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<document id="3495E295D609FAEC075CCAAF1A2F4089" ID-DOI="10.5281/zenodo.199403" ID-GBIF-Dataset="78e60046-26ff-4482-a1a7-924b76a8fc2d" ID-ISSN="1175-5326" ID-Zenodo-Dep="199403" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460335535356" checkinUser="plazi" docAuthor="Jr, Robert Murray Lasley, Mendoza, Jose Christopher E. &amp; Ng, Peter K. L." docDate="2010" docId="03B987ECFFEC3D2D70CBCAECFCB32AC8" docLanguage="en" docName="zt02684p013.pdf" docOrigin="Zootaxa 2684" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Guitonia troglophila Garth &amp; Iliffe 1992" docType="treatment" docVersion="8" lastPageNumber="4" masterDocId="FF80FF94FFEF3D2E705CCB4EFFA22E0C" masterDocTitle="Two new species of Guitonia Garth &amp; Iliffe, 1992 (Crustacea: Decapoda: Brachyura: Xanthidae) from the central and western Pacific Ocean" masterLastPageNumber="13" masterPageNumber="1" pageNumber="4" updateTime="1698259701857" updateUser="plazi">
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<mods:title id="A42C684120A6A089688F110AD1756150">Two new species of Guitonia Garth &amp; Iliffe, 1992 (Crustacea: Decapoda: Brachyura: Xanthidae) from the central and western Pacific Ocean</mods:title>
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<mods:namePart id="A59710299C866A108782B32CC191D35A">Jr, Robert Murray Lasley</mods:namePart>
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<mods:namePart id="00AD74D0B903E601A33A679C8D1535D8">Mendoza, Jose Christopher E.</mods:namePart>
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<mods:namePart id="6A692E1318D5590F82782DCE5ADADEA9">Ng, Peter K. L.</mods:namePart>
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<treatment id="03B987ECFFEC3D2D70CBCAECFCB32AC8" ID-DOI="http://doi.org/10.5281/zenodo.6209913" ID-GBIF-Taxon="119405245" ID-Zenodo-Dep="6209913" LSID="urn:lsid:plazi:treatment:03B987ECFFEC3D2D70CBCAECFCB32AC8" httpUri="http://treatment.plazi.org/id/03B987ECFFEC3D2D70CBCAECFCB32AC8" lastPageNumber="4" pageId="3" pageNumber="4">
<subSubSection id="C30A6571FFEC3D2D70CBCAECFED92FD3" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BAF36FAFFEC3D2D70CBCAECFD2D2FB0" blockId="3.[151,655,418,479]" box="[151,655,418,444]" pageId="3" pageNumber="4">
<heading id="D0E78196FFEC3D2D70CBCAECFD2D2FB0" bold="true" box="[151,655,418,444]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<taxonomicName id="4C104D79FFEC3D2D70CBCAECFD2D2FB0" ID-CoL="3HLFY" authority="Garth &amp; Iliffe, 1992" authorityName="Garth &amp; Iliffe" authorityYear="1992" box="[151,655,418,444]" class="Malacostraca" family="Xanthidae" genus="Guitonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="troglophila">
<emphasis id="B964EAE8FFEC3D2D70CBCAECFD2D2FB0" bold="true" box="[151,655,418,444]" pageId="3" pageNumber="4">
<emphasis id="B964EAE8FFEC3D2D70CBCAECFE2D2FB0" bold="true" box="[151,399,418,444]" italics="true" pageId="3" pageNumber="4">Guitonia troglophila</emphasis>
<bibRefCitation id="EF814B0BFFEC3D2D71CACAECFD2D2FB0" author="Garth" box="[406,655,418,444]" pageId="3" pageNumber="4" refString="Garth, J. S. &amp; Iliffe, T. M. (1992) Guitonia troglophila, a new genus and species of anchialine crab from the Galapagos, Isla Santa Cruz, Grieta de Caleta la Torta (Crustacea: Decapoda: Brachyura). Proceedings of the Biological Society of Washington, 105 (2), 310 - 316." type="journal article" year="1992">Garth &amp; Iliffe, 1992</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAF36FAFFEC3D2D70CBCA8BFED92FD3" blockId="3.[151,655,418,479]" box="[151,379,453,479]" pageId="3" pageNumber="4">
(
<figureCitation id="132B2A7FFFEC3D2D70C3CA8BFF4E2FD3" box="[159,236,453,479]" captionStart="FIGURE 1" captionStartId="4.[151,255,1576,1600]" captionTargetBox="[151,1436,271,1551]" captionTargetId="figure@4.[151,1436,271,1552]" captionTargetPageId="4" captionText="FIGURE 1. Guitonia leimomi n. sp., holotype male, 15.5 × 9.7 mm (ZRC 2010.0277), Hawaiian Is., A, dorsal view, B, frontal view; Guitonia paulayi n. sp., holotype male, 11.2 × 7.3 mm (ZRC 2010.0278), Guam, C, dorsal view, D, frontal view; Guitonia troglophila Garth &amp; Iliffe, 1992, holotype female, 14.2 × 9.7 mm (LACM CR 19871671), Galápagos, E, dorsal view, F, frontal view." httpUri="https://zenodo.org/record/199404/files/figure.png" pageId="3" pageNumber="4">Figs. 1</figureCitation>
E, F, 3G, H)
</paragraph>
</subSubSection>
<subSubSection id="C30A6571FFEC3D2D70CBC95EFB8F2C2B" box="[151,1069,528,551]" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="8BAF36FAFFEC3D2D70CBC95EFB8F2C2B" blockId="3.[151,1069,528,551]" box="[151,1069,528,551]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="AB0011D4FFEC3D2D70CBC95EFB8F2C2B" box="[151,1069,528,551]" pageId="3" pageNumber="4">
<taxonomicName id="4C104D79FFEC3D2D70CBC95EFD252C2B" ID-CoL="3HLFY" authority="Garth &amp; Iliffe, 1992: 311" authorityName="Garth &amp; Iliffe" authorityPageNumber="311" authorityYear="1992" box="[151,647,528,551]" class="Malacostraca" family="Xanthidae" genus="Guitonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="troglophila">
<emphasis id="B964EAE8FFEC3D2D70CBC95EFED12C2B" box="[151,371,528,551]" italics="true" pageId="3" pageNumber="4">Guitonia troglophila</emphasis>
<treatmentCitation id="0AB110EBFFEC3D2D7126C95EFD252C2B" author="Garth" box="[378,647,528,551]" page="311" pageId="3" pageNumber="4" year="1992">
<bibRefCitation id="EF814B0BFFEC3D2D7126C95EFDF62C2B" author="Garth" box="[378,596,528,551]" pageId="3" pageNumber="4" refString="Garth, J. S. &amp; Iliffe, T. M. (1992) Guitonia troglophila, a new genus and species of anchialine crab from the Galapagos, Isla Santa Cruz, Grieta de Caleta la Torta (Crustacea: Decapoda: Brachyura). Proceedings of the Biological Society of Washington, 105 (2), 310 - 316." type="journal article" year="1992">Garth &amp; Iliffe, 1992</bibRefCitation>
: 311
</treatmentCitation>
</taxonomicName>
, figs. 1, 2. — Ng
<emphasis id="B964EAE8FFEC3D2D7319C95EFCD92C2B" box="[837,891,528,551]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
2008: 202 (list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30A6571FFEC3D2D70CBC91BFBAF2C98" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="8BAF36FAFFEC3D2D70CBC91BFBAF2C98" blockId="3.[151,1437,597,1220]" pageId="3" pageNumber="4">
<emphasis id="B964EAE8FFEC3D2D70CBC91BFE322C63" bold="true" box="[151,400,597,623]" pageId="3" pageNumber="4">Material examined.</emphasis>
<typeStatus id="54AB8858FFEC3D2D71C5C91BFDAA2C63" box="[409,520,597,623]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
female, 14.2 × 9.7 mm (
<collectionCode id="ED01AE3FFFEC3D2D736FC91BFC292C63" LSID="urn:lsid:biocol.org:col:34802" box="[819,907,597,623]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="3" pageNumber="4">LACM</collectionCode>
CR 1987.167.1), anchialine cave, Grieta de Caleta la Torta, Santa Cruz Is., Galápagos Is., coll. T. M. Iliffe,
<date id="FFAE103AFFEC3D2D7322C934FBAB2C98" box="[894,1033,634,660]" pageId="3" pageNumber="4" value="1987-06-08">8 Jun. 1987</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A6571FFEC3D2D709AC9D1FC7F2D96" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BAF36FAFFEC3D2D709AC9D1FC7F2D96" blockId="3.[151,1437,597,1220]" pageId="3" pageNumber="4">
<emphasis id="B964EAE8FFEC3D2D709AC9D1FEE42CB5" bold="true" box="[198,326,671,697]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
Anterolateral margin with teeth broad, lobate. Anterior regions of carapace with slight transverse rows of granules, almost indiscernible, not basally conjoined. Corneas, eyestalks reduced. Area lateral to the orbits with 2 granular ridges linking at level of exorbital angle, first anterolateral tooth, enclosing small, depressed area. Frontal margin angled toward posterior laterally, medial notch relatively large. Chelae unequal, dimorphic; dactylus of major chela with large basal tooth; outer surface of fixed finger, dactylus of minor chela granulated. Ambulatory legs relatively long, slender; distal end of merus of fourth ambulatory leg almost reaching last anterolateral tooth when apposed against carapace.
</paragraph>
</subSubSection>
<subSubSection id="C30A6571FFEC3D2D709AC8EBFAB62A93" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8BAF36FAFFEC3D2D709AC8EBFAB62A93" blockId="3.[151,1437,597,1220]" pageId="3" pageNumber="4">
<emphasis id="B964EAE8FFEC3D2D709AC8EBFEE22DB3" bold="true" box="[198,320,933,959]" pageId="3" pageNumber="4">Remarks.</emphasis>
It is noteworthy to note that in the original description of
<taxonomicName id="4C104D79FFEC3D2D73B3C8EBFA3E2DB3" authority="Garth &amp; Iliffe (1992: 310)" authorityName="Garth &amp; Iliffe" authorityPageNumber="310" authorityYear="1992" box="[1007,1436,933,959]" class="Malacostraca" family="Xanthidae" genus="Guitonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAE8FFEC3D2D73B3C8EBFBF52DB2" box="[1007,1111,933,958]" italics="true" pageId="3" pageNumber="4">Guitonia</emphasis>
,
<bibRefCitation id="EF814B0BFFEC3D2D743AC8EBFA3E2DB3" author="Garth" box="[1126,1436,933,959]" pageId="3" pageNumber="4" refString="Garth, J. S. &amp; Iliffe, T. M. (1992) Guitonia troglophila, a new genus and species of anchialine crab from the Galapagos, Isla Santa Cruz, Grieta de Caleta la Torta (Crustacea: Decapoda: Brachyura). Proceedings of the Biological Society of Washington, 105 (2), 310 - 316." type="journal article" year="1992">Garth &amp; Iliffe (1992: 310)</bibRefCitation>
</taxonomicName>
attribute a unique character to the genus: “Orbits failing to close externally at level of postorbital (first anterolateral) tooth, instead communicating by means of shallow trough or gutter with second anterolateral tooth.” Indeed, the figures (
<bibRefCitation id="EF814B0BFFEC3D2D7182CF5BFC2E2A23" author="Garth" box="[478,908,1045,1071]" pageId="3" pageNumber="4" refString="Garth, J. S. &amp; Iliffe, T. M. (1992) Guitonia troglophila, a new genus and species of anchialine crab from the Galapagos, Isla Santa Cruz, Grieta de Caleta la Torta (Crustacea: Decapoda: Brachyura). Proceedings of the Biological Society of Washington, 105 (2), 310 - 316." type="journal article" year="1992">Garth &amp; Iliffe 1992: 312, fig. 1B, C</bibRefCitation>
) show this character in
<taxonomicName id="4C104D79FFEC3D2D74F5CF5BFAEC2A22" box="[1193,1358,1045,1070]" class="Malacostraca" family="Xanthidae" genus="Guitonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="troglophila">
<emphasis id="B964EAE8FFEC3D2D74F5CF5BFAEC2A22" box="[1193,1358,1045,1070]" italics="true" pageId="3" pageNumber="4">G. troglophila</emphasis>
</taxonomicName>
. A reexamination of the
<typeStatus id="54AB8858FFEC3D2D7125CF74FE0E2A58" box="[377,428,1082,1108]" pageId="3" pageNumber="4">type</typeStatus>
specimen of
<taxonomicName id="4C104D79FFEC3D2D7214CF74FD482A5F" box="[584,746,1082,1107]" class="Malacostraca" family="Xanthidae" genus="Guitonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="troglophila">
<emphasis id="B964EAE8FFEC3D2D7214CF74FD482A5F" box="[584,746,1082,1107]" italics="true" pageId="3" pageNumber="4">G. troglophila</emphasis>
</taxonomicName>
, however, shows that this is not the case, and the orbits are typical of most xanthids (
<figureCitation id="132B2A7FFFEC3D2D719DCF2EFDA12A76" box="[449,515,1120,1146]" captionStart="FIGURE 1" captionStartId="4.[151,255,1576,1600]" captionTargetBox="[151,1436,271,1551]" captionTargetId="figure@4.[151,1436,271,1552]" captionTargetPageId="4" captionText="FIGURE 1. Guitonia leimomi n. sp., holotype male, 15.5 × 9.7 mm (ZRC 2010.0277), Hawaiian Is., A, dorsal view, B, frontal view; Guitonia paulayi n. sp., holotype male, 11.2 × 7.3 mm (ZRC 2010.0278), Guam, C, dorsal view, D, frontal view; Guitonia troglophila Garth &amp; Iliffe, 1992, holotype female, 14.2 × 9.7 mm (LACM CR 19871671), Galápagos, E, dorsal view, F, frontal view." httpUri="https://zenodo.org/record/199404/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
F). The original descriptions and figures are therefore incorrect in this respect. The orbital structures of
<taxonomicName id="4C104D79FFEC3D2D71E4CFCBFD932A92" box="[440,561,1157,1182]" class="Malacostraca" family="Xanthidae" genus="Guitonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="paulayi" status="sp. nov.">
<emphasis id="B964EAE8FFEC3D2D71E4CFCBFD932A92" box="[440,561,1157,1182]" italics="true" pageId="3" pageNumber="4">G. paulayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2575793FFEC3D2D7264CFCBFDD82A93" box="[568,634,1157,1183]" pageId="3" pageNumber="4" rank="species">
<emphasis id="B964EAE8FFEC3D2D7264CFCBFDD82A93" bold="true" box="[568,634,1157,1183]" pageId="3" pageNumber="4">n. sp.</emphasis>
</taxonomicNameLabel>
, and
<taxonomicName id="4C104D79FFEC3D2D72E6CFCBFC9B2A92" box="[698,825,1157,1182]" class="Malacostraca" family="Xanthidae" genus="Guitonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="leimomi" status="sp. nov.">
<emphasis id="B964EAE8FFEC3D2D72E6CFCBFC9B2A92" box="[698,825,1157,1182]" italics="true" pageId="3" pageNumber="4">G. leimomi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2575793FFEC3D2D731CCFCBFC202A93" box="[832,898,1157,1183]" pageId="3" pageNumber="4" rank="species">
<emphasis id="B964EAE8FFEC3D2D731CCFCBFC202A93" bold="true" box="[832,898,1157,1183]" pageId="3" pageNumber="4">n. sp.</emphasis>
</taxonomicNameLabel>
, are also typical of most xanthids.
</paragraph>
</subSubSection>
<subSubSection id="C30A6571FFEC3D2D709ACFE4FCB32AC8" box="[198,785,1194,1220]" pageId="3" pageNumber="4" type="distribution">
<paragraph id="8BAF36FAFFEC3D2D709ACFE4FCB32AC8" blockId="3.[151,1437,597,1220]" box="[198,785,1194,1220]" pageId="3" pageNumber="4">
<emphasis id="B964EAE8FFEC3D2D709ACFE4FEC52AC8" bold="true" box="[198,359,1194,1220]" pageId="3" pageNumber="4">Distribution.</emphasis>
Known only from the Galápagos Is.
</paragraph>
</subSubSection>
</treatment>
</document>