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<mods:title id="CC330F21476546D24BF69BEEA35E55C4">Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XX. Barbaracurus feti sp. n. from Somaliland (Buthidae)</mods:title>
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<figureCitation id="133958A0FFFDFF8FFBF2FB3B598EA0B7" box="[1042,1158,1181,1205]" captionStart-0="Figures 14" captionStart-1="Figures 514" captionStart-2="Figures 1527" captionStart-3="Figures 2835" captionStart-4="Figures 3637" captionStart-5="Figures 3844" captionStart-6="Figures 4546" captionStartId-0="4.[115,192,1981,2003]" captionStartId-1="5.[117,194,1956,1978]" captionStartId-2="6.[116,193,1029,1051]" captionStartId-3="7.[116,193,1516,1538]" captionStartId-4="8.[113,190,2009,2031]" captionStartId-5="9.[115,192,1924,1946]" captionStartId-6="10.[114,191,1981,2003]" captionTargetBox-0="[121,1471,247,1958]" captionTargetBox-1="[113,1474,244,1938]" captionTargetBox-2="[117,1470,244,1004]" captionTargetBox-3="[117,1469,270,1496]" captionTargetBox-4="[113,1474,234,1993]" captionTargetBox-5="[113,1466,237,1904]" captionTargetBox-6="[113,1474,234,1971]" captionTargetId-0="figure-58@4.[113,1474,234,1970]" captionTargetId-1="figure-7@5.[113,1474,244,1944]" captionTargetId-2="figure-483@6.[113,1474,234,1019]" captionTargetId-3="figure-242@7.[113,1474,234,1502]" captionTargetId-4="figure-17@8.[110,1474,234,1993]" captionTargetId-5="figure-8@9.[113,1474,231,1912]" captionTargetId-6="figure-17@10.[113,1474,234,1973]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionText-0="Figures 14: Barbaracurus feti sp. n. Figures 12. Holotype male, dorsal (1) and ventral (2) views. Figures 34. Paratype female, dorsal (3) and ventral (4) views. Scale bar: 10 mm." captionText-1="Figures 514: Barbaracurus feti sp. n. Figures 5, 7. Holotype male, carapace and tergites IIII (5) and sternopectinal region and sternites IIIV (7). Figures 6, 8, 914. Paratype female, carapace and tergites IIII (6), sternopectinal region and sternite III (8), distal segments of right legs IIV, retrolateral views (912), left chelicera, dorsal (13) and ventral (14) views." captionText-2="Figures 1527: Barbaracurus feti sp. n. Figures 1516. Paratype female. Pedipalp chela, dorsal (15), and external (16) views. Figures 1727. Holotype male. Pedipalp chela, dorsal (17), external (18), and ventrointernal (19) views.Pedipalp patella, dorsal (20), external (21), and ventral (22) views. Pedipalp femur and trochanter, internal (23), ventral (24) and dorsal (25) views. Pedipalp chela, movable (26) and fixed (27) fingers dentate margin. The trichobothrial pattern is indicated in Figures 1621, 23, and 25 (white circles)." captionText-3="Figures 2835: Barbaracurus feti sp. n. Figures 28, 3032. Holotype male, metasoma V and telson lateral view (28), metasoma and telson, lateral (30), dorsal (31) and ventral (32) views. Figures 29, 3335. Paratype female, metasoma V and telson lateral view (29), metasoma and telson, lateral (33), dorsal (34) ventral and sternite VII (35) views. Scale bar: 10 mm (3032, 3335)." captionText-4="Figures 3637: Barbaracurus feti sp. n., in vivo habitus. Male holotype (36) and female paratype (37)." captionText-5="Figures 3844: Barbaracurus feti sp. n. Figures 3842: Right hemispermatophore of male holotype. Figure 38. Whole hemispermatophore, convex view. Scale bar: 1 mm. Figures 3942. Capsule in convex compressed (39), convex (40), anterior (41) and posterior (42) views. Scale bar: 500 μm. Figures 4344: Mitotic metaphase (43) and ideogram (44) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values) of male holotype. Arrows show one extra large odd chromosome. Scale bar: 5 μm." captionText-6="Figures 4546: Figure 45. Map showing confirmed distribution of Barbaracurus spp. For exact position of Rugay village, the type locality of Barbaracurus feti sp. n. see fig. 84 in Kovařík et al., 2019. Figure 46. Barbaracurus feti sp. n., type locality." figureDoi-0="http://doi.org/10.5281/zenodo.4648236" figureDoi-1="http://doi.org/10.5281/zenodo.4648238" figureDoi-2="http://doi.org/10.5281/zenodo.4648244" figureDoi-3="http://doi.org/10.5281/zenodo.4648246" figureDoi-4="http://doi.org/10.5281/zenodo.4648248" figureDoi-5="http://doi.org/10.5281/zenodo.4648250" figureDoi-6="http://doi.org/10.5281/zenodo.4648252" httpUri-0="https://zenodo.org/record/4648236/files/figure.png" httpUri-1="https://zenodo.org/record/4648238/files/figure.png" httpUri-2="https://zenodo.org/record/4648244/files/figure.png" httpUri-3="https://zenodo.org/record/4648246/files/figure.png" httpUri-4="https://zenodo.org/record/4648248/files/figure.png" httpUri-5="https://zenodo.org/record/4648250/files/figure.png" httpUri-6="https://zenodo.org/record/4648252/files/figure.png" pageId="3" pageNumber="2">Figs. 146</figureCitation>
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No. 1528)
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<specimenCount id="9D048FACFFFDFF8FFC6FFA5B5EC3A217" box="[911,971,1533,1557]" pageId="3" pageNumber="2" type="generic">1juv.</specimenCount>
(
<typeStatus id="54B9FA87FFFDFF8FFC39FA5B594DA217" box="[985,1093,1533,1557]" pageId="3" pageNumber="2" type="paratype">paratypes</typeStatus>
), leg. F. Kovařík,
<collectionCode id="ED13DCE0FFFDFF8FFAE4FA5B584FA217" box="[1284,1351,1533,1557]" pageId="3" pageNumber="2">FKCP</collectionCode>
,
<collectionCode id="ED13DCE0FFFDFF8FFAB1FA5B5891A217" box="[1361,1433,1533,1557]" pageId="3" pageNumber="2">GLPC</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C31817AEFFFDFF8FFCD0F99B5E96A297" pageId="3" pageNumber="2" type="etymology">
<paragraph id="8BBD4425FFFDFF8FFCD0F99B5E96A297" blockId="3.[816,1475,1597,1685]" pageId="3" pageNumber="2">
ETYMOLOGY. The specific epithet honors Victor Fet (Marshall University,
<collectingCountry id="F31504B5FFFDFF8FFC4DF9FB5EECA277" box="[941,996,1629,1653]" name="United States of America" pageId="3" pageNumber="2">USA</collectingCountry>
) for his friendship and lifelong dedication to scorpions.
</paragraph>
</subSubSection>
<subSubSection id="C31817AEFFFDFF8AFCD0F91B5DFFA1A8" lastPageId="6" lastPageNumber="5" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="8BBD4425FFFDFF8AFCD0F91B5DFFA1A8" blockId="3.[816,1474,1725,2038]" lastBlockId="6.[113,772,1170,1450]" lastPageId="6" lastPageNumber="5" pageId="3" pageNumber="2">
DIAGNOSIS. Total length of adult male
<quantity id="4CFAE9C0FFFDFF8FFB25F91B583BA2D7" box="[1221,1331,1725,1749]" metricMagnitude="-2" metricUnit="m" metricValue="3.125" pageId="3" pageNumber="2" unit="mm" value="31.25">31.25 mm</quantity>
, adult female
<quantity id="4CFAE9C0FFFDFF8FFCD0F97B5EA9A2F7" box="[816,929,1757,1781]" metricMagnitude="-2" metricUnit="m" metricValue="3.88" pageId="3" pageNumber="2" unit="mm" value="38.8">38.80 mm</quantity>
. Coloration pale yellow to light orange, chelicerae yellow without reticulation. Pedipalp chela manus wider in male than female, chela length/width ratio
<quantity id="4CFAE9C0FFFDFF8FFAD4F8BB588BA337" box="[1332,1411,1821,1845]" metricMagnitude="-1" metricUnit="m" metricValue="1.11252" pageId="3" pageNumber="2" unit="in" value="4.38">4.38 in</quantity>
male and
<quantity id="4CFAE9C0FFFDFF8FFC84F89B5EBCA357" box="[868,948,1853,1877]" metricMagnitude="-1" metricUnit="m" metricValue="1.4097" pageId="3" pageNumber="2" unit="in" value="5.55">5.55 in</quantity>
female; proximal margins of pedipalp fingers of female straight, of male weakly undulate with very small gap with fingers closed (
<figureCitation id="133958A0FFFDFF8FFBDAF8DB5982A397" box="[1082,1162,1917,1941]" captionStart="Figures 1527" captionStartId="6.[116,193,1029,1051]" captionTargetBox="[117,1470,244,1004]" captionTargetId="figure-483@6.[113,1474,234,1019]" captionTargetPageId="6" captionText="Figures 1527: Barbaracurus feti sp. n. Figures 1516. Paratype female. Pedipalp chela, dorsal (15), and external (16) views. Figures 1727. Holotype male. Pedipalp chela, dorsal (17), external (18), and ventrointernal (19) views.Pedipalp patella, dorsal (20), external (21), and ventral (22) views. Pedipalp femur and trochanter, internal (23), ventral (24) and dorsal (25) views. Pedipalp chela, movable (26) and fixed (27) fingers dentate margin. The trichobothrial pattern is indicated in Figures 1621, 23, and 25 (white circles)." figureDoi="http://doi.org/10.5281/zenodo.4648244" httpUri="https://zenodo.org/record/4648244/files/figure.png" pageId="3" pageNumber="2">Fig. 18</figureCitation>
); dentate margin of movable finger armed with 6 rows of granules, and a short apical row of 3 denticles (
<figureCitation id="133958A0FFFDFF8FFC32F81B592BA3D7" box="[978,1059,1981,2005]" captionStart="Figures 1527" captionStartId="6.[116,193,1029,1051]" captionTargetBox="[117,1470,244,1004]" captionTargetId="figure-483@6.[113,1474,234,1019]" captionTargetPageId="6" captionText="Figures 1527: Barbaracurus feti sp. n. Figures 1516. Paratype female. Pedipalp chela, dorsal (15), and external (16) views. Figures 1727. Holotype male. Pedipalp chela, dorsal (17), external (18), and ventrointernal (19) views.Pedipalp patella, dorsal (20), external (21), and ventral (22) views. Pedipalp femur and trochanter, internal (23), ventral (24) and dorsal (25) views. Pedipalp chela, movable (26) and fixed (27) fingers dentate margin. The trichobothrial pattern is indicated in Figures 1621, 23, and 25 (white circles)." figureDoi="http://doi.org/10.5281/zenodo.4648244" httpUri="https://zenodo.org/record/4648244/files/figure.png" pageId="3" pageNumber="2">Fig. 26</figureCitation>
); most proximal granule row with one external accessory granule. Pectines with 2527 teeth in both sexes. Hemispermatophore basal lobe a short oblique lamina, length ca. half of basal width of posterior lobe. Metasoma narrow, metasoma V length/width ratio
<quantity id="4CFAE9C0FFF8FF8AFDDCFB745F86A0E8" box="[572,654,1234,1258]" metricMagnitude="-2" metricUnit="m" metricValue="6.4008" pageId="6" pageNumber="5" unit="in" value="2.52">2.52 in</quantity>
male and
<quantity id="4CFAE9C0FFF8FF8AFF91FB545DC9A108" box="[113,193,1266,1290]" metricMagnitude="-2" metricUnit="m" metricValue="6.7564" pageId="6" pageNumber="5" unit="in" value="2.66">2.66 in</quantity>
female; metasoma I with 10 carinae, IIIV with 8 carinae. Telson setose, bearing numerous long macrosetae and short, pointed subaculear tubercle; vesicle smooth, elongate, ellipsoidal or weakly pyriform in lateral profile, telson length/ depth ratio
<quantity id="4CFAE9C0FFF8FF8AFF09FAD45C3BA188" box="[233,307,1394,1418]" metricMagnitude="-2" metricUnit="m" metricValue="8.2296" pageId="6" pageNumber="5" unit="in" value="3.24">3.24 in</quantity>
both sexes; aculeus slender, curved, shorter than vesicle.
</paragraph>
</subSubSection>
<caption id="DF7D14ADFFFAFF88FF93F81B5CF2A3ED" ID-DOI="http://doi.org/10.5281/zenodo.4648236" ID-Zenodo-Dep="4648236" httpUri="https://zenodo.org/record/4648236/files/figure.png" pageId="4" pageNumber="3" startId="4.[115,192,1981,2003]" targetBox="[121,1471,247,1958]" targetPageId="4">
<paragraph id="8BBD4425FFFAFF88FF93F81B5CF2A3ED" blockId="4.[115,1473,1981,2032]" pageId="4" pageNumber="3">
<emphasis id="B9769837FFFAFF88FF93F81B5DE2A3D0" bold="true" box="[115,234,1981,2003]" pageId="4" pageNumber="3">Figures 14</emphasis>
:
<taxonomicName id="4C023FA6FFFAFF88FF16F81B5CAAA3D0" authority="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre, 2019" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre" authorityYear="2019" box="[246,418,1981,2002]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="feti" status="sp. nov.">
<emphasis id="B9769837FFFAFF88FF16F81B5CAAA3D0" box="[246,418,1981,2002]" italics="true" pageId="4" pageNumber="3">Barbaracurus feti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245254CFFFAFF88FE47F81B5CD5A3D0" box="[423,477,1981,2002]" pageId="4" pageNumber="3" rank="species">
<emphasis id="B9769837FFFAFF88FE47F81B5CB6A3D0" bold="true" box="[423,446,1981,2002]" pageId="4" pageNumber="3">sp</emphasis>
.
<emphasis id="B9769837FFFAFF88FE2AF81B5CDFA3D0" bold="true" box="[458,471,1981,2002]" pageId="4" pageNumber="3">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B9769837FFFAFF88FE03F81B5F52A3D0" bold="true" box="[483,602,1981,2003]" pageId="4" pageNumber="3">Figures 12</emphasis>
. Holotype male, dorsal (1) and ventral (2) views.
<emphasis id="B9769837FFFAFF88FBDAF81B59B9A3D0" bold="true" box="[1082,1201,1981,2003]" pageId="4" pageNumber="3">Figures 34</emphasis>
. Paratype female, dorsal (3) and ventral (4) views. Scale bar: 10 mm.
</paragraph>
</caption>
<caption id="DF7D14ADFFFBFF89FF95F8025EAFA3F6" ID-DOI="http://doi.org/10.5281/zenodo.4648238" ID-Zenodo-Dep="4648238" httpUri="https://zenodo.org/record/4648238/files/figure.png" pageId="5" pageNumber="4" startId="5.[117,194,1956,1978]" targetBox="[113,1474,244,1938]" targetPageId="5">
<paragraph id="8BBD4425FFFBFF89FF95F8025EAFA3F6" blockId="5.[117,1475,1956,2036]" pageId="5" pageNumber="4">
<emphasis id="B9769837FFFBFF89FF95F8025DF2A3BB" bold="true" box="[117,250,1956,1978]" pageId="5" pageNumber="4">Figures 514</emphasis>
:
<taxonomicName id="4C023FA6FFFBFF89FEE8F8025CBDA3BB" authority="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre, 2019" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre" authorityYear="2019" box="[264,437,1956,1977]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="feti" status="sp. nov.">
<emphasis id="B9769837FFFBFF89FEE8F8025CBDA3BB" box="[264,437,1956,1977]" italics="true" pageId="5" pageNumber="4">Barbaracurus feti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245254CFFFBFF89FE5EF8025CFCA3B8" box="[446,500,1956,1978]" pageId="5" pageNumber="4" rank="species">
<emphasis id="B9769837FFFBFF89FE5EF8025CDBA3BB" bold="true" box="[446,467,1956,1977]" pageId="5" pageNumber="4">sp</emphasis>
.
<emphasis id="B9769837FFFBFF89FE01F8025CE6A3BB" bold="true" box="[481,494,1956,1977]" pageId="5" pageNumber="4">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B9769837FFFBFF89FE1CF8025F54A3BB" bold="true" box="[508,604,1956,1978]" pageId="5" pageNumber="4">Figures 5</emphasis>
,
<emphasis id="B9769837FFFBFF89FD8AF8025F7EA3BB" bold="true" box="[618,630,1956,1977]" pageId="5" pageNumber="4">7</emphasis>
. Holotype male, carapace and tergites IIII (5) and sternopectinal region and sternites IIIV (7).
<emphasis id="B9769837FFFBFF89FF37F8675C3DA3D4" bold="true" box="[215,309,1985,2007]" pageId="5" pageNumber="4">Figures 6</emphasis>
,
<emphasis id="B9769837FFFBFF89FEA0F8675C44A3D4" bold="true" box="[320,332,1985,2006]" pageId="5" pageNumber="4">8</emphasis>
,
<emphasis id="B9769837FFFBFF89FEB7F8675C8FA3D4" bold="true" box="[343,391,1985,2006]" pageId="5" pageNumber="4">914</emphasis>
. Paratype female, carapace and tergites IIII (6), sternopectinal region and sternite III (8), distal segments of right legs IIV, retrolateral views (912), left chelicera, dorsal (13) and ventral (14) views.
</paragraph>
</caption>
<caption id="DF7D14ADFFF8FF8AFF94FBA35921A070" ID-DOI="http://doi.org/10.5281/zenodo.4648244" ID-Zenodo-Dep="4648244" httpUri="https://zenodo.org/record/4648244/files/figure.png" pageId="6" pageNumber="5" startId="6.[116,193,1029,1051]" targetBox="[117,1470,244,1004]" targetPageId="6">
<paragraph id="8BBD4425FFF8FF8AFF94FBA35921A070" blockId="6.[116,1474,1029,1138]" pageId="6" pageNumber="5">
<emphasis id="B9769837FFF8FF8AFF94FBA35C0AA018" bold="true" box="[116,258,1029,1051]" pageId="6" pageNumber="5">Figures 1527</emphasis>
:
<taxonomicName id="4C023FA6FFF8FF8AFEEEFBA35CB1A018" authority="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre, 2019" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre" authorityYear="2019" box="[270,441,1029,1050]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="feti" status="sp. nov.">
<emphasis id="B9769837FFF8FF8AFEEEFBA35CB1A018" box="[270,441,1029,1050]" italics="true" pageId="6" pageNumber="5">Barbaracurus feti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245254CFFF8FF8AFE5DFBA05CFBA019" box="[445,499,1030,1051]" pageId="6" pageNumber="5" rank="species">
<emphasis id="B9769837FFF8FF8AFE5DFBA05CDCA019" bold="true" box="[445,468,1030,1051]" pageId="6" pageNumber="5">sp</emphasis>
.
<emphasis id="B9769837FFF8FF8AFE00FBA05CE5A019" bold="true" box="[480,493,1030,1051]" pageId="6" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B9769837FFF8FF8AFE19FBA35F8FA018" bold="true" box="[505,647,1029,1051]" pageId="6" pageNumber="5">Figures 1516</emphasis>
. Paratype female. Pedipalp chela, dorsal (15), and external (16) views.
<emphasis id="B9769837FFF8FF8AFACEFBA358B4A018" bold="true" box="[1326,1468,1029,1051]" pageId="6" pageNumber="5">Figures 1727</emphasis>
. Holotype male. Pedipalp chela, dorsal (17), external (18), and ventrointernal (19) views. Pedipalp patella, dorsal (20), external (21), and ventral (22) views. Pedipalp femur and trochanter, internal (23), ventral (24) and dorsal (25) views. Pedipalp chela, movable (26) and fixed (27) fingers dentate margin. The trichobothrial pattern is indicated in Figures 1621, 23, and 25 (white circles).
</paragraph>
</caption>
<subSubSection id="C31817AEFFF8FF87FF91FA745CACA081" lastPageId="11" lastPageNumber="10" pageId="6" pageNumber="5" type="description">
<paragraph id="8BBD4425FFF8FF8AFF91FA745CECA329" blockId="6.[113,772,1490,2026]" pageId="6" pageNumber="5">
DESCRIPTION. Total length of adults 31.25 (male)
<quantity id="4CFAE9C0FFF8FF8AFD77FA745E0BA1E8" box="[663,771,1490,1514]" metricMagnitude="-2" metricUnit="m" metricValue="3.88" pageId="6" pageNumber="5" unit="mm" value="38.8">38.80 mm</quantity>
(female). Measurements of the carapace, telson, segments of the metasoma and pedipalps are given in
<tableCitation id="C680719EFFF8FF8AFDD2F9B45F8BA228" box="[562,643,1554,1578]" captionStart="Table 1" captionStartId="3.[114,170,1063,1084]" captionTargetId="graphics-430@3.[117,1470,238,1048]" captionTargetPageId="3" captionText="Table 1. Comparative measurements of adults of Barbaracurus feti sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/DF7D14ADFFFDFF8FFF92FB815C73A05F" pageId="6" pageNumber="5" tableUuid="DF7D14ADFFFDFF8FFF92FB815C73A05F">Table 1</tableCitation>
. Base color is pale yellow to light orange (
<figureCitation id="133958A0FFF8FF8AFE52F9945F10A248" box="[434,536,1586,1610]" captionStart="Figures 14" captionStartId="4.[115,192,1981,2003]" captionTargetBox="[121,1471,247,1958]" captionTargetId="figure-58@4.[113,1474,234,1970]" captionTargetPageId="4" captionText="Figures 14: Barbaracurus feti sp. n. Figures 12. Holotype male, dorsal (1) and ventral (2) views. Figures 34. Paratype female, dorsal (3) and ventral (4) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4648236" httpUri="https://zenodo.org/record/4648236/files/figure.png" pageId="6" pageNumber="5">Figs. 14</figureCitation>
), metasoma V is dark. Chelicerae are yellow without reticulation (
<figureCitation id="133958A0FFF8FF8AFD8CF9F45FFFA268" box="[620,759,1618,1642]" captionStart="Figures 514" captionStartId="5.[117,194,1956,1978]" captionTargetBox="[113,1474,244,1938]" captionTargetId="figure-7@5.[113,1474,244,1944]" captionTargetPageId="5" captionText="Figures 514: Barbaracurus feti sp. n. Figures 5, 7. Holotype male, carapace and tergites IIII (5) and sternopectinal region and sternites IIIV (7). Figures 6, 8, 914. Paratype female, carapace and tergites IIII (6), sternopectinal region and sternite III (8), distal segments of right legs IIV, retrolateral views (912), left chelicera, dorsal (13) and ventral (14) views." figureDoi="http://doi.org/10.5281/zenodo.4648238" httpUri="https://zenodo.org/record/4648238/files/figure.png" pageId="6" pageNumber="5">Figs. 1314</figureCitation>
).
<emphasis id="B9769837FFF8FF8AFF91F9D45C4CA288" box="[113,324,1650,1674]" italics="true" pageId="6" pageNumber="5">Sexual dimorphism</emphasis>
minor, adult male with wider pedipalp chela than female, pedipalp chela length/width ratio
<quantity id="4CFAE9C0FFF8FF8AFD34F9345D8EA2C8" metricMagnitude="-1" metricUnit="m" metricValue="1.11252" pageId="6" pageNumber="5" unit="in" value="4.38">4.38 in</quantity>
male and
<quantity id="4CFAE9C0FFF8FF8AFF1EF9145C43A2C8" box="[254,331,1714,1738]" metricMagnitude="-1" metricUnit="m" metricValue="1.4097" pageId="6" pageNumber="5" unit="in" value="5.55">5.55 in</quantity>
female; pedipalps fingers are shorter in males, the fingers of females are straight, while those of males are weakly undulate proximally; no sexual difference in length and width of metasomal segments.
</paragraph>
<paragraph id="8BBD4425FFF8FF8AFF91F8945ECAA0E8" blockId="6.[113,772,1490,2026]" lastBlockId="6.[816,1475,1170,2026]" pageId="6" pageNumber="5">
<emphasis id="B9769837FFF8FF8AFF91F8945DDEA348" bold="true" box="[113,214,1842,1866]" pageId="6" pageNumber="5">Pedipalp</emphasis>
(
<figureCitation id="133958A0FFF8FF8AFF11F8945C75A348" box="[241,381,1842,1866]" captionStart="Figures 1527" captionStartId="6.[116,193,1029,1051]" captionTargetBox="[117,1470,244,1004]" captionTargetId="figure-483@6.[113,1474,234,1019]" captionTargetPageId="6" captionText="Figures 1527: Barbaracurus feti sp. n. Figures 1516. Paratype female. Pedipalp chela, dorsal (15), and external (16) views. Figures 1727. Holotype male. Pedipalp chela, dorsal (17), external (18), and ventrointernal (19) views.Pedipalp patella, dorsal (20), external (21), and ventral (22) views. Pedipalp femur and trochanter, internal (23), ventral (24) and dorsal (25) views. Pedipalp chela, movable (26) and fixed (27) fingers dentate margin. The trichobothrial pattern is indicated in Figures 1621, 23, and 25 (white circles)." figureDoi="http://doi.org/10.5281/zenodo.4648244" httpUri="https://zenodo.org/record/4648244/files/figure.png" pageId="6" pageNumber="5">Figs. 1527</figureCitation>
). Pedipalp mostly very sparsely hirsute, but more densely so on ventral surface of movable finger. Femur granulated, with five granulose carinae. Patella almost smooth with seven granulose carinae developed. Chela smooth with traces of carinae visible; fingers long, curved; movable finger with 6 granule rows and short apical row of 4 denticles on dentate margins, the most proximal row with one external and no internal accessory granule; fixed finger with 7 granule rows.
</paragraph>
<paragraph id="8BBD4425FFF8FF8AFCD0FB54583CA1E8" blockId="6.[816,1475,1170,2026]" pageId="6" pageNumber="5">
<emphasis id="B9769837FFF8FF8AFCD0FB545E96A108" bold="true" box="[816,926,1266,1290]" pageId="6" pageNumber="5">Carapace</emphasis>
(
<figureCitation id="133958A0FFF8FF8AFC5EFB545930A108" box="[958,1080,1266,1290]" captionStart="Figures 514" captionStartId="5.[117,194,1956,1978]" captionTargetBox="[113,1474,244,1938]" captionTargetId="figure-7@5.[113,1474,244,1944]" captionTargetPageId="5" captionText="Figures 514: Barbaracurus feti sp. n. Figures 5, 7. Holotype male, carapace and tergites IIII (5) and sternopectinal region and sternites IIIV (7). Figures 6, 8, 914. Paratype female, carapace and tergites IIII (6), sternopectinal region and sternite III (8), distal segments of right legs IIV, retrolateral views (912), left chelicera, dorsal (13) and ventral (14) views." figureDoi="http://doi.org/10.5281/zenodo.4648238" httpUri="https://zenodo.org/record/4648238/files/figure.png" pageId="6" pageNumber="5">Figs. 56</figureCitation>
). Slightly trapezoidal (narrower anteriorly) and slightly longer than wide; anterior margin concave, with some short microsetae. Carination absent. Median and posterior lateral furrows wide and deep, others vestigial to absent. Tegument densely and coarsely granulose. Median eyes large and raised; four or five pairs of lateral eyes: three major ocelli aligned along each anterolateral corner, plus two minor ocelli that may be vestigial or absent.
</paragraph>
<paragraph id="8BBD4425FFF8FF8AFCD0FA545985A388" blockId="6.[816,1475,1170,2026]" pageId="6" pageNumber="5">
<emphasis id="B9769837FFF8FF8AFCD0FA545EA0A208" bold="true" box="[816,936,1522,1546]" pageId="6" pageNumber="5">Mesosoma</emphasis>
(
<figureCitation id="133958A0FFF8FF8AFC5FFA545926A208" box="[959,1070,1522,1546]" captionStart="Figures 14" captionStartId="4.[115,192,1981,2003]" captionTargetBox="[121,1471,247,1958]" captionTargetId="figure-58@4.[113,1474,234,1970]" captionTargetPageId="4" captionText="Figures 14: Barbaracurus feti sp. n. Figures 12. Holotype male, dorsal (1) and ventral (2) views. Figures 34. Paratype female, dorsal (3) and ventral (4) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4648236" httpUri="https://zenodo.org/record/4648236/files/figure.png" pageId="6" pageNumber="5">Figs. 14</figureCitation>
,
<figureCitation id="133958A0FFF8FF8AFBA1FA545964A208" box="[1089,1132,1522,1546]" captionStart="Figures 514" captionStartId="5.[117,194,1956,1978]" captionTargetBox="[113,1474,244,1938]" captionTargetId="figure-7@5.[113,1474,244,1944]" captionTargetPageId="5" captionText="Figures 514: Barbaracurus feti sp. n. Figures 5, 7. Holotype male, carapace and tergites IIII (5) and sternopectinal region and sternites IIIV (7). Figures 6, 8, 914. Paratype female, carapace and tergites IIII (6), sternopectinal region and sternite III (8), distal segments of right legs IIV, retrolateral views (912), left chelicera, dorsal (13) and ventral (14) views." figureDoi="http://doi.org/10.5281/zenodo.4648238" httpUri="https://zenodo.org/record/4648238/files/figure.png" pageId="6" pageNumber="5">58</figureCitation>
). Tergites IVI bear a single conspicuous median carina; tergite VII bears five well-defined carinae (median, submedians and laterals), which are long and serrate to crenulate.All tergites densely and coarsely granulose mainly on posterior parts. Sternum
<typeStatus id="54B9FA87FFF8FF8AFB47F9D559DDA289" box="[1191,1237,1651,1675]" pageId="6" pageNumber="5">type</typeStatus>
1, triangular in shape; medial depression large. Pectines extending to around a quarter of sternite V in male and a quarter of sternite IV in female. Tooth count
<quantity id="4CFAE9C0FFF8FF8AFC5EF9745917A2E8" box="[958,1055,1746,1770]" metricMagnitude="-1" metricUnit="m" metricValue="6.476999999999999" metricValueMax="6.604" metricValueMin="6.35" pageId="6" pageNumber="5" unit="in" value="25.5" valueMax="26.0" valueMin="25.0">2526 in</quantity>
male and juvenile, and
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female. Pectines with 3 marginal lamellae and 89 middle lamellae. Sternites lacking carinae, surfaces smooth and sparsely setose. Posterior margin of sternite V without smooth median patch in both sexes. Sternite VII with four well-defined carinae, which are long and serrate to crenulate.
</paragraph>
<paragraph id="8BBD4425FFF8FF8BFCD0F8345FCCA3E8" blockId="6.[816,1475,1170,2026]" lastBlockId="7.[113,771,1618,2026]" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5">
<emphasis id="B9769837FFF8FF8AFCD0F834591FA3A8" bold="true" box="[816,1047,1938,1962]" pageId="6" pageNumber="5">Hemispermatophore</emphasis>
(
<figureCitation id="133958A0FFF8FF8AFBC6F83459AEA3A8" box="[1062,1190,1938,1962]" captionStart="Figures 3844" captionStartId="9.[115,192,1924,1946]" captionTargetBox="[113,1466,237,1904]" captionTargetId="figure-8@9.[113,1474,231,1912]" captionTargetPageId="9" captionText="Figures 3844: Barbaracurus feti sp. n. Figures 3842: Right hemispermatophore of male holotype. Figure 38. Whole hemispermatophore, convex view. Scale bar: 1 mm. Figures 3942. Capsule in convex compressed (39), convex (40), anterior (41) and posterior (42) views. Scale bar: 500 μm. Figures 4344: Mitotic metaphase (43) and ideogram (44) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values) of male holotype. Arrows show one extra large odd chromosome. Scale bar: 5 μm." figureDoi="http://doi.org/10.5281/zenodo.4648250" httpUri="https://zenodo.org/record/4648250/files/figure.png" pageId="6" pageNumber="5">Figs. 3842</figureCitation>
). Flagelliform. Trunk long, narrow, gradually widening basally. Capsule short, its length measured from basal lobe only 11% of trunk length. Flagellum narrow, slightly tapered, pars recta 45% of trunk length, pars reflecta is truncated in the available material. Sperm hemiduct with 2 elongated lobes, posterior lobe longer with broad base, subtriangular with distinct carina, apex blunt; anterior lobe shorter, constricted at base, separated from posterior lobe by small notch or aperture at base. Basal lobe a short lamina, prominently projecting from convex surface of capsule, oriented along oblique, transverse axis from anterior base of posterior lobe. Lamina length is about half of the basal width of the posterior lobe, similar to the shorter laminae of
<taxonomicName id="4C023FA6FFF9FF8BFD8BF8D55E0BA388" baseAuthorityName="Borelli" baseAuthorityYear="1902" box="[619,771,1906,1930]" class="Arachnida" family="Buthidae" genus="Babycurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="zambonellii">
<emphasis id="B9769837FFF9FF8BFD8BF8D55F73A388" box="[619,635,1907,1930]" italics="true" pageId="7" pageNumber="6">B</emphasis>
.
<emphasis id="B9769837FFF9FF8BFD69F8D45E0BA388" box="[649,771,1906,1930]" italics="true" pageId="7" pageNumber="6">zambonellii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C023FA6FFF9FF8BFF42F8355C25A3A8" authorityName="Somaliland" baseAuthorityName="Hirst" baseAuthorityYear="1907" box="[162,301,1938,1962]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="somalicus">
<emphasis id="B9769837FFF9FF8BFF42F8355DBAA3A8" box="[162,178,1939,1962]" italics="true" pageId="7" pageNumber="6">B</emphasis>
.
<emphasis id="B9769837FFF9FF8BFF24F8345C25A3A8" box="[196,301,1938,1962]" italics="true" pageId="7" pageNumber="6">somalicus</emphasis>
</taxonomicName>
(cf. figs. 2728, 35
<quantity id="4CFAE9C0FFF9FF8BFDF8F8345F5EA3A8" box="[536,598,1938,1962]" metricMagnitude="-1" metricUnit="m" metricValue="9.144" pageId="7" pageNumber="6" unit="in" value="36.0">36 in</quantity>
:
<bibRefCitation id="EF9339D4FFF9FF8BFD84F8345DA2A3C8" author="KOVARIK" pageId="7" pageNumber="6" pagination="1 - 41" refId="ref4813" refString="KOVARIK, F., G. LOWE &amp; F. STAHLAVSKY. 2018. Review of the genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), with description of Barbaracurus gen. n. and two new species from Oman and Yemen. Euscorpius, 267: 1 - 41." type="journal article" year="2018">Kovařík et al., 2018</bibRefCitation>
). Notably, these three allopatric species are distributed adjacently along the coastal region of the Horn of Africa.
</paragraph>
<caption id="DF7D14ADFFF9FF8BFF94FA4A5936A239" ID-DOI="http://doi.org/10.5281/zenodo.4648246" ID-Zenodo-Dep="4648246" httpUri="https://zenodo.org/record/4648246/files/figure.png" pageId="7" pageNumber="6" startId="7.[116,193,1516,1538]" targetBox="[117,1469,270,1496]" targetPageId="7">
<paragraph id="8BBD4425FFF9FF8BFF94FA4A5936A239" blockId="7.[116,1474,1516,1596]" pageId="7" pageNumber="6">
<emphasis id="B9769837FFF9FF8BFF94FA4A5C0BA203" bold="true" box="[116,259,1516,1538]" pageId="7" pageNumber="6">Figures 2835</emphasis>
:
<taxonomicName id="4C023FA6FFF9FF8BFEF1FA4A5CB5A203" authority="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre, 2019" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre" authorityYear="2019" box="[273,445,1516,1537]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="feti" status="sp. nov.">
<emphasis id="B9769837FFF9FF8BFEF1FA4A5CB5A203" box="[273,445,1516,1537]" italics="true" pageId="7" pageNumber="6">Barbaracurus feti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245254CFFF9FF8BFE23FA4A5CF2A203" box="[451,506,1516,1537]" pageId="7" pageNumber="6" rank="species">
<emphasis id="B9769837FFF9FF8BFE23FA4A5CD2A203" bold="true" box="[451,474,1516,1537]" pageId="7" pageNumber="6">sp</emphasis>
.
<emphasis id="B9769837FFF9FF8BFE07FA4A5CFCA203" bold="true" box="[487,500,1516,1537]" pageId="7" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B9769837FFF9FF8BFDE1FA4A5F64A203" bold="true" box="[513,620,1516,1538]" pageId="7" pageNumber="6">Figures 28</emphasis>
,
<emphasis id="B9769837FFF9FF8BFD99FA4A5FBDA203" bold="true" box="[633,693,1516,1537]" pageId="7" pageNumber="6">3032</emphasis>
. Holotype male, metasoma V and telson lateral view (28), metasoma and telson, lateral (30), dorsal (31) and ventral (32) views.
<emphasis id="B9769837FFF9FF8BFDDEF9AE5FA1A21C" bold="true" box="[574,681,1544,1566]" pageId="7" pageNumber="6">Figures 29</emphasis>
,
<emphasis id="B9769837FFF9FF8BFD55F9AF5FF9A21C" bold="true" box="[693,753,1545,1566]" pageId="7" pageNumber="6">3335</emphasis>
. Paratype female, metasoma V and telson lateral view (29), metasoma and telson, lateral (33), dorsal (34) ventral and sternite VII (35) views. Scale bar: 10 mm (3032, 3335).
</paragraph>
</caption>
<paragraph id="8BBD4425FFF9FF8BFCD0F9F45E91A30B" blockId="7.[816,1475,1618,2026]" pageId="7" pageNumber="6">
<emphasis id="B9769837FFF9FF8BFCD0F9F45E6DA268" bold="true" box="[816,869,1618,1642]" pageId="7" pageNumber="6">Legs</emphasis>
(
<figureCitation id="133958A0FFF9FF8BFC93F9F45EEEA268" box="[883,998,1618,1642]" captionStart="Figures 514" captionStartId="5.[117,194,1956,1978]" captionTargetBox="[113,1474,244,1938]" captionTargetId="figure-7@5.[113,1474,244,1944]" captionTargetPageId="5" captionText="Figures 514: Barbaracurus feti sp. n. Figures 5, 7. Holotype male, carapace and tergites IIII (5) and sternopectinal region and sternites IIIV (7). Figures 6, 8, 914. Paratype female, carapace and tergites IIII (6), sternopectinal region and sternite III (8), distal segments of right legs IIV, retrolateral views (912), left chelicera, dorsal (13) and ventral (14) views." figureDoi="http://doi.org/10.5281/zenodo.4648238" httpUri="https://zenodo.org/record/4648238/files/figure.png" pageId="7" pageNumber="6">Figs. 912</figureCitation>
). Tarsomeres bearing two rows of macrosetae on their ventral surface and numerous macrosetae on other surfaces; bristle combs absent. Femur bearing only solitary macrosetae. Femur surface coarsely granulose, femur and patella with carinae developed. Moderate tibial spurs present on leg IV.
</paragraph>
<paragraph id="8BBD4425FFF9FF87FCD0F8B45FE4A5A1" blockId="7.[816,1475,1618,2026]" lastBlockId="11.[113,772,235,1155]" lastPageId="11" lastPageNumber="10" pageId="7" pageNumber="6">
<emphasis id="B9769837FFF9FF8BFCD0F8B45925A328" bold="true" box="[816,1069,1810,1834]" pageId="7" pageNumber="6">Metasoma and telson</emphasis>
(
<figureCitation id="133958A0FFF9FF8BFBA1F8B459C1A328" box="[1089,1225,1810,1834]" captionStart="Figures 2835" captionStartId="7.[116,193,1516,1538]" captionTargetBox="[117,1469,270,1496]" captionTargetId="figure-242@7.[113,1474,234,1502]" captionTargetPageId="7" captionText="Figures 2835: Barbaracurus feti sp. n. Figures 28, 3032. Holotype male, metasoma V and telson lateral view (28), metasoma and telson, lateral (30), dorsal (31) and ventral (32) views. Figures 29, 3335. Paratype female, metasoma V and telson lateral view (29), metasoma and telson, lateral (33), dorsal (34) ventral and sternite VII (35) views. Scale bar: 10 mm (3032, 3335)." figureDoi="http://doi.org/10.5281/zenodo.4648246" httpUri="https://zenodo.org/record/4648246/files/figure.png" pageId="7" pageNumber="6">Figs. 2835</figureCitation>
). Segments IIV with granulate, completely developed carinae, segment V with carinae indicated in both sexes. The carinae are composed of minute, rounded, equal-sized, evenly spaced granules. The first metasomal segment has a total of 10 carinae, the second through fourth segments have eight carinae, and the fifth segment has five indicated carinae. All metasomal segments are very sparsely granulated. Metasoma is very sparsely hirsute. Telson smooth with only a weakly indicated ventral carina and a dense cover of long setae mainly on the ventral surface. Subaculear tubercle short and pointed. Vesicle elongate, ellipsoidal or pyriform, telson length/depth ratio
<quantity id="4CFAE9C0FFF5FF87FD34FECD5D8EA5A1" metricMagnitude="-2" metricUnit="m" metricValue="8.2296" pageId="11" pageNumber="10" unit="in" value="3.24">3.24 in</quantity>
both sexes. Aculeus slender, curved, shorter than vesicle.
</paragraph>
<caption id="DF7D14ADFFF6FF84FF91F87F595AA3ED" ID-DOI="http://doi.org/10.5281/zenodo.4648248" ID-Zenodo-Dep="4648248" box="[113,1106,2009,2031]" httpUri="https://zenodo.org/record/4648248/files/figure.png" pageId="8" pageNumber="7" startId="8.[113,190,2009,2031]" targetBox="[113,1474,234,1993]" targetPageId="8">
<paragraph id="8BBD4425FFF6FF84FF91F87F595AA3ED" blockId="8.[113,1106,2009,2031]" box="[113,1106,2009,2031]" pageId="8" pageNumber="7">
<emphasis id="B9769837FFF6FF84FF91F87F5C08A3EC" bold="true" box="[113,256,2009,2031]" pageId="8" pageNumber="7">Figures 3637</emphasis>
:
<taxonomicName id="4C023FA6FFF6FF84FEEDF87F5CB1A3EC" authority="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre, 2019" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre" authorityYear="2019" box="[269,441,2009,2030]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="feti" status="sp. nov.">
<emphasis id="B9769837FFF6FF84FEEDF87F5CB1A3EC" box="[269,441,2009,2030]" italics="true" pageId="8" pageNumber="7">Barbaracurus feti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245254CFFF6FF84FE5EF87F5CFBA3ED" box="[446,499,2009,2031]" pageId="8" pageNumber="7" rank="species">
<emphasis id="B9769837FFF6FF84FE5EF87F5CDDA3EC" bold="true" box="[446,469,2009,2030]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="B9769837FFF6FF84FE01F87F5CE6A3EC" bold="true" box="[481,494,2009,2030]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
, in vivo habitus. Male holotype (36) and female paratype (37).
</paragraph>
</caption>
<caption id="DF7D14ADFFF7FF85FF93F8225936A3F2" ID-DOI="http://doi.org/10.5281/zenodo.4648250" ID-Zenodo-Dep="4648250" httpUri="https://zenodo.org/record/4648250/files/figure.png" pageId="9" pageNumber="8" startId="9.[115,192,1924,1946]" targetBox="[113,1466,237,1904]" targetPageId="9">
<paragraph id="8BBD4425FFF7FF85FF93F8225936A3F2" blockId="9.[115,1473,1924,2032]" pageId="9" pageNumber="8">
<emphasis id="B9769837FFF7FF85FF93F8225C0AA39B" bold="true" box="[115,258,1924,1946]" pageId="9" pageNumber="8">Figures 3844</emphasis>
:
<taxonomicName id="4C023FA6FFF7FF85FEEFF8225CB3A39B" authority="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre, 2019" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre" authorityYear="2019" box="[271,443,1924,1945]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="feti" status="sp. nov.">
<emphasis id="B9769837FFF7FF85FEEFF8225CB3A39B" box="[271,443,1924,1945]" italics="true" pageId="9" pageNumber="8">Barbaracurus feti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245254CFFF7FF85FE20F8225CFFA398" box="[448,503,1924,1946]" pageId="9" pageNumber="8" rank="species">
<emphasis id="B9769837FFF7FF85FE20F8225CDFA39B" bold="true" box="[448,471,1924,1945]" pageId="9" pageNumber="8">sp</emphasis>
.
<emphasis id="B9769837FFF7FF85FE03F8225CF8A39B" bold="true" box="[483,496,1924,1945]" pageId="9" pageNumber="8">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B9769837FFF7FF85FE1DF8225F84A39B" bold="true" box="[509,652,1924,1946]" pageId="9" pageNumber="8">Figures 3842</emphasis>
: Right hemispermatophore of male holotype.
<emphasis id="B9769837FFF7FF85FBA8F82259A2A39B" bold="true" box="[1096,1194,1924,1946]" pageId="9" pageNumber="8">Figure 38</emphasis>
. Whole hemispermatophore, convex view. Scale bar: 1 mm.
<emphasis id="B9769837FFF7FF85FE7EF8075F25A3B4" bold="true" box="[414,557,1953,1975]" pageId="9" pageNumber="8">Figures 3942</emphasis>
. Capsule in convex compressed (39), convex (40), anterior (41) and posterior (42) views. Scale bar: 500 μm.
<emphasis id="B9769837FFF7FF85FF12F81B5C89A3D1" bold="true" box="[242,385,1981,2003]" pageId="9" pageNumber="8">Figures 4344</emphasis>
: Mitotic metaphase (43) and ideogram (44) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values) of male holotype. Arrows show one extra large odd chromosome. Scale bar: 5 μm.
</paragraph>
</caption>
<caption id="DF7D14ADFFF4FF86FF92F81B5981A3F2" ID-DOI="http://doi.org/10.5281/zenodo.4648252" ID-Zenodo-Dep="4648252" httpUri="https://zenodo.org/record/4648252/files/figure.png" pageId="10" pageNumber="9" startId="10.[114,191,1981,2003]" targetBox="[113,1474,234,1971]" targetPageId="10">
<paragraph id="8BBD4425FFF4FF86FF92F81B5981A3F2" blockId="10.[114,1472,1981,2032]" pageId="10" pageNumber="9">
<emphasis id="B9769837FFF4FF86FF92F81B5C08A3D0" bold="true" box="[114,256,1981,2003]" pageId="10" pageNumber="9">Figures 4546</emphasis>
:
<emphasis id="B9769837FFF4FF86FEEDF81B5C66A3D0" bold="true" box="[269,366,1981,2003]" pageId="10" pageNumber="9">Figure 45</emphasis>
. Map showing confirmed distribution of
<taxonomicName id="4C023FA6FFF4FF86FD17F81B5E76A3D0" authorityName="Kovarik, Lowe et Stahlavsky" authorityYear="2018" box="[759,894,1981,2002]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9769837FFF4FF86FD17F81B5E76A3D0" box="[759,894,1981,2002]" italics="true" pageId="10" pageNumber="9">Barbaracurus</emphasis>
</taxonomicName>
spp. For exact position of Rugay village, the type locality of
<taxonomicName id="4C023FA6FFF4FF86FF92F87C5C16A3ED" authority="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre, 2019" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre" authorityYear="2019" box="[114,286,2010,2031]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="feti" status="sp. nov.">
<emphasis id="B9769837FFF4FF86FF92F87C5C16A3ED" box="[114,286,2010,2031]" italics="true" pageId="10" pageNumber="9">Barbaracurus feti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245254CFFF4FF86FEC3F87C5C51A3ED" box="[291,345,2010,2031]" pageId="10" pageNumber="9" rank="species">
<emphasis id="B9769837FFF4FF86FEC3F87C5C32A3ED" bold="true" box="[291,314,2010,2031]" pageId="10" pageNumber="9">sp</emphasis>
.
<emphasis id="B9769837FFF4FF86FEA6F87C5C5BA3ED" bold="true" box="[326,339,2010,2031]" pageId="10" pageNumber="9">n</emphasis>
.
</taxonomicNameLabel>
see fig. 84 in
<bibRefCitation id="EF9339D4FFF4FF86FE00F87C5FAAA3ED" author="KOVARIK" box="[480,674,2010,2032]" pageId="10" pageNumber="9" pagination="1 - 18" refId="ref4696" refString="KOVARIK, F., G. LOWE &amp; T. MAZUCH. 2019. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XIX. Pandiborellius meidensis (Karsch, 1879) and Pandinurus fulvipes sp. n. (Scorpionidae) from Somaliland. Euscorpius, 275: 1 - 18." type="journal article" year="2019">Kovařík et al., 2019</bibRefCitation>
.
<emphasis id="B9769837FFF4FF86FD4BF87F5E05A3EC" bold="true" box="[683,781,2009,2031]" pageId="10" pageNumber="9">Figure 46</emphasis>
.
<taxonomicName id="4C023FA6FFF4FF86FCF9F87C5ECDA3ED" authority="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre, 2019" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre" authorityYear="2019" box="[793,965,2010,2031]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="feti" status="sp. nov.">
<emphasis id="B9769837FFF4FF86FCF9F87C5ECDA3ED" box="[793,965,2010,2031]" italics="true" pageId="10" pageNumber="9">Barbaracurus feti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245254CFFF4FF86FC2AF87C5EF7A3ED" box="[970,1023,2010,2031]" pageId="10" pageNumber="9" rank="species">
<emphasis id="B9769837FFF4FF86FC2AF87C5EE9A3ED" bold="true" box="[970,993,2010,2031]" pageId="10" pageNumber="9">sp</emphasis>
.
<emphasis id="B9769837FFF4FF86FC0DF87C5EF2A3ED" bold="true" box="[1005,1018,2010,2031]" pageId="10" pageNumber="9">n</emphasis>
.
</taxonomicNameLabel>
, type locality.
</paragraph>
</caption>
<paragraph id="8BBD4425FFF5FF87FF91FE0D5CACA081" blockId="11.[113,772,235,1155]" pageId="11" pageNumber="10">
<emphasis id="B9769837FFF5FF87FF91FE0D5DE1A5C1" bold="true" box="[113,233,427,451]" pageId="11" pageNumber="10">Karyotype</emphasis>
(
<figureCitation id="133958A0FFF5FF87FF1EFE0D5C8DA5C1" box="[254,389,427,451]" captionStart="Figures 3844" captionStartId="9.[115,192,1924,1946]" captionTargetBox="[113,1466,237,1904]" captionTargetId="figure-8@9.[113,1474,231,1912]" captionTargetPageId="9" captionText="Figures 3844: Barbaracurus feti sp. n. Figures 3842: Right hemispermatophore of male holotype. Figure 38. Whole hemispermatophore, convex view. Scale bar: 1 mm. Figures 3942. Capsule in convex compressed (39), convex (40), anterior (41) and posterior (42) views. Scale bar: 500 μm. Figures 4344: Mitotic metaphase (43) and ideogram (44) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values) of male holotype. Arrows show one extra large odd chromosome. Scale bar: 5 μm." figureDoi="http://doi.org/10.5281/zenodo.4648250" httpUri="https://zenodo.org/record/4648250/files/figure.png" pageId="11" pageNumber="10">Figs. 4344</figureCitation>
). We analyzed the male
<typeStatus id="54B9FA87FFF5FF87FD40FE0D5E08A5C1" box="[672,768,427,451]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
. The diploid number of this specimen is 23 chromosomes. The lengths of first two chromosomes are similar and these two chromosomes correspond to the first chromosome pair that is distinctly longer (over 10% of the diploid set) than the remaining chromosomes. The third chromosome also has a distinctive length (7.67% of the diploid set) compared to the following chromosomes that gradually decrease in length from 5.23 to 1.92% of the diploid set (
<figureCitation id="133958A0FFF5FF87FE9AFD0D5CC1A6C1" box="[378,457,683,707]" captionStart="Figures 3844" captionStartId="9.[115,192,1924,1946]" captionTargetBox="[113,1466,237,1904]" captionTargetId="figure-8@9.[113,1474,231,1912]" captionTargetPageId="9" captionText="Figures 3844: Barbaracurus feti sp. n. Figures 3842: Right hemispermatophore of male holotype. Figure 38. Whole hemispermatophore, convex view. Scale bar: 1 mm. Figures 3942. Capsule in convex compressed (39), convex (40), anterior (41) and posterior (42) views. Scale bar: 500 μm. Figures 4344: Mitotic metaphase (43) and ideogram (44) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values) of male holotype. Arrows show one extra large odd chromosome. Scale bar: 5 μm." figureDoi="http://doi.org/10.5281/zenodo.4648250" httpUri="https://zenodo.org/record/4648250/files/figure.png" pageId="11" pageNumber="10">Fig. 44</figureCitation>
). This, the third chromosome, is probably the cause of the odd diploid number in this species (2n=23) as an effect of fusions/fissions of this chromosome. These types of chromosomal rearrangements are frequently detected in buthid scorpions with holocentric chromosomes (e. g.
<bibRefCitation id="EF9339D4FFF5FF87FF4EFCED5C71A761" author="MATTOS, V. F." box="[174,377,843,867]" pageId="11" pageNumber="10" pagination="121 - 136" refId="ref5032" refString="MATTOS, V. F., D. M. CELLA, L. S. CARVALHO, D. M. CANDIDO &amp; M. C. SCHNEIDER. 2013. High chromosome variability and the presence of multivalent associations in buthid scorpions. Chromosome Research, 21: 121 - 136." type="journal article" year="2013">Mattos et al., 2013</bibRefCitation>
). Unfortunately, we did not observed meiosis in
<taxonomicName id="4C023FA6FFF5FF87FF3FFCCD5C15A781" authority="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre, 2019" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre" authorityYear="2019" box="[223,285,875,899]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="feti" status="sp. nov.">
<emphasis id="B9769837FFF5FF87FF3FFCCD5C15A781" box="[223,285,875,899]" italics="true" pageId="11" pageNumber="10">B. feti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245254CFFF5FF87FEC1FCCD5C53A780" box="[289,347,875,899]" pageId="11" pageNumber="10" rank="species">
<emphasis id="B9769837FFF5FF87FEC1FCCD5C32A781" bold="true" box="[289,314,875,899]" pageId="11" pageNumber="10">sp</emphasis>
.
<emphasis id="B9769837FFF5FF87FEA5FCCD5C5CA781" bold="true" box="[325,340,875,899]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
This is the reason why we cannot specify the possible presence of multivalent association as was already described in
<taxonomicName id="4C023FA6FFF5FF87FEE6FC0D5F3AA7C1" box="[262,562,939,963]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="sofomarensis">
<emphasis id="B9769837FFF5FF87FEE6FC0D5F3AA7C1" box="[262,562,939,963]" italics="true" pageId="11" pageNumber="10">Barbaracurus sofomarensis</emphasis>
</taxonomicName>
(2n=22) (
<bibRefCitation id="EF9339D4FFF5FF87FD4BFC0D5DE6A7E0" author="KOVARIK" pageId="11" pageNumber="10" pagination="1 - 31" refId="ref4754" refString="KOVARIK, F., G. LOWE, M. SEITER, J. PLISKOVA &amp; F. STAHLAVSKY. 2015. Scorpions of Ethiopia (Arachnida: Scorpiones), Part II. Genus Babycurus Karsch, 1886 (Buthidae), with description of two new species. Euscorpius, 196: 1 - 31." type="journal article" year="2015">Kovařík et al., 2015</bibRefCitation>
). These two species have similar karyotypes with different numbers of considerably longer chromosomes. In contrast to this, the karyotypes of
<taxonomicName id="4C023FA6FFF5FF87FE0AFBAD5E0BA021" baseAuthorityName="Borelli" baseAuthorityYear="1902" box="[490,771,1035,1059]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="zambonellii">
<emphasis id="B9769837FFF5FF87FE0AFBAD5E0BA021" box="[490,771,1035,1059]" italics="true" pageId="11" pageNumber="10">Barbaracurus zambonellii</emphasis>
</taxonomicName>
(2n=26) and
<taxonomicName id="4C023FA6FFF5FF87FF1DFB8D5F0CA041" authorityName="Somaliland" baseAuthorityName="Hirst" baseAuthorityYear="1907" box="[253,516,1067,1091]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="somalicus">
<emphasis id="B9769837FFF5FF87FF1DFB8D5F0CA041" box="[253,516,1067,1091]" italics="true" pageId="11" pageNumber="10">Barbaracurus somalicus</emphasis>
</taxonomicName>
(2n=36) contain higher numbers of chromosomes and all of them gradually decrease in length (
<bibRefCitation id="EF9339D4FFF5FF87FF21FBCD5C90A081" author="KOVARIK" box="[193,408,1131,1155]" pageId="11" pageNumber="10" pagination="1 - 41" refId="ref4813" refString="KOVARIK, F., G. LOWE &amp; F. STAHLAVSKY. 2018. Review of the genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), with description of Barbaracurus gen. n. and two new species from Oman and Yemen. Euscorpius, 267: 1 - 41." type="journal article" year="2018">Kovařík et al., 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31817AEFFF5FF87FF91FB0D5CD3A201" pageId="11" pageNumber="10" type="discussion">
<paragraph id="8BBD4425FFF5FF87FF91FB0D5CD3A201" blockId="11.[113,772,1195,1539]" pageId="11" pageNumber="10">
COMMENTS ON LOCALITIES AND LIFE STRATEGY. The
<typeStatus id="54B9FA87FFF5FF87FD35FB0D5E0BA0C1" box="[725,771,1195,1219]" pageId="11" pageNumber="10">type</typeStatus>
locality, 18SE is rocky semi-desert (
<figureCitation id="133958A0FFF5FF87FDEBFB6D5F55A0E1" box="[523,605,1227,1251]" captionStart="Figures 4546" captionStartId="10.[114,191,1981,2003]" captionTargetBox="[113,1474,234,1971]" captionTargetId="figure-17@10.[113,1474,234,1973]" captionTargetPageId="10" captionText="Figures 4546: Figure 45. Map showing confirmed distribution of Barbaracurus spp. For exact position of Rugay village, the type locality of Barbaracurus feti sp. n. see fig. 84 in Kovařík et al., 2019. Figure 46. Barbaracurus feti sp. n., type locality." figureDoi="http://doi.org/10.5281/zenodo.4648252" httpUri="https://zenodo.org/record/4648252/files/figure.png" pageId="11" pageNumber="10">Fig. 46</figureCitation>
and fig.
<quantity id="4CFAE9C0FFF5FF87FD28FB6D5E0BA0E1" box="[712,771,1227,1251]" metricMagnitude="0" metricUnit="m" metricValue="1.0668" pageId="11" pageNumber="10" unit="in" value="42.0">42 in</quantity>
<bibRefCitation id="EF9339D4FFF5FF87FF91FB4D5C5AA101" author="KOVARIK" box="[113,338,1259,1283]" pageId="11" pageNumber="10" pagination="1 - 18" refId="ref4696" refString="KOVARIK, F., G. LOWE &amp; T. MAZUCH. 2019. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XIX. Pandiborellius meidensis (Karsch, 1879) and Pandinurus fulvipes sp. n. (Scorpionidae) from Somaliland. Euscorpius, 275: 1 - 18." type="journal article" year="2019">Kovařík et al., 2019</bibRefCitation>
). The
<typeStatus id="54B9FA87FFF5FF87FE79FB4D5CD9A101" box="[409,465,1259,1283]" pageId="11" pageNumber="10">types</typeStatus>
of
<taxonomicName id="4C023FA6FFF5FF87FE1AFB4D5FB3A101" authority="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre, 2019" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský &amp; Hurre" authorityYear="2019" box="[506,699,1259,1283]" class="Arachnida" family="Buthidae" genus="Barbaracurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="feti" status="sp. nov.">
<emphasis id="B9769837FFF5FF87FE1AFB4D5FB3A101" box="[506,699,1259,1283]" italics="true" pageId="11" pageNumber="10">Barbaracurus feti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245254CFFF5FF87FD24FB4D5E0BA100" box="[708,771,1259,1283]" pageId="11" pageNumber="10" rank="species">
<emphasis id="B9769837FFF5FF87FD24FB4D5FD5A101" bold="true" box="[708,733,1259,1283]" pageId="11" pageNumber="10">sp</emphasis>
.
<emphasis id="B9769837FFF5FF87FD0EFB4D5FF5A101" bold="true" box="[750,765,1259,1283]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
were recorded at night during UV collecting together with
<taxonomicName id="4C023FA6FFF5FF87FF91FA8D5DE2A140" box="[113,234,1323,1346]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B9769837FFF5FF87FF91FA8D5DE2A140" box="[113,234,1323,1346]" italics="true" pageId="11" pageNumber="10">Hottentotta</emphasis>
</taxonomicName>
sp. and
<taxonomicName id="4C023FA6FFF5FF87FEA6FA8D5E08A141" authority="(Karsch, 1879)" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[326,768,1323,1347]" class="Arachnida" family="Scorpionidae" genus="Pandiborellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="meidensis">
<emphasis id="B9769837FFF5FF87FEA6FA8D5F5CA141" box="[326,596,1323,1347]" italics="true" pageId="11" pageNumber="10">Pandiborellius meidensis</emphasis>
(Karsch, 1879)
</taxonomicName>
. The first author (F.K.) visited the locality on
<date id="FFBC62E5FFF5FF87FD8BFAEC5DAEA181" pageId="11" pageNumber="10" value="2018-08-24" valueMax="2018-08-25" valueMin="2018-08-24">2425 August 2018</date>
and recorded maximum daytime temperatures of 38 ºC and a minimum nighttime temperature of 32 ºC. The recorded humidity was between 20% (minimum at day) and 33% (maximum at night). More information about the locality is available in
<bibRefCitation id="EF9339D4FFF5FF87FF14FA4D5CDFA201" author="KOVARIK" box="[244,471,1515,1539]" pageId="11" pageNumber="10" pagination="1 - 18" refId="ref4696" refString="KOVARIK, F., G. LOWE &amp; T. MAZUCH. 2019. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XIX. Pandiborellius meidensis (Karsch, 1879) and Pandinurus fulvipes sp. n. (Scorpionidae) from Somaliland. Euscorpius, 275: 1 - 18." type="journal article" year="2019">Kovařík et al. (2019)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>