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<document id="D6B5AADF5BD64882C77776D9DD1FC7B4" ID-DOI="10.11646/phytotaxa.247.1.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="13678782" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725411541331" checkinUser="felipe" docAuthor="Zeng, Si-Jin, Zou, Long-Hai, Wang, Pan, Hong, Wen-Jun, Zhang, Guo-Qiang, Chen, Li-Jun &amp; Zhuang, Xue-Ying" docDate="2016" docId="03B1CD73FFB3EF06FD80F95D81D3D4F0" docLanguage="en" docName="phytotaxa.247.1.3.pdf" docOrigin="Phytotaxa 247 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.247.1.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Fordiophyton chenii S. Jin Zeng &amp; X. Y. Zhuang. A 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="57" masterDocId="FF88B50BFFBAEF0AFD08FFA98102D75D" masterDocTitle="Preliminary phylogeny of Fordiophyton (Melastomataceae), with the description of two new species" masterLastPageNumber="61" masterPageNumber="45" pageNumber="54" updateTime="1725431212836" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FC4B40804FF36ABD7DFC20A10500F274">Preliminary phylogeny of Fordiophyton (Melastomataceae), with the description of two new species</mods:title>
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<taxonomicName id="4C1807E6FFB3EF03FD80F95D83B5D052" ID-CoL="6JHG4" authority="S. Jin Zeng &amp; X. Y. Zhuang" authorityName="S. Jin Zeng &amp; X. Y. Zhuang. A" authorityYear="2016" box="[136,695,1780,1807]" class="Magnoliopsida" family="Melastomataceae" genus="Fordiophyton" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="54" phylum="Tracheophyta" rank="species" species="chenii" status="sp. nov.">
<emphasis id="B96CA077FFB3EF03FD80F95D8074D052" bold="true" box="[136,374,1780,1807]" italics="true" pageId="9" pageNumber="54">Fordiophyton chenii</emphasis>
S. Jin Zeng &amp; X. Y. Zhuang
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,
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(ṪKṖÜË), (
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,
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)
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.
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(
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(ĭ‼),
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(
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), on damp slopes of ravines in broad-leaved forests,
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. Jin Zeng et al.
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(
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;
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,
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, HK,
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,
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)
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.
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<paragraph id="8BA77C65FFB3EF03FD80F8218259D09C" blockId="9.[136,1452,1854,1985]" pageId="9" pageNumber="54">
Diagnosis: The new species
<taxonomicName id="4C1807E6FFB3EF03FC9EF821835BD0C0" authorityName="S. Jin Zeng &amp; X. Y. Zhuang. A" authorityYear="2016" box="[406,601,1928,1949]" class="Magnoliopsida" family="Melastomataceae" genus="Fordiophyton" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="54" phylum="Tracheophyta" rank="species" species="chenii">
<emphasis id="B96CA077FFB3EF03FC9EF821835BD0C0" box="[406,601,1928,1949]" italics="true" pageId="9" pageNumber="54">Fordiophyton chenii</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C1807E6FFB3EF03FFCBF821824BD0C0" box="[707,841,1928,1949]" class="Magnoliopsida" family="Melastomataceae" genus="Fordiophyton" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="54" phylum="Tracheophyta" rank="species" species="cordifolium">
<emphasis id="B96CA077FFB3EF03FFCBF821824BD0C0" box="[707,841,1928,1949]" italics="true" pageId="9" pageNumber="54">F. cordifolium</emphasis>
</taxonomicName>
but differs in its very short stems, rosette leaves, hypanthia with densely glandular trichomes and shorter stamens with yellow anthers.
</paragraph>
<caption id="DF672CEDFFB0EF00FD80F8088272D0A3" ID-DOI="http://doi.org/10.5281/zenodo.13678810" ID-Zenodo-Dep="13678810" httpUri="https://zenodo.org/record/13678810/files/figure.png" pageId="10" pageNumber="55" startId="10.[136,229,1953,1974]" targetBox="[194,1405,217,1904]" targetPageId="10" targetType="figure">
<paragraph id="8BA77C65FFB0EF00FD80F8088272D0A3" blockId="10.[136,1452,1953,2046]" pageId="10" pageNumber="55">
<emphasis id="B96CA077FFB0EF00FD80F80881FDD0EB" bold="true" box="[136,255,1953,1974]" pageId="10" pageNumber="55">FIGURE 7.</emphasis>
Drawing of
<taxonomicName id="4C1807E6FFB0EF00FC75F808827DD0EB" authority="S. Jin Zeng &amp; X. Y. Zhuang. A" authorityName="S. Jin Zeng &amp; X. Y. Zhuang. A" authorityYear="2016" box="[381,895,1953,1974]" class="Magnoliopsida" family="Melastomataceae" genus="Fordiophyton" kingdom="Plantae" order="Myrtales" pageId="10" pageNumber="55" phylum="Tracheophyta" rank="species" species="chenii">
<emphasis id="B96CA077FFB0EF00FC75F8088341D0EB" box="[381,579,1953,1974]" italics="true" pageId="10" pageNumber="55">Fordiophyton chenii</emphasis>
S. Jin Zeng &amp; X. Y. Zhuang. A
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: flowering plant; B: flower bud, front view. C: pistil and stamens, side view; D: flower, front view; E: petal; F: longer stamens, front view and back view; G: shorter stamens, front view and back view; H: ovary, vertical view and transverse view. Drawn by Pei-Wen Zhang.
</paragraph>
</caption>
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<paragraph id="8BA77C65FFB1EF01FD80F8088220D087" blockId="11.[136,1452,1953,2010]" pageId="11" pageNumber="56">
<emphasis id="B96CA077FFB1EF01FD80F80881FCD0EB" bold="true" box="[136,254,1953,1974]" pageId="11" pageNumber="56">FIGURE 8.</emphasis>
Comparison of
<taxonomicName id="4C1807E6FFB1EF01FC92F808835CD0EB" authorityName="S. Jin Zeng &amp; X. Y. Zhuang. A" authorityYear="2016" box="[410,606,1953,1974]" class="Magnoliopsida" family="Melastomataceae" genus="Fordiophyton" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="56" phylum="Tracheophyta" rank="species" species="chenii">
<emphasis id="B96CA077FFB1EF01FC92F808835CD0EB" box="[410,606,1953,1974]" italics="true" pageId="11" pageNumber="56">Fordiophyton chenii</emphasis>
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and
<taxonomicName id="4C1807E6FFB1EF01FF85F8088216D0EB" box="[653,788,1953,1974]" class="Magnoliopsida" family="Melastomataceae" genus="Fordiophyton" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="56" phylum="Tracheophyta" rank="species" species="cordifolium">
<emphasis id="B96CA077FFB1EF01FF85F8088216D0EB" box="[653,788,1953,1974]" italics="true" pageId="11" pageNumber="56">F. cordifolium</emphasis>
</taxonomicName>
. AE:
<taxonomicName id="4C1807E6FFB1EF01FE5FF80882A8D0EB" authorityName="S. Jin Zeng &amp; X. Y. Zhuang. A" authorityYear="2016" box="[855,938,1953,1974]" class="Magnoliopsida" family="Melastomataceae" genus="Fordiophyton" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="56" phylum="Tracheophyta" rank="species" species="chenii">
<emphasis id="B96CA077FFB1EF01FE5FF80882A8D0EB" box="[855,938,1953,1974]" italics="true" pageId="11" pageNumber="56">F. chenii</emphasis>
</taxonomicName>
; FI:
<taxonomicName id="4C1807E6FFB1EF01FEEDF808856ED0EB" box="[997,1132,1953,1974]" class="Magnoliopsida" family="Melastomataceae" genus="Fordiophyton" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="56" phylum="Tracheophyta" rank="species" species="cordifolium">
<emphasis id="B96CA077FFB1EF01FEEDF808856ED0EB" box="[997,1132,1953,1974]" italics="true" pageId="11" pageNumber="56">F. cordifolium</emphasis>
</taxonomicName>
; A, F: the plants
<emphasis id="B96CA077FFB1EF01F81AF808844CD0EB" box="[1298,1358,1953,1974]" italics="true" pageId="11" pageNumber="56">in situ</emphasis>
; B, C, G: inflorescences; D, H: flower buds; E, I: an opening flower, side view.
</paragraph>
</caption>
<paragraph id="8BA77C65FFB6EF06FD80FF0983B5D5EC" blockId="12.[136,1452,160,941]" pageId="12" pageNumber="57">
<collectionCode id="ED09E4A0FFB6EF06FD80FF098199D7E4" box="[136,155,160,185]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="12" pageNumber="57" type="Herbarium">A</collectionCode>
perennial herb, succulent, with raphides both in vegetative and floral parts (
<figureCitation id="132360E0FFB6EF06FEF8FF098537D7E4" box="[1008,1077,160,185]" captionStart="FIGURE 5" captionStartId="7.[136,229,1983,2004]" captionTargetBox="[151,1436,996,1958]" captionTargetId="figure-267@7.[151,1436,996,1958]" captionTargetPageId="7" captionText="FIGURE 5. Raphides in the leaf, peduncle and ovary. AC: Fordiophyton chenii; D: F. peperomiifolium. A: leaf blade; B: peduncle; C: ovary; D: petiole. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13678804" httpUri="https://zenodo.org/record/13678804/files/figure.png" pageId="12" pageNumber="57">Fig. 5</figureCitation>
,
<collectionCode id="ED09E4A0FFB6EF06F937FF098557D7E4" box="[1087,1109,160,185]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="12" pageNumber="57" type="Herbarium">A</collectionCode>
<collectionCode id="ED09E4A0FFB6EF06F96DFF09857AD7E4" box="[1125,1144,160,185]" country="Denmark" name="University of Copenhagen" pageId="12" pageNumber="57" type="Herbarium">C</collectionCode>
). Stem stoloniferous,
<quantity id="4CE0D180FFB6EF06F865FF0981AFD780" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="3.0" metricValueMin="1.6" pageId="12" pageNumber="57" unit="cm" value="2.3" valueMax="3.0" valueMin="1.6">1.63 cm</quantity>
, internodes not distinct. Leaves in a basal or sub-basal rosette; petiole
<quantity id="4CE0D180FFB6EF06FEEDFF6D854BD780" box="[997,1097,196,221]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="1.8" metricValueMin="0.8" pageId="12" pageNumber="57" unit="cm" value="13.0" valueMax="18.0" valueMin="8.0">818 cm</quantity>
long, 4-sided, narrow-winged, sparsely glandular trichomes when young, glabrescent later; leaf blade ovate or ovate-elliptic, 913 ×
<quantity id="4CE0D180FFB6EF06F801FF41846CD65C" box="[1289,1390,232,257]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="12" pageNumber="57" unit="cm" value="10.5" valueMax="12.0" valueMin="9.0">912 cm</quantity>
, base cordate, apex acute, margin entire, membranaceous or slightly fleshy, glabrous; secondary veins 4 on each side of midvein, conspicuous; tertiary veins numerous, parallel, and connecting secondary veins; glabrous on both surfaces; abaxial veins typically purple. Inflorescences terminal, umbellate panicles, each branch 512-flowered, subsessile; peduncle
<quantity id="4CE0D180FFB6EF06FDFCFED1806ED6CC" box="[244,364,376,401]" metricMagnitude="-1" metricUnit="m" metricValue="1.625" metricValueMax="2.45" metricValueMin="0.8" pageId="12" pageNumber="57" unit="cm" value="16.25" valueMax="24.5" valueMin="8.0">824.5 cm</quantity>
, hirsute on nodes, but glabrescent later; bracts oblong, with dense glandular trichomes, caducous. Hypanthia funnel-shaped,
<quantity id="4CE0D180FFB6EF06FCBBFE358332D6E8" box="[435,560,412,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="12" pageNumber="57" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">1012 mm</quantity>
, with dense glandular trichomes; calyx lobes ovate-triangle, 5 ×
<quantity id="4CE0D180FFB6EF06F81BFE358470D6E8" box="[1299,1394,412,437]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="12" pageNumber="57" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
with dense glandular trichomes. Petals pink or purplish, oblong, ca. 10 ×
<quantity id="4CE0D180FFB6EF06FE74FE6982BCD684" box="[892,958,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="57" unit="mm" value="8.0">8 mm</quantity>
, apex pinkish and with 1 glandular trichome at tip. Stamens 8, 4 long and 4 short arranged in 2 whorls. Longer stamens ca.
<quantity id="4CE0D180FFB6EF06F902FE4D855FD6A0" box="[1034,1117,484,509]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="57" unit="mm" value="22.0">22 mm</quantity>
; anthers oblong-linear, ca.
<quantity id="4CE0D180FFB6EF06F898FE4D81B4D57C" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="57" unit="mm" value="12.0">12 mm</quantity>
, bases with an obtuse fork, connective bases inflated and shorter than that of anther (
<figureCitation id="132360E0FFB6EF06F972FDA185BDD57C" box="[1146,1215,520,545]" captionStart="FIGURE 2" captionStartId="2.[136,229,1930,1951]" captionTargetBox="[151,1436,190,1905]" captionTargetId="figure-15@2.[151,1436,190,1906]" captionTargetPageId="2" captionText="FIGURE 2. Stamens of four Fordiophyton species. A, B: F. chenii; C, D: F. cordifolium; E, F: F. huizhouense; G, H: F. peperomiifolium. A, C, F, H: bases of the longer stamens, side view; B, D: shorter stamens, back view; E, G: bases of the longer stamens, front view. The red arrow represents the anther, and the blue arrow represents the connective." figureDoi="http://doi.org/10.5281/zenodo.13678790" httpUri="https://zenodo.org/record/13678790/files/figure.png" pageId="12" pageNumber="57">Fig. 2</figureCitation>
,
<collectionCode id="ED09E4A0FFB6EF06F9C3FDA185E1D57C" box="[1227,1251,520,545]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="12" pageNumber="57" type="Herbarium">A</collectionCode>
). Shorter stamens ca.
<quantity id="4CE0D180FFB6EF06FDB9FD8581FAD518" box="[177,248,556,581]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="57" unit="mm" value="9.0">9 mm</quantity>
; anthers oblong, ca.
<quantity id="4CE0D180FFB6EF06FCEFFD85832ED518" box="[487,556,556,581]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="57" unit="mm" value="4.0">4 mm</quantity>
, yellow, base obtuse fork, connective base slightly inflated (
<figureCitation id="132360E0FFB6EF06F9ECFD858429D518" box="[1252,1323,556,581]" captionStart="FIGURE 2" captionStartId="2.[136,229,1930,1951]" captionTargetBox="[151,1436,190,1905]" captionTargetId="figure-15@2.[151,1436,190,1906]" captionTargetPageId="2" captionText="FIGURE 2. Stamens of four Fordiophyton species. A, B: F. chenii; C, D: F. cordifolium; E, F: F. huizhouense; G, H: F. peperomiifolium. A, C, F, H: bases of the longer stamens, side view; B, D: shorter stamens, back view; E, G: bases of the longer stamens, front view. The red arrow represents the anther, and the blue arrow represents the connective." figureDoi="http://doi.org/10.5281/zenodo.13678790" httpUri="https://zenodo.org/record/13678790/files/figure.png" pageId="12" pageNumber="57">Fig. 2</figureCitation>
,
<collectionCode id="ED09E4A0FFB6EF06F831FD85844CD518" box="[1337,1358,556,581]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="12" pageNumber="57" type="Herbarium">B</collectionCode>
). Ovary half-inferior, ovate, apex truncate and with a membranous crown; placentas shortly stalked (
<figureCitation id="132360E0FFB6EF06F98BFDF985C5D534" box="[1155,1223,592,617]" captionStart="FIGURE 6" captionStartId="8.[136,228,1953,1974]" captionTargetBox="[216,1371,190,1929]" captionTargetId="figure-15@8.[216,1371,190,1929]" captionTargetPageId="8" captionText="FIGURE 6. Transections of ovaries of four Fordiophyton species. A, B: F. chenii; C, D: F. cordifolium; E, F: F. huizhouense; G, H: F. peperomiifolium; A, C, E, G: general view of ovaries; B, F: close-up of a locule; D, H: close-up of the placenta. CO: ovary column; HY: hypanthium wall; OW: ovary wall; OV: ovule; PL: placenta; SP: sporangia; ST: stalk of placenta; VB: vascular bundle. Scale bars = 250 μm." figureDoi="http://doi.org/10.5281/zenodo.13678806" httpUri="https://zenodo.org/record/13678806/files/figure.png" pageId="12" pageNumber="57">Fig. 6</figureCitation>
,
<collectionCode id="ED09E4A0FFB6EF06F9DAFDF985E5D534" box="[1234,1255,592,617]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="12" pageNumber="57" type="Herbarium">B</collectionCode>
). Capsule funnel-shaped, apex truncate, ca.
<quantity id="4CE0D180FFB6EF06FCB9FDDD8329D5D0" box="[433,555,628,653]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="12" pageNumber="57" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">1215 mm</quantity>
long,
<quantity id="4CE0D180FFB6EF06FF7AFDDD83D2D5D0" box="[626,720,628,653]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="12" pageNumber="57" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
in diameter. Seeds many, light brown, ca.
<quantity id="4CE0D180FFB6EF06F9A7FDDD85F3D5D0" box="[1199,1265,628,653]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="57" unit="mm" value="1.0">1 mm</quantity>
long. Flowering in JanuaryFebruary, fruiting in FebruaryMarch.
</paragraph>
</subSubSection>
<subSubSection id="C3022FEEFFB6EF06FDB5FD158291D5A4" pageId="12" pageNumber="57" type="distribution">
<paragraph id="8BA77C65FFB6EF06FDB5FD158291D5A4" blockId="12.[136,1452,160,941]" pageId="12" pageNumber="57">
<emphasis id="B96CA077FFB6EF06FDB5FD1580EDD588" bold="true" box="[189,495,700,725]" pageId="12" pageNumber="57">Distribution and habitat:</emphasis>
<taxonomicName id="4C1807E6FFB6EF06FCF1FD1583E2D588" authorityName="S. Jin Zeng &amp; X. Y. Zhuang. A" authorityYear="2016" box="[505,736,700,725]" class="Magnoliopsida" family="Melastomataceae" genus="Fordiophyton" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="57" phylum="Tracheophyta" rank="species" species="chenii">
<emphasis id="B96CA077FFB6EF06FCF1FD1583E2D588" box="[505,736,700,725]" italics="true" pageId="12" pageNumber="57">Fordiophyton chenii</emphasis>
</taxonomicName>
grows in damp locations in ravines with broad-leaved forests between
<quantity id="4CE0D180FFB6EF06FDE4FD498031D5A4" box="[236,307,736,761]" metricMagnitude="2" metricUnit="m" metricValue="3.27" pageId="12" pageNumber="57" unit="m" value="327.0">327 m</quantity>
and
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in Jiangmen,
<collectingRegion id="49DCB287FFB6EF06FF46FD49823FD5A4" box="[590,829,736,761]" country="China" name="Guangdong" pageId="12" pageNumber="57">Guangdong Province</collectingRegion>
,
<collectingCountry id="F30F3CF5FFB6EF06FE41FD498292D5A4" box="[841,912,736,761]" name="China" pageId="12" pageNumber="57">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3022FEEFFB6EF06FDB5FCAD81D3D4F0" pageId="12" pageNumber="57" type="etymology">
<paragraph id="8BA77C65FFB6EF06FDB5FCAD83E9D41C" blockId="12.[136,1452,160,941]" pageId="12" pageNumber="57">
<emphasis id="B96CA077FFB6EF06FDB5FCAD804AD440" bold="true" box="[189,328,772,797]" pageId="12" pageNumber="57">Etymology:</emphasis>
The specific epithet is derived from the Chinese botanist and Professor Cheih Chen, who made great contributions to the taxonomy of Chinese melastomes.
</paragraph>
<paragraph id="8BA77C65FFB6EF06FDB5FCE581D3D4F0" blockId="12.[136,1452,160,941]" pageId="12" pageNumber="57">
<emphasis id="B96CA077FFB6EF06FDB5FCE580B8D438" bold="true" box="[189,442,844,869]" pageId="12" pageNumber="57">Conservation status:</emphasis>
Based on its narrow geographic range and small and isolated populations with few individuals,
<taxonomicName id="4C1807E6FFB6EF06FC1AFCD980F7D4D4" authorityName="S. Jin Zeng &amp; X. Y. Zhuang. A" authorityYear="2016" box="[274,501,880,905]" class="Magnoliopsida" family="Melastomataceae" genus="Fordiophyton" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="57" phylum="Tracheophyta" rank="species" species="chenii">
<emphasis id="B96CA077FFB6EF06FC1AFCD980F7D4D4" box="[274,501,880,905]" italics="true" pageId="12" pageNumber="57">Fordiophyton chenii</emphasis>
</taxonomicName>
should be considered endangered (EN) according to the IUCN red list criteria (
<bibRefCitation id="EF890194FFB6EF06F86DFCD981C7D4F0" author="IUCN" pageId="12" pageNumber="57" refId="ref7907" refString="IUCN (2015) The IUCN red list of threatened species, version 2015.2. IUCN Red List Unit, Cambridge U. K. Available from: http: // www. iucnredlist. org / (accessed 21 July 2015)" type="url" year="2015">IUCN 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>