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<document id="015B8F8BA5875C0570E95F344E3844E2" ID-CLB-Dataset="22395" ID-DOI="10.5852/ejt.2015.136" ID-GBIF-Dataset="2bc5e28e-c045-49d7-aaf2-5f5be028eb21" ID-ISSN="2118-9773" ID-Zenodo-Dep="3785302" ID-ZooBank="530F395F-97A9-46F1-957C-8E57B9C3ACD9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588614102535" checkinUser="carolina" docAuthor="Smith, Robin James &amp; Kamiya, Takahiro" docDate="2015" docId="03BB878CBC5E1B4E9862D93C02E92F2A" docLanguage="en" docName="ejt-2015-136.pdf.imf" docOrigin="European Journal of Taxonomy 136" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pseudocandona falcula Smith &amp; Kamiya 2015, sp. nov." docType="treatment" docUuid="0138AA0F-95A0-4AED-AF47-9DD1CCCB63C1" docUuidSource="ZooBank" docVersion="8" lastPageNumber="23" masterDocId="FF82FFF4BC4F1B589A39D837031E2919" masterDocTitle="Four new species of the subfamily Candoninae (Crustacea, Ostracoda) from freshwater habitats in Japan" masterLastPageNumber="34" masterPageNumber="1" pageNumber="18" updateTime="1698838423492" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="77FE17FF98B8A43A70D4FA01C733D105">Four new species of the subfamily Candoninae (Crustacea, Ostracoda) from freshwater habitats in Japan</mods:title>
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<mods:namePart id="932CCA2B9FABF3C731BE30762B7FF890">Smith, Robin James</mods:namePart>
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<mods:affiliation id="F432175E6D7222156FB4A1C13B7C0992">Lake Biwa Museum, 1091 Oroshimo, Kusatsu, Shiga 525 - 0001, Japan. Email: smith @ lbm. go. jp (corresponding author) &amp; urn: lsid: zoobank. org: author: D 63 C 5 DD 6 - C 0 BB- 43 EA- 8 D 2 F- 98134 CC 29188</mods:affiliation>
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<mods:namePart id="A73720897938069F969839829D469E0D">Kamiya, Takahiro</mods:namePart>
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<mods:affiliation id="88044EADAF14B5F5B526DA8AAB437047">College of Science and Engineering, School of Natural System, University of Kanazawa, Kakuma, Kanazawa 920 - 1192, Japan. &amp; urn: lsid: zoobank. org: author: 5 FB 5 FC 58 - 0 FBF- 45 DA- 9 FDC- 7569 BDA 1 FDF 9</mods:affiliation>
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<mods:title id="E08C7D535E9180D5EA7435D2D32617F6">European Journal of Taxonomy</mods:title>
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<mods:date id="7D6FB10FC8C7E0EDAA2DF7123C37ADC6">2015</mods:date>
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<mods:number id="07C916ACFF4D731E3ED666097ED14FFE">2015-09-07</mods:number>
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<treatment id="03BB878CBC5E1B4E9862D93C02E92F2A" ID-DOI="http://doi.org/10.5281/zenodo.3794659" ID-GBIF-Taxon="163887691" ID-Zenodo-Dep="3794659" LSID="urn:lsid:zoobank.org:act:0138AA0F-95A0-4AED-AF47-9DD1CCCB63C1" httpUri="http://treatment.plazi.org/id/03BB878CBC5E1B4E9862D93C02E92F2A" lastPageId="22" lastPageNumber="23" pageId="17" pageNumber="18">
<subSubSection id="C3086511BC5E1B499862D93C00C7283F" box="[603,985,267,294]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BAD369ABC5E1B499862D93C00C7283F" blockId="17.[351,1236,267,367]" box="[603,985,267,294]" pageId="17" pageNumber="18">
<heading id="D0E581F6BC5E1B499862D93C00C7283F" box="[603,985,267,294]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
<taxonomicName id="4C124D19BC5E1B499862D93C0069283F" authority="Smith &amp; Kamiya, 2015" authorityName="Smith &amp; Kamiya" authorityYear="2015" box="[603,887,267,294]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="falcula" status="sp. nov.">
<emphasis id="B966EA88BC5E1B499862D93C0069283F" bold="true" box="[603,887,267,294]" italics="true" pageId="17" pageNumber="18">Pseudocandona falcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557F3BC5E1B499947D93B00C7283F" box="[894,985,268,294]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C3086511BC5E1B499B66D90700842876" pageId="17" pageNumber="18" type="description">
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<uri id="FF833A98BC5E1B499B66D90707CA2852" box="[351,1236,304,331]" pageId="17" pageNumber="18">
urn:lsid:zoobank.org:act:
<uuid id="FFB40C4FBC5E1B499847D90707CA2852" box="[638,1236,304,331]" pageId="17" pageNumber="18">0138AA0F-95A0-4AED-AF47-9DD1CCCB63C1</uuid>
</uri>
</paragraph>
<paragraph id="8BAD369ABC5E1B4998A0D96200842876" blockId="17.[351,1236,267,367]" box="[665,922,341,367]" pageId="17" pageNumber="18">
<figureCitation id="13292A1FBC5E1B4998A0D96201E62876" box="[665,760,341,367]" captionStart="Fig" captionStartId="9.[189,232,1895,1921]" captionTargetBox="[193,1395,564,1847]" captionTargetId="figure@9.[189,1399,559,1854]" captionTargetPageId="9" captionText="Fig. 6. A. Pseudocandona becca sp. nov. hemipenis (LBM 1430006264). B. Pseudocandona atmeta sp. nov. hemipenis (LBM 1430006270). C. Pseudocandona falcula sp. nov. hemipenis (LBM 1430006275). D. Pseudocandona tenuirostris Hiruta &amp; Mawatari, 2013 hemipenis. E. Undulacandona colymba sp. nov. hemipenis (LBM 1430006279)." figureDoi="http://doi.org/10.5281/zenodo.3785316" httpUri="https://zenodo.org/record/3785316/files/figure.png" pageId="17" pageNumber="18">Figs 6C</figureCitation>
,
<figureCitation id="13292A1FBC5E1B49993DD962000F2876" box="[772,785,341,367]" captionStart="Fig" captionStartId="12.[189,232,1789,1815]" captionTargetBox="[245,1342,265,1748]" captionTargetId="figure@12.[245,1342,265,1748]" captionTargetPageId="12" captionText="Fig. 7. AB. Pseudocandona atmeta sp. nov. A. Dorsal view of male carapace, anterior to left (LBM 1430006272). B. Detail of external surface of male right valve (LBM 1430006272). CF. Pseudocandona falcula sp. nov. C. Right view of female carapace (LBM 1430006278). D. Dorsal view of female carapace, anterior to left (LBM 1430006278). E. Internal view of female left valve (LBM 1430006277). F. Internal view of female right valve (LBM 1430006277). GH. Candona candida (O.F. Müller, 1776) G. Internal view of female left valve. H. Dorsal view of female carapace, anterior to left. Scale bar: A, CF = 500 µm; B = 134 µm; GH = 412 µm." figureDoi="http://doi.org/10.5281/zenodo.3785318" httpUri="https://zenodo.org/record/3785318/files/figure.png" pageId="17" pageNumber="18">7</figureCitation>
CF, 1113
</paragraph>
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<subSubSection id="C3086511BC5E1B499A84D9AF00D728AB" box="[189,969,407,434]" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8BAD369ABC5E1B499A84D9AF00D728AB" blockId="17.[189,969,407,434]" box="[189,969,407,434]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="AB0211B4BC5E1B499A84D9AF00D728AB" box="[189,969,407,434]" pageId="17" pageNumber="18">
<taxonomicName id="4C124D19BC5E1B499A84D9AF02A728A8" ID-CoL="1e7147f5-7393-4995-be0d-169d4e5ca6f0" authorityName="in Delorme" authorityYear="1970" baseAuthorityName="Hartwig" baseAuthorityYear="1899" box="[189,441,408,434]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarsi">
<emphasis id="B966EA88BC5E1B499A84D9AF02A728A8" box="[189,441,408,434]" italics="true" pageId="17" pageNumber="18">Pseudocandona sarsi</emphasis>
</taxonomicName>
<treatmentCitation id="0AB3108BBC5E1B499BEFD9A0001B28AB" author="Smith R. J. &amp; Janz H." box="[470,773,407,434]" httpUri="http://treatment.plazi.org/id/6530807DEC07FF8BAFC8FF2AFCF6FC52" page="2909" pageId="17" pageNumber="18" year="2008">
<bibRefCitation id="EF834B6BBC5E1B499BEFD9A0001B28AB" author="Smith R. J. &amp; Janz H." box="[470,773,407,434]" pageId="17" pageNumber="18" pagination="2865 - 2922" refId="ref13871" refString="Smith R. J. &amp; Janz H. 2008. Recent species of the Family Candonidae (Ostracoda, Crustacea) from the ancient Lake Biwa, Central Japan. Journal of Natural History 42 (45 - 46): 2865 - 2922. http: // dx. doi. org / 10.1080 / 00222930802361030" type="journal article" year="2008">Smith &amp; Janz 2008: 2909</bibRefCitation>
</treatmentCitation>
, figs 17KL, 25.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3086511BC5E1B499A84D9ED07EB2A77" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8BAD369ABC5E1B499A84D9ED022828ED" blockId="17.[189,310,474,500]" box="[189,310,474,500]" pageId="17" pageNumber="18">
<heading id="D0E581F6BC5E1B499A84D9ED022828ED" bold="true" box="[189,310,474,500]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B966EA88BC5E1B499A84D9ED022828ED" bold="true" box="[189,310,474,500]" pageId="17" pageNumber="18">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD369ABC5E1B499A84DA3D07EB2A77" blockId="17.[189,1399,522,879]" pageId="17" pageNumber="18">
Hinge straight, sloping to anterior region. Maximum height at posterior third. Posterior margin more inflated than anterior margin, both margins more or less evenly rounded. Calcified inner lamella narrow, wider anteriorly than posteriorly. Dorsal view compressed, with rounded anterior and posterior ends. Male antenna with sub-divided second endopodal segment, and with well-developed t2 and t
<specimenCount id="9D14FD13BC5E1B499F1CDA4E06682B8B" box="[1317,1398,632,659]" pageId="17" pageNumber="18" type="male">3 male</specimenCount>
bristles. Female antennal claw G2 approximately half length of claw G3. Mandible with 3+1+beta setae on second segment of palp, and with long gamma seta and very short alpha and beta setae; beta shorter than alpha. Walking leg with long d1 seta, and with e, and f setae shorter than next segment respectively. Seventh limb with five segments, terminating with long h2 and h3 setae of approximately equal length, and short, reflexed and curled h1 seta. Hemipenis with curved, relatively short M-process, outer lobe (a) large, and extending significantly beyond medial lobe (h), inner lobe (b) angular and folded.
</paragraph>
</subSubSection>
<subSubSection id="C3086511BC5E1B499A84DBAF02DB2D1E" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BAD369ABC5E1B499A84DBAF025A2AAB" blockId="17.[189,324,920,946]" box="[189,324,920,946]" pageId="17" pageNumber="18">
<heading id="D0E581F6BC5E1B499A84DBAF025A2AAB" bold="true" box="[189,324,920,946]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B966EA88BC5E1B499A84DBAF025A2AAB" bold="true" box="[189,324,920,946]" pageId="17" pageNumber="18">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD369ABC5E1B499A84DBFF02DB2D1E" blockId="17.[189,1398,967,1031]" pageId="17" pageNumber="18">
From the Latin
<taxonomicName id="4C124D19BC5E1B499B4EDBF002D62AF8" authorityName="Smith &amp; Kamiya" authorityYear="2015" box="[375,456,967,993]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="falcula">
<emphasis id="B966EA88BC5E1B499B4EDBF002D62AF8" box="[375,456,967,993]" italics="true" pageId="17" pageNumber="18">falcula</emphasis>
</taxonomicName>
, meaning a “sickle” or “scythe”, and referring to the shape of the M-process of the male sexual organ.
</paragraph>
</subSubSection>
<subSubSection id="C3086511BC5E1B499A84DC1800A42CE6" pageId="17" pageNumber="18" type="materials_examined">
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<heading id="D0E581F6BC5E1B499A84DC1802722D50" bold="true" box="[189,364,1071,1097]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B966EA88BC5E1B499A84DC1802722D50" bold="true" box="[189,364,1071,1097]" pageId="17" pageNumber="18">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B7A3CC7BC5E1B499AD3DC6807BD2DDB" ID-GBIF-Occurrence="2610058624" collectingDate="2004-09-27" collectionCode="LBM" pageId="17" pageNumber="18" specimenCode="LBM 1430006275" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="8BAD369ABC5E1B499AD3DC6802422D60" blockId="17.[189,1398,1119,1219]" box="[234,348,1119,1145]" pageId="17" pageNumber="18">
<emphasis id="B966EA88BC5E1B499AD3DC6802422D60" bold="true" box="[234,348,1119,1145]" pageId="17" pageNumber="18">
<typeStatus id="54A98838BC5E1B499AD3DC6802422D60" box="[234,348,1119,1145]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAD369ABC5E1B499A84DCB307BD2DDB" blockId="17.[189,1398,1119,1219]" pageId="17" pageNumber="18">
<specimenCount id="9D14FD13BC5E1B499A84DCB303CD2D87" box="[189,211,1156,1182]" pageId="17" pageNumber="18" type="male"></specimenCount>
(
<specimenCode id="DBB49EE1BC5E1B499ADCDCB302D62D87" box="[229,456,1155,1182]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="17" pageNumber="18">LBM 1430006275</specimenCode>
), dissected with appendages sealed in a glass slide and valves stored dry in a micropalaeontological cavity slide. Collected from the type locality on
<date id="FFAC105ABC5E1B499E38DC9F07802DDB" box="[1025,1182,1192,1218]" pageId="17" pageNumber="18" value="2004-09-27">
<collectingDate id="EFE8E9B2BC5E1B499E38DC9F07802DDB" box="[1025,1182,1192,1218]" pageId="17" pageNumber="18" value="2004-09-27">27 Sep. 2004</collectingDate>
</date>
.
</paragraph>
</materialsCitation>
<paragraph id="8BAD369ABC5E1B499AD3DCDC024A2C1C" blockId="17.[189,1398,1259,1359]" box="[234,340,1259,1285]" pageId="17" pageNumber="18">
<emphasis id="B966EA88BC5E1B499AD3DCDC024A2C1C" bold="true" box="[234,340,1259,1285]" pageId="17" pageNumber="18">
<typeStatus id="54A98838BC5E1B499AD3DCDC024A2C1C" box="[234,340,1259,1285]" pageId="17" pageNumber="18" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAD369ABC5E1B499A84DD2707BD2C56" blockId="17.[189,1398,1259,1359]" pageId="17" pageNumber="18">
<specimenCount id="9D14FD13BC5E1B499A84DD2703CD2C33" box="[189,211,1296,1322]" pageId="17" pageNumber="18" type="female"></specimenCount>
(
<specimenCode id="DBB49EE1BC5E1B499ADCDD2702D62C32" box="[229,456,1296,1323]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="17" pageNumber="18">LBM 1430006276</specimenCode>
), dissected with appendages sealed in a glass slide and valves stored dry in a micropalaeontological cavity slide. Collected from the
<typeStatus id="54A98838BC5E1B49997ADD0200682C56" box="[835,886,1333,1359]" pageId="17" pageNumber="18">type</typeStatus>
locality on
<date id="FFAC105ABC5E1B499E38DD0207802C56" box="[1025,1182,1333,1359]" pageId="17" pageNumber="18" value="2004-09-27">
<collectingDate id="EFE8E9B2BC5E1B499E38DD0207802C56" box="[1025,1182,1333,1359]" pageId="17" pageNumber="18" value="2004-09-27">27 Sep. 2004</collectingDate>
</date>
.
</paragraph>
<paragraph id="8BAD369ABC5E1B499AD3DD4F02792C8B" blockId="17.[189,1398,1400,1536]" box="[234,359,1400,1426]" pageId="17" pageNumber="18">
<emphasis id="B966EA88BC5E1B499AD3DD4F02792C8B" bold="true" box="[234,359,1400,1426]" pageId="17" pageNumber="18">
<typeStatus id="54A98838BC5E1B499AD3DD4F02792C8B" box="[234,359,1400,1426]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAD369ABC5E1B499A84DDAB00A42CE6" blockId="17.[189,1398,1400,1536]" pageId="17" pageNumber="18">
<specimenCount id="9D14FD13BC5E1B499A84DDAB03F52CAE" box="[189,235,1436,1463]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED03AE5FBC5E1B499AC5DDAB02232CAF" box="[252,317,1436,1462]" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="17" pageNumber="18">LBM</collectionCode>
1430006277), dissected with appendages sealed in a glass slide and valves stored dry in a micropalaeontological cavity slide.
<specimenCount id="9D14FD13BC5E1B49986FDDF601612CC2" box="[598,639,1473,1499]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED03AE5FBC5E1B4998B5DDF601D32CC2" box="[652,717,1473,1499]" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="17" pageNumber="18">LBM</collectionCode>
1430006278), whole, stored dry in a micropalaeontological cavity slide. All collected from the
<typeStatus id="54A98838BC5E1B499863DDD101932F19" box="[602,653,1510,1536]" pageId="17" pageNumber="18">type</typeStatus>
locality on
<date id="FFAC105ABC5E1B499921DDD200AB2CE6" box="[792,949,1509,1535]" pageId="17" pageNumber="18" value="2004-09-27">27 Sep. 2004</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3086511BC5E1B499A84DE1F06452F8E" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BAD369ABC5E1B499A84DE1F02432F5B" blockId="17.[189,349,1576,1602]" box="[189,349,1576,1602]" pageId="17" pageNumber="18">
<heading id="D0E581F6BC5E1B499A84DE1F02432F5B" bold="true" box="[189,349,1576,1602]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B966EA88BC5E1B499A84DE1F02432F5B" bold="true" box="[189,349,1576,1602]" pageId="17" pageNumber="18">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD369ABC5E1B499A84DE6F06452F8E" blockId="17.[189,1398,1624,1687]" pageId="17" pageNumber="18">
Sarugamori, Higashidori, Shimokita District,
<collectingRegion id="49D6F878BC5E1B4998F1DE6F00BA2F6B" box="[712,932,1624,1650]" country="Japan" name="Aomori" pageId="17" pageNumber="18">Aomori Prefecture</collectingRegion>
,
<collectingCountry id="F305760ABC5E1B499994DE6E00EE2F6A" box="[941,1008,1625,1651]" name="Japan" pageId="17" pageNumber="18">Japan</collectingCountry>
(
<geoCoordinate id="EE26505DBC5E1B4999C4DE6F07B72F6A" box="[1021,1193,1624,1651]" degrees="41" direction="north" minutes="16" orientation="latitude" pageId="17" pageNumber="18" precision="1" seconds="00.6" value="41.266834">41º1600.6” N</geoCoordinate>
,
<geoCoordinate id="EE26505DBC5E1B499E8BDE6F06772F6A" box="[1202,1385,1624,1651]" degrees="141" direction="east" minutes="22" orientation="longitude" pageId="17" pageNumber="18" precision="1" seconds="38.4" value="141.37733">141º2238.4” E</geoCoordinate>
). Boggy, marshy area with little standing water, lots of reeds, and sodden ground. Locality 4 on
<figureCitation id="13292A1FBC5E1B499F36DE4A064B2F8E" box="[1295,1365,1661,1687]" captionStart="Fig" captionStartId="1.[189,232,2000,2026]" captionTargetBox="[189,1399,737,1960]" captionTargetId="figure@1.[190,1397,739,1959]" captionTargetPageId="1" captionText="Fig. 1. Map of the main islands of Japan, showing the localities mentioned in the text." figureDoi="http://doi.org/10.5281/zenodo.3785304" httpUri="https://zenodo.org/record/3785304/files/figure.png" pageId="17" pageNumber="18">Fig. 1</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3086511BC5E1B499A84DEF7000B2E13" pageId="17" pageNumber="18" type="key">
<paragraph id="8BAD369ABC5E1B499A84DEF702E72FC3" blockId="17.[189,505,1728,1754]" box="[189,505,1728,1754]" pageId="17" pageNumber="18">
<heading id="D0E581F6BC5E1B499A84DEF702E72FC3" bold="true" box="[189,505,1728,1754]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B966EA88BC5E1B499A84DEF702E72FC3" bold="true" box="[189,505,1728,1754]" pageId="17" pageNumber="18">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD369ABC5E1B499A84DEC7000B2E13" blockId="17.[189,789,1776,1802]" box="[189,789,1776,1802]" pageId="17" pageNumber="18">
<specimenCount id="9D14FD13BC5E1B499A84DEC703F72E13" box="[189,233,1776,1802]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, from the
<typeStatus id="54A98838BC5E1B499B5ADEC702882E13" box="[355,406,1776,1802]" pageId="17" pageNumber="18">type</typeStatus>
locality, collected
<date id="FFAC105ABC5E1B49984ADEC7000E2E13" box="[627,784,1776,1802]" pageId="17" pageNumber="18" value="2004-09-27">27 Sep. 2004</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3086511BC5E1B4E9A84DF0507122881" lastPageId="22" lastPageNumber="23" pageId="17" pageNumber="18" type="description">
<paragraph id="8BAD369ABC5E1B499A84DF0502502E55" blockId="17.[189,334,1842,1868]" box="[189,334,1842,1868]" pageId="17" pageNumber="18">
<heading id="D0E581F6BC5E1B499A84DF0502502E55" bold="true" box="[189,334,1842,1868]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B966EA88BC5E1B499A84DF0502502E55" bold="true" box="[189,334,1842,1868]" pageId="17" pageNumber="18">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD369ABC5E1B499A84DF5406692EF2" blockId="17.[189,1399,1890,2027]" pageId="17" pageNumber="18">
Carapace (
<figureCitation id="13292A1FBC5E1B499B02DF54029A2E64" box="[315,388,1891,1917]" captionStart="Fig" captionStartId="12.[189,232,1789,1815]" captionTargetBox="[245,1342,265,1748]" captionTargetId="figure@12.[245,1342,265,1748]" captionTargetPageId="12" captionText="Fig. 7. AB. Pseudocandona atmeta sp. nov. A. Dorsal view of male carapace, anterior to left (LBM 1430006272). B. Detail of external surface of male right valve (LBM 1430006272). CF. Pseudocandona falcula sp. nov. C. Right view of female carapace (LBM 1430006278). D. Dorsal view of female carapace, anterior to left (LBM 1430006278). E. Internal view of female left valve (LBM 1430006277). F. Internal view of female right valve (LBM 1430006277). GH. Candona candida (O.F. Müller, 1776) G. Internal view of female left valve. H. Dorsal view of female carapace, anterior to left. Scale bar: A, CF = 500 µm; B = 134 µm; GH = 412 µm." figureDoi="http://doi.org/10.5281/zenodo.3785318" httpUri="https://zenodo.org/record/3785318/files/figure.png" pageId="17" pageNumber="18">Figs 7</figureCitation>
CF, 11AB) length
<quantity id="4CEA9B7FBC5E1B499844DF5400082E64" box="[637,790,1891,1917]" metricMagnitude="-1" metricUnit="m" metricValue="9.275" metricValueMax="9.37" metricValueMin="9.18" pageId="17" pageNumber="18" unit="mm" value="927.5" valueMax="937.0" valueMin="918.0">918937 µm</quantity>
, height
<quantity id="4CEA9B7FBC5E1B49994FDF55070E2E64" box="[886,1040,1890,1917]" metricMagnitude="-1" metricUnit="m" metricValue="5.145" metricValueMax="5.33" metricValueMin="4.96" pageId="17" pageNumber="18" unit="mm" value="514.5" valueMax="533.0" valueMin="496.0">496533 µm</quantity>
. Maximum height at posterior third, hinge straight, sloping towards evenly rounded anterior margin. Posterior margin more inflated than anterior margin, and evenly curved. Ventral margin slightly concave. Inner calcified lamella relatively narrow, wider anteriorly than posteriorly. Six small adductor muscle scars in tight formation
</paragraph>
<paragraph id="8BAD369ABC5D1B4A9A84D93B00322850" blockId="18.[189,1399,267,329]" pageId="18" pageNumber="19">at mid-height, anterior of mid-length. Indistinct dorsal scars near dorsal margin. Female very similar to male. Colour, white. Surface covered with stiff setae.</paragraph>
<paragraph id="8BAD369ABC5D1B4A9A84D946008C2B03" blockId="18.[189,1399,369,538]" pageId="18" pageNumber="19">
Antennule with seven articulated segments (
<figureCitation id="13292A1FBC5D1B4A98EFD946005D2892" box="[726,835,369,395]" captionStart="Fig" captionStartId="19.[189,232,1860,1886]" captionTargetBox="[209,1379,269,1812]" captionTargetId="figure@19.[209,1379,265,1819]" captionTargetPageId="19" captionText="Fig. 11. Pseudocandona falcula sp. nov. A. Internal view of male left valve (LBM 1430006275). B. Internal view of female left valve (LBM 1430006276). C. Male antennule (LBM 1430006275). D. Male antenna (LBM 1430006275). E. Detail of male antenna (LBM 1430006275). F. Detail of female antenna (LBM 1430006276). G. Male mandibular coxa (LBM 1430006275). Scale bar: AB = 543 µm; CD, G = 151 µm; EF = 82 µm." figureDoi="http://doi.org/10.5281/zenodo.3785326" httpUri="https://zenodo.org/record/3785326/files/figure.png" pageId="18" pageNumber="19">Fig. 11C</figureCitation>
). First segment large, supporting two setae on dorsal edge and two long setae on ventral-apical corner. Second and third segments quadrate each with one dorsal-apical seta. Fourth and fifth segments each with two long dorsal-apical setae and one short ventral-apical seta. Sixth segment with two long, one mid-length and one short apical setae. Final segment with two long and one short setae, and aesthetasc ya.
</paragraph>
<paragraph id="8BAD369ABC5D1B4A9A84DA76013E2A28" blockId="18.[189,1399,577,817]" pageId="18" pageNumber="19">
Male antenna with second endopodite segment sub-divided (
<figureCitation id="13292A1FBC5D1B4A99A9DA7600FC2B42" box="[912,994,577,603]" captionStart="Fig" captionStartId="19.[189,232,1860,1886]" captionTargetBox="[209,1379,269,1812]" captionTargetId="figure@19.[209,1379,265,1819]" captionTargetPageId="19" captionText="Fig. 11. Pseudocandona falcula sp. nov. A. Internal view of male left valve (LBM 1430006275). B. Internal view of female left valve (LBM 1430006276). C. Male antennule (LBM 1430006275). D. Male antenna (LBM 1430006275). E. Detail of male antenna (LBM 1430006275). F. Detail of female antenna (LBM 1430006276). G. Male mandibular coxa (LBM 1430006275). Scale bar: AB = 543 µm; CD, G = 151 µm; EF = 82 µm." figureDoi="http://doi.org/10.5281/zenodo.3785326" httpUri="https://zenodo.org/record/3785326/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
DE). Setae t2 and t3 represented by male bristles, both similar in morphology, but with t2 longer than t3, both terminating with small, triangular process. Seta t4 tiny, protruding from near base of t3. Setae z2 and z3 very short, z1 represented by well-developed claw, slightly shorter than claw G2. Claw G1 less than half length of claw G2. Claw Gm on final segment half the length of claw GM. Female antennal claw G2 just under half length of G1, z1 stout and claw-like, less than half length of G2, setae z2 and z3 very short. Claw Gm more than half length of claw GM (
<figureCitation id="13292A1FBC5D1B4A9B92DB20010D2A28" box="[427,531,791,817]" captionStart="Fig" captionStartId="19.[189,232,1860,1886]" captionTargetBox="[209,1379,269,1812]" captionTargetId="figure@19.[209,1379,265,1819]" captionTargetPageId="19" captionText="Fig. 11. Pseudocandona falcula sp. nov. A. Internal view of male left valve (LBM 1430006275). B. Internal view of female left valve (LBM 1430006276). C. Male antennule (LBM 1430006275). D. Male antenna (LBM 1430006275). E. Detail of male antenna (LBM 1430006275). F. Detail of female antenna (LBM 1430006276). G. Male mandibular coxa (LBM 1430006275). Scale bar: AB = 543 µm; CD, G = 151 µm; EF = 82 µm." figureDoi="http://doi.org/10.5281/zenodo.3785326" httpUri="https://zenodo.org/record/3785326/files/figure.png" pageId="18" pageNumber="19">Fig. 11F</figureCitation>
).
</paragraph>
<paragraph id="8BAD369ABC5D1B4A9A84DB6E02C32D75" blockId="18.[189,1399,857,1132]" pageId="18" pageNumber="19">
Mandibular palp (
<figureCitation id="13292A1FBC5D1B4A9BAFDB6E02F22A6A" box="[406,492,857,883]" captionStart="Fig" captionStartId="20.[189,232,1860,1886]" captionTargetBox="[219,1369,269,1819]" captionTargetId="figure@20.[219,1369,265,1819]" captionTargetPageId="20" captionText="Fig. 12. Pseudocandona falcula sp. nov. A. Male mandibular palp (LBM 1430006275). B. Detail of alpha and beta setae of mandibular palp (LBM 1430006275). C. Male maxillula (setae on endites not drawn) (LBM 1430006275). D. Male left fifth limb palp (LBM 1430006275). E. Male right fifth limb palp (LBM 1430006275). F. Female fifth limb (LBM 1430006276). G. Male sixth limb (LBM 1430006275). H. Male seventh limb (LBM 1430006275). Scale bar: AC = 82 µm; DH = 151 µm." figureDoi="http://doi.org/10.5281/zenodo.3785328" httpUri="https://zenodo.org/record/3785328/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
AB) with four segments. Alpha seta of first segment short, and with very narrow, flagellum-like end. Inner edge of second segment with 3+1+beta arrangement of setae; seta beta shorter than alpha. Outer edge of second segment with two apical setae. Third segment with three long sub-apical setae on outer edge, and three long and one short setae arranged along apical edge; outermost gamma seta, long and without obvious setules. Final segment with robust seta on outer edge, thick claw-like seta in mid-apical position and three shorter setae on inner apical edge. Number of setae on branchial plate not observed. Mandibular coxa (
<figureCitation id="13292A1FBC5D1B4A98D2DC18004D2D50" box="[747,851,1071,1097]" captionStart="Fig" captionStartId="19.[189,232,1860,1886]" captionTargetBox="[209,1379,269,1812]" captionTargetId="figure@19.[209,1379,265,1819]" captionTargetPageId="19" captionText="Fig. 11. Pseudocandona falcula sp. nov. A. Internal view of male left valve (LBM 1430006275). B. Internal view of female left valve (LBM 1430006276). C. Male antennule (LBM 1430006275). D. Male antenna (LBM 1430006275). E. Detail of male antenna (LBM 1430006275). F. Detail of female antenna (LBM 1430006276). G. Male mandibular coxa (LBM 1430006275). Scale bar: AB = 543 µm; CD, G = 151 µm; EF = 82 µm." figureDoi="http://doi.org/10.5281/zenodo.3785326" httpUri="https://zenodo.org/record/3785326/files/figure.png" pageId="18" pageNumber="19">Fig. 11G</figureCitation>
) with five well-developed teeth plus two much smaller, spine-like teeth.
</paragraph>
<paragraph id="8BAD369ABC5D1B4A9A84DCA307132C00" blockId="18.[189,1399,1172,1306]" pageId="18" pageNumber="19">
Maxillula (
<figureCitation id="13292A1FBC5D1B4A9B07DCA302B82DB7" box="[318,422,1172,1198]" captionStart="Fig" captionStartId="20.[189,232,1860,1886]" captionTargetBox="[219,1369,269,1819]" captionTargetId="figure@20.[219,1369,265,1819]" captionTargetPageId="20" captionText="Fig. 12. Pseudocandona falcula sp. nov. A. Male mandibular palp (LBM 1430006275). B. Detail of alpha and beta setae of mandibular palp (LBM 1430006275). C. Male maxillula (setae on endites not drawn) (LBM 1430006275). D. Male left fifth limb palp (LBM 1430006275). E. Male right fifth limb palp (LBM 1430006275). F. Female fifth limb (LBM 1430006276). G. Male sixth limb (LBM 1430006275). H. Male seventh limb (LBM 1430006275). Scale bar: AC = 82 µm; DH = 151 µm." figureDoi="http://doi.org/10.5281/zenodo.3785328" httpUri="https://zenodo.org/record/3785328/files/figure.png" pageId="18" pageNumber="19">Fig. 12C</figureCitation>
) palp first segment with three setae on apical outer margin, and one apical seta offset towards inner edge. Second segment with slightly stepped apical margin, with outer part more distal than inner part. Outer part of apical margin with two long and one short setae, and inner part with three mid-length setae. Branchial plate with morphology typical of subfamily.
</paragraph>
<paragraph id="8BAD369ABC5D1B4A9A84DD7606772C66" blockId="18.[189,1399,1345,1407]" pageId="18" pageNumber="19">
Fifth limb male palps asymmetrical (
<figureCitation id="13292A1FBC5D1B4A985CDD7601A82C42" box="[613,694,1345,1371]" captionStart="Fig" captionStartId="20.[189,232,1860,1886]" captionTargetBox="[219,1369,269,1819]" captionTargetId="figure@20.[219,1369,265,1819]" captionTargetPageId="20" captionText="Fig. 12. Pseudocandona falcula sp. nov. A. Male mandibular palp (LBM 1430006275). B. Detail of alpha and beta setae of mandibular palp (LBM 1430006275). C. Male maxillula (setae on endites not drawn) (LBM 1430006275). D. Male left fifth limb palp (LBM 1430006275). E. Male right fifth limb palp (LBM 1430006275). F. Female fifth limb (LBM 1430006276). G. Male sixth limb (LBM 1430006275). H. Male seventh limb (LBM 1430006275). Scale bar: AC = 82 µm; DH = 151 µm." figureDoi="http://doi.org/10.5281/zenodo.3785328" httpUri="https://zenodo.org/record/3785328/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
DE). Right palp rounded, helmet-shaped, more inflated than left. Left palp almost sub-triangular distally with angular outer margin, and almost straight inner margin.
</paragraph>
<paragraph id="8BAD369ABC5D1B4A9A84DD9007C52F11" blockId="18.[189,1399,1447,1544]" pageId="18" pageNumber="19">
Fifth limb of female (
<figureCitation id="13292A1FBC5D1B4A9B9FDD9001182CD8" box="[422,518,1447,1473]" captionStart="Fig" captionStartId="20.[189,232,1860,1886]" captionTargetBox="[219,1369,269,1819]" captionTargetId="figure@20.[219,1369,265,1819]" captionTargetPageId="20" captionText="Fig. 12. Pseudocandona falcula sp. nov. A. Male mandibular palp (LBM 1430006275). B. Detail of alpha and beta setae of mandibular palp (LBM 1430006275). C. Male maxillula (setae on endites not drawn) (LBM 1430006275). D. Male left fifth limb palp (LBM 1430006275). E. Male right fifth limb palp (LBM 1430006275). F. Female fifth limb (LBM 1430006276). G. Male sixth limb (LBM 1430006275). H. Male seventh limb (LBM 1430006275). Scale bar: AC = 82 µm; DH = 151 µm." figureDoi="http://doi.org/10.5281/zenodo.3785328" httpUri="https://zenodo.org/record/3785328/files/figure.png" pageId="18" pageNumber="19">Fig.12F</figureCitation>
) with long b and d setae, and one long a seta on basis. Endite with approximately nine apical setae, and four sub-apical setae on inner edge. Palp (endopodite) terminating with three setae on unequal lengths. Branchial plate consisting of two rays, one shorter and narrower than other.
</paragraph>
<paragraph id="8BAD369ABC5D1B4A9A84DE0706682F88" blockId="18.[189,1399,1584,1682]" pageId="18" pageNumber="19">
Sixth limb (
<figureCitation id="13292A1FBC5D1B4A9B72DE0702A82F53" box="[331,438,1584,1610]" captionStart="Fig" captionStartId="20.[189,232,1860,1886]" captionTargetBox="[219,1369,269,1819]" captionTargetId="figure@20.[219,1369,265,1819]" captionTargetPageId="20" captionText="Fig. 12. Pseudocandona falcula sp. nov. A. Male mandibular palp (LBM 1430006275). B. Detail of alpha and beta setae of mandibular palp (LBM 1430006275). C. Male maxillula (setae on endites not drawn) (LBM 1430006275). D. Male left fifth limb palp (LBM 1430006275). E. Male right fifth limb palp (LBM 1430006275). F. Female fifth limb (LBM 1430006276). G. Male sixth limb (LBM 1430006275). H. Male seventh limb (LBM 1430006275). Scale bar: AC = 82 µm; DH = 151 µm." figureDoi="http://doi.org/10.5281/zenodo.3785328" httpUri="https://zenodo.org/record/3785328/files/figure.png" pageId="18" pageNumber="19">Fig. 12G</figureCitation>
) five-segmented, with first segment bearing d1 seta. Setae e and f of second and third segments respectively, both about half length of next respective segment. Fourth segment with g seta reaching to end of fifth segment. Fifth segment with short h1 seta, tiny h3 seta and well-developed claw h2.
</paragraph>
<paragraph id="8BAD369ABC5D1B4A9A84DE8E06692E03" blockId="18.[189,1399,1721,1818]" pageId="18" pageNumber="19">
Seventh limb with five segments (
<figureCitation id="13292A1FBC5D1B4A986EDE8E01D82FCA" box="[599,710,1721,1747]" captionStart="Fig" captionStartId="20.[189,232,1860,1886]" captionTargetBox="[219,1369,269,1819]" captionTargetId="figure@20.[219,1369,265,1819]" captionTargetPageId="20" captionText="Fig. 12. Pseudocandona falcula sp. nov. A. Male mandibular palp (LBM 1430006275). B. Detail of alpha and beta setae of mandibular palp (LBM 1430006275). C. Male maxillula (setae on endites not drawn) (LBM 1430006275). D. Male left fifth limb palp (LBM 1430006275). E. Male right fifth limb palp (LBM 1430006275). F. Female fifth limb (LBM 1430006276). G. Male sixth limb (LBM 1430006275). H. Male seventh limb (LBM 1430006275). Scale bar: AC = 82 µm; DH = 151 µm." figureDoi="http://doi.org/10.5281/zenodo.3785328" httpUri="https://zenodo.org/record/3785328/files/figure.png" pageId="18" pageNumber="19">Fig. 12H</figureCitation>
). First segment with long dp seta, and medium-length d1 and d2 setae. Second and third segments with no setae. Fourth segment with long g seta. Fifth segment approximately quadrate, h2 and h3 long and of approximately equal length, h1 short, hook-like and reflexed.
</paragraph>
<paragraph id="8BAD369ABC5D1B4A9A84DF7506022EF2" blockId="18.[189,1399,1858,2027]" pageId="18" pageNumber="19">
Caudal ramus (
<figureCitation id="13292A1FBC5D1B4A9B5EDF7502AA2E45" box="[359,436,1858,1884]" captionStart="Fig" captionStartId="21.[189,232,1895,1921]" captionTargetBox="[189,1399,548,1855]" captionTargetId="figure@21.[189,1399,544,1855]" captionTargetPageId="21" captionText="Fig. 13. Pseudocandona falcula sp. nov. A. Male caudal ramus (LBM 1430006275). B. Male caudal ramus attachment (LBM 1430006275). C. Female caudal ramus and genital lobe (LBM 1430006277), black triangle indicates rounded lobe (see text for details). D. Hemipenis (M-process shaded grey) (LBM 1430006275). Scale bar = 151 µm." figureDoi="http://doi.org/10.5281/zenodo.3785330" httpUri="https://zenodo.org/record/3785330/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
AC) with inflated base, tapering distally, slightly curved to straight. Claw Ga straight proximally, curving distally. Claw Gp slightly shorter and thinner than Ga. Seta sa short, seta sp long, about 50% length of claw Gp. Caudal ramus attachment of male (
<figureCitation id="13292A1FBC5D1B4A9992DFBE070F2EBD" box="[939,1041,1929,1956]" captionStart="Fig" captionStartId="21.[189,232,1895,1921]" captionTargetBox="[189,1399,548,1855]" captionTargetId="figure@21.[189,1399,544,1855]" captionTargetPageId="21" captionText="Fig. 13. Pseudocandona falcula sp. nov. A. Male caudal ramus (LBM 1430006275). B. Male caudal ramus attachment (LBM 1430006275). C. Female caudal ramus and genital lobe (LBM 1430006277), black triangle indicates rounded lobe (see text for details). D. Hemipenis (M-process shaded grey) (LBM 1430006275). Scale bar = 151 µm." figureDoi="http://doi.org/10.5281/zenodo.3785330" httpUri="https://zenodo.org/record/3785330/files/figure.png" pageId="18" pageNumber="19">Fig. 13B</figureCitation>
) with one posterior branch and two anterior branches on dorsal-most half. Female caudal ramus attachment (
<figureCitation id="13292A1FBC5D1B4A9E09DF9A07892EDE" box="[1072,1175,1965,1991]" captionStart="Fig" captionStartId="21.[189,232,1895,1921]" captionTargetBox="[189,1399,548,1855]" captionTargetId="figure@21.[189,1399,544,1855]" captionTargetPageId="21" captionText="Fig. 13. Pseudocandona falcula sp. nov. A. Male caudal ramus (LBM 1430006275). B. Male caudal ramus attachment (LBM 1430006275). C. Female caudal ramus and genital lobe (LBM 1430006277), black triangle indicates rounded lobe (see text for details). D. Hemipenis (M-process shaded grey) (LBM 1430006275). Scale bar = 151 µm." figureDoi="http://doi.org/10.5281/zenodo.3785330" httpUri="https://zenodo.org/record/3785330/files/figure.png" pageId="18" pageNumber="19">Fig. 13C</figureCitation>
) with one posterior branch and two anterior branches, lowest of which curves to posterior edge of female genital lobe.
</paragraph>
<caption id="DF6D6612BC5C1B4B9A84DF7301582EF2" ID-DOI="http://doi.org/10.5281/zenodo.3785326" ID-Zenodo-Dep="3785326" httpUri="https://zenodo.org/record/3785326/files/figure.png" pageId="19" pageNumber="20" startId="19.[189,232,1860,1886]" targetBox="[209,1379,269,1812]" targetPageId="19">
<paragraph id="8BAD369ABC5C1B4B9A84DF7301582EF2" blockId="19.[189,1399,1860,2027]" pageId="19" pageNumber="20">
<emphasis id="B966EA88BC5C1B4B9A84DF7302032E47" bold="true" box="[189,285,1860,1886]" pageId="19" pageNumber="20">Fig. 11.</emphasis>
<taxonomicName id="4C124D19BC5C1B4B9B10DF73015C2E47" authority="Smith &amp; Kamiya, 2015" authorityName="Smith &amp; Kamiya" authorityYear="2015" box="[297,578,1860,1886]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="falcula" status="sp. nov.">
<emphasis id="B966EA88BC5C1B4B9B10DF73015C2E47" box="[297,578,1860,1886]" italics="true" pageId="19" pageNumber="20">Pseudocandona falcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557F3BC5C1B4B9876DF7301B12E47" box="[591,687,1860,1886]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B966EA88BC5C1B4B9885DF7301CF2E47" bold="true" box="[700,721,1860,1886]" pageId="19" pageNumber="20">A</emphasis>
. Internal view of male left valve (LBM 1430006275).
<emphasis id="B966EA88BC5C1B4B9A84DF5003CF2E98" bold="true" box="[189,209,1895,1921]" pageId="19" pageNumber="20">B</emphasis>
. Internal view of female left valve (LBM 1430006276).
<emphasis id="B966EA88BC5C1B4B99A9DF5000BB2E98" bold="true" box="[912,933,1895,1921]" pageId="19" pageNumber="20">C</emphasis>
. Male antennule (LBM 1430006275).
<emphasis id="B966EA88BC5C1B4B9A84DFBD03CC2EBD" bold="true" box="[189,210,1930,1956]" pageId="19" pageNumber="20">D</emphasis>
. Male antenna (LBM 1430006275).
<emphasis id="B966EA88BC5C1B4B9843DFBD01902EBD" bold="true" box="[634,654,1930,1956]" pageId="19" pageNumber="20">E</emphasis>
. Detail of male antenna (LBM 1430006275).
<emphasis id="B966EA88BC5C1B4B9EA4DFBD07B12EBD" bold="true" box="[1181,1199,1930,1956]" pageId="19" pageNumber="20">F</emphasis>
. Detail of female antenna (LBM 1430006276).
<emphasis id="B966EA88BC5C1B4B9820DF9A012E2EDE" bold="true" box="[537,560,1965,1991]" pageId="19" pageNumber="20">G</emphasis>
. Male mandibular coxa (LBM 1430006275). Scale bar: AB = 543 µm; CD, G = 151 µm; EF = 82 µm.
</paragraph>
</caption>
<caption id="DF6D6612BC5B1B4C9A84DF7307B92EF2" ID-DOI="http://doi.org/10.5281/zenodo.3785328" ID-Zenodo-Dep="3785328" httpUri="https://zenodo.org/record/3785328/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,232,1860,1886]" targetBox="[219,1369,269,1819]" targetPageId="20">
<paragraph id="8BAD369ABC5B1B4C9A84DF7307B92EF2" blockId="20.[189,1399,1860,2027]" pageId="20" pageNumber="21">
<emphasis id="B966EA88BC5B1B4C9A84DF7302092E47" bold="true" box="[189,279,1860,1886]" pageId="20" pageNumber="21">Fig. 12.</emphasis>
<taxonomicName id="4C124D19BC5B1B4C9B25DF7301332E47" authority="Smith &amp; Kamiya, 2015" authorityName="Smith &amp; Kamiya" authorityYear="2015" box="[284,557,1860,1886]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="falcula" status="sp. nov.">
<emphasis id="B966EA88BC5B1B4C9B25DF7301332E47" box="[284,557,1860,1886]" italics="true" pageId="20" pageNumber="21">Pseudocandona falcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557F3BC5B1B4C980ADF7301952E47" box="[563,651,1860,1886]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B966EA88BC5B1B4C98A9DF7301BB2E47" bold="true" box="[656,677,1860,1886]" pageId="20" pageNumber="21">A</emphasis>
. Male mandibular palp (LBM 1430006275).
<emphasis id="B966EA88BC5B1B4C9E94DF7307DF2E47" bold="true" box="[1197,1217,1860,1886]" pageId="20" pageNumber="21">B</emphasis>
. Detail of alpha and beta setae of mandibular palp (LBM 1430006275).
<emphasis id="B966EA88BC5B1B4C9974DF50007C2E98" bold="true" box="[845,866,1895,1921]" pageId="20" pageNumber="21">C</emphasis>
. Male maxillula (setae on endites not drawn) (LBM 1430006275).
<emphasis id="B966EA88BC5B1B4C9B86DFBD02CA2EBD" bold="true" box="[447,468,1930,1956]" pageId="20" pageNumber="21">D</emphasis>
. Male left fifth limb palp (LBM 1430006275).
<emphasis id="B966EA88BC5B1B4C9E23DFBD07302EBD" bold="true" box="[1050,1070,1930,1956]" pageId="20" pageNumber="21">E</emphasis>
. Male right fifth limb palp (LBM 1430006275).
<emphasis id="B966EA88BC5B1B4C9B8EDF9902D72ED1" bold="true" box="[439,457,1966,1992]" pageId="20" pageNumber="21">F</emphasis>
. Female fifth limb (LBM 1430006276).
<emphasis id="B966EA88BC5B1B4C9999DF9A00A92EDE" bold="true" box="[928,951,1965,1991]" pageId="20" pageNumber="21">G</emphasis>
. Male sixth limb (LBM 1430006275).
<emphasis id="B966EA88BC5B1B4C9A84DFE603CA2EF2" bold="true" box="[189,212,2001,2027]" pageId="20" pageNumber="21">H</emphasis>
. Male seventh limb (LBM 1430006275). Scale bar: AC = 82 µm; DH = 151 µm.
</paragraph>
</caption>
<paragraph id="8BAD369ABC5A1B4D9A84D93C06422896" blockId="21.[189,1398,267,399]" pageId="21" pageNumber="22">
Hemipenis (
<figureCitation id="13292A1FBC5A1B4D9B72D93C02AB283C" box="[331,437,267,293]" captionStart="Fig" captionStartId="21.[189,232,1895,1921]" captionTargetBox="[189,1399,548,1855]" captionTargetId="figure@21.[189,1399,544,1855]" captionTargetPageId="21" captionText="Fig. 13. Pseudocandona falcula sp. nov. A. Male caudal ramus (LBM 1430006275). B. Male caudal ramus attachment (LBM 1430006275). C. Female caudal ramus and genital lobe (LBM 1430006277), black triangle indicates rounded lobe (see text for details). D. Hemipenis (M-process shaded grey) (LBM 1430006275). Scale bar = 151 µm." figureDoi="http://doi.org/10.5281/zenodo.3785330" httpUri="https://zenodo.org/record/3785330/files/figure.png" pageId="21" pageNumber="22">Fig. 13D</figureCitation>
) outer lobe (a) with truncated, sub-quadrate distal margin, extending beyond other two lobes. Medial lobe (h) with unevenly curved, rounded distal margin. Inner lobe (b) folded, triangular on inner edge, one part tongue-shaped, other quadrate. M-process small, rounded proximally, strongly curved and hook-like with lobe on inner edge. Bursa copulatrix (e) elongate with long finger distally.
</paragraph>
<caption id="DF6D6612BC5A1B4D9A84DF5001482EF2" ID-DOI="http://doi.org/10.5281/zenodo.3785330" ID-Zenodo-Dep="3785330" httpUri="https://zenodo.org/record/3785330/files/figure.png" pageId="21" pageNumber="22" startId="21.[189,232,1895,1921]" targetBox="[189,1399,548,1855]" targetPageId="21">
<paragraph id="8BAD369ABC5A1B4D9A84DF5001482EF2" blockId="21.[189,1399,1895,2027]" pageId="21" pageNumber="22">
<emphasis id="B966EA88BC5A1B4D9A84DF5002022E98" bold="true" box="[189,284,1895,1921]" pageId="21" pageNumber="22">Fig. 13.</emphasis>
<taxonomicName id="4C124D19BC5A1B4D9B1CDF5001252E98" authority="Smith &amp; Kamiya, 2015" authorityName="Smith &amp; Kamiya" authorityYear="2015" box="[293,571,1895,1921]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="falcula" status="sp. nov.">
<emphasis id="B966EA88BC5A1B4D9B1CDF5001252E98" box="[293,571,1895,1921]" italics="true" pageId="21" pageNumber="22">Pseudocandona falcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557F3BC5A1B4D987CDF5F01BC2E9B" box="[581,674,1896,1922]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B966EA88BC5A1B4D9892DF5001DE2E98" bold="true" box="[683,704,1895,1921]" pageId="21" pageNumber="22">A</emphasis>
. Male caudal ramus (LBM 1430006275).
<emphasis id="B966EA88BC5A1B4D9E86DF5007CD2E98" bold="true" box="[1215,1235,1895,1921]" pageId="21" pageNumber="22">B</emphasis>
. Male caudal ramus attachment (LBM 1430006275).
<emphasis id="B966EA88BC5A1B4D98AADFBD01B62EBD" bold="true" box="[659,680,1930,1956]" pageId="21" pageNumber="22">C</emphasis>
. Female caudal ramus and genital lobe (LBM 1430006277), black triangle indicates rounded lobe (see text for details).
<emphasis id="B966EA88BC5A1B4D9959DF99006B2ED1" bold="true" box="[864,885,1966,1992]" pageId="21" pageNumber="22">D</emphasis>
. Hemipenis (M-process shaded grey) (LBM 1430006275). Scale bar = 151 µm.
</paragraph>
</caption>
<paragraph id="8BAD369ABC591B4E9A84D93C07122881" blockId="22.[189,1399,267,409]" pageId="22" pageNumber="23">
Female genital lobe (
<figureCitation id="13292A1FBC591B4E9B82D93C0138283C" box="[443,550,267,293]" captionStart="Fig" captionStartId="21.[189,232,1895,1921]" captionTargetBox="[189,1399,548,1855]" captionTargetId="figure@21.[189,1399,544,1855]" captionTargetPageId="21" captionText="Fig. 13. Pseudocandona falcula sp. nov. A. Male caudal ramus (LBM 1430006275). B. Male caudal ramus attachment (LBM 1430006275). C. Female caudal ramus and genital lobe (LBM 1430006277), black triangle indicates rounded lobe (see text for details). D. Hemipenis (M-process shaded grey) (LBM 1430006275). Scale bar = 151 µm." figureDoi="http://doi.org/10.5281/zenodo.3785330" httpUri="https://zenodo.org/record/3785330/files/figure.png" pageId="22" pageNumber="23">Fig. 13C</figureCitation>
) with prominent projection with rounded apex. NB morphology noted before cover slip added. In material examined, lobes lightly sclerotized, with tendency to deform and break apart with addition of cover slip. In particular, rounded lobe (marked with triangle on
<figureCitation id="13292A1FBC591B4E9F3CD96F0673286B" box="[1285,1389,344,370]" captionStart="Fig" captionStartId="21.[189,232,1895,1921]" captionTargetBox="[189,1399,548,1855]" captionTargetId="figure@21.[189,1399,544,1855]" captionTargetPageId="21" captionText="Fig. 13. Pseudocandona falcula sp. nov. A. Male caudal ramus (LBM 1430006275). B. Male caudal ramus attachment (LBM 1430006275). C. Female caudal ramus and genital lobe (LBM 1430006277), black triangle indicates rounded lobe (see text for details). D. Hemipenis (M-process shaded grey) (LBM 1430006275). Scale bar = 151 µm." figureDoi="http://doi.org/10.5281/zenodo.3785330" httpUri="https://zenodo.org/record/3785330/files/figure.png" pageId="22" pageNumber="23">Fig. 13C</figureCitation>
) less prominent and more spread out laterally after addition of cover slip.
</paragraph>
</subSubSection>
<subSubSection id="C3086511BC591B4E9A84D9F400CD2DC1" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BAD369ABC591B4E9A84D9F4023128C4" blockId="22.[189,303,451,477]" box="[189,303,451,477]" pageId="22" pageNumber="23">
<heading id="D0E581F6BC591B4E9A84D9F4023128C4" bold="true" box="[189,303,451,477]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B966EA88BC591B4E9A84D9F4023128C4" bold="true" box="[189,303,451,477]" pageId="22" pageNumber="23">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD369ABC591B4E9A84D9C201272B9B" blockId="22.[189,1399,501,642]" pageId="22" pageNumber="23">
As noted above, the female genital lobe was noticeably deformed in shape in dissection slides after the cover slip had been lowered into position. The female genital lobe of candonids is widely used for the discrimination of species, but as noted by
<bibRefCitation id="EF834B6BBC591B4E98DBDA7607422B42" author="Namiotko T. &amp; Danielopol D. L." box="[738,1116,577,603]" pageId="22" pageNumber="23" pagination="109 - 125" refId="ref13537" refString="Namiotko T. &amp; Danielopol D. L. 2004. Review of the eremita species-group of the Pseudocandona Kaufmann (Ostracoda, Crustacea), with the description of a new species. Revista Espanola de Micropaleontologia 36: 109 - 125." type="journal article" year="2004">Namiotko &amp; Danielopol (2004)</bibRefCitation>
, this feature is prone to deformation in dissection slides.
</paragraph>
<paragraph id="8BAD369ABC591B4E9A84DA9B07A12AE0" blockId="22.[189,1399,684,1018]" pageId="22" pageNumber="23">
<taxonomicName id="4C124D19BC591B4E9A84DA9B02CF2BDF" authority="Smith &amp; Kamiya, 2015" authorityName="Smith &amp; Kamiya" authorityYear="2015" box="[189,465,684,710]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="falcula" status="sp. nov.">
<emphasis id="B966EA88BC591B4E9A84DA9B02CF2BDF" box="[189,465,684,710]" italics="true" pageId="22" pageNumber="23">Pseudocandona falcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557F3BC591B4E9BE3DA9A01282BDE" box="[474,566,685,711]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="4C124D19BC591B4E98E8DA9B078A2BDF" authority="(Hartwig, 1899)" authorityName="in Delorme" authorityYear="1970" baseAuthorityName="Hartwig" baseAuthorityYear="1899" box="[721,1172,684,710]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sarsi">
<emphasis id="B966EA88BC591B4E98E8DA9B00D02BDF" box="[721,974,684,710]" italics="true" pageId="22" pageNumber="23">Pseudocandona sarsi</emphasis>
(Hartwig, 1899)
</taxonomicName>
in the shape of the carapace in lateral view, although the dorsal view of
<taxonomicName id="4C124D19BC591B4E9923DAE300712BF4" authorityName="in Delorme" authorityYear="1970" baseAuthorityName="Hartwig" baseAuthorityYear="1899" box="[794,879,724,749]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sarsi">
<emphasis id="B966EA88BC591B4E9923DAE300712BF4" box="[794,879,724,749]" italics="true" pageId="22" pageNumber="23">P. sarsi</emphasis>
</taxonomicName>
is slightly wider and more pointed anteriorly and posteriorly. The male fifth limb palps are similar in both species, but the right palp of
<taxonomicName id="4C124D19BC591B4E9E82DACE02102A20" authority="Smith &amp; Kamiya, 2015" authorityName="Smith &amp; Kamiya" authorityYear="2015" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="falcula" status="sp. nov.">
<emphasis id="B966EA88BC591B4E9E82DACE02102A20" italics="true" pageId="22" pageNumber="23">Pseudocandona falcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557F3BC591B4E9B2FDB17026F2A23" box="[278,369,800,826]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
is more rounded on the outer edge than that of
<taxonomicName id="4C124D19BC591B4E99A7DB1700EA2A20" authorityName="in Delorme" authorityYear="1970" baseAuthorityName="Hartwig" baseAuthorityYear="1899" box="[926,1012,800,825]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sarsi">
<emphasis id="B966EA88BC591B4E99A7DB1700EA2A20" box="[926,1012,800,825]" italics="true" pageId="22" pageNumber="23">P. sarsi</emphasis>
</taxonomicName>
(which has a more angular outer edge). The seventh limbs of the two species are also similar, both with a short reflexed h1 seta on the final segment. Differences in the hemipenes include a larger outer lobe (a) in
<taxonomicName id="4C124D19BC591B4E9E59DB5B06682A9F" authority="Smith &amp; Kamiya, 2015" authorityName="Smith &amp; Kamiya" authorityYear="2015" box="[1120,1398,876,902]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="falcula" status="sp. nov.">
<emphasis id="B966EA88BC591B4E9E59DB5B06682A9F" box="[1120,1398,876,902]" italics="true" pageId="22" pageNumber="23">Pseudocandona falcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557F3BC591B4E9A84DBA402042AB4" box="[189,282,915,941]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
, which clearly extends beyond the distal edge of the medial lobe (h). In
<taxonomicName id="4C124D19BC591B4E9E59DBA407AB2AB5" authorityName="in Delorme" authorityYear="1970" baseAuthorityName="Hartwig" baseAuthorityYear="1899" box="[1120,1205,915,940]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sarsi">
<emphasis id="B966EA88BC591B4E9E59DBA407AB2AB5" box="[1120,1205,915,940]" italics="true" pageId="22" pageNumber="23">P. sarsi</emphasis>
</taxonomicName>
the outer lobe is more offset to the outer edge of the hemipenes, has a more angular outer edge, and only slightly extends beyond the medial lobe. The M-processes of the two species are also different in shape.
</paragraph>
<paragraph id="8BAD369ABC591B4E9A84DC1300CD2DC1" blockId="22.[189,1399,1060,1240]" pageId="22" pageNumber="23">
The females of this species closely resemble the female specimens of
<taxonomicName id="4C124D19BC591B4E99C9DC1207582D27" authorityName="in Delorme" authorityYear="1970" baseAuthorityName="Hartwig" baseAuthorityYear="1899" box="[1008,1094,1061,1086]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sarsi">
<emphasis id="B966EA88BC591B4E99C9DC1207582D27" box="[1008,1094,1061,1086]" italics="true" pageId="22" pageNumber="23">P. sarsi</emphasis>
</taxonomicName>
reported from Lake Biwa (locality 8 on
<figureCitation id="13292A1FBC591B4E9B66DC7D02BF2D7D" box="[351,417,1098,1124]" captionStart="Fig" captionStartId="1.[189,232,2000,2026]" captionTargetBox="[189,1399,737,1960]" captionTargetId="figure@1.[190,1397,739,1959]" captionTargetPageId="1" captionText="Fig. 1. Map of the main islands of Japan, showing the localities mentioned in the text." figureDoi="http://doi.org/10.5281/zenodo.3785304" httpUri="https://zenodo.org/record/3785304/files/figure.png" pageId="22" pageNumber="23">Fig. 1</figureCitation>
) by
<bibRefCitation id="EF834B6BBC591B4E9BEEDC7D01D02D7D" author="Smith R. J. &amp; Janz H." box="[471,718,1098,1125]" pageId="22" pageNumber="23" pagination="2865 - 2922" refId="ref13871" refString="Smith R. J. &amp; Janz H. 2008. Recent species of the Family Candonidae (Ostracoda, Crustacea) from the ancient Lake Biwa, Central Japan. Journal of Natural History 42 (45 - 46): 2865 - 2922. http: // dx. doi. org / 10.1080 / 00222930802361030" type="journal article" year="2008">Smith &amp; Janz (2008)</bibRefCitation>
. As no males were found from Lake Biwa, Smith &amp; Janzs (
<emphasis id="B966EA88BC591B4E9AFEDC4602062D93" box="[199,280,1137,1162]" italics="true" pageId="22" pageNumber="23">op. cit.</emphasis>
) identification was tentative. Further investigation indicates that the carapaces and appendages of females from Lake Biwa very closely match material from the
<typeStatus id="54A98838BC591B4E99FCDCA000E62DA8" box="[965,1016,1175,1201]" pageId="22" pageNumber="23">type</typeStatus>
locality, and it is concluded that the specimens from Lake Biwa are
<taxonomicName id="4C124D19BC591B4E9865DC8A006E2DCE" authority="Smith &amp; Kamiya, 2015" authorityName="Smith &amp; Kamiya" authorityYear="2015" box="[604,880,1213,1239]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="falcula" status="sp. nov.">
<emphasis id="B966EA88BC591B4E9865DC8A006E2DCE" box="[604,880,1213,1239]" italics="true" pageId="22" pageNumber="23">Pseudocandona falcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557F3BC591B4E9941DC8900CD2DC1" box="[888,979,1214,1240]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3086511BC591B4E9A84DD3502E92F2A" pageId="22" pageNumber="23" type="biology_ecology">
<paragraph id="8BAD369ABC591B4E9A84DD3502EC2C05" blockId="22.[189,498,1282,1308]" box="[189,498,1282,1308]" pageId="22" pageNumber="23">
<heading id="D0E581F6BC591B4E9A84DD3502EC2C05" bold="true" box="[189,498,1282,1308]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B966EA88BC591B4E9A84DD3502EC2C05" bold="true" box="[189,498,1282,1308]" pageId="22" pageNumber="23">Ecology and distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD369ABC591B4E9A84DD0402E92F2A" blockId="22.[189,1399,1331,1588]" pageId="22" pageNumber="23">
The type locality of this species is the same as for
<taxonomicName id="4C124D19BC591B4E9960DD04076D2C54" authority="Smith &amp; Kamiya, 2015" authorityName="Smith &amp; Kamiya" authorityYear="2015" box="[857,1139,1331,1357]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="atmeta" status="sp. nov.">
<emphasis id="B966EA88BC591B4E9960DD04076D2C54" box="[857,1139,1331,1357]" italics="true" pageId="22" pageNumber="23">Pseudocandona atmeta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557F3BC591B4E9EB8DD0307FD2C57" box="[1153,1251,1332,1358]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
(see below
<taxonomicName id="4C124D19BC591B4E9A84DD6D02CE2C6D" authority="Smith &amp; Kamiya, 2015" authorityName="Smith &amp; Kamiya" authorityYear="2015" box="[189,464,1370,1396]" class="Ostracoda" family="Candonidae" genus="Pseudocandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="atmeta" status="sp. nov.">
<emphasis id="B966EA88BC591B4E9A84DD6D02CE2C6D" box="[189,464,1370,1396]" italics="true" pageId="22" pageNumber="23">Pseudocandona atmeta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557F3BC591B4E9BEEDD6D012C2C6D" box="[471,562,1370,1396]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
for a description of that site). In Lake Biwa, the specimens were found at the waters edge in the root mat of a willow tree (
<bibRefCitation id="EF834B6BBC591B4E9918DDB707152C83" author="Smith R. J. &amp; Janz H." box="[801,1035,1408,1435]" pageId="22" pageNumber="23" pagination="2865 - 2922" refId="ref13871" refString="Smith R. J. &amp; Janz H. 2008. Recent species of the Family Candonidae (Ostracoda, Crustacea) from the ancient Lake Biwa, Central Japan. Journal of Natural History 42 (45 - 46): 2865 - 2922. http: // dx. doi. org / 10.1080 / 00222930802361030" type="journal article" year="2008">Smith &amp; Janz 2008</bibRefCitation>
). To date, only a small number of females have been collected from Lake Biwa, in samples collected during
<date id="FFAC105ABC591B4E9E69DD9007D72CD8" box="[1104,1225,1446,1473]" pageId="22" pageNumber="23" value="2005-04">Apr. 2005</date>
. It hasnt been collected since this time despite numerous samples taken at the same locality, and this may indicate that it was a temporary occurrence or it is rare in Lake Biwa. So far, this species is only known from Lake Biwa and the
<typeStatus id="54A98838BC591B4E9B66DE2D028C2F2D" box="[351,402,1562,1588]" pageId="22" pageNumber="23">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>