plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/98/87/039887FDFFD6FFB8783B1CB27293FE65.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

321 lines
38 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="DB776A28A7B3D0F7B0E430F9C7FC1A85" ID-CLB-Dataset="57159" ID-DOI="10.31610/zsr/2019.28.1.24" ID-GBIF-Dataset="2b690c17-f7fc-4c69-83d1-1d9047c876c9" ID-ISSN="2410-0226" ID-Zenodo-Dep="10124208" ID-ZooBank="urn:lsid:zoobank.org:pub:D82459C6-A090-4203-8D28-8BE062383D2B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1688479652846" checkinUser="felipe" docAuthor="Shamshev, I. V." docDate="2019" docId="039887FDFFD6FFB8783B1CB27293FE65" docLanguage="en" docName="ZoosystRoss.28.1.24-41.pdf" docOrigin="Zoosystematica Rossica (Zoosyst. Rossica) 28 (1)" docSource="http://dx.doi.org/10.31610/zsr/2019.28.1.24" docStyle="DocumentStyle:8B7E50B51175314E898B46063364B751.1:ZoosystRoss.2019-.journal_article" docStyleId="8B7E50B51175314E898B46063364B751" docStyleName="ZoosystRoss.2019-.journal_article" docStyleVersion="1" docTitle="Empis shebalinensis Shamshev 2019, sp. nov." docType="treatment" docVersion="7" lastPageNumber="41" masterDocId="FFA1FF85FFD8FFA97B121C0A7019FF88" masterDocTitle="Six new species of the genus Empis (Diptera: Empididae) from the Altai Mountains of Russia" masterLastPageNumber="41" masterPageNumber="24" pageNumber="38" updateTime="1700049859112" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="CF2466FEA1881348D07DC31072DE8D20" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E0D3844B4EBAC9B973B4A63C4D72C142">
<mods:title id="D82C2F8BB1E27F7242218030980E27B4">Six new species of the genus Empis (Diptera: Empididae) from the Altai Mountains of Russia</mods:title>
</mods:titleInfo>
<mods:name id="2151E4D0644F55AFDEF6BE0D5D0182CA" type="personal">
<mods:role id="C04E6DC107908858A8E791C6365DF170">
<mods:roleTerm id="99362CE047AD6AB67FC0609042A6004E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9A527E5F26F2FEA8B45F0AC689A6A062">Shamshev, I. V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="E5A1EFA527735E1A927686085485CB62">text</mods:typeOfResource>
<mods:relatedItem id="BD2957E0967BB991BDBADF7A70A89D68" type="host">
<mods:titleInfo id="23DB0D2CB69D62096E65C17510F952B6">
<mods:title id="DB158B3EC5192F81731B05140592E79A">Zoosystematica Rossica</mods:title>
</mods:titleInfo>
<mods:part id="A2EFE1A7DFB698EAC0B1AD584E000FBA">
<mods:date id="69F028FBA068FD584255E6E39E85D95D">2019</mods:date>
<mods:detail id="66DC0F8CDC42F1AB8F1D4C854D1D986A" type="series">
<mods:title id="ED994783539FE10F2402BE0C3F22A434">Zoosyst. Rossica</mods:title>
</mods:detail>
<mods:detail id="2E7D417B4978F204C3BA8C14DCCE0A5A" type="pubDate">
<mods:number id="F57F27E9E84EECD5A80B98DFC417983D">2019-04-18</mods:number>
</mods:detail>
<mods:detail id="ED0481041336D3AB427B5356F07783EB" type="volume">
<mods:number id="DE5C35ECC1D0F7A064861EC351B88C67">28</mods:number>
</mods:detail>
<mods:detail id="E823FF3A4E43997FC9ACCEEE57D9A0BF" type="issue">
<mods:number id="A90A60572C90902828075DAACE8FFC58">1</mods:number>
</mods:detail>
<mods:extent id="EBD56C1B5ACB01A21E4BE3F37F0D2E56" unit="page">
<mods:start id="777E86B57A0AF76A90462E2B7300B10E">24</mods:start>
<mods:end id="44CE39CDE824798D454BF82A60028DE8">41</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="80E9F959628EE78AE631FB1AB6E10929">
<mods:url id="728B68AFB2DB7915CAB37C560E7C786E">http://dx.doi.org/10.31610/zsr/2019.28.1.24</mods:url>
</mods:location>
<mods:classification id="8DFE48B5A5D02C366067F9CC262135F5">journal article</mods:classification>
<mods:identifier id="C208B94015FAE96257A8E598879C19E0" type="CLB-Dataset">57159</mods:identifier>
<mods:identifier id="4E55E10C2333517A2D0B1FD8E571021F" type="DOI">10.31610/zsr/2019.28.1.24</mods:identifier>
<mods:identifier id="421B60D4C89D7CE0185502E5ADE8B8A7" type="GBIF-Dataset">2b690c17-f7fc-4c69-83d1-1d9047c876c9</mods:identifier>
<mods:identifier id="17FE2F75E2DEEE05E401637EEFEBF722" type="ISSN">2410-0226</mods:identifier>
<mods:identifier id="C2739E2560C51D1EB2BFDB8F9AD95E45" type="Zenodo-Dep">10124208</mods:identifier>
<mods:identifier id="E22A4D2EC8904DE84294139956C9F66A" type="ZooBank">urn:lsid:zoobank.org:pub:D82459C6-A090-4203-8D28-8BE062383D2B</mods:identifier>
</mods:mods>
<treatment id="039887FDFFD6FFB8783B1CB27293FE65" ID-DOI="http://doi.org/10.5281/zenodo.8114723" ID-GBIF-Taxon="210810425" ID-Zenodo-Dep="8114723" LSID="urn:lsid:plazi:treatment:039887FDFFD6FFB8783B1CB27293FE65" httpUri="http://treatment.plazi.org/id/039887FDFFD6FFB8783B1CB27293FE65" lastPageId="17" lastPageNumber="41" pageId="14" pageNumber="38">
<subSubSection id="C32B6560FFD6FFA7783B1CB274B8FF5A" box="[809,1185,184,211]" pageId="14" pageNumber="38" type="nomenclature">
<paragraph id="8B8E36EBFFD6FFA7783B1CB274B8FF5A" blockId="14.[809,1185,184,211]" box="[809,1185,184,211]" pageId="14" pageNumber="38">
<heading id="D0C68187FFD6FFA7783B1CB274B8FF5A" box="[809,1185,184,211]" fontSize="11" level="2" pageId="14" pageNumber="38" reason="2">
<emphasis id="B945EAF9FFD6FFA7783B1CB274B8FF5A" bold="true" box="[809,1185,184,211]" pageId="14" pageNumber="38">
<taxonomicName id="4C314D68FFD6FFA7783B1CB2742CFF5B" authority="Shamshev, 2019" authorityName="Shamshev" authorityYear="2019" box="[809,1077,184,211]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="38" phylum="Arthropoda" rank="species" species="shebalinensis" status="sp. nov.">
<emphasis id="B945EAF9FFD6FFA7783B1CB2742CFF5B" bold="true" box="[809,1077,184,211]" italics="true" pageId="14" pageNumber="38">Empis shebalinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765782FFD6FFA77F2E1CB274B8FF5A" box="[1084,1185,184,210]" pageId="14" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32B6560FFD6FFB8783B1CD67293FE65" lastPageId="17" lastPageNumber="41" pageId="14" pageNumber="38" type="description">
<paragraph id="8B8E36EBFFD6FFA7783B1CD673D5FF7F" blockId="14.[809,972,219,247]" box="[809,972,219,247]" pageId="14" pageNumber="38">
(
<figureCitation id="130A2A6EFFD6FFA778201CD673DAFF7F" box="[818,963,219,247]" captionStart="Figs16–18" captionStartId="15.[159,212,1832,1856]" captionTargetBox="[234,1359,196,1803]" captionTargetId="figure-22@15.[165,1422,181,1807]" captionTargetPageId="15" captionText="Figs16–18. Empis shebalinensis sp.nov., male. 16, habitus, lateral view (holotype); 17, postabdomen, lateral view; 18, terminalia, lateral view. Abbreviations: cerc, cercus; ej apod, ejaculatory apodeme; epand, epandrium; hypd, hypandrium; ph, phallus. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135545" httpUri="https://zenodo.org/record/10135545/files/figure.png" pageId="14" pageNumber="38">Figs 16–18</figureCitation>
)
</paragraph>
<paragraph id="8B8E36EBFFD6FFA7785D1D1F74BEFE39" blockId="14.[809,1429,277,1927]" pageId="14" pageNumber="38">
<materialsCitation id="3B593CB6FFD6FFA7785D1D1F74BEFE39" ID-GBIF-Occurrence="4144857312" collectingDate="2014-07-04" collectingDateMax="2014-07-04" collectingDateMin="2014-07-03" collectionCode="ZIN" collectorName="I. Shamshev" country="Russia" elevation="1325" latitude="51.1" location="Topuchaya" longLatPrecision="1092" longitude="85.61667" pageId="14" pageNumber="38" specimenCode="INS_DIP_0000756" specimenCount="1" stateProvince="Republic of Altai" typeStatus="holotype">
<emphasis id="B945EAF9FFD6FFA7785D1D1F73ACFEA6" box="[847,949,277,302]" italics="true" pageId="14" pageNumber="38">
<typeStatus id="548A8849FFD6FFA7785D1D1F73ACFEA6" box="[847,949,277,302]" pageId="14" pageNumber="38" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCount id="9D37FD62FFD6FFA778D11D1F7419FEA6" box="[963,1024,277,302]" pageId="14" pageNumber="38" type="male">Male</specimenCount>
, “
<collectingCountry id="F326767BFFD6FFA77F051D1C7498FEA6" box="[1047,1153,278,302]" name="Russia" pageId="14" pageNumber="38">RUSSIA</collectingCountry>
, Altai Rep. [=
<collectingRegion id="49F5F809FFD6FFA77E3C1D1F739AFEC7" country="Russia" name="Altay" pageId="14" pageNumber="38">Republic of Altai</collectingRegion>
], / Shebalino distr.,
<locationDeviation id="9A3C2FC9FFD6FFA77F751D3C74DEFEC6" box="[1127,1223,310,335]" pageId="14" pageNumber="38">
<quantity id="4CC99B0EFFD6FFA77F751D3C7487FEC7" box="[1127,1182,310,335]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="38" unit="km" value="5.0">5 km</quantity>
SE
</locationDeviation>
vill. /
<location id="8EEE6030FFD6FFA77E001D3C7588FEC7" LSID="urn:lsid:plazi:treatment:039887FDFFD6FFB8783B1CB27293FE65:8EEE6030FFD6FFA77E001D3C7588FEC7" box="[1298,1425,310,335]" country="Russia" latitude="51.1" longLatPrecision="1092" longitude="85.61667" name="Topuchaya" pageId="14" pageNumber="38" stateProvince="Republic of Altai">Topuchaya</location>
,
<quantity id="4CC99B0EFFD6FFA7783B1D5C7366FEE7" box="[809,895,342,367]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="14" pageNumber="38" unit="m" value="1325.0">
<elevation id="001CD1D8FFD6FFA7783B1D5C7366FEE7" box="[809,895,342,367]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="14" pageNumber="38" unit="m" value="1325.0">1325 m</elevation>
</quantity>
, /
<geoCoordinate id="EE05502CFFD6FFA7788F1D5C73E0FEE7" box="[925,1017,342,367]" degrees="51" direction="north" minutes="06" orientation="latitude" pageId="14" pageNumber="38" precision="925" value="51.1">51°06'N</geoCoordinate>
<geoCoordinate id="EE05502CFFD6FFA778ED1D5C7444FEE7" box="[1023,1117,342,367]" degrees="85" direction="east" minutes="37" orientation="longitude" pageId="14" pageNumber="38" precision="925" value="85.61667">85°37'E</geoCoordinate>
,
<collectingDate id="EFCBE9C3FFD6FFA77F7B1D5D74E3FEE7" box="[1129,1274,343,367]" pageId="14" pageNumber="38" value="2014-07-04">
3–
<date id="FF8F102BFFD6FFA77F911D5D74E3FEE7" box="[1155,1274,343,367]" pageId="14" pageNumber="38" value="2014-07-04">4.07.2014</date>
</collectingDate>
, leg.
<collectorName id="26C4533DFFD6FFA77E211D527347FE18" pageId="14" pageNumber="38">I. Shamshev</collectorName>
”, “
<taxonomicName id="4C314D68FFD6FFA778921D7274FFFE18" authority="Shamshev" authorityName="Shamshev" authorityYear="2019" box="[896,1254,375,400]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="38" phylum="Arthropoda" rank="species" species="shebalinensis" status="sp. nov.">
<emphasis id="B945EAF9FFD6FFA778921D72747FFE18" box="[896,1126,375,400]" italics="true" pageId="14" pageNumber="38">Empis shebalinensis</emphasis>
Shamshev
</taxonomicName>
,
<taxonomicNameLabel id="A2765782FFD6FFA77FE61D72752AFE07" box="[1268,1331,376,399]" pageId="14" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
[red label]” (
<collectionCode id="ED20AE2EFFD6FFA7787F1D9273BBFE38" box="[877,930,408,432]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="14" pageNumber="38" type="Museum">ZIN</collectionCode>
,
<specimenCode id="DB979E90FFD6FFA778BC1D927483FE39" box="[942,1178,408,433]" pageId="14" pageNumber="38">INS_DIP_0000756</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B8E36EBFFD6FFA7785D1DB0757EFDBB" blockId="14.[809,1429,277,1927]" pageId="14" pageNumber="38">
<materialsCitation id="3B593CB6FFD6FFA7785D1DB073C2FE7A" ID-GBIF-Occurrence="4144857317" collectingDate="2014-07-04" collectionCode="ZIN, INS" collectorName="I. Shamshev" country="Russia" elevation="1325" latitude="51.1" location="Russia" longLatPrecision="1092" longitude="85.61667" pageId="14" pageNumber="38" specimenCount="12" specimenCount-female="8" specimenCount-male="4" stateProvince="Altay" typeStatus="paratype">
<emphasis id="B945EAF9FFD6FFA7785D1DB073D8FE59" box="[847,961,442,465]" italics="true" pageId="14" pageNumber="38">
<typeStatus id="548A8849FFD6FFA7785D1DB073D8FE59" box="[847,961,442,465]" pageId="14" pageNumber="38" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<emphasis id="B945EAF9FFD6FFA778C11DB37434FE59" bold="true" box="[979,1069,441,465]" pageId="14" pageNumber="38">
<collectingCountry id="F326767BFFD6FFA778C11DB37431FE59" box="[979,1064,441,465]" name="Russia" pageId="14" pageNumber="38">Russia</collectingCountry>
:
</emphasis>
<specimenCount id="9D37FD62FFD6FFA77F2B1DB37492FE59" box="[1081,1163,440,465]" pageId="14" pageNumber="38" type="male">1 male</specimenCount>
,
<specimenCount id="9D37FD62FFD6FFA77F8E1DB37514FE59" box="[1180,1293,440,465]" pageId="14" pageNumber="38" type="female">8 females</specimenCount>
, same data as for holotype;
</materialsCitation>
<materialsCitation id="3B593CB6FFD6FFA778F11DD0750EFE7A" ID-GBIF-Occurrence="4144857320" box="[995,1303,473,498]" collectingDate="2014-07-04" collectionCode="ZIN, INS" collectorName="I. Shamshev" country="Russia" elevation="1325" latitude="51.1" location="Russia" longLatPrecision="1092" longitude="85.61667" pageId="14" pageNumber="38" specimenCount="12" specimenCount-female="8" specimenCount-male="4" stateProvince="Altay" typeStatus="paratype">
<specimenCount id="9D37FD62FFD6FFA778F11DD07423FE7A" box="[995,1082,473,498]" pageId="14" pageNumber="38" type="male">2 males</specimenCount>
(
<specimenCount id="9D37FD62FFD6FFA77F5F1DD0748EFE7A" box="[1101,1175,473,498]" pageId="14" pageNumber="38" type="male">1 male</specimenCount>
dissected)
</materialsCitation>
,
<materialsCitation id="3B593CB6FFD6FFA77E311DD3757AFDBB" ID-GBIF-Occurrence="4144857311" collectingDate="2005-06-28" collectingDateMax="2005-06-29" collectingDateMin="2005-06-28" collectionCode="ZIN" collectorName="D. Kostrov" country="Russia" location="Tigirek Vill" municipality="Krasnoshchekovskiy Distr" pageId="14" pageNumber="38" specimenCount="5" specimenCount-female="5" stateProvince="Altai Terr." typeStatus="paratype">
<specimenCount id="9D37FD62FFD6FFA77E311DD37589FE7A" box="[1315,1424,473,498]" pageId="14" pageNumber="38" type="female">5 females</specimenCount>
,
<emphasis id="B945EAF9FFD6FFA7783B1DF07385FD9A" box="[809,924,506,531]" italics="true" pageId="14" pageNumber="38">
<collectingRegion id="49F5F809FFD6FFA7783B1DF07385FD9A" box="[809,924,506,531]" pageId="14" pageNumber="38">Altai Terr.</collectingRegion>
</emphasis>
,
<collectingMunicipality id="6BEAAC91FFD6FFA778BB1DF074C4FD9B" box="[937,1245,506,531]" pageId="14" pageNumber="38">Krasnoshchekovskiy Distr</collectingMunicipality>
., environs of
<location id="8EEE6030FFD6FFA77E601DF07382FDBB" LSID="urn:lsid:plazi:treatment:039887FDFFD6FFB8783B1CB27293FE65:8EEE6030FFD6FFA77E601DF07382FDBB" country="Russia" municipality="Krasnoshchekovskiy Distr" name="Tigirek Vill" pageId="14" pageNumber="38" stateProvince="Altai Terr.">Tigirek Vill</location>
.,
<date id="FF8F102BFFD6FFA778BB1E107442FDBB" box="[937,1115,538,563]" pageId="14" pageNumber="38" value="2005-06-28" valueMax="2005-06-29" valueMin="2005-06-28">
<collectingDate id="EFCBE9C3FFD6FFA778BB1E107442FDBB" box="[937,1115,538,563]" pageId="14" pageNumber="38" value="2005-06-28" valueMax="2005-06-29" valueMin="2005-06-28">28–29.VI.2005</collectingDate>
</date>
,
<collectorName id="26C4533DFFD6FFA77F771E1674FDFDBB" box="[1125,1252,540,563]" pageId="14" pageNumber="38">D. Kostrov</collectorName>
leg. (
<collectionCode id="ED20AE2EFFD6FFA77E331E117543FDBB" box="[1313,1370,539,563]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="14" pageNumber="38" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8E36EBFFD6FFA7785D1E347390FCA6" blockId="14.[809,1429,277,1927]" pageId="14" pageNumber="38">
<emphasis id="B945EAF9FFD6FFA7785D1E3473D2FDDF" box="[847,971,574,599]" italics="true" pageId="14" pageNumber="38">Diagnosis</emphasis>
. Mid-sized (body length about
<quantity id="4CC99B0EFFD6FFA7783B1E6A7391FDF3" box="[809,904,608,636]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="38" unit="mm" value="4.5">4.5 mm</quantity>
) blackish brown species with fairly slender legs lacking strong setae, and with black halters. Male: scutum and abdomen velvet-brown, wings brownish. Female: scutum with four brownish vittae, abdomen greyish pruinose, wings faintly infuscate.
</paragraph>
<paragraph id="8B8E36EBFFD6FFA7785D1F33744FFC12" blockId="14.[809,1429,277,1927]" pageId="14" pageNumber="38">
<emphasis id="B945EAF9FFD6FFA7785D1F3373C6FCDA" box="[847,991,825,850]" italics="true" pageId="14" pageNumber="38">Description</emphasis>
.
<emphasis id="B945EAF9FFD6FFA778FC1F327434FCDA" box="[1006,1069,824,850]" italics="true" pageId="14" pageNumber="38">Male</emphasis>
(
<figureCitation id="130A2A6EFFD6FFA77F2D1F327484FCDB" box="[1087,1181,823,851]" captionStart="Figs16–18" captionStartId="15.[159,212,1832,1856]" captionTargetBox="[234,1359,196,1803]" captionTargetId="figure-22@15.[165,1422,181,1807]" captionTargetPageId="15" captionText="Figs16–18. Empis shebalinensis sp.nov., male. 16, habitus, lateral view (holotype); 17, postabdomen, lateral view; 18, terminalia, lateral view. Abbreviations: cerc, cercus; ej apod, ejaculatory apodeme; epand, epandrium; hypd, hypandrium; ph, phallus. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135545" httpUri="https://zenodo.org/record/10135545/files/figure.png" pageId="14" pageNumber="38">Fig. 16</figureCitation>
). Body length
<quantity id="4CC99B0EFFD6FFA77E4B1F3D7391FCFF" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.6" metricValueMin="4.5" pageId="14" pageNumber="38" unit="mm" value="4.55" valueMax="4.6" valueMin="4.5">4.5– 4.6 mm</quantity>
(
<typeStatus id="548A8849FFD6FFA7788B1F567414FCFE" box="[921,1037,860,886]" pageId="14" pageNumber="38" type="holotype">holotype</typeStatus>
,
<quantity id="4CC99B0EFFD6FFA77F081F517464FCFF" box="[1050,1149,859,887]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="14" pageNumber="38" unit="mm" value="4.6">4.6 mm</quantity>
), wing length 3.9–4.0 mm (
<typeStatus id="548A8849FFD6FFA7787A1F8A73C5FC12" box="[872,988,896,922]" pageId="14" pageNumber="38" type="holotype">holotype</typeStatus>
, 4.0 mm).
</paragraph>
<paragraph id="8B8E36EBFFD6FFA7785D1FAE7558FA50" blockId="14.[809,1429,277,1927]" pageId="14" pageNumber="38">Head black. Eyes holoptic, upper ommatidia considerably enlarged. Frons reduced to small subtriangular space above antennae, blackish brown, bare. Face broad, short, much shorter than virtual frons, greyish pruinose, bare. Occiput velvet blackish brown, covered with numerous long thin black setae. Ocellar tubercle concolorous with occiput, with two long thin setae and several hair-like setulae. Antenna black; scape and pedicel subequally short, both with scattered minute setulae; postpedicel moderately long, 2.5–3 times as long as wide, with straight margins; stylus short, postpedicel nearly 2.5 times as long as stylus. Proboscis rather short; labrum brown, thin, only slightly longer than head height; palpus long, black, slender, with scattered black setulae.</paragraph>
<paragraph id="8B8E36EBFFD6FFB9785D19E872F0FE21" blockId="14.[809,1429,277,1927]" lastBlockId="16.[144,764,185,1926]" lastPageId="16" lastPageNumber="40" pageId="14" pageNumber="38">Thorax black, dull, brown pruinose, with black setation; scutum (in dorsal view) dark brown, without vittae. Prosternum bare. Proepisternum and antepronotum with subequally long thin setae. Postpronotal lobe with one long thin seta and several similar shorter setae. Mesonotal setation: one hardly differentiated presutural supra-alar (sometimes indistinguishable from numerous surrounding shorter setae), 3–4 notopleurals, one postsutural supra-alar (with several setulae anteriorly), one postalar and four scutellars (apical pair long, cruciate; lateral pair only slightly shorter); in addition, notopleuron with numerous setulae anteriorly; acrostichals 1–2-serial, scattered, short, thin, lacking on prescutellar depression; dorsocentrals multiserial and similar to acrostichals on presutural part, becoming less numerous and longer behind suture. Laterotergite with numerous setae. Anterior and posterior spiracles black.</paragraph>
<caption id="DF4E6663FFD7FFA67B8D1B227288F808" ID-DOI="http://doi.org/10.5281/zenodo.10135545" ID-Zenodo-Dep="10135545" httpUri="https://zenodo.org/record/10135545/files/figure.png" pageId="15" pageNumber="39" startId="15.[159,212,1832,1856]" targetBox="[234,1359,196,1803]" targetPageId="15" targetType="figure">
<paragraph id="8B8E36EBFFD7FFA67B8D1B227288F808" blockId="15.[159,1444,1832,1921]" pageId="15" pageNumber="39">
<emphasis id="B945EAF9FFD7FFA67B8D1B22713EF8C8" bold="true" box="[159,295,1832,1856]" pageId="15" pageNumber="39">Figs 16–18.</emphasis>
<taxonomicName id="4C314D68FFD7FFA67A3E1B237214F8C9" authority="Shamshev, 2019" authorityName="Shamshev" authorityYear="2019" box="[300,525,1832,1857]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="39" phylum="Arthropoda" rank="species" species="shebalinensis" status="sp. nov.">
<emphasis id="B945EAF9FFD7FFA67A3E1B237214F8C9" box="[300,525,1832,1857]" italics="true" pageId="15" pageNumber="39">Empis shebalinensis</emphasis>
</taxonomicName>
<emphasis id="B945EAF9FFD7FFA679001B237275F8C9" bold="true" box="[530,620,1833,1857]" pageId="15" pageNumber="39">
<taxonomicNameLabel id="A2765782FFD7FFA679001B237275F8C9" box="[530,620,1833,1857]" pageId="15" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="B945EAF9FFD7FFA679AB1B2272CDF8C8" bold="true" box="[697,724,1832,1856]" pageId="15" pageNumber="39">16</emphasis>
, habitus, lateral view (holotype);
<emphasis id="B945EAF9FFD7FFA67F5D1B227471F8C8" bold="true" box="[1103,1128,1832,1856]" pageId="15" pageNumber="39">17</emphasis>
, postabdomen, lateral view;
<emphasis id="B945EAF9FFD7FFA67B8D1B4270A3F8E8" bold="true" box="[159,186,1864,1888]" pageId="15" pageNumber="39">18</emphasis>
, terminalia, lateral view. Abbreviations:
<emphasis id="B945EAF9FFD7FFA679861B4372D8F8E8" box="[660,705,1865,1888]" italics="true" pageId="15" pageNumber="39">cerc</emphasis>
, cercus;
<emphasis id="B945EAF9FFD7FFA678351B437362F8E9" box="[807,891,1864,1889]" italics="true" pageId="15" pageNumber="39">ej apod</emphasis>
, ejaculatory apodeme;
<emphasis id="B945EAF9FFD7FFA67F941B4274D5F8E9" box="[1158,1228,1864,1889]" italics="true" pageId="15" pageNumber="39">epand</emphasis>
, epandrium;
<emphasis id="B945EAF9FFD7FFA67E771B427584F8E9" box="[1381,1437,1864,1889]" italics="true" pageId="15" pageNumber="39">hypd</emphasis>
, hypandrium;
<emphasis id="B945EAF9FFD7FFA67A2B1B62714CF809" box="[313,341,1896,1921]" italics="true" pageId="15" pageNumber="39">ph</emphasis>
, phallus. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B8E36EBFFC8FFB97BA71DB971BFFAC2" blockId="16.[144,764,185,1926]" pageId="16" pageNumber="40">Legs fairly slender, entirely black, with black setation. Coxae and trochanters with simple setae. Fore femur with anteroventral and posteroventral setae mostly minute, somewhat longer toward base of femur; fore femur clothed in moderately long setae posteriorly, without spinules anteriorly. Fore tibia without strong setae (besides circlet of short subapicals), pubescent ventrally. Fore basitarsus slender, with a short anterior seta closer to base; tarsomeres 1–4 pubescent ventrally, with circlet of short subapicals. Mid femur covered only with short setulae. Mid tibia with a short anteroventral seta near middle. Hind femur fairly slender in basal half, somewhat broader in apical half, with some short thin anterodorsal and dorsal setae toward base and minute anteroventral setae (except several somewhat longer subapicals). Hind tibia slightly evenly thickened toward apex, with some short to moderately long (closer to apex) thin anterodorsal and posterodorsal setae; posteroapical comb with one very short seta. Hind basitarsus somewhat thickened but slightly narrower than hind tibia at apex, covered with short setulae (except as noted); tarsomeres 1–4 with anteroventral and posteroventral spinule-like setae. Claws long.</paragraph>
<paragraph id="8B8E36EBFFC8FFB97BA7195E723FF904" blockId="16.[144,764,185,1926]" pageId="16" pageNumber="40">
Wing membrane almost uniformly brownish, somewhat paler near base; veins brownish. Pterostigma brownish, broad. Basal costal seta short, black. Veins
<emphasis id="B945EAF9FFC8FFB97A9419CA71BDFA68" box="[390,420,1472,1504]" italics="true" pageId="16" pageNumber="40">
R
<subScript id="17B534AEFFC8FFB97A8919D871BDFA68" attach="left" box="[411,420,1490,1504]" fontSize="6" pageId="16" pageNumber="40">5</subScript>
</emphasis>
and
<emphasis id="B945EAF9FFC8FFB97AF519CA7210FA68" box="[487,521,1472,1504]" italics="true" pageId="16" pageNumber="40">
M
<subScript id="17B534AEFFC8FFB9791219D87210FA68" attach="left" box="[512,521,1490,1504]" fontSize="6" pageId="16" pageNumber="40">1</subScript>
</emphasis>
parallel near wing margin; radial fork acute; anal vein complete, well-sclerotised throughout. Cell
<emphasis id="B945EAF9FFC8FFB9792C1A0D727DF9A9" box="[574,612,1543,1569]" italics="true" pageId="16" pageNumber="40">dm</emphasis>
short, with elongate apex. Anal angle almost right, subsequently anal lobe well-developed. Calypter brown, brownish fringed. Halter brown.
</paragraph>
<paragraph id="8B8E36EBFFC8FFB97BA71A9F747AFD2C" blockId="16.[144,764,185,1926]" lastBlockId="16.[809,1430,184,1926]" pageId="16" pageNumber="40">
Abdomen with tergites blackish brown, sternites somewhat greyer, with black hair-like setation. Tergites covered with very short setae dorsally and longer setae laterally (setae on tergites 1–3 the longest); sternites with numerous long setae. Sclerites of segment 8 separated; tergite 8 narrow, entire, with scattered setulae; sternite 8 narrow (
<figureCitation id="130A2A6EFFC8FFB978201CB2738BFF5B" box="[818,914,184,211]" captionStart="Figs16–18" captionStartId="15.[159,212,1832,1856]" captionTargetBox="[234,1359,196,1803]" captionTargetId="figure-22@15.[165,1422,181,1807]" captionTargetPageId="15" captionText="Figs16–18. Empis shebalinensis sp.nov., male. 16, habitus, lateral view (holotype); 17, postabdomen, lateral view; 18, terminalia, lateral view. Abbreviations: cerc, cercus; ej apod, ejaculatory apodeme; epand, epandrium; hypd, hypandrium; ph, phallus. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135545" httpUri="https://zenodo.org/record/10135545/files/figure.png" pageId="16" pageNumber="40">Fig. 17</figureCitation>
). Terminalia (
<figureCitation id="130A2A6EFFC8FFB97F541CB274BAFF5B" box="[1094,1187,184,211]" captionStart="Figs16–18" captionStartId="15.[159,212,1832,1856]" captionTargetBox="[234,1359,196,1803]" captionTargetId="figure-22@15.[165,1422,181,1807]" captionTargetPageId="15" captionText="Figs16–18. Empis shebalinensis sp.nov., male. 16, habitus, lateral view (holotype); 17, postabdomen, lateral view; 18, terminalia, lateral view. Abbreviations: cerc, cercus; ej apod, ejaculatory apodeme; epand, epandrium; hypd, hypandrium; ph, phallus. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135545" httpUri="https://zenodo.org/record/10135545/files/figure.png" pageId="16" pageNumber="40">Fig. 18</figureCitation>
) small, black. Cerci separated from each other and from epandrium; cercus rather large, at base nearly as broad as epandrial lamella, with deep excision posteriorly, covered with scattered moderately long brownish setae along upper margin; upper lobe digitiform, pointed, spine-like at tip; lower lobe subtriangular, narrow, slightly extending beyond apex of epandrial lamella. Epandrial lamella subrectangular (in lateral view), with moderately long setae scattered over subapical part. Hypandrium entirely sclerotised, rather subtriangular (in ventral view), with two short setae. Phallus almost entirely hidden, thin, evenly arcuate.
</paragraph>
<paragraph id="8B8E36EBFFC8FFB9785D1EA7747BFAE5" blockId="16.[809,1430,184,1926]" pageId="16" pageNumber="40">
<emphasis id="B945EAF9FFC8FFB9785D1EA773BEFD4F" box="[847,935,685,711]" italics="true" pageId="16" pageNumber="40">Female</emphasis>
. Body length
<quantity id="4CC99B0EFFC8FFB97F451EA674EBFD4F" box="[1111,1266,684,712]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.2" metricValueMin="4.8" pageId="16" pageNumber="40" unit="mm" value="5.0" valueMax="5.2" valueMin="4.8">4.8–5.2 mm</quantity>
; wing length
<quantity id="4CC99B0EFFC8FFB9783B1EDB73DCFD63" box="[809,965,721,747]" metricMagnitude="-3" metricUnit="m" metricValue="4.199999999999999" metricValueMax="4.3" metricValueMin="4.1" pageId="16" pageNumber="40" unit="mm" value="4.199999999999999" valueMax="4.3" valueMin="4.1">4.1–4.3 mm</quantity>
. Similar to male but thorax and abdomen more greyish. Frons moderately broad, in upper part nearly as broad as distance between inner margins of posterior ocelli, almost parallel-sided, slightly widened just above antennae, densely brownish grey pruinose, with marginal setulae. Thorax with somewhat stronger setae; scutum (in dorsal view) with four indistinct brownish vittae. Fore femur with shorter setae posteriorly; fore tibia and fore tarsomeres 1–4 covered with simple setulae ventrally. Mid tibia with two anterodorsal, two posterodorsal, 1–2 anteroventral, and 1–2 posteroventral setae (besides circlet of subapicals). Hind femur usually with a moderately long strong anteroventral seta in about apical fourth; hind tibia with stronger setae; hind basitarsus slender. Wing membrane only faintly infuscate. Abdomen with shorter setation; cercus long, slender, with scattered minute setulae.
</paragraph>
<paragraph id="8B8E36EBFFC8FFB8785D197272BDFF7E" blockId="16.[809,1430,184,1926]" lastBlockId="17.[159,779,184,494]" lastPageId="17" lastPageNumber="41" pageId="16" pageNumber="40">
<emphasis id="B945EAF9FFC8FFB9785D197273FCFA1A" box="[847,997,1400,1426]" italics="true" pageId="16" pageNumber="40">Comparison</emphasis>
. The new species resembles
<taxonomicName id="4C314D68FFC8FFB97E5B1972740EFA3E" authority="Collin, 1926" authorityName="Collin" authorityYear="1926" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="40" phylum="Arthropoda" rank="species" species="bazini">
<emphasis id="B945EAF9FFC8FFB97E5B1972736DFA3D" italics="true" pageId="16" pageNumber="40">Empis bazini</emphasis>
Collin, 1926
</taxonomicName>
(known from
<collectingCountry id="F326767BFFC8FFB97FD819917589FA3D" box="[1226,1424,1435,1461]" name="Czech Republic" pageId="16" pageNumber="40">Czech Republic</collectingCountry>
, the French Alps and southern
<collectingCountry id="F326767BFFC8FFB97FB219B57503FA51" box="[1184,1306,1471,1497]" name="Germany" pageId="16" pageNumber="40">Germany</collectingCountry>
), which is a single species of the subgenus
<taxonomicName id="4C314D68FFC8FFB97FA119E87382F9A9" authority="Collin, 1926" authorityName="Collin" authorityYear="1926" class="Insecta" family="Empididae" genus="Rhadinempis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAF9FFC8FFB97FA119E8754EFA74" box="[1203,1367,1506,1532]" italics="true" pageId="16" pageNumber="40">Rhadinempis</emphasis>
Collin, 1926
</taxonomicName>
. However,
<taxonomicName id="4C314D68FFC8FFB97F361A0D7489F9A8" authorityName="Collin" authorityYear="1926" box="[1060,1168,1542,1568]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="40" phylum="Arthropoda" rank="species" species="bazini">
<emphasis id="B945EAF9FFC8FFB97F361A0D7489F9A8" box="[1060,1168,1542,1568]" italics="true" pageId="16" pageNumber="40">E. bazini</emphasis>
</taxonomicName>
differs from the new species primarily in the absence of acrostichal setae, very short postpedicel (scarcely twice as long as broad) and very long stylus (three times as long as postpedicel). At the same time, these characters are used currently to distinguish
<taxonomicName id="4C314D68FFC8FFB97FEB1AB37324F97E" authorityName="Collin" authorityYear="1926" class="Insecta" family="Empididae" genus="Rhadinempis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAF9FFC8FFB97FEB1AB37324F97E" italics="true" pageId="16" pageNumber="40">Rhadinempis</emphasis>
</taxonomicName>
from other subgenera of
<taxonomicName id="4C314D68FFC8FFB97F931AD774D5F97E" baseAuthorityName="Shamshev" baseAuthorityYear="2001" box="[1153,1228,1757,1782]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAF9FFC8FFB97F931AD774D5F97E" box="[1153,1228,1757,1782]" italics="true" pageId="16" pageNumber="40">Empis</emphasis>
</taxonomicName>
. Thus, the new species cannot be included in
<taxonomicName id="4C314D68FFC8FFB97F8D1B0A755AF892" authorityName="Collin" authorityYear="1926" box="[1183,1347,1792,1818]" class="Insecta" family="Empididae" genus="Rhadinempis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAF9FFC8FFB97F8D1B0A755AF892" box="[1183,1347,1792,1818]" italics="true" pageId="16" pageNumber="40">Rhadinempis</emphasis>
</taxonomicName>
or the current concept of this group requires a revision. Additionally,
<taxonomicName id="4C314D68FFC8FFB978C81B437455F8EA" authorityName="Collin" authorityYear="1926" box="[986,1100,1864,1890]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="40" phylum="Arthropoda" rank="species" species="bazini">
<emphasis id="B945EAF9FFC8FFB978C81B437455F8EA" box="[986,1100,1864,1890]" italics="true" pageId="16" pageNumber="40">E. bazini</emphasis>
</taxonomicName>
and
<taxonomicName id="4C314D68FFC8FFB97F841B437578F8EA" authority="Shamshev, 2019" authorityName="Shamshev" authorityYear="2019" box="[1174,1377,1864,1890]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="40" phylum="Arthropoda" rank="species" species="shebalinensis" status="sp. nov.">
<emphasis id="B945EAF9FFC8FFB97F841B437578F8EA" box="[1174,1377,1864,1890]" italics="true" pageId="16" pageNumber="40">E. shebalinensis</emphasis>
</taxonomicName>
<emphasis id="B945EAF9FFC8FFB97E7D1B427379F80E" bold="true" pageId="16" pageNumber="40">
<taxonomicNameLabel id="A2765782FFC8FFB97E7D1B427379F80E" pageId="16" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are very similar to species of the subgenus
<taxonomicName id="4C314D68FFC9FFB87B8D1CB371DCFF5C" authority="Schiner, 1860" authorityName="Schiner" authorityYear="1860" box="[159,453,184,212]" class="Insecta" family="Empididae" genus="Holoclera" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAF9FFC9FFB87B8D1CB3710FFF5B" box="[159,278,185,211]" italics="true" pageId="17" pageNumber="41">Holoclera</emphasis>
Schiner, 1860
</taxonomicName>
of the genus
<taxonomicName id="4C314D68FFC9FFB879711CB3717EFF7E" authority="Meigen, 1822" authorityName="Meigen" authorityYear="1822" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAF9FFC9FFB879711CB370AEFF7E" italics="true" pageId="17" pageNumber="41">Rhamphomyia</emphasis>
Meigen, 1822
</taxonomicName>
(
<bibRefCitation id="EFA04B1AFFC9FFB87A641CD6728EFF7E" author="Bartak M. &amp; Kubik S." box="[374,663,220,246]" pageId="17" pageNumber="41" pagination="385 - 407" refId="ref10573" refString="Bartak M. &amp; Kubik S. 2012. A review of the Palaearctic species of Rhamphomyia subgenus Holoclera (Diptera: Empididae) with description of 5 new species. Revue Suisse de Zoologie, 119: 385 - 407. https: // doi. org / 10.5962 / bhl. part. 150199" type="journal article" year="2012">Barták &amp; Kubík, 2012</bibRefCitation>
).
</paragraph>
<paragraph id="8B8E36EBFFC9FFB87BD71CF571BFFEB4" blockId="17.[159,779,184,494]" pageId="17" pageNumber="41">
<emphasis id="B945EAF9FFC9FFB87BD71CF5715EFE91" box="[197,327,255,281]" italics="true" pageId="17" pageNumber="41">Etymology</emphasis>
. The epithet refers to the
<typeStatus id="548A8849FFC9FFB8796D1D0A72A1FE92" box="[639,696,256,282]" pageId="17" pageNumber="41">type</typeStatus>
locality of the new species.
</paragraph>
<paragraph id="8B8E36EBFFC9FFB87BD61D4F71FEFE0C" blockId="17.[159,779,184,494]" pageId="17" pageNumber="41">
<emphasis id="B945EAF9FFC9FFB87BD61D4F7144FED7" box="[196,349,325,351]" italics="true" pageId="17" pageNumber="41">Distribution</emphasis>
. Palaearctic:
<collectingCountry id="F326767BFFC9FFB8791C1D4F7278FED7" box="[526,609,325,351]" name="Russia" pageId="17" pageNumber="41">Russia</collectingCountry>
(Republic of
<collectingRegion id="49F5F809FFC9FFB87B8D1D6370C6FE0B" box="[159,223,361,387]" country="Russia" name="Altay" pageId="17" pageNumber="41">Altai</collectingRegion>
and
<collectingRegion id="49F5F809FFC9FFB87A091D637142FE0B" box="[283,347,361,387]" country="Russia" name="Altay" pageId="17" pageNumber="41">Altai</collectingRegion>
Territory).
</paragraph>
<paragraph id="8B8E36EBFFC9FFB87BD61D867293FE65" blockId="17.[159,779,184,494]" pageId="17" pageNumber="41">
<emphasis id="B945EAF9FFC9FFB87BD61D86727CFE2E" box="[196,613,396,422]" italics="true" pageId="17" pageNumber="41">Habitat and seasonal occurrence</emphasis>
. The species was collected by sweeping on a montane meadow (
<quantity id="4CC99B0EFFC9FFB87BB51DD87111FE65" box="[167,264,466,494]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="17" pageNumber="41" unit="m" value="1325.0">1325 m</quantity>
); collecting dates refer to June.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink