name id="F3D502D539173DCFF0C167C94D535663" type="personal">
<mods:role id="5CC2F626D4C6BA0DB6B720DBEF3B5A80">
<mods:roleTerm id="87315FC338D935CA13E64546A6CEC0E9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5D2E4459E2541B8E2288132BA5283E46">Devries, Phil</mods:namePart>
</mods:name>
<mods:name id="883DB7755F663B340C966EFAEC4D8654" type="personal">
<mods:role id="B272E807B7F0160F121DE681FD6F651D">
<mods:roleTerm id="940E40F6A251E46CABE68A163A33045D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F941248964968152BF3A8523972F3D83">Simonsen, Thomas J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="F01C18B2803A6A99DFDE62DF1FF5AE0E">text</mods:typeOfResource>
<mods:relatedItem id="9AC631800D08ECA76D931704B7548141" type="host">
<mods:titleInfo id="B68A264EF5CE6286F2A935BD70AB875C">
<mods:title id="C44CC5671FFC2C0D8982412CD9753FEC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E3746938E55C9D503709963B8C74F1CD">
<mods:date id="DC6CA9592D6AB3A7187D4D957210C8F7">2017</mods:date>
<mods:detail id="8C37C0E6E21CCDE3FB12590023D63737" type="volume">
<mods:number id="DEBC0B0D7202FE8D744F40770337825E">4258</mods:number>
</mods:detail>
<mods:detail id="C9CD85C59018D051DA2425B3B5EA7F94" type="issue">
<mods:number id="4709820267C17DE38A1B200CE490F8DC">3</mods:number>
</mods:detail>
<mods:extent id="1F2087EB09AEAFFCC72E95DB4A7D1EE5" unit="page">
<mods:start id="750FFCE4ECB0CD511618F3C95FC9B6C2">201</mods:start>
<mods:end id="6467486BFBC4C990682F05B2290E682C">237</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B1B30C1ED0C529C9951B40090269AE54">journal article</mods:classification>
<mods:identifier id="80848268699BA5D69C017BE78DB2727B" type="CLB-Dataset">33146</mods:identifier>
<mods:identifier id="3ECA3EA9DBE267A1E4E4032B45FE44C8" type="DOI">10.11646/zootaxa.4258.3.1</mods:identifier>
<mods:identifier id="796B58DB49406A5943674D5A9CD77D17" type="GBIF-Dataset">97942778-cef4-4cf6-b4e4-c077e633daa8</mods:identifier>
<mods:identifier id="1A5A86079D02E501CC287926EFBF4AC9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A485BE0A69AAFCDCA65B61C616F4DFAB" type="Zenodo-Dep">569729</mods:identifier>
<mods:identifier id="5E68D679938335A8E3662A8FC8836F9F" type="ZooBank">DE89E162-B255-474C-A824-20DFA551DBBD</mods:identifier>
</mods:mods>
<treatment id="03BB87F4FFB8FF9DE1FF22EFB810F926" ID-DOI="http://doi.org/10.5281/zenodo.6017401" ID-GBIF-Taxon="128810407" ID-Zenodo-Dep="6017401" LSID="urn:lsid:plazi:treatment:03BB87F4FFB8FF9DE1FF22EFB810F926" httpUri="http://treatment.plazi.org/id/03BB87F4FFB8FF9DE1FF22EFB810F926" lastPageNumber="221" pageId="20" pageNumber="221">
<subSubSection id="C3086569FFB8FF9DE1FF22EFBBD3FF2F" pageId="20" pageNumber="221" type="nomenclature">
<paragraph id="8BAD36E2FFB8FF9DE1FF22EFB800FF4E" blockId="20.[151,637,151,211]" box="[151,637,151,178]" pageId="20" pageNumber="221">
<heading id="D0E5818EFFB8FF9DE1FF22EFB800FF4E" bold="true" box="[151,637,151,178]" fontSize="11" level="1" pageId="20" pageNumber="221" reason="1">
<emphasis id="B966EAF0FFB8FF9DE1FF22EFB800FF4E" bold="true" box="[151,637,151,178]" pageId="20" pageNumber="221">
<taxonomicName id="4C124D61FFB8FF9DE1FF22EFBB80FF4E" authority="Penz &amp; DeVries" authorityName="Penz &amp; DeVries" box="[151,509,151,178]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="caelestis" status="sp. nov.">
<emphasis id="B966EAF0FFB8FF9DE1FF22EFBB50FF4D" bold="true" box="[151,301,151,177]" italics="true" pageId="20" pageNumber="221">Bia caelestis</emphasis>
Penz &amp; DeVries
</taxonomicName>
,
<taxonomicNameLabel id="A255578BFFB8FF9DE36122E0B800FF4E" box="[521,637,152,178]" pageId="20" pageNumber="221" rank="species">NEW SP.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD36E2FFB8FF9DE1FF22C2BBD3FF2F" blockId="20.[151,637,151,211]" box="[151,430,186,211]" pageId="20" pageNumber="221">
(
<figureCitation id="13292A67FFB8FF9DE1F722C2BA99FF2F" box="[159,228,186,211]" captionStart="FIGURE 8" captionStartId="21.[151,250,1583,1605]" captionTargetBox="[200,1359,368,1530]" captionTargetId="figure@21.[151,1436,342,1561]" captionTargetPageId="21" captionText="FIGURE 8. Adults in dorsal and ventral views. Taxon names appear below each set. Bia caelestis Penz &amp; DeVries, new sp.: a, male holotype, Brazil, Acre, Mâncio Lima (DZUP); b, male paratype, Brazil, Amazonas, São Paulo de Olivença (MfN); c, male paratype, Brazil, Amazonas, São Paulo de Olivença (CMNH); d, female paratype, Brazil, Amazonas, São Paulo de Olivença (CMNH). Bia pucallpa Casagrande &amp; Penz, new sp.: e, male holotype, Peru, Peru, Pasco, Rio Palcazu, Santa Rosa (MHNL). Intermediate between caelestis and actorion: f, male, Colombia, Amazonas, Puerto Nariño (LACM)." httpUri="https://zenodo.org/record/569738/files/figure.png" pageId="20" pageNumber="221">Figs 8</figureCitation>
ad, 10k, 11e, 12)
</paragraph>
</subSubSection>
<subSubSection id="C3086569FFB8FF9DE1FF237ABBCDFDE1" pageId="20" pageNumber="221" type="diagnosis">
<paragraph id="8BAD36E2FFB8FF9DE1FF237ABBCDFDE1" blockId="20.[151,1437,258,1754]" pageId="20" pageNumber="221">
<emphasis id="B966EAF0FFB8FF9DE1FF237ABB6CFEE7" bold="true" box="[151,273,258,283]" pageId="20" pageNumber="221">Diagnosis.</emphasis>
Defined by the following combination of characters: (1) MF DFW white apical ocelli large, proportionately larger than in other species. (2) MF DFW orange band extremely diffuse and narrow, barely visible in some specimens. (3) M DFW iridescent band from anal margin to M2, decreasing in width and intensity from posterior to anterior ends, and blending with the orange band. F DFW blue iridescence somewhat narrow and slightly faded. (4) M DFW androconial organ on Cu-CuA2 pale, contrasting scale color of surrounding area. (5) DHW discal androconial pad light brown to chocolate-brown, similar to or lighter than associated hairpencil. (6) M DHW discal hairpencil light brown to dark brown. (7) F VFW ripple pattern similar to that of M (but only two females were examined).
</paragraph>
</subSubSection>
<subSubSection id="C3086569FFB8FF9DE1AF2050BA9DFADE" pageId="20" pageNumber="221" type="description">
<paragraph id="8BAD36E2FFB8FF9DE1AF2050BA9DFADE" blockId="20.[151,1437,258,1754]" pageId="20" pageNumber="221">
<emphasis id="B966EAF0FFB8FF9DE1AF2050BB24FDBD" bold="true" box="[199,345,552,577]" pageId="20" pageNumber="221">Description.</emphasis>
Male FW length
<quantity id="4CEA9B07FFB8FF9DE3482050B8BBFDBC" box="[544,710,552,577]" metricMagnitude="-2" metricUnit="m" metricValue="2.585" metricValueMax="2.65" metricValueMin="2.52" pageId="20" pageNumber="221" unit="mm" value="25.85" valueMax="26.5" valueMin="25.2">25.226.5 mm</quantity>
, n=3; female FW length 29.7, n=2. HEAD: As described for the genus. THORAX: Vestiture, wing venation and shape as described for the genus. DFW and DHW mostly brown in both sexes; female paler than male, female DHW usually paler than DFW. Sexually dimorphic, but sharing basic color pattern components. DFW with two large white apical ocelli below Rs3 and Rs4; one dark brown, diffuse ocellus below M1; a thin and highly diffuse orange crescent band (more than in all other taxa and barely visible in some specimens) variable in position between sexes (female band slightly more proximal than male band); male with an iridescent blue patch from DFW tornus to cell M3 or above, sometimes reaching the Rs veins (
<figureCitation id="13292A67FFB8FF9DE452217DBF02FCE2" box="[1338,1407,773,798]" captionStart="FIGURE 8" captionStartId="21.[151,250,1583,1605]" captionTargetBox="[200,1359,368,1530]" captionTargetId="figure@21.[151,1436,342,1561]" captionTargetPageId="21" captionText="FIGURE 8. Adults in dorsal and ventral views. Taxon names appear below each set. Bia caelestis Penz &amp; DeVries, new sp.: a, male holotype, Brazil, Acre, Mâncio Lima (DZUP); b, male paratype, Brazil, Amazonas, São Paulo de Olivença (MfN); c, male paratype, Brazil, Amazonas, São Paulo de Olivença (CMNH); d, female paratype, Brazil, Amazonas, São Paulo de Olivença (CMNH). Bia pucallpa Casagrande &amp; Penz, new sp.: e, male holotype, Peru, Peru, Pasco, Rio Palcazu, Santa Rosa (MHNL). Intermediate between caelestis and actorion: f, male, Colombia, Amazonas, Puerto Nariño (LACM)." httpUri="https://zenodo.org/record/569738/files/figure.png" pageId="20" pageNumber="221">Fig. 8</figureCitation>
a), wide at tornus and becoming narrower and more diffuse in cells above; female with blue patch more proximal and less intensely iridescent than that in male. DHW plain brown, anal region pale. VFW with ripple pattern that fades posteriorly; ocelli mirror the dorsal side, with ocellus below M1 being the most conspicuous, and having a purple pupil. VHW with ripple pattern that does not differ between sexes; some of the ripple pattern has silvery white color; distal wing edge with a yellow-orange line followed by a brown line; one simple, yellow-orange ocellus always present below Sc+R, others may be present below Rs and M1, irregular ocellus with linear pupil below CuA1 always present; tail darkening towards tip. Androconial organ on the DFW CuA-CuA1-CuA2 intersection pale, a sparse cluster of modified, hair-like scales is located posteriorly to it; DHW with a cream-colored androconial pad inside the discal cell and brown associated hairpencil; posterior hairpencil between 1A+2A and 3A cream-colored. Male and female legs as described for the genus. ABDOMEN: General morphology as described for the genus. Male genitalia (
<figureCitation id="13292A67FFB8FF9DE08C26E5B848FB49" box="[484,565,1181,1206]" captionStart="FIGURE 11" captionStartId="25.[151,250,1869,1891]" captionTargetBox="[184,1395,202,1838]" captionTargetId="figure@25.[174,1410,193,1842]" captionTargetPageId="25" captionText="FIGURE 11. Genitalia. a, male actorion, Guyana (dissection Penz 12 - 01, CMNH); b, female actorion, Guyana (dissection Penz 10 - 02, LACM; scale bar different from male illustrations); c, male decaerulea, Lectotype, Brazil, Pará, Faro, (dissection Miller 9047, MfN); d, male rebeli, Peru, San Martín (dissection Penz 10 - 08, USNM); e, male caelestis, paratype, Brazil, Amazonas, São Paulo de Olivença (dissection Penz 10 - 29, CMNH); f, male pucallpa, Peru, Pucallpa (USUSNM 2041410); g, male peruana, Peru, Huanuco, Rio Pichis (dissection Penz 09 - 06, FLMNH)." httpUri="https://zenodo.org/record/569741/files/figure.png" pageId="20" pageNumber="221">Fig. 11</figureCitation>
e): general morphology as described for the genus; tegumen short; uncus long; valva tip narrowing to a round point distally and with a short spiny projection (lateral view), tip slightly bent outward (ventral view); gnathos medium-wide, evenly arched (lateral view). Female genitalia: as described for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C3086569FFB8FF9DE1AF2755B84AFA96" pageId="20" pageNumber="221" type="etymology">
<paragraph id="8BAD36E2FFB8FF9DE1AF2755B84AFA96" blockId="20.[151,1437,258,1754]" pageId="20" pageNumber="221">
<emphasis id="B966EAF0FFB8FF9DE1AF2755BB33FABA" bold="true" box="[199,334,1325,1350]" pageId="20" pageNumber="221">Etymology.</emphasis>
Latin for the color blue, the name
<taxonomicName id="4C124D61FFB8FF9DE3BA2755B94FFAB9" box="[722,818,1325,1349]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="caelestis">
<emphasis id="B966EAF0FFB8FF9DE3BA2755B94FFAB9" box="[722,818,1325,1349]" italics="true" pageId="20" pageNumber="221">caelestis</emphasis>
</taxonomicName>
refers to the fact that males of this species have a large dorsal forewing blue iridescent band.
</paragraph>
</subSubSection>
<subSubSection id="C3086569FFB8FF9DE1AF270DB8B3F9DE" pageId="20" pageNumber="221" type="materials_examined">
<paragraph id="8BAD36E2FFB8FF9DE1AF270DB8B3F9DE" blockId="20.[151,1437,258,1754]" pageId="20" pageNumber="221">
<emphasis id="B966EAF0FFB8FF9DE1AF270DBB09FA72" bold="true" box="[199,372,1397,1422]" pageId="20" pageNumber="221">Type material.</emphasis>
Holotype M (
<figureCitation id="13292A67FFB8FF9DE37A270DB829FA71" box="[530,596,1397,1422]" captionStart="FIGURE 8" captionStartId="21.[151,250,1583,1605]" captionTargetBox="[200,1359,368,1530]" captionTargetId="figure@21.[151,1436,342,1561]" captionTargetPageId="21" captionText="FIGURE 8. Adults in dorsal and ventral views. Taxon names appear below each set. Bia caelestis Penz &amp; DeVries, new sp.: a, male holotype, Brazil, Acre, Mâncio Lima (DZUP); b, male paratype, Brazil, Amazonas, São Paulo de Olivença (MfN); c, male paratype, Brazil, Amazonas, São Paulo de Olivença (CMNH); d, female paratype, Brazil, Amazonas, São Paulo de Olivença (CMNH). Bia pucallpa Casagrande &amp; Penz, new sp.: e, male holotype, Peru, Peru, Pasco, Rio Palcazu, Santa Rosa (MHNL). Intermediate between caelestis and actorion: f, male, Colombia, Amazonas, Puerto Nariño (LACM)." httpUri="https://zenodo.org/record/569738/files/figure.png" pageId="20" pageNumber="221">Fig. 8</figureCitation>
a), deposited at DZUP, four labels separated by // and transcribed verbatim: HOLOTYPUS //
<date id="FFAC1022FFB8FF9DE00427E1B866FA4D" box="[364,539,1433,1457]" pageId="20" pageNumber="221" value="2013-04-20" valueMax="2013-04-27" valueMin="2013-04-20">2027-IV-2013</date>
. RIO MOA. P[AR]Q[UE]. NACIONAL. SERRA DO DIVISOR (SEDE).
<materialsCitation id="3B7A3CBFFFB8FF9DE1FF27B8BB26FA24" ID-GBIF-Occurrence="1499315714" box="[151,347,1472,1496]" collectionCode="MANCIO" location="LIMA" pageId="20" pageNumber="221" specimenCount="1" stateProvince="LIMA">
<collectionCode id="ED03AE27FFB8FF9DE1FF27B8BB74FA24" box="[151,265,1472,1496]" pageId="20" pageNumber="221">MANCIO</collectionCode>
<collectingRegion id="49D6F800FFB8FF9DE07827B8BB26FA24" box="[272,347,1472,1496]" country="Peru" name="Lima" pageId="20" pageNumber="221">LIMA</collectingRegion>
</materialsCitation>
.
<materialsCitation id="3B7A3CBFFFB8FF9DE00D27B8B858FA24" ID-GBIF-Occurrence="1499315722" box="[357,549,1472,1496]" country="Brazil" location="ACRE" pageId="20" pageNumber="221" specimenCount="1" stateProvince="ACRE">
<collectingRegion id="49D6F800FFB8FF9DE00D27B8BBCEFA24" box="[357,435,1472,1496]" country="Brazil" name="Acre" pageId="20" pageNumber="221">ACRE</collectingRegion>
.
<collectingCountry id="F3057672FFB8FF9DE0D627B8B858FA24" box="[446,549,1472,1496]" name="Brazil" pageId="20" pageNumber="221">BRASIL</collectingCountry>
</materialsCitation>
. MIELKE, CASAGRANDE, CARNEIRO, DIAS &amp; DOLIBAINA LEG. 7o 26 52” S. 73o 39 55” W // DZ 32.849 // Holotypus
<taxonomicName id="4C124D61FFB8FF9DE3B7279FBEF7FA03" authority="Penz &amp; DeVries, 2017" authorityName="Penz &amp; DeVries" authorityYear="2017" box="[735,1162,1510,1535]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="caelestis">
<emphasis id="B966EAF0FFB8FF9DE3B7279FB90FFA02" box="[735,882,1510,1534]" italics="true" pageId="20" pageNumber="221">Bia caelestis</emphasis>
Penz &amp; DeVries, 2017
</taxonomicName>
. Paratypes are listed in Appendix, and
<figureCitation id="13292A67FFB8FF9DE02B2472BBFFF9DF" box="[323,386,1546,1571]" captionStart="FIGURE 8" captionStartId="21.[151,250,1583,1605]" captionTargetBox="[200,1359,368,1530]" captionTargetId="figure@21.[151,1436,342,1561]" captionTargetPageId="21" captionText="FIGURE 8. Adults in dorsal and ventral views. Taxon names appear below each set. Bia caelestis Penz &amp; DeVries, new sp.: a, male holotype, Brazil, Acre, Mâncio Lima (DZUP); b, male paratype, Brazil, Amazonas, São Paulo de Olivença (MfN); c, male paratype, Brazil, Amazonas, São Paulo de Olivença (CMNH); d, female paratype, Brazil, Amazonas, São Paulo de Olivença (CMNH). Bia pucallpa Casagrande &amp; Penz, new sp.: e, male holotype, Peru, Peru, Pasco, Rio Palcazu, Santa Rosa (MHNL). Intermediate between caelestis and actorion: f, male, Colombia, Amazonas, Puerto Nariño (LACM)." httpUri="https://zenodo.org/record/569738/files/figure.png" pageId="20" pageNumber="221">Fig. 8</figureCitation>
bd show paratypes M and F.
</paragraph>
</subSubSection>
<subSubSection id="C3086569FFB8FF9DE1AF2456B9ECF9BB" box="[199,913,1582,1607]" pageId="20" pageNumber="221" type="distribution">
<paragraph id="8BAD36E2FFB8FF9DE1AF2456B9ECF9BB" blockId="20.[151,1437,258,1754]" box="[199,913,1582,1607]" pageId="20" pageNumber="221">
<emphasis id="B966EAF0FFB8FF9DE1AF2456B8F2F9BB" bold="true" box="[199,655,1582,1607]" pageId="20" pageNumber="221">Distribution and examined specimens.</emphasis>
<figureCitation id="13292A67FFB8FF9DE3FD2456B898F9BB" box="[661,741,1582,1607]" captionStart="FIGURE 12" captionStartId="26.[151,250,1021,1043]" captionTargetBox="[154,1414,195,985]" captionTargetId="figure@26.[151,1435,193,1000]" captionTargetPageId="26" captionText="FIGURE 12. Map showing collection localities of Bia species and subspecies (see Appendix). Some close localities are represented by one single dot. Dubious label data, localities that could not be accurately identified or were incomplete (e. g., Brazil, Pará) were not placed on map." httpUri="https://zenodo.org/record/569742/files/figure.png" pageId="20" pageNumber="221">Fig. 12</figureCitation>
and Appendix.
</paragraph>
</subSubSection>
<subSubSection id="C3086569FFB8FF9DE1AF242AB810F926" pageId="20" pageNumber="221" type="discussion">
<paragraph id="8BAD36E2FFB8FF9DE1AF242AB810F926" blockId="20.[151,1437,258,1754]" pageId="20" pageNumber="221">
<emphasis id="B966EAF0FFB8FF9DE1AF242ABB46F997" bold="true" box="[199,315,1618,1643]" pageId="20" pageNumber="221">Remarks.</emphasis>
One male specimen from
<collectingCountry id="F3057672FFB8FF9DE311242AB897F997" box="[633,746,1618,1643]" name="Colombia" pageId="20" pageNumber="221">Colombia</collectingCountry>
, Puerto
<collectingRegion id="49D6F800FFB8FF9DE226242AB9E1F997" box="[846,924,1618,1643]" country="Colombia" name="Narino" pageId="20" pageNumber="221">Narino</collectingRegion>
appears intermediate between
<taxonomicName id="4C124D61FFB8FF9DE460242BBF15F997" box="[1288,1384,1619,1643]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="caelestis">
<emphasis id="B966EAF0FFB8FF9DE460242BBF15F997" box="[1288,1384,1619,1643]" italics="true" pageId="20" pageNumber="221">caelestis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C124D61FFB8FF9DE1FF240FBA8BF972" box="[151,246,1655,1678]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="actorion">
<emphasis id="B966EAF0FFB8FF9DE1FF240FBA8BF972" box="[151,246,1655,1678]" italics="true" pageId="20" pageNumber="221">actorion</emphasis>
</taxonomicName>
(
<figureCitation id="13292A67FFB8FF9DE06D240EBB34F973" box="[261,329,1654,1679]" captionStart="FIGURE 8" captionStartId="21.[151,250,1583,1605]" captionTargetBox="[200,1359,368,1530]" captionTargetId="figure@21.[151,1436,342,1561]" captionTargetPageId="21" captionText="FIGURE 8. Adults in dorsal and ventral views. Taxon names appear below each set. Bia caelestis Penz &amp; DeVries, new sp.: a, male holotype, Brazil, Acre, Mâncio Lima (DZUP); b, male paratype, Brazil, Amazonas, São Paulo de Olivença (MfN); c, male paratype, Brazil, Amazonas, São Paulo de Olivença (CMNH); d, female paratype, Brazil, Amazonas, São Paulo de Olivença (CMNH). Bia pucallpa Casagrande &amp; Penz, new sp.: e, male holotype, Peru, Peru, Pasco, Rio Palcazu, Santa Rosa (MHNL). Intermediate between caelestis and actorion: f, male, Colombia, Amazonas, Puerto Nariño (LACM)." httpUri="https://zenodo.org/record/569738/files/figure.png" pageId="20" pageNumber="221">Fig. 8</figureCitation>
f). This specimen has well-developed dorsal forewing white apical ocelli and a diffuse and narrow orange band, but the blue iridescent band on the forewing tornus terminates sharply at CuA2, being clearly smaller than that of a typical
<taxonomicName id="4C124D61FFB8FF9DE0EC24B9BB98F925" box="[388,485,1729,1753]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="caelestis">
<emphasis id="B966EAF0FFB8FF9DE0EC24B9BB98F925" box="[388,485,1729,1753]" italics="true" pageId="20" pageNumber="221">caelestis</emphasis>
</taxonomicName>
(
<figureCitation id="13292A67FFB8FF9DE09B24B9B848F926" box="[499,565,1729,1754]" captionStart="FIGURE 8" captionStartId="21.[151,250,1583,1605]" captionTargetBox="[200,1359,368,1530]" captionTargetId="figure@21.[151,1436,342,1561]" captionTargetPageId="21" captionText="FIGURE 8. Adults in dorsal and ventral views. Taxon names appear below each set. Bia caelestis Penz &amp; DeVries, new sp.: a, male holotype, Brazil, Acre, Mâncio Lima (DZUP); b, male paratype, Brazil, Amazonas, São Paulo de Olivença (MfN); c, male paratype, Brazil, Amazonas, São Paulo de Olivença (CMNH); d, female paratype, Brazil, Amazonas, São Paulo de Olivença (CMNH). Bia pucallpa Casagrande &amp; Penz, new sp.: e, male holotype, Peru, Peru, Pasco, Rio Palcazu, Santa Rosa (MHNL). Intermediate between caelestis and actorion: f, male, Colombia, Amazonas, Puerto Nariño (LACM)." httpUri="https://zenodo.org/record/569738/files/figure.png" pageId="20" pageNumber="221">Fig. 8</figureCitation>
ac).
</paragraph>
</subSubSection>
</treatment>
</document>