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<mods:title id="3371395E66872599BC06FAB1432D4AB3">Six new species and three new records of infaunal alpheid shrimps from the genera Leptalpheus Williams, 1965 and Fenneralpheus Felder &amp; Manning, 1986 (Crustacea, Decapoda)</mods:title>
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<subSubSection id="C3086548FF96FF8AFF7F207DFE4C9C6A" box="[151,491,293,319]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BAD36C3FF96FF8AFF7F207DFE4C9C6A" blockId="2.[151,491,293,319]" box="[151,491,293,319]" pageId="2" pageNumber="3">
<heading id="D0E581AFFF96FF8AFF7F207DFE4C9C6A" bold="true" box="[151,491,293,319]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<taxonomicName id="4C124D40FF96FF8AFF7F207DFE4C9C6A" ID-CoL="84PMC" authority="Williams, 1965" authorityName="Williams" authorityYear="1965" box="[151,491,293,319]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF96FF8AFF7F207DFE4C9C6A" bold="true" box="[151,491,293,319]" pageId="2" pageNumber="3">
<emphasis id="B966EAD1FF96FF8AFF7F207DFE8C9C6A" bold="true" box="[151,299,293,319]" italics="true" pageId="2" pageNumber="3">Leptalpheus</emphasis>
<bibRefCitation id="EF834B32FF96FF8AFEDA207DFE4C9C6A" author="Williams" box="[306,491,293,319]" pageId="2" pageNumber="3" refString="Williams A. B. (1965) A new genus and species of snapping shrimp (Decapoda, Alpheidae) from the southeastern United States. Crustaceana, 9, 192 - 198." type="journal article" year="1965">Williams, 1965</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3086548FF96FF8AFF7F2034FB7D988E" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="8BAD36C3FF96FF8AFF7F2034FE499CD4" blockId="2.[151,494,363,386]" box="[151,494,363,386]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="AB0211EDFF96FF8AFF7F2034FE499CD4" box="[151,494,363,386]" pageId="2" pageNumber="3">
<taxonomicName id="4C124D40FF96FF8AFF7F2034FE4D9CD4" ID-CoL="84PMC" authority="Williams, 1965: 193" authorityName="Williams" authorityPageNumber="193" authorityYear="1965" box="[151,490,363,386]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF96FF8AFF7F2034FEB49CD7" box="[151,275,364,386]" italics="true" pageId="2" pageNumber="3">Leptalpheus</emphasis>
<treatmentCitation id="0AB310D2FF96FF8AFEF12033FE4D9CD4" author="Williams" box="[281,490,363,385]" page="193" pageId="2" pageNumber="3" year="1965">
<bibRefCitation id="EF834B32FF96FF8AFEF12033FE109CD4" author="Williams" box="[281,439,363,385]" pageId="2" pageNumber="3" refString="Williams A. B. (1965) A new genus and species of snapping shrimp (Decapoda, Alpheidae) from the southeastern United States. Crustaceana, 9, 192 - 198." type="journal article" year="1965">Williams, 1965</bibRefCitation>
: 193
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="8BAD36C3FF96FF8AFF7F20F7FA8B9873" blockId="2.[151,1437,430,1859]" pageId="2" pageNumber="3">
<emphasis id="B966EAD1FF96FF8AFF7F20F7FE219C92" bold="true" box="[151,390,430,455]" pageId="2" pageNumber="3">Emended diagnosis.</emphasis>
Carapace glabrous, not setose; frontal margin without orbital teeth, rounded or with more or less developed median rostral projection; small orbital crests present or absent; anterolateral suture present; pterygostomial angle rounded; branchiostegial margin with narrow ventral lip; posterior margin with deep cardiac notch. Sixth abdominal somite with posteroventral articulated plate. Telson with two pairs of spiniform setae dorsally; posterior margin rounded, with two pairs of posterolateral spiniform setae; anal tubercles more or less developed. Eyes concealed in dorsal and lateral views; anteromesial margin bluntly projecting. Antennular peduncles sometimes flattened dorsoventrally; ventromesial carina of first article with strong tooth; stylocerite appressed or not, rarely exceeding distal margin of first article; second article varying from shorter to much longer than wide; lateral antennular flagellum with short secondary ramus. Antenna with basicerite bearing distoventral tooth; scaphocerite with distolateral tooth; carpocerite exceeding scaphocerite. Mouthparts not especially modified; mandible with two-articulated palp; first maxilliped with elongate palp and broad caridean lobe; second maxilliped with elongate epipod. Third maxilliped with lateral plate on coxa more or less produced dorsally; tip of ultimate article unarmed. First pereiopods (chelipeds) very unequal in size and asymmetrical in shape, carried flexed; major cheliped elongate, with enlarged, elongate chela; merus depressed ventrally, with margins smooth, rugose or furnished with tubercles; carpus short, more or less cup-shaped; palm ventromesially depressed, smooth or with row or fields of tubercles; fingers with variously shaped armature, without tooth-fossa system; adhesive discs present or absent. Second pereiopod with carpus composed of four or five articles. Third pereiopod and fourth pereiopods strongly compressed laterally; ischium with or without spiniform seta ventrolaterally; carpus with distoventral spiniform seta; propodus with spiniform setae on ventral margin; dactylus simple, conical. Fifth pereiopod not compressed, shorter and more slender than third and fourth pereiopods; propodus with at least three setal rows distolaterally. Male second pleopod with appendix masculina exceeding appendix interna; female second pleopod with appendix interna only. Uropod with lateral lobe of protopod bifid distally; exopod distally rounded or truncate; endopod sometimes produced into posterior filament; diaeresis with deep incision and large triangular tooth near mesial margin of exopod. Gill/exopod formula: 5 pleurobranchs (P15), 1 arthrobranch (Mxp3), 2 lobe-like epipods (Mxp12), 0 podobranchs, 5 mastigobranchs (Mxp3, P14), 5 setobranchs (P15), 3 exopods (Mxp13).
</paragraph>
<paragraph id="8BAD36C3FF96FF8AFF2F246AFB7D988E" blockId="2.[151,1437,430,1859]" pageId="2" pageNumber="3">
<emphasis id="B966EAD1FF96FF8AFF2F246AFE34981E" bold="true" box="[199,403,1330,1355]" pageId="2" pageNumber="3">Species included.</emphasis>
<taxonomicName id="4C124D40FF96FF8AFE73246BFC97981F" ID-CoL="8533J" authority="Williams, 1965" authorityName="Williams" authorityYear="1965" box="[411,816,1330,1355]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="forceps">
<emphasis id="B966EAD1FF96FF8AFE73246BFDDB981E" box="[411,636,1331,1355]" italics="true" pageId="2" pageNumber="3">Leptalpheus forceps</emphasis>
<bibRefCitation id="EF834B32FF96FF8AFD6C246AFC97981F" author="Williams" box="[644,816,1330,1354]" pageId="2" pageNumber="3" refString="Williams A. B. (1965) A new genus and species of snapping shrimp (Decapoda, Alpheidae) from the southeastern United States. Crustaceana, 9, 192 - 198." type="journal article" year="1965">Williams, 1965</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54A98861FF96FF8AFCD7246AFCD6981F" box="[831,881,1330,1354]" pageId="2" pageNumber="3">type</typeStatus>
species),
<taxonomicName id="4C124D40FF96FF8AFC09246CFADE981F" ID-CoL="8533K" authority="Banner &amp; Banner, 1974" authorityName="Banner &amp; Banner" authorityYear="1974" box="[993,1401,1330,1355]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B966EAD1FF96FF8AFC09246CFBC3981E" box="[993,1124,1331,1355]" italics="true" pageId="2" pageNumber="3">L. pacificus</emphasis>
<bibRefCitation id="EF834B32FF96FF8AFB83246BFADE981F" author="Banner" box="[1131,1401,1330,1354]" pageId="2" pageNumber="3" refString="Banner, A. H. &amp; Banner, D. M. (1974) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean, part XVII: additional notes on the Hawaiian alpheids: new species, subspecies and some nomenclatorial changes. Pacific Science, 28, 423 - 437." type="journal article" year="1974">Banner &amp; Banner, 1974</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C124D40FF96FF8AFA6D246CFD8C983B" ID-CoL="85TJQ" authority="Rios &amp; Carvacho, 1983" authorityName="Rios &amp; Carvacho" authorityYear="1983" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B966EAD1FF96FF8AFA6D246CFEAA983A" italics="true" pageId="2" pageNumber="3">L. mexicanus</emphasis>
<bibRefCitation id="EF834B32FF96FF8AFEFF240EFD8C983B" author="Rios" box="[279,555,1366,1390]" pageId="2" pageNumber="3" refString="Rios R. &amp; Carvacho A. (1983) Caridean shrimps of the Gulf of California III. Leptalpheus mexicanus, new species (Crustacea, Decapoda, Alpheidae). Journal of Crustacean Biology, 3, 306 - 313." type="journal article" year="1983">Ríos &amp; Carvacho, 1983</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C124D40FF96FF8AFDD12400FB84983B" ID-CoL="85TJB" authority="Dworschak &amp; Coelho, 1999" authorityName="Dworschak &amp; Coelho" authorityYear="1999" box="[569,1059,1366,1391]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="axianassae">
<emphasis id="B966EAD1FF96FF8AFDD12400FD71983A" box="[569,726,1368,1391]" italics="true" pageId="2" pageNumber="3">L. axianassae</emphasis>
<bibRefCitation id="EF834B32FF96FF8AFD08240EFB84983B" author="Dworschak" box="[736,1059,1366,1390]" pageId="2" pageNumber="3" refString="Dworschak, P. C. &amp; Coelho, V. R. (1999) On two alpheids from Araca (Sao Paulo, Brazil) with a description of a new species of Leptalpheus (Decapoda: Caridea: Alpheidae). Annalen des Naturhistorischen Museums in Wien, 101 B, 475 - 488." type="journal article" year="1999">Dworschak &amp; Coelho, 1999</bibRefCitation>
</taxonomicName>
(? =
<taxonomicName id="4C124D40FF96FF8AFB8E2400FE8298C7" authority="Ramos-Porto &amp; Souza, 1994" authorityName="Ramos-Porto &amp; Souza" authorityYear="1994" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="petronii">
<emphasis id="B966EAD1FF96FF8AFB8E2400FB78983A" box="[1126,1247,1368,1391]" italics="true" pageId="2" pageNumber="3">L. petronii</emphasis>
<bibRefCitation id="EF834B32FF96FF8AFB00240FFE8298C7" author="Ramos-Porto" pageId="2" pageNumber="3" refString="Ramos-Porto, M. &amp; Souza, S. T. de (1994) Descricao de Leptalpheus petronii sp. n. (Decapoda, Alpheidae) para o Brasil. In: Bonecker, S. L. C. (ed.), Resumos do XX Congresso Brasiliero de Zoologia, 24 - 29 de julho 1994, Universidade Federal do Rio de Janeiro, p. 22." type="journal article" year="1994">Ramos-Porto &amp; Souza, 1994</bibRefCitation>
</taxonomicName>
,
<emphasis id="B966EAD1FF96FF8AFED92424FE7098C6" box="[305,471,1403,1427]" italics="true" pageId="2" pageNumber="3">nomen dubium</emphasis>
, see Anker
<emphasis id="B966EAD1FF96FF8AFDB72424FD3F98C6" box="[607,664,1403,1427]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
2006 for discussion),
<taxonomicName id="4C124D40FF96FF8AFC7C2424FADD98C7" ID-CoL="85TJK" authority="Anker, Vera Caripe &amp; Lira, 2006" authorityName="Anker, Vera Caripe &amp; Lira" authorityYear="2006" box="[916,1402,1402,1427]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="felderi">
<emphasis id="B966EAD1FF96FF8AFC7C2424FC5C98C6" box="[916,1019,1403,1427]" italics="true" pageId="2" pageNumber="3">L. felderi</emphasis>
<bibRefCitation id="EF834B32FF96FF8AFBEB2422FADD98C7" author="Anker" box="[1027,1402,1402,1426]" pageId="2" pageNumber="3" refString="Anker, A., Vera Caripe, J. A. &amp; Lira, C. (2006) Description of a new species of commensal alpheid shrimp (Crustacea, Decapoda) from the southern Caribbean Sea. Zoosystema, 28, 683 - 702." type="journal article" year="2006">Anker, Vera Caripe &amp; Lira, 2006</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C124D40FF96FF8AFA6D2424FE0098E3" ID-CoL="85TJT" authority="Anker, 2008" authorityName="Anker" authorityYear="2008" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="pierrenoeli">
<emphasis id="B966EAD1FF96FF8AFA6D2424FEB598E3" italics="true" pageId="2" pageNumber="3">L. pierrenoeli</emphasis>
<bibRefCitation id="EF834B32FF96FF8AFEF124C6FE0098E3" author="Anker" box="[281,423,1438,1462]" pageId="2" pageNumber="3" refString="Anker A. (2008) The alpheid shrimp genus Leptalpheus Williams, 1965 in the southwestern Caribbean Sea, with description of one new species from Panama (Crustacea: Decapoda). Zoosystema, 30, 781 - 794." type="journal article" year="2008">Anker, 2008</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C124D40FF96FF8AFE5924F8FCF298E3" ID-CoL="85TJH" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[433,853,1438,1463]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B966EAD1FF96FF8AFE5924F8FDFF98E3" box="[433,600,1438,1463]" italics="true" pageId="2" pageNumber="3">L. denticulatus</emphasis>
<bibRefCitation id="EF834B32FF96FF8AFDB624C6FCF298E3" author="Anker" box="[606,853,1438,1462]" pageId="2" pageNumber="3" refString="Anker A, Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin, 2009</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C124D40FF96FF8AFC8824F8FB5C98E3" ID-CoL="85TJJ" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[864,1275,1438,1463]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dworschaki">
<emphasis id="B966EAD1FF96FF8AFC8824F8FC5998E3" box="[864,1022,1438,1463]" italics="true" pageId="2" pageNumber="3">L. dworschaki</emphasis>
<bibRefCitation id="EF834B32FF96FF8AFBED24C6FB5C98E3" author="Anker" box="[1029,1275,1438,1462]" pageId="2" pageNumber="3" refString="Anker A, Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin, 2009</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C124D40FF96FF8AFAEE24F8FA3B98E3" ID-CoL="85TJM" box="[1286,1436,1438,1463]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="marginalis" status="sp. nov.">
<emphasis id="B966EAD1FF96FF8AFAEE24F8FA3B98E3" box="[1286,1436,1438,1463]" italics="true" pageId="2" pageNumber="3">L. marginalis</emphasis>
</taxonomicName>
<emphasis id="B966EAD1FF96FF8AFF7F249BFF57988E" bold="true" box="[151,240,1475,1499]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="A25557AAFF96FF8AFF7F249BFF57988E" box="[151,240,1475,1499]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C124D40FF96FF8AFF16249CFEC0988E" ID-CoL="85TJC" box="[254,359,1476,1499]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">
<emphasis id="B966EAD1FF96FF8AFF16249CFEC0988E" box="[254,359,1476,1499]" italics="true" pageId="2" pageNumber="3">L. azuero</emphasis>
</taxonomicName>
<emphasis id="B966EAD1FF96FF8AFE86249BFE60988E" bold="true" box="[366,455,1475,1499]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="A25557AAFF96FF8AFE86249BFE60988E" box="[366,455,1475,1499]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C124D40FF96FF8AFE3D249CFDD1988E" ID-CoL="85TJR" box="[469,630,1475,1499]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="penicillatus" status="sp. nov.">
<emphasis id="B966EAD1FF96FF8AFE3D249CFDD1988E" box="[469,630,1475,1499]" italics="true" pageId="2" pageNumber="3">L. penicillatus</emphasis>
</taxonomicName>
<emphasis id="B966EAD1FF96FF8AFD94249BFD71988E" bold="true" box="[636,726,1475,1499]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="A25557AAFF96FF8AFD94249BFD71988E" box="[636,726,1475,1499]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C124D40FF96FF8AFD0C249CFCDF988E" ID-CoL="85TJL" box="[740,888,1475,1499]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="hendrickxi" status="sp. nov.">
<emphasis id="B966EAD1FF96FF8AFD0C249CFCDF988E" box="[740,888,1475,1499]" italics="true" pageId="2" pageNumber="3">L. hendrickxi</emphasis>
</taxonomicName>
<emphasis id="B966EAD1FF96FF8AFC97249BFC7F988E" bold="true" box="[895,984,1475,1499]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="A25557AAFF96FF8AFC97249BFC7F988E" box="[895,984,1475,1499]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C124D40FF96FF8AFC0E249CFBDD988E" ID-CoL="85TJD" box="[998,1146,1475,1499]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bicristatus" status="sp. nov.">
<emphasis id="B966EAD1FF96FF8AFC0E249CFBDD988E" box="[998,1146,1475,1499]" italics="true" pageId="2" pageNumber="3">L. bicristatus</emphasis>
</taxonomicName>
<emphasis id="B966EAD1FF96FF8AFB69249BFB7D988E" bold="true" box="[1153,1242,1475,1499]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="A25557AAFF96FF8AFB69249BFB7D988E" box="[1153,1242,1475,1499]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3086548FF96FF8AFF2F24BEFF4F9B3F" pageId="2" pageNumber="3" type="distribution">
<paragraph id="8BAD36C3FF96FF8AFF2F24BEFF4F9B3F" blockId="2.[151,1437,430,1859]" pageId="2" pageNumber="3">
<emphasis id="B966EAD1FF96FF8AFF2F24BEFEC798AA" bold="true" box="[199,352,1510,1535]" pageId="2" pageNumber="3">Distribution.</emphasis>
Pantropical except for East Atlantic. Indo-West Pacific:
<collectingCountry id="F3057653FF96FF8AFC0824BEFBCE98AB" box="[992,1129,1510,1534]" name="Madagascar" pageId="2" pageNumber="3">Madagascar</collectingCountry>
,
<collectingCountry id="F3057653FF96FF8AFB9C24BEFB7198AB" box="[1140,1238,1510,1534]" name="Vietnam" pageId="2" pageNumber="3">Vietnam</collectingCountry>
,
<collectingCountry id="F3057653FF96FF8AFB0A24BEFAC498AB" box="[1250,1379,1510,1534]" name="Philippines" pageId="2" pageNumber="3">Philippines</collectingCountry>
,
<collectingCountry id="F3057653FF96FF8AFA8624BEFA3F98AB" box="[1390,1432,1510,1534]" name="Fiji" pageId="2" pageNumber="3">Fiji</collectingCountry>
,
<collectingCountry id="F3057653FF96FF8AFF7F2752FEFB9B77" box="[151,348,1546,1570]" name="French Polynesia" pageId="2" pageNumber="3">French Polynesia</collectingCountry>
, Hawaii. East Pacific:
<collectingCountry id="F3057653FF96FF8AFDB32752FD149B77" box="[603,691,1546,1570]" name="Mexico" pageId="2" pageNumber="3">Mexico</collectingCountry>
,
<collectingCountry id="F3057653FF96FF8AFD572752FC929B77" box="[703,821,1546,1570]" name="Nicaragua" pageId="2" pageNumber="3">Nicaragua</collectingCountry>
,
<collectingCountry id="F3057653FF96FF8AFCA82752FC199B77" box="[832,958,1546,1570]" name="Costa Rica" pageId="2" pageNumber="3">Costa Rica</collectingCountry>
,
<collectingCountry id="F3057653FF96FF8AFC212753FB839B77" box="[969,1060,1547,1570]" name="Panama" pageId="2" pageNumber="3">Panama</collectingCountry>
,
<collectingCountry id="F3057653FF96FF8AFBD92752FB049B77" box="[1073,1187,1546,1570]" name="Colombia" pageId="2" pageNumber="3">Colombia</collectingCountry>
. West Atlantic: southeastern
<collectingCountry id="F3057653FF96FF8AFF182776FE8F9B13" box="[240,296,1582,1606]" name="United States of America" pageId="2" pageNumber="3">USA</collectingCountry>
(North Carolina, Florida, Louisiana),
<collectingCountry id="F3057653FF96FF8AFD0A2776FCFF9B13" box="[738,856,1582,1606]" name="Venezuela" pageId="2" pageNumber="3">Venezuela</collectingCountry>
,
<collectingCountry id="F3057653FF96FF8AFC8E2776FC089B13" box="[870,943,1582,1606]" name="Aruba" pageId="2" pageNumber="3">Aruba</collectingCountry>
,
<collectingCountry id="F3057653FF96FF8AFC552777FBBF9B13" box="[957,1048,1583,1606]" name="Panama" pageId="2" pageNumber="3">Panama</collectingCountry>
,
<collectingCountry id="F3057653FF96FF8AFBCE2776FB019B13" box="[1062,1190,1582,1606]" name="Costa Rica" pageId="2" pageNumber="3">Costa Rica</collectingCountry>
,
<collectingCountry id="F3057653FF96FF8AFB5D2776FB5E9B13" box="[1205,1273,1582,1606]" name="Brazil" pageId="2" pageNumber="3">Brazil</collectingCountry>
(Sergipe, São Paulo).
</paragraph>
</subSubSection>
<subSubSection id="C3086548FF96FF8AFF2F272FFC399A17" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BAD36C3FF96FF8AFF2F272FFC399A17" blockId="2.[151,1437,430,1859]" pageId="2" pageNumber="3">
<emphasis id="B966EAD1FF96FF8AFF2F272FFE9C9BDA" bold="true" box="[199,315,1655,1679]" pageId="2" pageNumber="3">Remarks.</emphasis>
The increasing morphological diversity of
<taxonomicName id="4C124D40FF96FF8AFCC0272EFC169BDB" box="[808,945,1654,1678]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF96FF8AFCC0272EFC169BDB" box="[808,945,1654,1678]" italics="true" pageId="2" pageNumber="3">Leptalpheus</emphasis>
</taxonomicName>
, especially in the proportions of the antennular peduncles, the shape and length of the stylocerite, and the armature of the major chela fingers, calls for a phylogenetic investigation of this genus. It is possible that some morphologically and genetically well-defined clades will be assigned to new subgenera or even genera. However, in the absence of molecular data for many taxa and in the presence of further undescribed species (
<bibRefCitation id="EF834B32FF96FF8AFD6F265EFCF59A4B" author="Felder" box="[647,850,1798,1823]" pageId="2" pageNumber="3" refString="Felder, D. L., Nates, S. F. &amp; Robles, R. R. (2003) Hurricane Mitch: impacts of bioturbating crustaceans in shrimp ponds and adjacent estuaries of coastal Nicaragua. USGS Open File Report, OFR, 03 - 179, 1 - 47." type="book chapter" year="2003">
Felder
<emphasis id="B966EAD1FF96FF8AFD3F2650FCB79A4A" box="[727,784,1799,1823]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
2003
</bibRefCitation>
; Anker
<emphasis id="B966EAD1FF96FF8AFC422650FC459A4A" box="[938,994,1799,1823]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
2006; see also below), a subdivision of
<taxonomicName id="4C124D40FF96FF8AFF7F2673FE879A16" box="[151,288,1835,1859]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF96FF8AFF7F2673FE879A16" box="[151,288,1835,1859]" italics="true" pageId="2" pageNumber="3">Leptalpheus</emphasis>
</taxonomicName>
in either subgenera or genera is not possible at this time.
</paragraph>
</subSubSection>
</treatment>
</document>