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<document id="B6373DC141F9286AEFE2E2EBDE82586C" ID="10.11646/zootaxa.4193.2.10" ID-DOI="10.11646/zootaxa.4193.2.10" ID-GBIF-Dataset="c9b07cdc-abd3-44c7-8855-0dbd9bc8ae5f" ID-ISSN="1175-5326" ID-Zenodo-Dep="166979" ID-ZooBank="3D257D9C-7C01-4822-BF7F-6D8740CBEF84" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479284524079" checkinUser="plazi" docAuthor="Luz, David R., Williams, Kevin A. &amp; Bartholomay, Pedro R." docDate="2016" docId="03BB87C5FFDD245FFF7394A6FB8233DA" docLanguage="en" docName="zootaxa.4193.2.10.pdf" docOrigin="Zootaxa 4193 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dasymutilla paradoxa" docType="treatment" docVersion="9" lastPageNumber="362" masterDocId="FF82FFBDFFDC245EFFE49629FF863401" masterDocTitle="The mutillid wasps of the Dasymutilla paradoxa species-group (Hymenoptera, Mutillidae)" masterLastPageNumber="372" masterPageNumber="361" pageNumber="362" updateTime="1698393713946" updateUser="plazi">
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<mods:title id="72C3A2BD73A9818472772325F25E9030">The mutillid wasps of the Dasymutilla paradoxa species-group (Hymenoptera, Mutillidae)</mods:title>
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<mods:namePart id="F789300994EB8425D20B681FBF5A0901">Luz, David R.</mods:namePart>
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<mods:namePart id="0472FDC14081D193C7DD00251957A837">Williams, Kevin A.</mods:namePart>
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<mods:namePart id="C4942E2E9EAD7E6BD2668883AC12B828">Bartholomay, Pedro R.</mods:namePart>
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<mods:date id="4661ECA59C9C24B947D8688F9BC0A3E0">2016</mods:date>
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<treatment id="03BB87C5FFDD245FFF7394A6FB8233DA" ID-DOI="http://doi.org/10.5281/zenodo.5087175" ID-GBIF-Taxon="124736679" ID-Zenodo-Dep="5087175" LSID="urn:lsid:plazi:treatment:03BB87C5FFDD245FFF7394A6FB8233DA" httpUri="http://treatment.plazi.org/id/03BB87C5FFDD245FFF7394A6FB8233DA" lastPageNumber="362" pageId="1" pageNumber="362">
<subSubSection id="C3086558FFDD245FFF7394A6FCCE36AB" box="[151,840,655,682]" pageId="1" pageNumber="362" type="nomenclature">
<paragraph id="8BAD36D3FFDD245FFF7394A6FCCE36AB" blockId="1.[151,840,655,682]" box="[151,840,655,682]" pageId="1" pageNumber="362">
<heading id="D0E581BFFFDD245FFF7394A6FCCE36AB" bold="true" box="[151,840,655,682]" fontSize="11" level="1" pageId="1" pageNumber="362" reason="1">
<emphasis id="B966EAC1FFDD245FFF7394A6FCCE36AB" bold="true" box="[151,840,655,682]" pageId="1" pageNumber="362">
<taxonomicName id="4C124D50FFDD245FFF7394A6FE2536A8" box="[151,419,655,681]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="B966EAC1FFDD245FFF7394A6FE2536A8" bold="true" box="[151,419,655,681]" italics="true" pageId="1" pageNumber="362">Dasymutilla paradoxa</emphasis>
</taxonomicName>
species-group, new species-group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3086558FFDD245FFF7394FFFCB937A2" pageId="1" pageNumber="362" type="diagnosis">
<paragraph id="8BAD36D3FFDD245FFF7394FFFCB937A2" blockId="1.[151,1437,726,2011]" pageId="1" pageNumber="362">
<emphasis id="B966EAC1FFDD245FFF7394FFFE8C36EE" bold="true" box="[151,266,726,751]" pageId="1" pageNumber="362">Diagnosis</emphasis>
. FEMALE. The females of this species-group can be immediately separated from other
<taxonomicName id="4C124D50FFDD245FFB0B94FFFAFE36EE" box="[1263,1400,726,751]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC1FFDD245FFB0B94FFFAFE36EE" box="[1263,1400,726,751]" italics="true" pageId="1" pageNumber="362">Dasymutilla</emphasis>
</taxonomicName>
by their elongate mesosoma that lacks a scutellar scale; specifically, in lateral view, the propodeal height and length are subequal, while other
<taxonomicName id="4C124D50FFDD245FFE5F9537FDC23736" box="[443,580,798,823]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC1FFDD245FFE5F9537FDC23736" box="[443,580,798,823]" italics="true" pageId="1" pageNumber="362">Dasymutilla</emphasis>
</taxonomicName>
species-groups have the propodeal height greater than the length. They can be separated from
<taxonomicName id="4C124D50FFDD245FFE8C956BFDA2375A" box="[360,548,834,859]" class="Insecta" family="Mutillidae" genus="Traumatomutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC1FFDD245FFE8C956BFDA2375A" box="[360,548,834,859]" italics="true" pageId="1" pageNumber="362">Traumatomutilla</emphasis>
</taxonomicName>
species with a similar elongate mesosoma by their globose sub-cylindrical T1, by the four integumental spots on T2 and by the simple postero-lateral margins of the pygidium (bearing raised flanges in
<emphasis id="B966EAC1FFDD245FFEE895A3FE3C37A2" box="[268,442,906,931]" italics="true" pageId="1" pageNumber="362">Tramatomutilla</emphasis>
species with elongate mesosoma).
</paragraph>
</subSubSection>
<subSubSection id="C3086558FFDD245FFF239586FE7530C2" pageId="1" pageNumber="362" type="description">
<paragraph id="8BAD36D3FFDD245FFF239586FD503033" blockId="1.[151,1437,726,2011]" pageId="1" pageNumber="362">MALE. The males of this species-group can be immediately recognized by the axillae, which are elongated and contorted into lateral “arms” attached to the scutellum. Most of the species have a seta-filled pit on S2 that is anteriorly situated and all of the species have T2 marked with integumental spots or entirely yellowish, contrasting with the predominantly black to reddish metasoma.</paragraph>
<paragraph id="8BAD36D3FFDD245FFF239217FE7530C2" blockId="1.[151,1437,726,2011]" pageId="1" pageNumber="362">
<emphasis id="B966EAC1FFDD245FFF239217FEEA3056" bold="true" box="[199,364,1086,1111]" pageId="1" pageNumber="362">Included taxa</emphasis>
. This predominantly Neotropical species-group includes eight species:
<taxonomicName id="4C124D50FFDD245FFB7D9216FF54307A" authority="Mickel, 1938" authorityName="Mickel" authorityYear="1938" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="fasciventris">
<emphasis id="B966EAC1FFDD245FFB7D9216FABB3056" box="[1177,1341,1087,1111]" italics="true" pageId="1" pageNumber="362">D. fasciventris</emphasis>
<bibRefCitation id="EF834B22FFDD245FFAA19217FF54307A" author="Mickel" pageId="1" pageNumber="362" refString="Mickel, C. E. (1938) Synonymical notes and new species of Pseudomethoca and Dasymutilla (Mutillidae: Hymenoptera). Annals of the Entomological Society of America, 31, 147 - 156. http: // dx. doi. org / 10.1093 / aesa / 31.2.147" type="journal article" year="1938">Mickel, 1938</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C124D50FFDD245FFF3B924DFD02307A" authority="Manley &amp; Pitts, 2007" authorityName="Manley &amp; Pitts" authorityYear="2007" box="[223,644,1122,1147]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="B966EAC1FFDD245FFF3B924DFE04307A" box="[223,386,1124,1147]" italics="true" pageId="1" pageNumber="362">D. guanacaste</emphasis>
<bibRefCitation id="EF834B22FFDD245FFE6F924BFD02307A" author="Manley" box="[395,644,1122,1147]" pageId="1" pageNumber="362" refString="Manley, D. G. &amp; Pitts, J. P. (2007) Tropical and subtropical velvet ants of the genus Dasymutilla Ashmead (Hymenoptera: Mutillidae) with descriptions of 45 new species. Zootaxa, 1487, 1 - 128." type="journal article" year="2007">Manley &amp; Pitts, 2007</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C124D50FFDD245FFD76924DFB97307A" authority="Manley &amp; Pitts, 2007" authorityName="Manley &amp; Pitts" authorityYear="2007" box="[658,1041,1122,1147]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="naranjo">
<emphasis id="B966EAC1FFDD245FFD76924DFC89307A" box="[658,783,1124,1147]" italics="true" pageId="1" pageNumber="362">D. naranjo</emphasis>
<bibRefCitation id="EF834B22FFDD245FFCF3924BFB97307A" author="Manley" box="[791,1041,1122,1147]" pageId="1" pageNumber="362" refString="Manley, D. G. &amp; Pitts, J. P. (2007) Tropical and subtropical velvet ants of the genus Dasymutilla Ashmead (Hymenoptera: Mutillidae) with descriptions of 45 new species. Zootaxa, 1487, 1 - 128." type="journal article" year="2007">Manley &amp; Pitts, 2007</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C124D50FFDD245FFBFA924DFA11307A" authority="Gerstaecker, 1874" authorityName="Gerstaecker" authorityYear="1874" box="[1054,1431,1122,1147]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="paradoxa" status="comb. nov.">
<emphasis id="B966EAC1FFDD245FFBFA924DFB2B307A" box="[1054,1197,1122,1147]" italics="true" pageId="1" pageNumber="362">D. paradoxa</emphasis>
(
<bibRefCitation id="EF834B22FFDD245FFB59924BFA16307A" author="Gerstaecker" box="[1213,1424,1122,1147]" pageId="1" pageNumber="362" refString="Gerstaecker, A. (1874) Mutillarum Americae meridionalis indigenarum synopsis systematica et synonymica. Archiv fur Naturgeschichte, 40, 41 - 77, 299 - 328." type="journal article" year="1874">Gerstaecker, 1874</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A25557BAFFDD245FFF7392AFFE91309E" box="[151,279,1158,1183]" pageId="1" pageNumber="362">
<emphasis id="B966EAC1FFDD245FFF7392AFFE91309E" bold="true" box="[151,279,1158,1183]" pageId="1" pageNumber="362">comb. nov.</emphasis>
</taxonomicNameLabel>
;
<emphasis id="B966EAC1FFDD245FFEC192AEFE68309E" box="[293,494,1158,1183]" italics="true" pageId="1" pageNumber="362">
<taxonomicName id="4C124D50FFDD245FFEC192AEFE6C309E" box="[293,490,1158,1183]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="paraparadoxa" status="sp. nov.">D. paraparadoxa</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A25557BAFFDD245FFE1292AFFDC9309E" box="[502,591,1158,1183]" pageId="1" pageNumber="362" rank="species">
<emphasis id="B966EAC1FFDD245FFE1292AFFDC9309E" bold="true" box="[502,591,1158,1183]" pageId="1" pageNumber="362">sp. nov.</emphasis>
</taxonomicNameLabel>
;
<taxonomicName id="4C124D50FFDD245FFDBA92AEFCFA309E" authority="Cameron, 1895" authorityName="Cameron" authorityYear="1895" box="[606,892,1158,1183]" class="Insecta" family="Mutillidae" genus="Sphaerophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="phya" status="comb. nov.">
<emphasis id="B966EAC1FFDD245FFDBA92AEFD31309E" box="[606,695,1158,1183]" italics="true" pageId="1" pageNumber="362">D. phya</emphasis>
(Cameron, 1895)
</taxonomicName>
,
<taxonomicNameLabel id="A25557BAFFDD245FFC6E92AFFB8C309E" box="[906,1034,1158,1183]" pageId="1" pageNumber="362">
<emphasis id="B966EAC1FFDD245FFC6E92AFFB8C309E" bold="true" box="[906,1034,1158,1183]" pageId="1" pageNumber="362">comb. nov.</emphasis>
</taxonomicNameLabel>
;
<taxonomicName id="4C124D50FFDD245FFBFD92AEFAC0309E" authority="Cameron, 1895" authorityName="Cameron" authorityYear="1895" box="[1049,1350,1158,1183]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="relata">
<emphasis id="B966EAC1FFDD245FFBFD92AEFBFB309E" box="[1049,1149,1158,1183]" italics="true" pageId="1" pageNumber="362">D. relata</emphasis>
(Cameron, 1895)
</taxonomicName>
; and
<taxonomicName id="4C124D50FFDD245FFA6592AEFE6A30C2" authority="Manley &amp; Pitts, 2007" authorityName="Manley &amp; Pitts" authorityYear="2007" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="sleipniri">
<emphasis id="B966EAC1FFDD245FFA6592AEFF7230C2" italics="true" pageId="1" pageNumber="362">D. sleipniri</emphasis>
<bibRefCitation id="EF834B22FFDD245FFF1F9283FE6A30C2" author="Manley" box="[251,492,1194,1219]" pageId="1" pageNumber="362" refString="Manley, D. G. &amp; Pitts, J. P. (2007) Tropical and subtropical velvet ants of the genus Dasymutilla Ashmead (Hymenoptera: Mutillidae) with descriptions of 45 new species. Zootaxa, 1487, 1 - 128." type="journal article" year="2007">Manley &amp; Pitts, 2007</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3086558FFDD245FFF2392E7FCA730E6" box="[199,801,1230,1255]" pageId="1" pageNumber="362" type="biology_ecology">
<paragraph id="8BAD36D3FFDD245FFF2392E7FCA730E6" blockId="1.[151,1437,726,2011]" box="[199,801,1230,1255]" pageId="1" pageNumber="362">
<emphasis id="B966EAC1FFDD245FFF2392E7FE8030E6" bold="true" box="[199,262,1230,1255]" pageId="1" pageNumber="362">Host.</emphasis>
Unknown for all member of this species-group.
</paragraph>
</subSubSection>
<subSubSection id="C3086558FFDD245FFF2392DBFB8233DA" pageId="1" pageNumber="362" type="discussion">
<paragraph id="8BAD36D3FFDD245FFF2392DBFC6F3177" blockId="1.[151,1437,726,2011]" pageId="1" pageNumber="362">
<emphasis id="B966EAC1FFDD245FFF2392DBFEB2310A" bold="true" box="[199,308,1266,1291]" pageId="1" pageNumber="362">Remarks</emphasis>
. Members of the
<taxonomicName id="4C124D50FFDD245FFDE292DDFD11310A" box="[518,663,1266,1291]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="B966EAC1FFDD245FFDE292DDFD11310A" box="[518,663,1266,1291]" italics="true" pageId="1" pageNumber="362">D. paradoxa</emphasis>
</taxonomicName>
species-group have not been formally grouped in the past, but the axillar structure of these males and the elongate mesosoma and T1 of the females clearly link these taxa. Only one species,
<taxonomicName id="4C124D50FFDD245FFF129315FE1D3152" box="[246,411,1339,1363]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="fasciventris">
<emphasis id="B966EAC1FFDD245FFF129315FE1D3152" box="[246,411,1339,1363]" italics="true" pageId="1" pageNumber="362">D. fasciventris</emphasis>
</taxonomicName>
, is known from both sexes. Genitalia characters and illustrations were given by
<bibRefCitation id="EF834B22FFDD245FFACD9313FE9C3176" author="Manley" pageId="1" pageNumber="362" refString="Manley, D. G. &amp; Pitts, J. P. (2007) Tropical and subtropical velvet ants of the genus Dasymutilla Ashmead (Hymenoptera: Mutillidae) with descriptions of 45 new species. Zootaxa, 1487, 1 - 128." type="journal article" year="2007">Manley &amp; Pitts (2007)</bibRefCitation>
for
<taxonomicName id="4C124D50FFDD245FFEAD9376FE6C3177" box="[329,490,1375,1398]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="B966EAC1FFDD245FFEAD9376FE6C3177" box="[329,490,1375,1398]" italics="true" pageId="1" pageNumber="362">D. guanacaste</emphasis>
</taxonomicName>
,
<taxonomicName id="4C124D50FFDD245FFE1D9376FDF23177" box="[505,628,1375,1398]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="naranjo">
<emphasis id="B966EAC1FFDD245FFE1D9376FDF23177" box="[505,628,1375,1398]" italics="true" pageId="1" pageNumber="362">D. naranjo</emphasis>
</taxonomicName>
,
<taxonomicName id="4C124D50FFDD245FFD659376FCA33176" box="[641,805,1375,1399]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="fasciventris">
<emphasis id="B966EAC1FFDD245FFD659376FCA33176" box="[641,805,1375,1399]" italics="true" pageId="1" pageNumber="362">D. fasciventris</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C124D50FFDD245FFC869376FC673176" box="[866,993,1374,1399]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="sleipniri">
<emphasis id="B966EAC1FFDD245FFC869376FC673176" box="[866,993,1374,1399]" italics="true" pageId="1" pageNumber="362">D. sleipniri</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAD36D3FFDD245FFF2393ABFEC132BA" blockId="1.[151,1437,726,2011]" pageId="1" pageNumber="362">
<materialsCitation id="3B7A3C8EFFDD245FFF2393ABFEC132BA" ID-GBIF-Occurrence="1325928617" country="Panama" county="Such" location="Cambra" municipality="Cambra" pageId="1" pageNumber="362" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EF834B22FFDD245FFF2393ABFE7F319A" author="Cambra" box="[199,505,1410,1435]" pageId="1" pageNumber="362" refString="Cambra, R. A. &amp; Quintero, D. (1992) Velvet ants of Panama: distribution and systematics (Hymenoptera: Mutillidae). In: Quintero, D. &amp; Aiello, A. (Eds.), Insects of Panama and Mesoamerica: Selected Studies. Oxford University Press, Oxford, pp. 459 - 478." type="book chapter" year="1992">Cambra &amp; Quintero (1992)</bibRefCitation>
transferred
<taxonomicName id="4C124D50FFDD245FFD6493ADFD5C319A" box="[640,730,1410,1435]" class="Insecta" family="Mutillidae" genus="Sphaerophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="phya">
<emphasis id="B966EAC1FFDD245FFD6493ADFD5C319A" box="[640,730,1410,1435]" italics="true" pageId="1" pageNumber="362">D. phya</emphasis>
</taxonomicName>
and
<taxonomicName id="4C124D50FFDD245FFCF493ADFC00319A" box="[784,902,1410,1435]" class="Insecta" family="Mutillidae" genus="Sphaerophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="melissa">
<emphasis id="B966EAC1FFDD245FFCF493ADFC00319A" box="[784,902,1410,1435]" italics="true" pageId="1" pageNumber="362">D. melissa</emphasis>
</taxonomicName>
to
<taxonomicName id="4C124D50FFDD245FFC4E93ABFB7C319A" authority="Andre, 1901" authorityName="Andre" authorityYear="1901" box="[938,1274,1410,1435]" class="Insecta" family="Mutillidae" genus="Traumatomutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC1FFDD245FFC4E93ABFBE1319A" box="[938,1127,1410,1435]" italics="true" pageId="1" pageNumber="362">Traumatomutilla</emphasis>
<bibRefCitation id="EF834B22FFDD245FFB8A93ABFB7C319A" author="Andre" box="[1134,1274,1410,1435]" pageId="1" pageNumber="362" refString="Andre, E. (1901) Descriptions de quelques especes et varietes nouvelles de Mutilles d'Amerique appartenant au Musee Civique de Genes. Zeitschrift fur Systematische Hymenopterologie und Dipterologie, 1, 257 - 264." type="journal article" year="1901">André, 1901</bibRefCitation>
</taxonomicName>
and placed
<taxonomicName id="4C124D50FFDD245FFA6593ADFF6D31BE" class="Insecta" family="Mutillidae" genus="Sphaerophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="melissa">
<emphasis id="B966EAC1FFDD245FFA6593ADFF6D31BE" italics="true" pageId="1" pageNumber="362">D. melissa</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C124D50FFDD245FFE12938EFDC831BE" box="[502,590,1446,1471]" class="Insecta" family="Mutillidae" genus="Sphaerophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="phya">
<emphasis id="B966EAC1FFDD245FFE12938EFDC831BE" box="[502,590,1446,1471]" italics="true" pageId="1" pageNumber="362">D. phya</emphasis>
</taxonomicName>
based on the examination of the
<typeStatus id="54A98871FFDD245FFC59938EFC7E31BE" box="[957,1016,1447,1471]" pageId="1" pageNumber="362">types</typeStatus>
for both species.
<bibRefCitation id="EF834B22FFDD245FFB58938FFAE331BE" author="Cambra" box="[1212,1381,1446,1471]" pageId="1" pageNumber="362" refString="Cambra, R. A. (1997) Comparacion de la diversidad en la Sphaeropthalminae (Hymenoptera: Mutillidae) de Costa Rica y Panama, con notas sobre biologia. Scientia, 12 (2), 115 - 128." type="journal article" year="1997">
<location id="8ECD6008FFDD245FFB58938FFA9331BE" LSID="urn:lsid:plazi:treatment:03BB87C5FFDD245FFF7394A6FB8233DA:8ECD6008FFDD245FFB58938FFA9331BE" box="[1212,1301,1446,1471]" country="Panama" county="Such" municipality="Cambra" name="Cambra" pageId="1" pageNumber="362">Cambra</location>
(1997)
</bibRefCitation>
then transferred
<taxonomicName id="4C124D50FFDD245FFEFD93E5FEFB31E2" box="[281,381,1484,1507]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="icaris">
<emphasis id="B966EAC1FFDD245FFEFD93E5FEFB31E2" box="[281,381,1484,1507]" italics="true" pageId="1" pageNumber="362">D. icaris</emphasis>
</taxonomicName>
and within the comments regarding
<taxonomicName id="4C124D50FFDD245FFCC593E3FA9631E2" authority="Cameron, 1895" authorityName="Cameron" authorityYear="1895" box="[801,1296,1482,1507]" class="Insecta" family="Mutillidae" genus="Traumatomutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="dictynna">
<emphasis id="B966EAC1FFDD245FFCC593E3FBC131E2" box="[801,1095,1482,1507]" italics="true" pageId="1" pageNumber="362">Traumatomutilla dictynna</emphasis>
(Cameron, 1895)
</taxonomicName>
in the same paper, stated that sexual associations for
<taxonomicName id="4C124D50FFDD245FFD8293C6FD623206" box="[614,740,1518,1543]" class="Insecta" family="Mutillidae" genus="Traumatomutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="dictynna">
<emphasis id="B966EAC1FFDD245FFD8293C6FD623206" box="[614,740,1518,1543]" italics="true" pageId="1" pageNumber="362">T. dictynna</emphasis>
</taxonomicName>
,
<taxonomicName id="4C124D50FFDD245FFD1793C6FCC03206" box="[755,838,1518,1543]" class="Insecta" family="Mutillidae" genus="Sphaerophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="phya">
<emphasis id="B966EAC1FFDD245FFD1793C6FCC03206" box="[755,838,1518,1543]" italics="true" pageId="1" pageNumber="362">T. phya</emphasis>
</taxonomicName>
and
<taxonomicName id="4C124D50FFDD245FFC9A93C6FC5D3207" box="[894,987,1519,1542]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="icaris">
<emphasis id="B966EAC1FFDD245FFC9A93C6FC5D3207" box="[894,987,1519,1542]" italics="true" pageId="1" pageNumber="362">T. icaris</emphasis>
</taxonomicName>
were made in
<collectingCountry id="F3057643FFDD245FFB6093C6FB663207" box="[1156,1248,1519,1542]" name="Panama" pageId="1" pageNumber="362">Panama</collectingCountry>
.
<location id="8ECD6008FFDD245FFB0893C6FA883207" LSID="urn:lsid:plazi:treatment:03BB87C5FFDD245FFF7394A6FB8233DA:8ECD6008FFDD245FFB0893C6FA883207" box="[1260,1294,1519,1542]" country="Panama" county="Such" municipality="Cambra" name="No" pageId="1" pageNumber="362">No</location>
justification for the genus changes in either paper was provided and no diagnosis or description was given for the supposed males mentioned by
<bibRefCitation id="EF834B22FFDD245FFE63901FFDB1324E" author="Cambra" box="[391,567,1590,1615]" pageId="1" pageNumber="362" refString="Cambra, R. A. (1997) Comparacion de la diversidad en la Sphaeropthalminae (Hymenoptera: Mutillidae) de Costa Rica y Panama, con notas sobre biologia. Scientia, 12 (2), 115 - 128." type="journal article" year="1997">
<location id="8ECD6008FFDD245FFE63901FFE66324E" LSID="urn:lsid:plazi:treatment:03BB87C5FFDD245FFF7394A6FB8233DA:8ECD6008FFDD245FFE63901FFE66324E" box="[391,480,1590,1615]" country="Panama" county="Such" municipality="Cambra" name="Cambra" pageId="1" pageNumber="362">Cambra</location>
(1997)
</bibRefCitation>
.
<bibRefCitation id="EF834B22FFDD245FFDA1901FFCF8324E" author="Cambra" box="[581,894,1590,1615]" pageId="1" pageNumber="362" refString="Cambra, R. A. &amp; Quintero, D. (1992) Velvet ants of Panama: distribution and systematics (Hymenoptera: Mutillidae). In: Quintero, D. &amp; Aiello, A. (Eds.), Insects of Panama and Mesoamerica: Selected Studies. Oxford University Press, Oxford, pp. 459 - 478." type="book chapter" year="1992">
<location id="8ECD6008FFDD245FFDA1901FFD18324E" LSID="urn:lsid:plazi:treatment:03BB87C5FFDD245FFF7394A6FB8233DA:8ECD6008FFDD245FFDA1901FFD18324E" box="[581,670,1590,1615]" country="Panama" county="Such" municipality="Cambra" name="Cambra" pageId="1" pageNumber="362">Cambra</location>
&amp;
<location id="8ECD6008FFDD245FFD22901FFCAC324E" LSID="urn:lsid:plazi:treatment:03BB87C5FFDD245FFF7394A6FB8233DA:8ECD6008FFDD245FFD22901FFCAC324E" box="[710,810,1590,1615]" country="Panama" county="Such" municipality="Cambra" name="Quintero" pageId="1" pageNumber="362">Quintero</location>
(1992)
</bibRefCitation>
stated that a formal statement and conclusions supporting the changes made regarding
<taxonomicName id="4C124D50FFDD245FFD819075FD593272" box="[613,735,1626,1651]" class="Insecta" family="Mutillidae" genus="Sphaerophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="melissa">
<emphasis id="B966EAC1FFDD245FFD819075FD593272" box="[613,735,1626,1651]" italics="true" pageId="1" pageNumber="362">D. melissa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C124D50FFDD245FFCF89075FCFE3272" box="[796,888,1626,1651]" class="Insecta" family="Mutillidae" genus="Sphaerophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="phya">
<emphasis id="B966EAC1FFDD245FFCF89075FCFE3272" box="[796,888,1626,1651]" italics="true" pageId="1" pageNumber="362">D. phya</emphasis>
</taxonomicName>
would be published elsewhere.
<bibRefCitation id="EF834B22FFDD245FFB0B9073FA1A3272" author="Cambra" box="[1263,1436,1626,1651]" pageId="1" pageNumber="362" refString="Cambra, R. A. (1997) Comparacion de la diversidad en la Sphaeropthalminae (Hymenoptera: Mutillidae) de Costa Rica y Panama, con notas sobre biologia. Scientia, 12 (2), 115 - 128." type="journal article" year="1997">
<collectingMunicipality id="6BC9ACA9FFDD245FFB0B9073FACE3272" box="[1263,1352,1626,1651]" pageId="1" pageNumber="362">Cambra</collectingMunicipality>
(1997)
</bibRefCitation>
stated that the descriptions of the males were to be published in subsequent papers.
<collectingCounty id="62CC4E5FFFDD245FFBDE9057FBF43296" box="[1082,1138,1662,1687]" pageId="1" pageNumber="362">Such</collectingCounty>
papers, however, have not been published.
</materialsCitation>
</paragraph>
<paragraph id="8BAD36D3FFDD245FFF2390EFFB8233DA" blockId="1.[151,1437,726,2011]" pageId="1" pageNumber="362">
Although three of the included species were placed in the genus
<taxonomicName id="4C124D50FFDD245FFC5990EFFA9532DE" authority="Andre, 1901" authorityName="Andre" authorityYear="1901" box="[957,1299,1734,1759]" class="Insecta" family="Mutillidae" genus="Traumatomutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC1FFDD245FFC5990EFFBFF32DE" box="[957,1145,1734,1759]" italics="true" pageId="1" pageNumber="362">Traumatomutilla</emphasis>
<bibRefCitation id="EF834B22FFDD245FFB6090EFFA9532DE" author="Andre" box="[1156,1299,1734,1759]" pageId="1" pageNumber="362" refString="Andre, E. (1901) Descriptions de quelques especes et varietes nouvelles de Mutilles d'Amerique appartenant au Musee Civique de Genes. Zeitschrift fur Systematische Hymenopterologie und Dipterologie, 1, 257 - 264." type="journal article" year="1901">André, 1901</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="EF834B22FFDD245FFAA790EFFF653302" author="Cambra" pageId="1" pageNumber="362" refString="Cambra, R. A. (1997) Comparacion de la diversidad en la Sphaeropthalminae (Hymenoptera: Mutillidae) de Costa Rica y Panama, con notas sobre biologia. Scientia, 12 (2), 115 - 128." type="journal article" year="1997">Cambra (1997)</bibRefCitation>
, we place this species-group in the genus
<taxonomicName id="4C124D50FFDD245FFD5290C3FC733302" authority="Ashmead, 1899" authorityName="Ashmead" authorityYear="1899" box="[694,1013,1770,1795]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC1FFDD245FFD5290C3FCB93302" box="[694,831,1770,1795]" italics="true" pageId="1" pageNumber="362">Dasymutilla</emphasis>
<bibRefCitation id="EF834B22FFDD245FFCA290C3FC733302" author="Ashmead" box="[838,1013,1770,1795]" pageId="1" pageNumber="362" refString="Ashmead, W. H. (1899) Superfamilies in the Hymenoptera and generic synopses of the families Thynnidae, Myrmosidae, and Mutillidae. Journal of the New York Entomological Society, 7, 45 - 60." type="journal article" year="1899">Ashmead, 1899</bibRefCitation>
</taxonomicName>
for four reasons. All of the males and two females were treated as members of
<taxonomicName id="4C124D50FFDD245FFD8E9127FD753326" box="[618,755,1806,1831]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC1FFDD245FFD8E9127FD753326" box="[618,755,1806,1831]" italics="true" pageId="1" pageNumber="362">Dasymutilla</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF834B22FFDD245FFCFB9127FBA53326" author="Manley" box="[799,1059,1806,1831]" pageId="1" pageNumber="362" refString="Manley, D. G. &amp; Pitts, J. P. (2007) Tropical and subtropical velvet ants of the genus Dasymutilla Ashmead (Hymenoptera: Mutillidae) with descriptions of 45 new species. Zootaxa, 1487, 1 - 128." type="journal article" year="2007">Manley &amp; Pitts (2007)</bibRefCitation>
. All of the included species have black tibial spurs, a trait that is common in
<taxonomicName id="4C124D50FFDD245FFD60911BFC8B334A" box="[644,781,1842,1867]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC1FFDD245FFD60911BFC8B334A" box="[644,781,1842,1867]" italics="true" pageId="1" pageNumber="362">Dasymutilla</emphasis>
</taxonomicName>
and rare in
<taxonomicName id="4C124D50FFDD245FFC72911BFA1E334A" authority="Quintero &amp; Cambra, 1996" authorityName="Quintero &amp; Cambra" authorityYear="1996" box="[918,1432,1842,1867]" class="Insecta" family="Mutillidae" genus="Traumatomutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC1FFDD245FFC72911BFBD4334A" box="[918,1106,1842,1867]" italics="true" pageId="1" pageNumber="362">Traumatomutilla</emphasis>
(
<bibRefCitation id="EF834B22FFDD245FFB85911BFA16334A" author="Quintero" box="[1121,1424,1842,1867]" pageId="1" pageNumber="362" refString="Quintero, D. &amp; Cambra, R. A. (1996) Contribution a la sistematica de las Mutilidas (Hymenoptera) del Peru, en especial las de la Estacion Biologica BIOLAT, Rio Manu, Pakitza. In: Wilson, D. E. &amp; Sandoval, A. (Eds.), Manu: The Biodiversity of Southeastern Peru. Smithsonian Institution Press, Washington, D. C., pp. 327 - 357." type="book chapter" year="1996">Quintero &amp; Cambra, 1996</bibRefCitation>
)
</taxonomicName>
. The species-group is widely distributed in North America, which includes over 150
<taxonomicName id="4C124D50FFDD245FFBB4917FFB5F336E" box="[1104,1241,1878,1903]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC1FFDD245FFBB4917FFB5F336E" box="[1104,1241,1878,1903]" italics="true" pageId="1" pageNumber="362">Dasymutilla</emphasis>
</taxonomicName>
species and only one
<taxonomicName id="4C124D50FFDD245FFF2D9153FE033392" box="[201,389,1914,1939]" class="Insecta" family="Mutillidae" genus="Traumatomutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC1FFDD245FFF2D9153FE033392" box="[201,389,1914,1939]" italics="true" pageId="1" pageNumber="362">Traumatomutilla</emphasis>
</taxonomicName>
species (
<bibRefCitation id="EF834B22FFDD245FFE149153FD003392" author="Mickel" box="[496,646,1914,1939]" pageId="1" pageNumber="362" refString="Mickel, C. E. (1936) New species and records of nearctic mutillid wasps of the genus Dasymutilla (Hymenoptera). Annals of the Entomological Society of America, 29, 29 - 60. http: // dx. doi. org / 10.1093 / aesa / 29.1.29" type="journal article" year="1936">Mickel 1936</bibRefCitation>
,
<bibRefCitation id="EF834B22FFDD245FFD709153FC0F3392" author="Manley" box="[660,905,1914,1939]" pageId="1" pageNumber="362" refString="Manley, D. G. &amp; Pitts, J. P. (2007) Tropical and subtropical velvet ants of the genus Dasymutilla Ashmead (Hymenoptera: Mutillidae) with descriptions of 45 new species. Zootaxa, 1487, 1 - 128." type="journal article" year="2007">Manley &amp; Pitts 2007</bibRefCitation>
,
<bibRefCitation id="EF834B22FFDD245FFC739153FBD23392" author="Nonveiller" box="[919,1108,1914,1939]" pageId="1" pageNumber="362" refString="Nonveiller, G. (1990) Catalogue of the Mutillidae, Myrmosidae and Bradynobaenidae of the Neotropical Region including Mexico (Insecta, Hymenoptera). Hymenopterorum Catalogus (Nova Editio), 18. Den Haag, SPB Academic Publishing. 150 pp." type="book" year="1990">Nonveiller 1990</bibRefCitation>
). Finally, in an unplublished Bayesian phylogenetic reconstruction (Williams, 2012, pg. 47), three members of this species-group were recovered within
<taxonomicName id="4C124D50FFDD245FFEBF91EBFE6233DA" box="[347,484,1986,2011]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC1FFDD245FFEBF91EBFE6233DA" box="[347,484,1986,2011]" italics="true" pageId="1" pageNumber="362">Dasymutilla</emphasis>
</taxonomicName>
, as the sister group to
<taxonomicName id="4C124D50FFDD245FFD0491EDFC7933DA" authority="Smith, 1855" authorityName="Smith" authorityYear="1855" box="[736,1023,1986,2011]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B966EAC1FFDD245FFD0491EDFCDC33DA" box="[736,858,1986,2011]" italics="true" pageId="1" pageNumber="362">D. pulchra</emphasis>
(
<bibRefCitation id="EF834B22FFDD245FFC8C91EBFC7133DA" author="Smith" box="[872,1015,1986,2011]" pageId="1" pageNumber="362" refString="Smith, F. (1855) Catalogue of Hymenopterous Insects in the Collection of the British Museum. Part III. Mutillidae and Pompilidae. Trustees of the British Museum, London. [4] + 1 - 206. [plates A + plate 1 - 5.]" type="journal article" year="1855">Smith, 1855</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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