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<document id="9025C6827BD078DADEF91E779F336F1F" ID="10.11646/zootaxa.4258.3.1SLASH10916" ID-CLB-Dataset="33146" ID-DOI="10.11646/zootaxa.4258.3.1" ID-GBIF-Dataset="97942778-cef4-4cf6-b4e4-c077e633daa8" ID-ISSN="1175-5326" ID-Zenodo-Dep="569729" ID-ZooBank="DE89E162-B255-474C-A824-20DFA551DBBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493363817835" checkinUser="plazi" docAuthor="Penz, Carla M., Casagrande, Mirna M., Devries, Phil &amp; Simonsen, Thomas J." docDate="2017" docId="03BB87F4FFA0FF85E1FF22EFBE5DFCC1" docLanguage="en" docName="zootaxa.4258.3.1SLASH10916.pdf" docOrigin="Zootaxa 4258 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bia decaerulea subsp. decaerulea Weymer" docType="treatment" docVersion="7" lastPageNumber="213" masterDocId="FF82FF8CFFACFF89E1682278BA7DFFFC" masterDocTitle="Documenting diversity in the Amazonian butterfly genus Bia (Lepidoptera, Nymphalidae)" masterLastPageNumber="237" masterPageNumber="201" pageNumber="213" updateTime="1698439134094" updateUser="ExternalLinkService">
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<mods:title id="752C8ECD87E0B8B299D17F1159008C3C">Documenting diversity in the Amazonian butterfly genus Bia (Lepidoptera, Nymphalidae)</mods:title>
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<mods:namePart id="6BA78485D9A1B64832DC1D2234635059">Penz, Carla M.</mods:namePart>
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<mods:namePart id="F1E136B153BA3E899B6F401FD53AE6D4">Casagrande, Mirna M.</mods:namePart>
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<mods:namePart id="B85603DFCF174110ABDA7D2879AB4D14">Devries, Phil</mods:namePart>
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<mods:namePart id="0F277BFEC2EB3291866E6BD24ED986FC">Simonsen, Thomas J.</mods:namePart>
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<treatment id="03BB87F4FFA0FF85E1FF22EFBE5DFCC1" ID-DOI="http://doi.org/10.5281/zenodo.6017379" ID-GBIF-Taxon="128810415" ID-Zenodo-Dep="6017379" LSID="urn:lsid:plazi:treatment:03BB87F4FFA0FF85E1FF22EFBE5DFCC1" httpUri="http://treatment.plazi.org/id/03BB87F4FFA0FF85E1FF22EFBE5DFCC1" lastPageNumber="213" pageId="12" pageNumber="213">
<subSubSection id="C3086569FFA0FF85E1FF22EFBBD3FF2F" pageId="12" pageNumber="213" type="nomenclature">
<paragraph id="8BAD36E2FFA0FF85E1FF22EFB834FF4E" blockId="12.[151,585,151,211]" box="[151,585,151,178]" pageId="12" pageNumber="213">
<heading id="D0E5818EFFA0FF85E1FF22EFB834FF4E" bold="true" box="[151,585,151,178]" fontSize="11" level="1" pageId="12" pageNumber="213" reason="1">
<taxonomicName id="4C124D61FFA0FF85E1FF22EFB834FF4E" authority="Weymer" authorityName="Weymer" box="[151,585,151,178]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="subSpecies" species="decaerulea" subSpecies="decaerulea">
<emphasis id="B966EAF0FFA0FF85E1FF22EFB834FF4E" bold="true" box="[151,585,151,178]" pageId="12" pageNumber="213">
<emphasis id="B966EAF0FFA0FF85E1FF22EFBBA5FF4D" bold="true" box="[151,472,151,177]" italics="true" pageId="12" pageNumber="213">Bia decaerulea decaerulea</emphasis>
Weymer
</emphasis>
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<paragraph id="8BAD36E2FFA0FF85E1FF22C2BBD3FF2F" blockId="12.[151,585,151,211]" box="[151,430,186,211]" pageId="12" pageNumber="213">
(
<figureCitation id="13292A67FFA0FF85E1F722C2BA98FF2F" box="[159,229,186,211]" captionStart="FIGURE 4" captionStartId="11.[151,250,1848,1870]" captionTargetBox="[262,1339,234,1759]" captionTargetId="figure@11.[209,1367,195,1798]" captionTargetPageId="11" captionText="FIGURE 4. Adults in dorsal and ventral views. Taxon names appear below each set. Bia decaerulea decaerulea: a, male, Brazil, Amazonas, Manacapuru (CMNH); b, female, same locality (CMNH). Bia decaerulea cayana, Simonsen &amp; Penz, new ssp.: c, male holotype, French Guiana, Mana River (CMNH); d, male paratype, same locality (CMNH); e, male paratype, same locality (CMNH); f, female paratype, French Guiana, Oyapok River (CMNH). Bia decaerulea pallida Casagrande &amp; Penz, new ssp: g, male holotype, Brazil, Amazonas, Barcelos, Rio Padauari (DZUP); h, female paratype, Brazil, Amazonas, Barcelos, Rio Padauari (DZUP)." httpUri="https://zenodo.org/record/569733/files/figure.png" pageId="12" pageNumber="213">Figs 4</figureCitation>
ab, 10d, 11c, 12)
</paragraph>
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<subSubSection id="C3086569FFA0FF85E1FF237ABB94FD2C" pageId="12" pageNumber="213" type="description">
<paragraph id="8BAD36E2FFA0FF85E1FF237ABB94FD2C" blockId="12.[151,1437,258,829]" pageId="12" pageNumber="213">
<emphasis id="B966EAF0FFA0FF85E1FF237ABBD1FEE7" bold="true" box="[151,428,258,283]" pageId="12" pageNumber="213">Diagnostic description.</emphasis>
Defined by the following combination of characters: (1) MF DFW white apical ocelli medium-large. (2) MF DFW orange band somewhat diffuse, moderately wide (more than
<emphasis id="B966EAF0FFA0FF85E5F7235FBFE1FEC2" box="[1183,1436,294,318]" italics="true" pageId="12" pageNumber="213">
<taxonomicName id="4C124D61FFA0FF85E5F7235FBFEAFEC2" box="[1183,1431,294,318]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="subSpecies" species="decaerulea" subSpecies="pallida">B. decaerulea pallida</taxonomicName>
,
</emphasis>
less than
<taxonomicName id="4C124D61FFA0FF85E1962333BB8CFE9E" box="[254,497,331,355]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="subSpecies" species="decaerulea" subSpecies="cayana">
<emphasis id="B966EAF0FFA0FF85E1962333BB8CFE9E" box="[254,497,331,355]" italics="true" pageId="12" pageNumber="213">B. decaerulea cayana</emphasis>
</taxonomicName>
); orange scales broadly extended proximally along veins, often lining cross-veins at the distal edge of discal cell. (3) M DFW iridescent band generally absent, but the location of the band might be visible under the scope when the specimen is tilted (a sheen is visible near tornus), a faint blue color is sometimes present. F DFW blue iridescence well developed but variable; usually extended across the entire DC, diffuse anteriorly; usually visible at the CuA1-CuA2 intersection; below the discal cell, the iridescent area is wide but usually does not spread towards the tornus. (4) M DFW androconial scales on the CuA-CuA1-CuA2 intersection dark brown, matching scale color of surrounding area and intermingled with orange scales. (5) M DHW discal androconial pad dark brown, nearly black, always darker than associated hairpencil. (6) M DHW discal hairpencil dark brown, but lighter in color than discal androconial pad. (7) F VFW ripple pattern less dense than that of M, with the VFW postmedial area showing a more predominantly yellow color and with one or two pale markings sometimes present across DC.
</paragraph>
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<subSubSection id="C3086569FFA0FF85E1AF20A4B9ECFD09" box="[199,913,732,757]" pageId="12" pageNumber="213" type="distribution">
<paragraph id="8BAD36E2FFA0FF85E1AF20A4B9ECFD09" blockId="12.[151,1437,258,829]" box="[199,913,732,757]" pageId="12" pageNumber="213">
<emphasis id="B966EAF0FFA0FF85E1AF20A4B8F2FD09" bold="true" box="[199,655,732,757]" pageId="12" pageNumber="213">Distribution and examined specimens.</emphasis>
<figureCitation id="13292A67FFA0FF85E3FD20A4B898FD09" box="[661,741,732,757]" captionStart="FIGURE 12" captionStartId="26.[151,250,1021,1043]" captionTargetBox="[154,1414,195,985]" captionTargetId="figure@26.[151,1435,193,1000]" captionTargetPageId="26" captionText="FIGURE 12. Map showing collection localities of Bia species and subspecies (see Appendix). Some close localities are represented by one single dot. Dubious label data, localities that could not be accurately identified or were incomplete (e. g., Brazil, Pará) were not placed on map." httpUri="https://zenodo.org/record/569742/files/figure.png" pageId="12" pageNumber="213">Fig. 12</figureCitation>
and Appendix.
</paragraph>
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<subSubSection id="C3086569FFA0FF85E1AF2178BE5DFCC1" pageId="12" pageNumber="213" type="discussion">
<paragraph id="8BAD36E2FFA0FF85E1AF2178BE5DFCC1" blockId="12.[151,1437,258,829]" pageId="12" pageNumber="213">
<emphasis id="B966EAF0FFA0FF85E1AF2178BB46FCE5" bold="true" box="[199,315,768,793]" pageId="12" pageNumber="213">Remarks.</emphasis>
Some specimens from Guyana and Suriname (Appendix) seem to be intermediate between
<taxonomicName id="4C124D61FFA0FF85E4552179BFE6FCE4" box="[1341,1435,769,792]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="actorion">
<emphasis id="B966EAF0FFA0FF85E4552179BFE6FCE4" box="[1341,1435,769,792]" italics="true" pageId="12" pageNumber="213">actorion</emphasis>
</taxonomicName>
and
<taxonomicName id="4C124D61FFA0FF85E1AE215CBB3FFCC0" box="[198,322,804,828]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="decaerulea">
<emphasis id="B966EAF0FFA0FF85E1AE215CBB3FFCC0" box="[198,322,804,828]" italics="true" pageId="12" pageNumber="213">decaerulea</emphasis>
</taxonomicName>
(see explanation in
<taxonomicName id="4C124D61FFA0FF85E34C215DB894FCC0" box="[548,745,805,828]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="subSpecies" species="actorion" subSpecies="actorion">
<emphasis id="B966EAF0FFA0FF85E34C215DB894FCC0" box="[548,745,805,828]" italics="true" pageId="12" pageNumber="213">actorion actorion</emphasis>
</taxonomicName>
subspecies account above).
</paragraph>
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