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<document id="C7A980171CD459CBE68550AF3C91875E" ID-CLB-Dataset="57159" ID-DOI="10.31610/zsr/2019.28.1.24" ID-GBIF-Dataset="2b690c17-f7fc-4c69-83d1-1d9047c876c9" ID-ISSN="2410-0226" ID-Zenodo-Dep="10124208" ID-ZooBank="urn:lsid:zoobank.org:pub:D82459C6-A090-4203-8D28-8BE062383D2B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1688479652846" checkinUser="felipe" docAuthor="Shamshev, I. V." docDate="2019" docId="039887FDFFD4FFA77B821CB3712DF80E" docLanguage="en" docName="ZoosystRoss.28.1.24-41.pdf" docOrigin="Zoosystematica Rossica (Zoosyst. Rossica) 28 (1)" docSource="http://dx.doi.org/10.31610/zsr/2019.28.1.24" docStyle="DocumentStyle:8B7E50B51175314E898B46063364B751.1:ZoosystRoss.2019-.journal_article" docStyleId="8B7E50B51175314E898B46063364B751" docStyleName="ZoosystRoss.2019-.journal_article" docStyleVersion="1" docTitle="Empis barkalovi Shamshev 2019, sp. nov." docType="treatment" docVersion="7" lastPageNumber="38" masterDocId="FFA1FF85FFD8FFA97B121C0A7019FF88" masterDocTitle="Six new species of the genus Empis (Diptera: Empididae) from the Altai Mountains of Russia" masterLastPageNumber="41" masterPageNumber="24" pageNumber="36" updateTime="1700049859112" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="940436CBC50C21AD7E8847516C9EEC9E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="036568322517DC84F670B865EA339855">
<mods:title id="41A0282A2005716984ABBFD5BF21EEB2">Six new species of the genus Empis (Diptera: Empididae) from the Altai Mountains of Russia</mods:title>
</mods:titleInfo>
<mods:name id="7965585A3E75760ACDADD7ACBB248ED3" type="personal">
<mods:role id="ACF32DC2E4BB021D4C88FA7D8C7803DE">
<mods:roleTerm id="4CBCAA39C18412B79FCD03A5456846F1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="453E37030500014B8615F2218E09E636">Shamshev, I. V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="638C56C620E59D129FDDCFEEB59324A1">text</mods:typeOfResource>
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<mods:title id="E6C3C49B06E8068BEE08CBD5CDDAD36B">Zoosystematica Rossica</mods:title>
</mods:titleInfo>
<mods:part id="CCC76D59E6B79CFB7826ABC7A9336008">
<mods:date id="2880D4ED68F7DF32E5877A27BDAAD946">2019</mods:date>
<mods:detail id="59B1D97C50F59BD75DC670A44D4C5047" type="series">
<mods:title id="03167BA77B55DEC93D237B765DEB97E4">Zoosyst. Rossica</mods:title>
</mods:detail>
<mods:detail id="D0D156635E8A0D7E2985524451701F37" type="pubDate">
<mods:number id="D24FBFA53A4B6E51E4AE33AB840C5AF5">2019-04-18</mods:number>
</mods:detail>
<mods:detail id="FDB83CBD90DFB23F8FD781D0263E2193" type="volume">
<mods:number id="712BB66A4D48DBACB70513B794D71581">28</mods:number>
</mods:detail>
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<mods:number id="0A8EE6E44149B62231446E19FA47C6E1">1</mods:number>
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<mods:start id="A887300C5C7784CCAEEEB87902EA64FD">24</mods:start>
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<mods:location id="7349D056E4133BFF2BE8A75493BD00B3">
<mods:url id="023124CEC5F12016192C91DC8EF81359">http://dx.doi.org/10.31610/zsr/2019.28.1.24</mods:url>
</mods:location>
<mods:classification id="D0BE6AF5EDA3169FCD63D0C055981975">journal article</mods:classification>
<mods:identifier id="6541D4B17A45A6B8E397DC05B1D164E3" type="CLB-Dataset">57159</mods:identifier>
<mods:identifier id="D5C027F3145FB871B2ABC33F4711551E" type="DOI">10.31610/zsr/2019.28.1.24</mods:identifier>
<mods:identifier id="3D5A1C7CCBFA647ABED23F109C98073A" type="GBIF-Dataset">2b690c17-f7fc-4c69-83d1-1d9047c876c9</mods:identifier>
<mods:identifier id="B9FA4C24ABE9BE5DCECFBDAEDBED10F4" type="ISSN">2410-0226</mods:identifier>
<mods:identifier id="C7FA69FBA8A5162410BFCA61BCA2685C" type="Zenodo-Dep">10124208</mods:identifier>
<mods:identifier id="3F66E5DC4945219ECC38753C131480E6" type="ZooBank">urn:lsid:zoobank.org:pub:D82459C6-A090-4203-8D28-8BE062383D2B</mods:identifier>
</mods:mods>
<treatment id="039887FDFFD4FFA77B821CB3712DF80E" ID-DOI="http://doi.org/10.5281/zenodo.8114721" ID-GBIF-Taxon="210810426" ID-Zenodo-Dep="8114721" LSID="urn:lsid:plazi:treatment:039887FDFFD4FFA77B821CB3712DF80E" httpUri="http://treatment.plazi.org/id/039887FDFFD4FFA77B821CB3712DF80E" lastPageId="14" lastPageNumber="38" pageId="12" pageNumber="36">
<subSubSection id="C32B6560FFD4FFA57B821CB371CDFF5B" box="[144,468,184,211]" pageId="12" pageNumber="36" type="nomenclature">
<paragraph id="8B8E36EBFFD4FFA57B821CB371CDFF5B" blockId="12.[144,468,184,211]" box="[144,468,184,211]" pageId="12" pageNumber="36">
<heading id="D0C68187FFD4FFA57B821CB371CDFF5B" box="[144,468,184,211]" fontSize="11" level="2" pageId="12" pageNumber="36" reason="2">
<emphasis id="B945EAF9FFD4FFA57B821CB371CDFF5B" bold="true" box="[144,468,184,211]" pageId="12" pageNumber="36">
<taxonomicName id="4C314D68FFD4FFA57B821CB37170FF5B" ID-CoL="8BZSY" authority="Shamshev, 2019" authorityName="Shamshev" authorityYear="2019" box="[144,361,184,211]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="36" phylum="Arthropoda" rank="species" species="barkalovi" status="sp. nov.">
<emphasis id="B945EAF9FFD4FFA57B821CB37170FF5B" bold="true" box="[144,361,184,211]" italics="true" pageId="12" pageNumber="36">Empis barkalovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765782FFD4FFA57A621CB371CDFF5B" box="[368,468,185,211]" pageId="12" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32B6560FFD4FFA77B821CD6712DF80E" lastPageId="14" lastPageNumber="38" pageId="12" pageNumber="36" type="description">
<paragraph id="8B8E36EBFFD4FFA57B821CD6712CFF70" blockId="12.[144,309,220,248]" box="[144,309,220,248]" pageId="12" pageNumber="36">
(
<figureCitation id="130A2A6EFFD4FFA57B8B1CD67133FF70" box="[153,298,220,248]" captionStart="Figs 13–15" captionStartId="13.[159,211,1832,1856]" captionTargetBox="[202,1384,193,1779]" captionTargetId="figure-21@13.[165,1422,181,1808]" captionTargetPageId="13" captionText="Figs 13–15. Empis barkalovi sp. nov., male. 13, habitus, lateral view (holotype); 14, postabdomen, lateral view; 15, terminalia, lateral view. Abbreviations: cerc, cercus; ej apod, ejaculatory apodeme; epand, epandrium; hypd, hypandrium; ph, phallus. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135543" httpUri="https://zenodo.org/record/10135543/files/figure.png" pageId="12" pageNumber="36">Figs 13–15</figureCitation>
)
</paragraph>
<paragraph id="8B8E36EBFFD4FFA57BA71D1F7191FE58" blockId="12.[144,764,277,1927]" pageId="12" pageNumber="36">
<materialsCitation id="3B593CB6FFD4FFA57BA71D1F7191FE58" ID-GBIF-Occurrence="4144857304" collectingDate="1964-07-24" collectionCode="ZIN" country="Russia" latitude="49.816666" location="Chikhacheva" longLatPrecision="1101" longitude="89.76667" municipality="Male" pageId="12" pageNumber="36" specimenCode="INS_DIP_0000787" specimenCount="1" stateProvince="Republic of Altai" typeStatus="holotype">
<emphasis id="B945EAF9FFD4FFA57BA71D1F7102FEA6" box="[181,283,277,302]" italics="true" pageId="12" pageNumber="36">
<typeStatus id="548A8849FFD4FFA57BA71D1F7102FEA6" box="[181,283,277,302]" pageId="12" pageNumber="36" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCount id="9D37FD62FFD4FFA57A3B1D1F717FFEA6" box="[297,358,277,302]" pageId="12" pageNumber="36" type="male">Male</specimenCount>
,
<emphasis id="B945EAF9FFD4FFA57A631D1F71DAFEA5" bold="true" box="[369,451,277,301]" pageId="12" pageNumber="36">
<collectingCountry id="F326767BFFD4FFA57A631D1F71DAFEA5" box="[369,451,277,301]" name="Russia" pageId="12" pageNumber="36">Russia</collectingCountry>
</emphasis>
,
<emphasis id="B945EAF9FFD4FFA57AC21D1F7288FEA6" box="[464,657,277,302]" italics="true" pageId="12" pageNumber="36">
<collectingRegion id="49F5F809FFD4FFA57AC21D1F7288FEA6" box="[464,657,277,302]" country="Russia" name="Altay" pageId="12" pageNumber="36">Republic of Altai</collectingRegion>
</emphasis>
: “[in Cyrillic, Russian] khr. [= khrebet, ridge]
<location id="8EEE6030FFD4FFA579761D3F72E2FEC6" LSID="urn:lsid:plazi:treatment:039887FDFFD4FFA77B821CB3712DF80E:8EEE6030FFD4FFA579761D3F72E2FEC6" box="[612,763,309,334]" country="Russia" latitude="49.816666" longLatPrecision="1101" longitude="89.76667" municipality="Male" name="Chikhacheva" pageId="12" pageNumber="36" stateProvince="Republic of Altai">Chikhacheva</location>
[
<geoCoordinate id="EE05502CFFD4FFA57B841D5D70EFFEE6" box="[150,246,343,367]" degrees="49" direction="north" minutes="49" orientation="latitude" pageId="12" pageNumber="36" precision="925" value="49.816666">49°49'N</geoCoordinate>
<geoCoordinate id="EE05502CFFD4FFA57BEB1D5C714EFEE6" box="[249,343,342,367]" degrees="89" direction="east" minutes="46" orientation="longitude" pageId="12" pageNumber="36" precision="925" value="89.76667">89°46'E</geoCoordinate>
] / pass / yugo-vost. [= yugo-vostochnyy, south-eastern] Altai / Nartshuk
<date id="FF8F102BFFD4FFA579731D7C72F5FE07" box="[609,748,374,399]" pageId="12" pageNumber="36" value="1964-07-24">
<collectingDate id="EFCBE9C3FFD4FFA579731D7C72F5FE07" box="[609,748,374,399]" pageId="12" pageNumber="36" value="1964-07-24">24.VII.1964</collectingDate>
</date>
”, “
<taxonomicName id="4C314D68FFD4FFA57B881D9271CAFE38" authority="Shamshev" authorityName="Shamshev" authorityYear="2019" box="[154,467,407,432]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="36" phylum="Arthropoda" rank="species" species="barkalovi" status="sp. nov.">
<emphasis id="B945EAF9FFD4FFA57B881D92714DFE38" box="[154,340,407,432]" italics="true" pageId="12" pageNumber="36">Empis barkalovi</emphasis>
Shamshev
</taxonomicName>
,
<taxonomicNameLabel id="A2765782FFD4FFA57AF01D927239FE27" box="[482,544,408,431]" pageId="12" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
[red label]” (
<collectionCode id="ED20AE2EFFD4FFA579D31D9272EEFE38" box="[705,759,408,432]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="36" type="Museum">ZIN</collectionCode>
,
<specimenCode id="DB979E90FFD4FFA57B821DB27165FE58" box="[144,380,440,464]" pageId="12" pageNumber="36">INS_DIP_0000787</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B8E36EBFFD4FFA57BA71DD37213FCDE" blockId="12.[144,764,277,1927]" pageId="12" pageNumber="36">
<materialsCitation id="3B593CB6FFD4FFA57BA71DD372E5FDDA" ID-GBIF-Occurrence="4144857307" collectingDate="2008-07-09" collectingDateMax="2006-07-10" collectingDateMin="2006-07-06" collectionCode="SZMN" collectorName="V. Zinchenko" country="Russia" county="Kosh-Agach Distr." elevation="3000" latitude="49.566666" location="Ukok" longLatPrecision="1103" longitude="87.71667" municipality="Ukok" pageId="12" pageNumber="36" specimenCount="2" specimenCount-male="2" stateProvince="Republic of Altai" typeStatus="paratype">
<emphasis id="B945EAF9FFD4FFA57BA71DD3713EFE78" box="[181,295,473,496]" italics="true" pageId="12" pageNumber="36">
<typeStatus id="548A8849FFD4FFA57BA71DD3713EFE78" box="[181,295,473,496]" pageId="12" pageNumber="36" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<emphasis id="B945EAF9FFD4FFA57A201DD2719DFE78" bold="true" box="[306,388,472,496]" pageId="12" pageNumber="36">
<collectingCountry id="F326767BFFD4FFA57A201DD2719DFE78" box="[306,388,472,496]" name="Russia" pageId="12" pageNumber="36">Russia</collectingCountry>
</emphasis>
,
<emphasis id="B945EAF9FFD4FFA57A9D1DD27255FE79" box="[399,588,472,497]" italics="true" pageId="12" pageNumber="36">
<collectingRegion id="49F5F809FFD4FFA57A9D1DD27255FE79" box="[399,588,472,497]" pageId="12" pageNumber="36">Republic of Altai</collectingRegion>
</emphasis>
:
<specimenCount id="9D37FD62FFD4FFA5794A1DD372B4FE79" box="[600,685,472,497]" pageId="12" pageNumber="36" type="male">2 males</specimenCount>
,
<collectingCounty id="62EF4E67FFD4FFA579A41DD27131FD99" pageId="12" pageNumber="36">Kosh-Agach Distr.</collectingCounty>
, plateau
<location id="8EEE6030FFD4FFA57A851DF271C3FD99" LSID="urn:lsid:plazi:treatment:039887FDFFD4FFA77B821CB3712DF80E:8EEE6030FFD4FFA57A851DF271C3FD99" box="[407,474,504,529]" country="Russia" county="Kosh-Agach Distr." latitude="49.566666" longLatPrecision="1103" longitude="87.71667" municipality="Ukok" name="Ukok" pageId="12" pageNumber="36" stateProvince="Republic of Altai">Ukok</location>
, environs of
<location id="8EEE6030FFD4FFA5796A1DF270E1FDBA" LSID="urn:lsid:plazi:treatment:039887FDFFD4FFA77B821CB3712DF80E:8EEE6030FFD4FFA5796A1DF270E1FDBA" country="Russia" county="Kosh-Agach Distr." latitude="49.566666" longLatPrecision="1103" longitude="87.71667" municipality="Ukok" name="Muzdy-Bulak Lake" pageId="12" pageNumber="36" stateProvince="Republic of Altai">Muzdy-Bulak Lake</location>
, S slope of
<location id="8EEE6030FFD4FFA57A941E137286FDBA" LSID="urn:lsid:plazi:treatment:039887FDFFD4FFA77B821CB3712DF80E:8EEE6030FFD4FFA57A941E137286FDBA" box="[390,671,537,562]" country="Russia" county="Kosh-Agach Distr." latitude="49.566666" longLatPrecision="1103" longitude="87.71667" municipality="Ukok" name="Cholok-Chad Mountain" pageId="12" pageNumber="36" stateProvince="Republic of Altai">Cholok-Chad Mountain</location>
,
<quantity id="4CC99B0EFFD4FFA579BC1E1070F0FDD9" metricMagnitude="3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="12" pageNumber="36" unit="m" value="3000.0" valueMax="3100.0" valueMin="2900.0">
<elevation id="001CD1D8FFD4FFA579BC1E1070F0FDD9" metricMagnitude="3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="12" pageNumber="36" unit="m" value="3000.0" valueMax="3100.0" valueMin="2900.0">2900– 3100 m</elevation>
</quantity>
, 1 &amp;
<date id="FF8F102BFFD4FFA57A391E3071B5FDDA" box="[299,428,570,594]" pageId="12" pageNumber="36" value="2008-07-09">
<collectingDate id="EFCBE9C3FFD4FFA57A391E3071B5FDDA" box="[299,428,570,594]" pageId="12" pageNumber="36" value="2008-07-09">9.VII.2008</collectingDate>
</date>
,
<collectorName id="26C4533DFFD4FFA57AAB1E30724CFDDA" box="[441,597,569,594]" pageId="12" pageNumber="36">V. Zinchenko</collectorName>
leg. (
<collectionCode id="ED20AE2EFFD4FFA579851E3072F6FDDA" box="[663,751,570,594]" collectionName="Russia, Novosibirsk, Institute of Animal Systematics and Ecology, Siberian Zoological Museum" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="12" pageNumber="36">SZMN</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B593CB6FFD4FFA57B821E517226FD3C" ID-GBIF-Occurrence="4144857310" collectingDate="2006-07-06" collectingDateMax="2006-07-10" collectingDateMin="2006-07-06" collectionCode="ZIN" collectorName="T. Novgorodova" country="Russia" county="Agach Distr." elevation="2600" latitude="49.566666" location="Ukok" longLatPrecision="1103" longitude="87.71667" municipality="Ukok" pageId="12" pageNumber="36" specimenCount="1" specimenCount-male="1" stateProvince="Altay" typeStatus="paratype">
<specimenCount id="9D37FD62FFD4FFA57B821E5170F9FDFB" box="[144,224,602,627]" pageId="12" pageNumber="36" type="male">1 male</specimenCount>
, plateau
<location id="8EEE6030FFD4FFA57A5D1E50718BFDFB" LSID="urn:lsid:plazi:treatment:039887FDFFD4FFA77B821CB3712DF80E:8EEE6030FFD4FFA57A5D1E50718BFDFB" box="[335,402,602,627]" country="Russia" county="Agach Distr." latitude="49.566666" longLatPrecision="1103" longitude="87.71667" municipality="Ukok" name="Ukok" pageId="12" pageNumber="36" stateProvince="Altay">Ukok</location>
,
<locationDeviation id="9A3C2FC9FFD4FFA57AB21E517216FDFA" box="[416,527,602,627]" pageId="12" pageNumber="36">
<quantity id="4CC99B0EFFD4FFA57AB21E5171C4FDFB" box="[416,477,602,627]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="36" unit="km" value="8.0">8 km</quantity>
NE
</locationDeviation>
of
<location id="8EEE6030FFD4FFA579281E5070D9FD1B" LSID="urn:lsid:plazi:treatment:039887FDFFD4FFA77B821CB3712DF80E:8EEE6030FFD4FFA579281E5070D9FD1B" country="Russia" county="Agach Distr." latitude="49.566666" longLatPrecision="1103" longitude="87.71667" municipality="Ukok" name="Maytobe Mountain" pageId="12" pageNumber="36" stateProvince="Altay">Maytobe Mountain</location>
,
<geoCoordinate id="EE05502CFFD4FFA57BDE1E717132FD1A" box="[204,299,635,659]" degrees="49" direction="north" minutes="34" orientation="latitude" pageId="12" pageNumber="36" precision="925" value="49.566666">49°34'N</geoCoordinate>
<geoCoordinate id="EE05502CFFD4FFA57A231E717194FD1A" box="[305,397,635,659]" degrees="87" direction="east" minutes="43" orientation="longitude" pageId="12" pageNumber="36" precision="925" value="87.71667">87°43'E</geoCoordinate>
,
<quantity id="4CC99B0EFFD4FFA57A8B1E717226FD1A" box="[409,575,635,659]" metricMagnitude="3" metricUnit="m" metricValue="2.6" metricValueMax="2.8" metricValueMin="2.4" pageId="12" pageNumber="36" unit="m" value="2600.0" valueMax="2800.0" valueMin="2400.0">
<elevation id="001CD1D8FFD4FFA57A8B1E717226FD1A" box="[409,575,635,659]" metricMagnitude="3" metricUnit="m" metricValue="2.6" metricValueMax="2.8" metricValueMin="2.4" pageId="12" pageNumber="36" unit="m" value="2600.0" valueMax="2800.0" valueMin="2400.0">2400–2800 m</elevation>
</quantity>
,
<date id="FF8F102BFFD4FFA579581E7072EFFD1B" box="[586,758,634,659]" pageId="12" pageNumber="36" value="2006-07-06" valueMax="2006-07-10" valueMin="2006-07-06">
<collectingDate id="EFCBE9C3FFD4FFA579581E7072EFFD1B" box="[586,758,634,659]" pageId="12" pageNumber="36" value="2006-07-06" valueMax="2006-07-10" valueMin="2006-07-06">6–10.VII.2006</collectingDate>
</date>
,
<collectorName id="26C4533DFFD4FFA57B821E96715FFD3C" box="[144,326,667,692]" pageId="12" pageNumber="36">T. Novgorodova</collectorName>
leg. (dissected;
<collectionCode id="ED20AE2EFFD4FFA57AE91E967228FD3C" box="[507,561,668,692]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="36" type="Museum">ZIN</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B593CB6FFD4FFA579571E967296FD7D" ID-GBIF-Occurrence="4144857318" collectingDate="2008-07-02" collectionCode="SZMN" collectorName="A. Barkalov" country="Russia" county="Agach Distr." elevation="2450" latitude="50.53" location="Kuray Ridge" longLatPrecision="1103" longitude="87.71667" municipality="Ulagan Distr." pageId="12" pageNumber="36" specimenCount="2" specimenCount-male="2" stateProvince="Altay" typeStatus="paratype">
<specimenCount id="9D37FD62FFD4FFA579571E967285FD3C" box="[581,668,667,692]" pageId="12" pageNumber="36" type="male">2 males</specimenCount>
,
<collectingMunicipality id="6BEAAC91FFD4FFA579B51E9170CDFD5C" pageId="12" pageNumber="36">Ulagan Distr.</collectingMunicipality>
,
<location id="8EEE6030FFD4FFA57BCD1EB6716CFD5C" LSID="urn:lsid:plazi:treatment:039887FDFFD4FFA77B821CB3712DF80E:8EEE6030FFD4FFA57BCD1EB6716CFD5C" box="[223,373,699,724]" country="Russia" county="Agach Distr." latitude="50.53" longLatPrecision="1103" longitude="87.71667" municipality="Ulagan Distr." name="Kuray Ridge" pageId="12" pageNumber="36" stateProvince="Altay">Kuray Ridge</location>
,
<quantity id="4CC99B0EFFD4FFA57A931EB171C3FD5B" box="[385,474,699,724]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="12" pageNumber="36" unit="m" value="2450.0">
<elevation id="001CD1D8FFD4FFA57A931EB171C3FD5B" box="[385,474,699,724]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="12" pageNumber="36" unit="m" value="2450.0">2450 m</elevation>
</quantity>
,
<geoCoordinate id="EE05502CFFD4FFA57AF71EB1725CFD5B" box="[485,581,699,724]" degrees="50" direction="north" minutes="33" orientation="latitude" pageId="12" pageNumber="36" precision="925" value="50.55">50°33'N</geoCoordinate>
<geoCoordinate id="EE05502CFFD4FFA5795E1EB172B3FD5B" box="[588,682,699,724]" degrees="87" direction="east" minutes="75" orientation="longitude" pageId="12" pageNumber="36" precision="925" value="87-75">87°75'E</geoCoordinate>
, meadow,
<date id="FF8F102BFFD4FFA57BD31ED7715BFD7D" box="[193,322,733,757]" pageId="12" pageNumber="36" value="2008-07-02">
<collectingDate id="EFCBE9C3FFD4FFA57BD31ED7715BFD7D" box="[193,322,733,757]" pageId="12" pageNumber="36" value="2008-07-02">2.VII.2008</collectingDate>
</date>
,
<collectorName id="26C4533DFFD4FFA57A411ED771FBFD7D" box="[339,482,732,757]" pageId="12" pageNumber="36">A. Barkalov</collectorName>
leg. (
<collectionCode id="ED20AE2EFFD4FFA579381ED7729BFD7D" box="[554,642,733,757]" collectionName="Russia, Novosibirsk, Institute of Animal Systematics and Ecology, Siberian Zoological Museum" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="12" pageNumber="36">SZMN</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B593CB6FFD4FFA579881ED770E4FCBE" ID-GBIF-Occurrence="4144857302" collectingDate="2008-07-03" collectionCode="SZMN" collectorName="A. Barkalov" country="Russia" elevation="2450" latitude="50.55" location="Kuray Ridge" municipality="Ulagan Distr." pageId="12" pageNumber="36" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D37FD62FFD4FFA579881ED772EEFD7D" box="[666,759,732,757]" pageId="12" pageNumber="36" type="male">2 males</specimenCount>
, same locality, tundra,
<date id="FF8F102BFFD4FFA57AB01EF7723AFC9D" box="[418,547,765,789]" pageId="12" pageNumber="36" value="2008-07-03">
<collectingDate id="EFCBE9C3FFD4FFA57AB01EF7723AFC9D" box="[418,547,765,789]" pageId="12" pageNumber="36" value="2008-07-03">3.VII.2008</collectingDate>
</date>
,
<collectorName id="26C4533DFFD4FFA579271EF772DFFC9D" box="[565,710,764,789]" pageId="12" pageNumber="36">A. Barkalov</collectorName>
leg. (
<collectionCode id="ED20AE2EFFD4FFA57B8A1F1470E9FCBE" box="[152,240,798,822]" collectionName="Russia, Novosibirsk, Institute of Animal Systematics and Ecology, Siberian Zoological Museum" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="12" pageNumber="36">SZMN</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B593CB6FFD4FFA57A111F147213FCDE" ID-GBIF-Occurrence="4144857313" collectingDate="2008-06-29" collectingDateMax="2008-06-30" collectingDateMin="2008-06-29" collectionCode="ZIN" collectorName="A. Barkalov" country="Russia" elevation="2650" latitude="50.55" location="Kuray Ridge" municipality="Ulagan Distr." pageId="12" pageNumber="36" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D37FD62FFD4FFA57A111F147156FCBE" box="[259,335,797,822]" pageId="12" pageNumber="36" type="male">1 male</specimenCount>
, same locality,
<quantity id="4CC99B0EFFD4FFA57AE81F177286FCBD" box="[506,671,797,822]" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="12" pageNumber="36" unit="m" value="2650.0" valueMax="2800.0" valueMin="2500.0">
<elevation id="001CD1D8FFD4FFA57AE81F177286FCBD" box="[506,671,797,822]" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="12" pageNumber="36" unit="m" value="2650.0" valueMax="2800.0" valueMin="2500.0">2500–2800 m</elevation>
</quantity>
,
<date id="FF8F102BFFD4FFA579BB1F1470EAFCDE" pageId="12" pageNumber="36" value="2008-06-29" valueMax="2008-06-30" valueMin="2008-06-29">
<collectingDate id="EFCBE9C3FFD4FFA579BB1F1470EAFCDE" pageId="12" pageNumber="36" value="2008-06-29" valueMax="2008-06-30" valueMin="2008-06-29">29–30. VI.2008</collectingDate>
</date>
,
<collectorName id="26C4533DFFD4FFA57BEC1F347191FCDE" box="[254,392,829,854]" pageId="12" pageNumber="36">A. Barkalov</collectorName>
leg. (
<collectionCode id="ED20AE2EFFD4FFA57AD71F3471E7FCDE" box="[453,510,830,854]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="12" pageNumber="36">ZIN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B8E36EBFFD4FFA57BA71F6B72E0FBA5" blockId="12.[144,764,277,1927]" pageId="12" pageNumber="36">
<emphasis id="B945EAF9FFD4FFA57BA71F6B7128FCF2" box="[181,305,865,890]" italics="true" pageId="12" pageNumber="36">Diagnosis</emphasis>
. Large species, with body length about
<quantity id="4CC99B0EFFD4FFA57BF01F8E7136FC16" box="[226,303,900,926]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="36" unit="mm" value="7.0">7 mm</quantity>
; eyes separated by broad frons; occiput laterally with numerous long somewhat flattened setae; palpus black, clothed in numerous long hair-like setae; mesonotum and legs covered with long hair-like setae; scutum with four vittae.
</paragraph>
<paragraph id="8B8E36EBFFD4FFA57BA7183D71F0FB11" blockId="12.[144,764,277,1927]" pageId="12" pageNumber="36">
<emphasis id="B945EAF9FFD4FFA57BA7183D715CFBD8" box="[181,325,1079,1104]" italics="true" pageId="12" pageNumber="36">Description</emphasis>
.
<emphasis id="B945EAF9FFD4FFA57A47183C718DFBD8" box="[341,404,1078,1104]" italics="true" pageId="12" pageNumber="36">Male</emphasis>
(
<figureCitation id="130A2A6EFFD4FFA57AB5183C721CFBD9" box="[423,517,1078,1105]" captionStart="Figs 13–15" captionStartId="13.[159,211,1832,1856]" captionTargetBox="[202,1384,193,1779]" captionTargetId="figure-21@13.[165,1422,181,1808]" captionTargetPageId="13" captionText="Figs 13–15. Empis barkalovi sp. nov., male. 13, habitus, lateral view (holotype); 14, postabdomen, lateral view; 15, terminalia, lateral view. Abbreviations: cerc, cercus; ej apod, ejaculatory apodeme; epand, epandrium; hypd, hypandrium; ph, phallus. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135543" httpUri="https://zenodo.org/record/10135543/files/figure.png" pageId="12" pageNumber="36">Fig. 13</figureCitation>
). Body length 7.0–
<quantity id="4CC99B0EFFD4FFA57B82185070EAFBFD" box="[144,243,1114,1141]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="12" pageNumber="36" unit="mm" value="7.3">7.3 mm</quantity>
(
<typeStatus id="548A8849FFD4FFA57A1818507167FBFC" box="[266,382,1114,1140]" pageId="12" pageNumber="36" type="holotype">holotype</typeStatus>
,
<quantity id="4CC99B0EFFD4FFA57A80185071EEFBFD" box="[402,503,1114,1141]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="12" pageNumber="36" unit="mm" value="7.1">7.1 mm</quantity>
); wing length
<quantity id="4CC99B0EFFD4FFA579D2185370F5FB11" metricMagnitude="-3" metricUnit="m" metricValue="6.6" metricValueMax="6.7" metricValueMin="6.5" pageId="12" pageNumber="36" unit="mm" value="6.6" valueMax="6.7" valueMin="6.5">6.5– 6.7 mm</quantity>
(
<typeStatus id="548A8849FFD4FFA57BE918747176FB10" box="[251,367,1150,1176]" pageId="12" pageNumber="36" type="holotype">holotype</typeStatus>
,
<quantity id="4CC99B0EFFD4FFA57A69187771C5FB11" box="[379,476,1149,1177]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="12" pageNumber="36" unit="mm" value="6.6">6.6 mm</quantity>
).
</paragraph>
<paragraph id="8B8E36EBFFD4FFA57BA718A8718FF80F" blockId="12.[144,764,277,1927]" pageId="12" pageNumber="36">Head black. Eyes dichoptic, ommatidia of equal size. Frons broad, in upper part broader than distance between outer margins of posterior ocelli, densely dark brown pruinose, with numerous long marginal setae. Face broad, parallel-sided, largely greyish brown pruinose, narrowly shiny along lower margin, bare; clypeus shiny. Occiput somewhat convex laterally, densely brownish grey pruinose, clothed in dense, long, hair-like, black setae, usually with numerous flattened setae laterally (sometimes less distinct). Ocellar tubercle concolorous with occiput, with several subequally long hair-like setae. Antenna black; scape rather short, about 1.5 times as long as pedicel, with long setulae; postpedicel moderately long, 2.0–2.3 times as long as wide, with ventral margin slightly concave in subapical part; stylus about two-thirds as long as postpedicel. Proboscis moderately long; labrum brownish, nearly 1.5 times as long as head height; palpus long, black, with numerous long, hair-like, black setae.</paragraph>
<paragraph id="8B8E36EBFFD4FFA5785D1CB37488FD63" blockId="12.[809,1429,185,1926]" pageId="12" pageNumber="36">Thorax black, densely brownish grey pruinose (except as noted), with black setation; scutum with four distinct brown vittae (in dorsal view). Prosternum bare. Proepisternum with numerous long hair-like setae on lower part, bare on upper part. Antepronotum with numerous hair-like setae dorsally. Postpronotal lobe with numerous hair-like setae of different lengths. Mesonotal setation represented only by numerous long thin setae of subequal robustness; rows of acrostichal and dorsocentral setae separated by broad bare space; acrostichals arranged in 2–3 irregular rows, lacking on prescutellar depression; dorsocentrals multiserial, more numerous before suture. Laterotergite with numerous hair-like setae. Anterior and posterior spiracles black.</paragraph>
<paragraph id="8B8E36EBFFD4FFA5785D1EFF752BFB1F" blockId="12.[809,1429,185,1926]" pageId="12" pageNumber="36">Legs rather robust, entirely black to dark brown, with black hair-like setation; coxae faintly greyish pruinose, other parts of legs subshiny. Fore and mid femora of subequal thickness, hind femur somewhat broader; all femora whitish pubescent ventrally and clothed in long hair-like setae being denser and somewhat stronger on mid and hind femora ventrally. All tibiae with long hair-like setae dorsally and posteriorly (shorter on hind tibia); posteroapical comb with one short seta; all tarsomeres slender; fore tarsomeres 1–3 with longer setae dorsally and posteriorly.</paragraph>
<paragraph id="8B8E36EBFFD4FFA5785D18AB759CFA75" blockId="12.[809,1429,185,1926]" pageId="12" pageNumber="36">
Wing membrane hyaline (in some specimens slightly whitish in some view). Pterostigma pale yellow, narrow. Basal costal seta absent. Veins
<emphasis id="B945EAF9FFD4FFA5783B1907735FFAA6" box="[809,838,1293,1326]" italics="true" pageId="12" pageNumber="36">R 5</emphasis>
and
<emphasis id="B945EAF9FFD4FFA5789C190773B6FAA5" box="[910,943,1293,1325]" italics="true" pageId="12" pageNumber="36">M 1</emphasis>
somewhat divergent just before wing margin; anal vein usually mostly weakened and normally sclerotised only just before wing margin (sometimes normally sclerotised along entire length); radial fork acute. Cell
<emphasis id="B945EAF9FFD4FFA57F88199174A6FA3D" box="[1178,1215,1435,1461]" italics="true" pageId="12" pageNumber="36">dm</emphasis>
short. Anal angle acute, subsequently anal lobe well-developed. Calypter yellowish, brownish fringed. Halter yellow.
</paragraph>
<paragraph id="8B8E36EBFFD4FFA7785D1A0D71B8FDD4" blockId="12.[809,1429,185,1926]" lastBlockId="14.[144,764,185,1926]" lastPageId="14" lastPageNumber="38" pageId="12" pageNumber="36">
Abdomen entirely black, covered only with dense hair-like setae being longer on tergites laterally and on sternites; tergites subshiny, faintly greyish pruinose; sternites slightly denser pruinose. Segments 6 and 7 unmodified; sclerites of segment 8 separated; tergite 8 entire, narrow (in dorsal view), with several long setae; sternite 8 subrectangular (in lateral view), narrow, simple, with numerous long setae. Terminalia (
<figureCitation id="130A2A6EFFD4FFA57E381B2E7353F8EB" captionStart="Figs 13–15" captionStartId="13.[159,211,1832,1856]" captionTargetBox="[202,1384,193,1779]" captionTargetId="figure-21@13.[165,1422,181,1808]" captionTargetPageId="13" captionText="Figs 13–15. Empis barkalovi sp. nov., male. 13, habitus, lateral view (holotype); 14, postabdomen, lateral view; 15, terminalia, lateral view. Abbreviations: cerc, cercus; ej apod, ejaculatory apodeme; epand, epandrium; hypd, hypandrium; ph, phallus. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135543" httpUri="https://zenodo.org/record/10135543/files/figure.png" pageId="12" pageNumber="36">Figs 14, 15</figureCitation>
) small; cercus broadly brownish yellow, brownish near base; epandrial lamella and hypandrium brown. Cerci separated, appressed to epandrium; cercus unilobate, elongate oval (in lateral view) with pointed apex extending beyond apex of epandrial lamella, near base only slightly narrower than epandrial lamella; cercus with scattered dark setulae. Epandrial lamella rather subrectangular, with lower margin broadly ovate posteriorly, covered with thin black setae being denser and longer closer to lower margin. Hypandrium entire, narrow, rim-like, bare. Phallus mostly hidden, zigzag bent, rather thick, with beaker-shaped apex, sulcate in subapical part.
</paragraph>
<caption id="DF4E6663FFD5FFA47B8D1B227288F808" ID-DOI="http://doi.org/10.5281/zenodo.10135543" ID-Zenodo-Dep="10135543" httpUri="https://zenodo.org/record/10135543/files/figure.png" pageId="13" pageNumber="37" startId="13.[159,211,1832,1856]" targetBox="[202,1384,193,1779]" targetPageId="13" targetType="figure">
<paragraph id="8B8E36EBFFD5FFA47B8D1B227288F808" blockId="13.[159,1445,1832,1921]" pageId="13" pageNumber="37">
<emphasis id="B945EAF9FFD5FFA47B8D1B227133F8C8" bold="true" box="[159,298,1832,1856]" pageId="13" pageNumber="37">Figs 13–15.</emphasis>
<taxonomicName id="4C314D68FFD5FFA47A201B2371F3F8C9" authority="Shamshev, 2019" authorityName="Shamshev" authorityYear="2019" box="[306,490,1832,1857]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="37" phylum="Arthropoda" rank="species" species="barkalovi" status="sp. nov.">
<emphasis id="B945EAF9FFD5FFA47A201B2371F3F8C9" box="[306,490,1832,1857]" italics="true" pageId="13" pageNumber="37">Empis barkalovi</emphasis>
</taxonomicName>
<emphasis id="B945EAF9FFD5FFA47AE01B227256F8C8" bold="true" box="[498,591,1832,1856]" pageId="13" pageNumber="37">
<taxonomicNameLabel id="A2765782FFD5FFA47AE01B227256F8C8" box="[498,591,1832,1856]" pageId="13" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="B945EAF9FFD5FFA479B31B2272A4F8C8" bold="true" box="[673,701,1832,1856]" pageId="13" pageNumber="37">13</emphasis>
, habitus, lateral view (holotype);
<emphasis id="B945EAF9FFD5FFA47F541B227478F8C8" bold="true" box="[1094,1121,1832,1856]" pageId="13" pageNumber="37">14</emphasis>
, postabdomen, lateral view;
<emphasis id="B945EAF9FFD5FFA47B8D1B4270A2F8E8" bold="true" box="[159,187,1864,1888]" pageId="13" pageNumber="37">15</emphasis>
, terminalia, lateral view. Abbreviations:
<emphasis id="B945EAF9FFD5FFA479871B4372DBF8E8" box="[661,706,1865,1888]" italics="true" pageId="13" pageNumber="37">cerc</emphasis>
, cercus;
<emphasis id="B945EAF9FFD5FFA4783A1B437365F8E9" box="[808,892,1864,1889]" italics="true" pageId="13" pageNumber="37">ej apod</emphasis>
, ejaculatory apodeme;
<emphasis id="B945EAF9FFD5FFA47F951B4274D4F8E9" box="[1159,1229,1864,1889]" italics="true" pageId="13" pageNumber="37">epand</emphasis>
, epandrium;
<emphasis id="B945EAF9FFD5FFA47E741B427587F8E9" box="[1382,1438,1864,1889]" italics="true" pageId="13" pageNumber="37">hypd</emphasis>
, hypandrium;
<emphasis id="B945EAF9FFD5FFA47A2B1B62714CF809" box="[313,341,1896,1921]" italics="true" pageId="13" pageNumber="37">ph</emphasis>
, phallus. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B8E36EBFFD6FFA77BA71E6F71B9FDF7" blockId="14.[144,764,185,1926]" box="[181,416,613,640]" pageId="14" pageNumber="38">
<emphasis id="B945EAF9FFD6FFA77BA71E6F7114FDF7" box="[181,269,613,639]" italics="true" pageId="14" pageNumber="38">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8B8E36EBFFD6FFA77BA71E807246FC86" blockId="14.[144,764,185,1926]" pageId="14" pageNumber="38">
<emphasis id="B945EAF9FFD6FFA77BA71E807152FD2C" box="[181,331,650,676]" italics="true" pageId="14" pageNumber="38">Comparison</emphasis>
. The new species belongs to the species group known only from the eastern Palaearctic that has not been assigned yet to any subgenus of
<taxonomicName id="4C314D68FFD6FFA77A351EFF7242FC86" authority="(Shamshev, 2001)" baseAuthorityName="Shamshev" baseAuthorityYear="2001" box="[295,603,756,782]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAF9FFD6FFA77A351EFF716AFC86" box="[295,371,757,782]" italics="true" pageId="14" pageNumber="38">Empis</emphasis>
(
<bibRefCitation id="EFA04B1AFFD6FFA77A901EFE724AFC86" author="Shamshev I. V." box="[386,595,756,782]" pageId="14" pageNumber="38" pagination="195 - 227" refId="ref10735" refString="Shamshev I. V. 2001. Thirteen new species of the genus Empis Linnaeus (Diptera: Empididae) from Asiatic part of the Palaearctic region. An international Journal of dipterological Research, 12: 195 - 227." type="journal article" year="2001">Shamshev, 2001</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B8E36EBFFD6FFA77BA71F1271B6FA3E" blockId="14.[144,764,185,1926]" pageId="14" pageNumber="38">
One species of this group (
<taxonomicName id="4C314D68FFD6FFA779131F137145FCDE" authority="Shamshev, 2001" authorityName="Shamshev" authorityYear="2001" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="38" phylum="Arthropoda" rank="species" species="otchontengriensis">
<emphasis id="B945EAF9FFD6FFA779131F1372E2FCBA" box="[513,763,792,818]" italics="true" pageId="14" pageNumber="38">E. otchontengriensis</emphasis>
<bibRefCitation id="EFA04B1AFFD6FFA77B821F367145FCDE" author="Shamshev I. V." box="[144,348,828,854]" pageId="14" pageNumber="38" pagination="195 - 227" refId="ref10735" refString="Shamshev I. V. 2001. Thirteen new species of the genus Empis Linnaeus (Diptera: Empididae) from Asiatic part of the Palaearctic region. An international Journal of dipterological Research, 12: 195 - 227." type="journal article" year="2001">Shamshev, 2001</bibRefCitation>
</taxonomicName>
) has been known from Altai (including the Mongolian part). Both species inhabit similar biotopes occurring in the higher mountainous tundra and tundra-steppe (
<bibRefCitation id="EFA04B1AFFD6FFA779441FAD714FFC6D" author="Shamshev I. V. &amp; Barkalov A. V." pageId="14" pageNumber="38" pagination="319 - 323" refId="ref10855" refString="Shamshev I. V. &amp; Barkalov A. V. 2009. Fauna and zoogeographic characteristic of dance flies (Diptera, Empididae) of Mountain Altai. Euroasian entomological Journal, 8: 319 - 323. (In Russian, with English summary)." type="journal article" year="2009">Shamshev &amp; Barkalov, 2009</bibRefCitation>
). The new species can be readily distinguished from
<taxonomicName id="4C314D68FFD6FFA77A861FE5728FFB80" authorityName="Shamshev" authorityYear="2001" box="[404,662,1006,1032]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="38" phylum="Arthropoda" rank="species" species="otchontengriensis">
<emphasis id="B945EAF9FFD6FFA77A861FE5728FFB80" box="[404,662,1006,1032]" italics="true" pageId="14" pageNumber="38">E. otchontengriensis</emphasis>
</taxonomicName>
by the larger size (body
<quantity id="4CC99B0EFFD6FFA77A62181871F7FBA5" box="[368,494,1042,1069]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="7.3" metricValueMin="7.0" pageId="14" pageNumber="38" unit="mm" value="7.15" valueMax="7.3" valueMin="7.0">7–7.3 mm</quantity>
versus about
<quantity id="4CC99B0EFFD6FFA779B3181B72F6FBA5" box="[673,751,1041,1069]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="38" unit="mm" value="5.0">5 mm</quantity>
), black palpus (vs. yellow), entirely black to dark brown legs (vs. legs with yellowish hind basitarsus) and some other characters. Actually, the new species is most similar to
<taxonomicName id="4C314D68FFD6FFA77AF418A87108FB57" authority="Shamshev, 2001" authorityName="Shamshev" authorityYear="2001" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="38" phylum="Arthropoda" rank="species" species="keberlei">
<emphasis id="B945EAF9FFD6FFA77AF418A87284FB33" box="[486,669,1185,1211]" italics="true" pageId="14" pageNumber="38">Empis keberlei</emphasis>
<bibRefCitation id="EFA04B1AFFD6FFA779B818AB7108FB57" author="Shamshev I. V." pageId="14" pageNumber="38" pagination="195 - 227" refId="ref10735" refString="Shamshev I. V. 2001. Thirteen new species of the genus Empis Linnaeus (Diptera: Empididae) from Asiatic part of the Palaearctic region. An international Journal of dipterological Research, 12: 195 - 227." type="journal article" year="2001">Shamshev, 2001</bibRefCitation>
</taxonomicName>
described from
<collectingCountry id="F326767BFFD6FFA77AC518CF727FFB57" box="[471,614,1221,1247]" name="Uzbekistan" pageId="14" pageNumber="38">Uzbekistan</collectingCountry>
(
<bibRefCitation id="EFA04B1AFFD6FFA7796618CF70CAFA8B" author="Shamshev I. V." pageId="14" pageNumber="38" pagination="195 - 227" refId="ref10735" refString="Shamshev I. V. 2001. Thirteen new species of the genus Empis Linnaeus (Diptera: Empididae) from Asiatic part of the Palaearctic region. An international Journal of dipterological Research, 12: 195 - 227." type="journal article" year="2001">Shamshev, 2001</bibRefCitation>
). However,
<taxonomicName id="4C314D68FFD6FFA77A7618E371EEFA8A" authority="Shamshev, 2019" authorityName="Shamshev" authorityYear="2019" box="[356,503,1256,1283]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="38" phylum="Arthropoda" rank="species" species="barkalovi" status="sp. nov.">
<emphasis id="B945EAF9FFD6FFA77A7618E3716CFA8A" box="[356,373,1257,1282]" italics="true" pageId="14" pageNumber="38">E</emphasis>
.
<emphasis id="B945EAF9FFD6FFA77A9118E271EEFA8A" box="[387,503,1256,1282]" italics="true" pageId="14" pageNumber="38">barkalovi</emphasis>
</taxonomicName>
<emphasis id="B945EAF9FFD6FFA77AEC18E3727AFA8B" bold="true" box="[510,611,1257,1283]" pageId="14" pageNumber="38">
<taxonomicNameLabel id="A2765782FFD6FFA77AEC18E3727AFA8B" box="[510,611,1257,1283]" pageId="14" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C314D68FFD6FFA77B821907710AFAAE" authorityName="Shamshev" authorityYear="2001" box="[144,275,1292,1319]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="38" phylum="Arthropoda" rank="species" species="keberlei">
<emphasis id="B945EAF9FFD6FFA77B82190770B8FAAE" box="[144,161,1293,1318]" italics="true" pageId="14" pageNumber="38">E</emphasis>
.
<emphasis id="B945EAF9FFD6FFA77BA11906710AFAAE" box="[179,275,1292,1318]" italics="true" pageId="14" pageNumber="38">keberlei</emphasis>
</taxonomicName>
primarily in the presence of flattened setae on the occiput, black laterotergal setae (vs. pale in
<taxonomicName id="4C314D68FFD6FFA77BE6195E7160FAE6" authorityName="Shamshev" authorityYear="2001" box="[244,377,1364,1390]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="38" phylum="Arthropoda" rank="species" species="keberlei">
<emphasis id="B945EAF9FFD6FFA77BE6195E711CFAE5" box="[244,261,1364,1389]" italics="true" pageId="14" pageNumber="38">E</emphasis>
.
<emphasis id="B945EAF9FFD6FFA77A0B195E7160FAE6" box="[281,377,1364,1390]" italics="true" pageId="14" pageNumber="38">keberlei</emphasis>
</taxonomicName>
), black thoracic spiracles (vs. pale) and subshiny abdominal tergites (vs. densely whitish grey pruinose).
</paragraph>
<paragraph id="8B8E36EBFFD6FFA77BA719B57237F9CC" blockId="14.[144,764,185,1926]" pageId="14" pageNumber="38">
<emphasis id="B945EAF9FFD6FFA77BA719B5712EFA51" box="[181,311,1471,1497]" italics="true" pageId="14" pageNumber="38">Etymology</emphasis>
. The new species is named in honour of Russian entomologist Anatoly V. Barkalov (
<collectingRegion id="49F5F809FFD6FFA77B8A1A0C712DF9A9" box="[152,308,1542,1569]" country="Russia" name="Novosibirsk" pageId="14" pageNumber="38">Novosibirsk</collectingRegion>
) in recognition of his contribution to the knowledge of insects of
<collectingRegion id="49F5F809FFD6FFA77AF51A207233F9CC" box="[487,554,1578,1604]" country="Russia" name="Altay" pageId="14" pageNumber="38">Altai</collectingRegion>
.
</paragraph>
<paragraph id="8B8E36EBFFD6FFA77BA71A4470FBF904" blockId="14.[144,764,185,1926]" pageId="14" pageNumber="38">
<emphasis id="B945EAF9FFD6FFA77BA71A447157F9E0" box="[181,334,1614,1640]" italics="true" pageId="14" pageNumber="38">Distribution</emphasis>
. Palaearctic:
<collectingCountry id="F326767BFFD6FFA77AED1A44724BF9E0" box="[511,594,1614,1640]" name="Russia" pageId="14" pageNumber="38">Russia</collectingCountry>
(Republic of
<collectingRegion id="49F5F809FFD6FFA77B821A7B70CCF904" box="[144,213,1649,1676]" country="Russia" name="Altay" pageId="14" pageNumber="38">Altai</collectingRegion>
).
</paragraph>
<paragraph id="8B8E36EBFFD6FFA77BA71A9F712DF80E" blockId="14.[144,764,185,1926]" pageId="14" pageNumber="38">
<emphasis id="B945EAF9FFD6FFA77BA71A9F727BF927" box="[181,610,1685,1711]" italics="true" pageId="14" pageNumber="38">Habitat and seasonal occurrence</emphasis>
. According to the label data, the species occurs on meadows and in tundra of higher mountain areas (
<quantity id="4CC99B0EFFD6FFA779B71AD770F6F893" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="3.1" metricValueMin="2.4" pageId="14" pageNumber="38" unit="m" value="2750.0" valueMax="3100.0" valueMin="2400.0">2400– 3100 m</quantity>
). It was collected from late June until late July. One male is pinned together with a subequal in length specimen of
<taxonomicName id="4C314D68FFD6FFA77AB31B427236F8EA" box="[417,559,1864,1890]" class="Insecta" family="Therevidae" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="38" phylum="Arthropoda" rank="family">Therevidae</taxonomicName>
(
<taxonomicName id="4C314D68FFD6FFA779531B4270ABF80E" class="Insecta" family="Rhagionidae" genus="Symphoromyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="38" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B945EAF9FFD6FFA779531B4272E2F8EA" box="[577,763,1864,1890]" italics="true" pageId="14" pageNumber="38">Symphoromyia</emphasis>
sp.
</taxonomicName>
) as a prey.
</paragraph>
</subSubSection>
</treatment>
</document>
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