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<document id="F2296B8BDB6CE7B5EA469411BEFDAD56" ID-CLB-Dataset="46839" ID-DOI="10.11646/zootaxa.4072.5.1" ID-GBIF-Dataset="235dd6cd-8f4e-4d97-abe9-49048dee198a" ID-ISSN="1175-5326" ID-Zenodo-Dep="263020" ID-ZooBank="9C5743A5-6F97-471E-8F60-99A744193FE1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460359453096" checkinUser="plazi" docAuthor="Ray, C. Keith &amp; Armbruster, Jonathan W." docDate="2016" docId="039887CCFFDDFF9294F14B64FD0895AE" docLanguage="en" docName="zootaxa.4072.5.1.pdf" docOrigin="Zootaxa 4072 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aphanotorulus unicolor Steindachner 1908" docType="treatment" docVersion="8" lastPageNumber="526" masterDocId="FFA1FFB4FFCAFF8B94664C6BFF9C9008" masterDocTitle="The genera Isorineloricaria and Aphanotorulus (Siluriformes: Loricariidae) with description of a new species" masterLastPageNumber="539" masterPageNumber="501" pageNumber="524" updateTime="1698294258936" updateUser="ExternalLinkService">
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<mods:title id="2D5ACD984ABE20CA4CA9C6E86C0A6D31">The genera Isorineloricaria and Aphanotorulus (Siluriformes: Loricariidae) with description of a new species</mods:title>
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<mods:namePart id="5122E9061A5EA5710A9D669BF1D1AF18">Armbruster, Jonathan W.</mods:namePart>
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<treatment id="039887CCFFDDFF9294F14B64FD0895AE" ID-DOI="http://doi.org/10.5281/zenodo.6055320" ID-GBIF-Taxon="119409382" ID-Zenodo-Dep="6055320" LSID="urn:lsid:plazi:treatment:039887CCFFDDFF9294F14B64FD0895AE" httpUri="http://treatment.plazi.org/id/039887CCFFDDFF9294F14B64FD0895AE" lastPageId="25" lastPageNumber="526" pageId="23" pageNumber="524">
<subSubSection id="C32B6551FFDDFF9C94F14B64FF659743" pageId="23" pageNumber="524" type="nomenclature">
<paragraph id="8B8E36DAFFDDFF9C94F14B64FD209722" blockId="23.[151,700,1807,1867]" box="[151,700,1807,1834]" pageId="23" pageNumber="524">
<heading id="D0C681B6FFDDFF9C94F14B64FD209722" bold="true" box="[151,700,1807,1834]" fontSize="11" level="1" pageId="23" pageNumber="524" reason="1">
<taxonomicName id="4C314D59FFDDFF9C94F14B64FD209722" ID-CoL="FBNL" authority="Steindachner 1908" authorityName="Steindachner" authorityYear="1908" box="[151,700,1807,1834]" class="Actinopterygii" family="Loricariidae" genus="Aphanotorulus" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="524" phylum="Chordata" rank="species" species="unicolor">
<emphasis id="B945EAC8FFDDFF9C94F14B64FD209722" bold="true" box="[151,700,1807,1834]" pageId="23" pageNumber="524">
<emphasis id="B945EAC8FFDDFF9C94F14B64FE209721" bold="true" box="[151,444,1807,1833]" italics="true" pageId="23" pageNumber="524">Aphanotorulus unicolor</emphasis>
(
<bibRefCitation id="EFA04B2BFFDDFF9C95AC4B7BFD289722" author="Steindachner" box="[458,692,1808,1834]" pageId="23" pageNumber="524" refString="Steindachner, F. (1908) Uber zwei neue Siluroiden und zwei Curimatus - Arten, sowie uber eine Varietat von Ancistrus vittalus [sic, vittatus] aus dem Amazonasgebiete innerhalb Brasiliens. Anzeiger der Kaiserlichen Akademie der Wissenschaften, Mathematisch- Naturwissenschaftlichen Classe, 45, 163 - 168." type="journal article" year="1908">Steindachner 1908</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8E36DAFFDDFF9C94F14B59FF659743" blockId="23.[151,700,1807,1867]" box="[151,249,1842,1867]" pageId="23" pageNumber="524">
(
<figureCitation id="130A2A5FFFDDFF9C94F94B59FF6D9743" box="[159,241,1842,1867]" captionStart="FIGURE 14" captionStartId="24.[151,250,1828,1850]" captionTargetBox="[151,1434,193,1806]" captionTargetId="figure@24.[151,1435,193,1806]" captionTargetPageId="24" captionText="FIGURE 14. Lateral, dorsal, and ventral view of Aphanotorulus unicolor, ZMA 116640 (Aphanotorulus frankei holotype), 105.4 mm SL Photographs by K. S. Cummings. Scale = 1 cm." httpUri="https://zenodo.org/record/263033/files/figure.png" pageId="23" pageNumber="524">Fig. 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32B6551FFDDFF9294F14B11FDC89120" lastPageId="25" lastPageNumber="526" pageId="23" pageNumber="524" type="reference_group">
<paragraph id="8B8E36DAFFDDFF9C94F14B11FECD97A7" blockId="23.[151,1436,1914,2028]" pageId="23" pageNumber="524">
<treatmentCitationGroup id="AB2111F4FFDDFF9C94F14B11FECD97A7" pageId="23" pageNumber="524">
<taxonomicName id="4C314D59FFDDFF9C94F14B11FDF39798" ID-CoL="4K2YD" authority="Steindachner, 1908: 164" authorityName="Steindachner" authorityPageNumber="164" authorityYear="1908" box="[151,623,1914,1936]" family="Loricariidae" genus="Plecostomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="524" phylum="Chordata" rank="species" species="unicolor">
<emphasis id="B945EAC8FFDDFF9C94F14B11FEEE9798" box="[151,370,1914,1936]" italics="true" pageId="23" pageNumber="524">Plecostomus unicolor</emphasis>
<treatmentCitation id="0A9010CBFFDDFF9C951E4B11FDF39798" author="Steindachner" box="[376,623,1914,1936]" page="164" pageId="23" pageNumber="524" year="1908">
<bibRefCitation id="EFA04B2BFFDDFF9C951E4B11FDA19798" author="Steindachner" box="[376,573,1914,1936]" pageId="23" pageNumber="524" refString="Steindachner, F. (1908) Uber zwei neue Siluroiden und zwei Curimatus - Arten, sowie uber eine Varietat von Ancistrus vittalus [sic, vittatus] aus dem Amazonasgebiete innerhalb Brasiliens. Anzeiger der Kaiserlichen Akademie der Wissenschaften, Mathematisch- Naturwissenschaftlichen Classe, 45, 163 - 168." type="journal article" year="1908">Steindachner, 1908</bibRefCitation>
: 164
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548A8878FFDDFF9C961E4B10FD369798" box="[632,682,1915,1936]" pageId="23" pageNumber="524">Type</typeStatus>
locality: Rio Purús.
<typeStatus id="548A8878FFDDFF9C971F4B10FC7E9798" box="[889,994,1915,1936]" pageId="23" pageNumber="524" type="lectotype">Lectotype</typeStatus>
: NMW 44271: 2, designated by
<treatmentCitation id="0A9010CBFFDDFF9C91414B11FED197A7" author="Armbruster" page="254" pageId="23" pageNumber="524" year="1998">
<bibRefCitation id="EFA04B2BFFDDFF9C91414B11FED197A7" author="Armbruster" pageId="23" pageNumber="524" refString="Armbruster, J. W. (1998) Review of the loricariid catfish genus Aphanotorulus and redescription of A. unicolor (Teleostei: Siluriformes). Ichthyological Exploration of Freshwaters, 8, 253 - 262." type="journal article" year="1998" yearSuffix="b">Armbruster (1998b: 254)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8E36DAFFDDFF9C94F14BDCFD9797E4" blockId="23.[151,1436,1914,2028]" pageId="23" pageNumber="524">
<treatmentCitationGroup id="AB2111F4FFDDFF9C94F14BDCFD9797E4" pageId="23" pageNumber="524">
<taxonomicName id="4C314D59FFDDFF9C94F14BDCFDD497C5" ID-CoL="4K2X9" authority="Fowler, 1913: 571" authorityName="Fowler" authorityPageNumber="571" authorityYear="1913" box="[151,584,1975,1997]" family="Loricariidae" genus="Plecostomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="524" phylum="Chordata" rank="species" species="madeirae">
<emphasis id="B945EAC8FFDDFF9C94F14BDCFEE397C5" box="[151,383,1975,1997]" italics="true" pageId="23" pageNumber="524">Plecostomus madeirae</emphasis>
<treatmentCitation id="0A9010CBFFDDFF9C95EE4BDCFDD497C5" author="Fowler" box="[392,584,1975,1997]" page="571" pageId="23" pageNumber="524" year="1913">
<bibRefCitation id="EFA04B2BFFDDFF9C95EE4BDCFD8F97C5" author="Fowler" box="[392,531,1975,1997]" pageId="23" pageNumber="524" refString="Fowler, H. W. (1913) Fishes from the Madeira River, Brazil. Proceedings of the Academy of Natural Sciences, Philadelphia, 65, 517 - 579." type="journal article" year="1913">Fowler, 1913</bibRefCitation>
: 571
</treatmentCitation>
</taxonomicName>
, fig. 23.
<typeStatus id="548A8878FFDDFF9C96CE4BD3FD4697C5" box="[680,730,1976,1997]" pageId="23" pageNumber="524">Type</typeStatus>
locality: Madeira River, about
<quantity id="4CC99B3FFFDDFF9C90454BDCFB1597C5" box="[1059,1161,1975,1997]" metricMagnitude="5" metricUnit="m" metricValue="3.218688" pageId="23" pageNumber="524" unit="mi" value="200.0">200 miles</quantity>
east of W. long.
<geoCoordinate id="EE05501DFFDDFF9C91264BDCFA0497C5" box="[1344,1432,1975,1997]" direction="west" orientation="longitude" pageId="23" pageNumber="524" precision="925" value="-63.9">63º54'W</geoCoordinate>
,
<collectingCountry id="F326764AFFDDFF9C94A14BBDFE9497E4" box="[199,264,2006,2028]" name="Brazil" pageId="23" pageNumber="524">Brazil</collectingCountry>
.
<typeStatus id="548A8878FFDDFF9C95774BBDFEEF97E4" box="[273,371,2006,2028]" pageId="23" pageNumber="524" type="holotype">Holotype</typeStatus>
: ANSP 39312.
</treatmentCitationGroup>
</paragraph>
<caption id="DF4E6652FFD2FF9394F14B4FFC9A9750" httpUri="https://zenodo.org/record/263033/files/figure.png" pageId="24" pageNumber="525" targetBox="[151,1434,193,1806]" targetPageId="24">
<paragraph id="8B8E36DAFFD2FF9394F14B4FFC9A9750" blockId="24.[151,1436,1828,1880]" pageId="24" pageNumber="525">
<emphasis id="B945EAC8FFD2FF9394F14B4FFEBE9731" bold="true" box="[151,290,1828,1850]" pageId="24" pageNumber="525">FIGURE 14.</emphasis>
Lateral, dorsal, and ventral view of
<taxonomicName id="4C314D59FFD2FF9396C24B4FFC049732" ID-CoL="FBNL" box="[676,920,1828,1850]" class="Actinopterygii" family="Loricariidae" genus="Aphanotorulus" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="525" phylum="Chordata" rank="species" species="unicolor">
<emphasis id="B945EAC8FFD2FF9396C24B4FFC049732" box="[676,920,1828,1850]" italics="true" pageId="24" pageNumber="525">Aphanotorulus unicolor</emphasis>
</taxonomicName>
, ZMA 116640 (
<taxonomicName id="4C314D59FFD2FF9390224B4FFAB09732" ID-CoL="FBNJ" box="[1092,1324,1828,1850]" class="Actinopterygii" family="Loricariidae" genus="Aphanotorulus" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="525" phylum="Chordata" rank="species" species="frankei">
<emphasis id="B945EAC8FFD2FF9390224B4FFAB09732" box="[1092,1324,1828,1850]" italics="true" pageId="24" pageNumber="525">Aphanotorulus frankei</emphasis>
</taxonomicName>
holotype), 105.4 mm SL Photographs by K.S. Cummings. Scale = 1 cm.
</paragraph>
</caption>
<paragraph id="8B8E36DAFFD2FF9394F14B14FE9197BC" blockId="24.[151,1435,1919,2034]" pageId="24" pageNumber="525">
<treatmentCitationGroup id="AB2111F4FFD2FF9394F14B14FE9197BC" pageId="24" pageNumber="525">
<taxonomicName id="4C314D59FFD2FF9394F14B14FDBF979D" ID-CoL="4K2XS" authority="Pearson, 1924: 20" authorityName="Pearson" authorityPageNumber="20" authorityYear="1924" box="[151,547,1919,1942]" family="Loricariidae" genus="Plecostomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="525" phylum="Chordata" rank="species" species="popoi">
<emphasis id="B945EAC8FFD2FF9394F14B14FEC7979E" box="[151,347,1919,1942]" italics="true" pageId="24" pageNumber="525">Plecostomus popoi</emphasis>
<treatmentCitation id="0A9010CBFFD2FF9395024BEBFDBF979D" author="Pearson" box="[356,547,1919,1941]" page="20" pageId="24" pageNumber="525" year="1924">
<bibRefCitation id="EFA04B2BFFD2FF9395024BEBFE66979D" author="Pearson" box="[356,506,1919,1941]" pageId="24" pageNumber="525" refString="Pearson, N. E. (1924) The fishes of the eastern slope of the Andes. I. The fishes of the Rio Beni basin, Bolivia, collected by the Mulford expedition. Indiana University Studies, 11, 1 - 83, pls. 1 - 12." type="journal article" year="1924">Pearson, 1924</bibRefCitation>
: 20
</treatmentCitation>
</taxonomicName>
, pl. 3 (fig. 2).
<typeStatus id="548A8878FFD2FF9396A34BEBFD6B979D" box="[709,759,1920,1941]" pageId="24" pageNumber="525">Type</typeStatus>
locality:
<taxonomicName id="4C314D59FFD2FF9397384B14FC05979D" box="[862,921,1919,1941]" family="Loricariidae" genus="Plecostomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="525" phylum="Chordata" rank="species" species="popoi">Popoi</taxonomicName>
River, Upper Río Beni,
<collectingCountry id="F326764AFFD2FF9390C74B14FB73979D" box="[1185,1263,1919,1941]" name="Bolivia" pageId="24" pageNumber="525">Bolivia</collectingCountry>
.
<typeStatus id="548A8878FFD2FF93909D4B14FAC1979D" box="[1275,1373,1919,1941]" pageId="24" pageNumber="525" type="holotype">Holotype</typeStatus>
: CAS 77346.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8E36DAFFD2FF9394F14BD6FA1397FA" blockId="24.[151,1435,1919,2034]" pageId="24" pageNumber="525">
<treatmentCitationGroup id="AB2111F4FFD2FF9394F14BD6FA1397FA" infered="true" pageId="24" pageNumber="525">
<taxonomicName id="4C314D59FFD2FF9394F14BD6FD1397DB" ID-CoL="7836N" authority="La Monte, 1935: 1" authorityName="La Monte" authorityPageNumber="1" authorityYear="1935" box="[151,655,1981,2003]" family="Loricariidae" genus="Plecostomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="24" pageNumber="525" phylum="Chordata" rank="species" species="micropunctatus">
<emphasis id="B945EAC8FFD2FF9394F14BD6FE2297DB" box="[151,446,1981,2003]" italics="true" pageId="24" pageNumber="525">Plecostomus micropunctatus</emphasis>
<treatmentCitation id="0A9010CBFFD2FF9395A14BD5FD1397DB" author="La" box="[455,655,1981,2003]" page="1" pageId="24" pageNumber="525" year="1935">
<bibRefCitation id="EFA04B2BFFD2FF9395A14BD5FDE897DB" author="La" box="[455,628,1981,2003]" pageId="24" pageNumber="525" refString="La Monte, F. (1935) Fishes from Rio Jurua and Rio Purus, Brazilian Amazonas. American Museum Novitates, 784, 1 - 8." type="journal article" year="1935">La Monte, 1935</bibRefCitation>
: 1
</treatmentCitation>
</taxonomicName>
, fig. 1.
<typeStatus id="548A8878FFD2FF93968E4BD5FC8697DB" box="[744,794,1982,2003]" pageId="24" pageNumber="525">Type</typeStatus>
locality: Rio Purus: [...] in the vicinity of the mouth of Rio Macauhan, a tributary of Rio Iaco which, in turn, is a tributary of Rio Purus (
<geoCoordinate id="EE05501DFFD2FF9397B44BB7FBB597FA" box="[978,1065,2012,2034]" direction="west" orientation="longitude" pageId="24" pageNumber="525" precision="925" value="-70.25">70º15'W</geoCoordinate>
,
<geoCoordinate id="EE05501DFFD2FF9390524BB7FBE497FA" box="[1076,1144,2012,2034]" direction="south" orientation="latitude" pageId="24" pageNumber="525" precision="925" value="-7.5">7º30'S</geoCoordinate>
).
<typeStatus id="548A8878FFD2FF9390EC4BB7FB7097FA" box="[1162,1260,2012,2034]" pageId="24" pageNumber="525" type="holotype">Holotype</typeStatus>
: AMNH 12598.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8E36DAFFD3FF9294F14CF3FE5C90C4" blockId="25.[151,1437,152,296]" pageId="25" pageNumber="526">
<taxonomicName id="4C314D59FFD3FF9294F14CF3FDD890A6" authority="Fowler, 1940: 81" authorityName="Fowler" authorityPageNumber="81" authorityYear="1940" box="[151,580,152,174]" family="Loricariidae" genus="Plecostomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="526" phylum="Chordata" rank="species" species="chaparae">
<emphasis id="B945EAC8FFD3FF9294F14CF3FE1D90A6" box="[151,385,152,174]" italics="true" pageId="25" pageNumber="526">Plecostomus chaparae</emphasis>
Fowler, 1940: 81
</taxonomicName>
, figs. 3840.
<typeStatus id="548A8878FFD3FF9296BA4CF2FC9290A6" box="[732,782,153,174]" pageId="25" pageNumber="526">Type</typeStatus>
locality: Boca Chapare, Río Chimore, Cochabamba,
<collectingCountry id="F326764AFFD3FF92912D4CF3FA0590A6" box="[1355,1433,152,174]" name="Bolivia" pageId="25" pageNumber="526">Bolivia</collectingCountry>
.
<typeStatus id="548A8878FFD3FF9294A14CDDFEB590C4" box="[199,297,182,204]" pageId="25" pageNumber="526" type="holotype">Holotype</typeStatus>
: ANSP 69067.
</paragraph>
<paragraph id="8B8E36DAFFD3FF9294F14CBEFDC89120" blockId="25.[151,1437,152,296]" pageId="25" pageNumber="526">
<treatmentCitationGroup id="AB2111F4FFD3FF9294F14CBEFDC89120" infered="true" pageId="25" pageNumber="526">
<taxonomicName id="4C314D59FFD3FF9294F14CBEFD5290E3" ID-CoL="FBNJ" authority="Isbrucker &amp; Nijssen, 1983: 108" authorityName="Isbrucker &amp; Nijssen" authorityPageNumber="108" authorityYear="1983" box="[151,718,213,235]" class="Actinopterygii" family="Loricariidae" genus="Aphanotorulus" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="526" phylum="Chordata" rank="species" species="frankei">
<emphasis id="B945EAC8FFD3FF9294F14CBEFEE190E3" box="[151,381,213,235]" italics="true" pageId="25" pageNumber="526">Aphanotorulus frankei</emphasis>
<treatmentCitation id="0A9010CBFFD3FF9295E34CBEFD5290E3" author="Isbrucker" box="[389,718,213,235]" page="108" pageId="25" pageNumber="526" year="1983">
<bibRefCitation id="EFA04B2BFFD3FF9295E34CBEFD0690E3" author="Isbrucker" box="[389,666,213,235]" pageId="25" pageNumber="526" refString="Isbrucker, I. J. H. &amp; Nijssen, H. (1983) Aphanotorulus frankei, une espece et un genre nouveaux de poissons-chats cuirasses du Bassin du Rio Ucayali au Perou (Pisces, Siluriformes, Loricariidae). Revue francaise d'Aquariologie Herpetologie, 9 (for 1982), 105 - 110." type="journal article" year="1983">Isbrücker &amp; Nijssen, 1983</bibRefCitation>
: 108
</treatmentCitation>
</taxonomicName>
, figs. 12.
<typeStatus id="548A8878FFD3FF9297254CBDFCE990E3" box="[835,885,214,235]" pageId="25" pageNumber="526">Type</typeStatus>
locality: [...] du
<collectingCountry id="F326764AFFD3FF9290414CBEFBF990E3" box="[1063,1125,213,235]" name="Peru" pageId="25" pageNumber="526">Pérou</collectingCountry>
, Dept. Ucayali, Prov. Coronel Portillo, système du Rio Aguaytia. Rio Neshua (environ
<geoCoordinate id="EE05501DFFD3FF9297634C9FFCC89102" box="[773,852,244,266]" direction="south" orientation="latitude" pageId="25" pageNumber="526" precision="925" value="-8.6">08º36'S</geoCoordinate>
,
<geoCoordinate id="EE05501DFFD3FF9297394C9FFC259102" box="[863,953,244,266]" direction="west" orientation="longitude" pageId="25" pageNumber="526" precision="925" value="-74.833336">74º50'W</geoCoordinate>
),
<quantity id="4CC99B3FFFD3FF9297AC4C9FFB969102" box="[970,1034,244,266]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="526" unit="km" value="60.0">60 km</quantity>
S.
<collectingCountry id="F326764AFFD3FF9290424C9FFBA69102" box="[1060,1082,244,266]" name="Iceland" pageId="25" pageNumber="526">O</collectingCountry>
. de Pucallpa, le long de la route de Tingo Maria.
<typeStatus id="548A8878FFD3FF9295344D79FE289120" box="[338,436,274,296]" pageId="25" pageNumber="526" type="holotype">Holotype</typeStatus>
: ZMA 116.640.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32B6551FFD3FF9294F14D3EFB9A9166" box="[151,1030,341,366]" pageId="25" pageNumber="526" type="materials_examined">
<paragraph id="8B8E36DAFFD3FF9294F14D3EFB9A9166" blockId="25.[151,1437,341,1446]" box="[151,1030,341,366]" pageId="25" pageNumber="526">
<emphasis id="B945EAC8FFD3FF9294F14D3EFE1C9166" bold="true" box="[151,384,341,366]" pageId="25" pageNumber="526">Material examined.</emphasis>
See
<bibRefCitation id="EFA04B2BFFD3FF9295D04D3EFD6A9166" author="Armbruster" box="[438,758,341,366]" pageId="25" pageNumber="526" refString="Armbruster, J. W. &amp; Page, L. M. (1996) Redescription of Aphanotorulus (Teleostei: Loricariidae) with description of one new species, A. ammophilus, from the Rio Orinoco Basin. Copeia, 1996, 379 - 389. http: // dx. doi. org / 10.2307 / 1446854" type="journal article" year="1996">Armbruster and Page (1996)</bibRefCitation>
and
<bibRefCitation id="EFA04B2BFFD3FF92974B4D3EFB9F9166" author="Armbruster" box="[813,1027,341,366]" pageId="25" pageNumber="526" refString="Armbruster, J. W. (1998) Review of the loricariid catfish genus Aphanotorulus and redescription of A. unicolor (Teleostei: Siluriformes). Ichthyological Exploration of Freshwaters, 8, 253 - 262." type="journal article" year="1998">Armbruster (1998)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B6551FFD3FF9294A14D12FD9D92BA" pageId="25" pageNumber="526" type="diagnosis">
<paragraph id="8B8E36DAFFD3FF9294A14D12FD9D92BA" blockId="25.[151,1437,341,1446]" pageId="25" pageNumber="526">
<emphasis id="B945EAC8FFD3FF9294A14D12FEDD919A" bold="true" box="[199,321,377,402]" pageId="25" pageNumber="526">Diagnosis.</emphasis>
<taxonomicName id="4C314D59FFD3FF92952E4D12FDC99199" box="[328,597,377,401]" class="Actinopterygii" family="Loricariidae" genus="Aphanotorulus" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="526" phylum="Chordata" rank="species" species="unicolor">
<emphasis id="B945EAC8FFD3FF92952E4D12FDC99199" box="[328,597,377,401]" italics="true" pageId="25" pageNumber="526">Aphanotorulus unicolor</emphasis>
</taxonomicName>
differs from all other members of
<taxonomicName id="4C314D59FFD3FF9297BD4D12FB129199" box="[987,1166,377,401]" class="Actinopterygii" family="Loricariidae" genus="Isorineloricaria" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="526" phylum="Chordata" rank="genus">
<emphasis id="B945EAC8FFD3FF9297BD4D12FB129199" box="[987,1166,377,401]" italics="true" pageId="25" pageNumber="526">Isorineloricaria</emphasis>
</taxonomicName>
, except
<taxonomicName id="4C314D59FFD3FF92908A4D12FA099199" box="[1260,1429,377,401]" class="Actinopterygii" family="Loricariidae" genus="Aphanotorulus" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="526" phylum="Chordata" rank="species" species="ammophilus">
<emphasis id="B945EAC8FFD3FF92908A4D12FA099199" box="[1260,1429,377,401]" italics="true" pageId="25" pageNumber="526">A. ammophilus</emphasis>
</taxonomicName>
, by its numerous buccal papillae (vs. single, large papilla).
<taxonomicName id="4C314D59FFD3FF9297014DF6FBE791BD" box="[871,1147,413,437]" class="Actinopterygii" family="Loricariidae" genus="Aphanotorulus" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="526" phylum="Chordata" rank="species" species="unicolor">
<emphasis id="B945EAC8FFD3FF9297014DF6FBE791BD" box="[871,1147,413,437]" italics="true" pageId="25" pageNumber="526">Aphanotorulus unicolor</emphasis>
</taxonomicName>
is diagnosable from
<taxonomicName id="4C314D59FFD3FF9291E24DF5FEBC91D1" class="Actinopterygii" family="Loricariidae" genus="Aphanotorulus" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="526" phylum="Chordata" rank="species" species="ammophilus">
<emphasis id="B945EAC8FFD3FF9291E24DF5FEBC91D1" italics="true" pageId="25" pageNumber="526">A. ammophilus</emphasis>
</taxonomicName>
by the absence of a ridge on the compound pterotic (vs. ridge present in
<taxonomicName id="4C314D59FFD3FF9290324DAAFB6791D1" box="[1108,1275,449,473]" class="Actinopterygii" family="Loricariidae" genus="Aphanotorulus" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="526" phylum="Chordata" rank="species" species="ammophilus">
<emphasis id="B945EAC8FFD3FF9290324DAAFB6791D1" box="[1108,1275,449,473]" italics="true" pageId="25" pageNumber="526">A. ammophilus</emphasis>
</taxonomicName>
). The urohyal is also uniquely modified in
<taxonomicName id="4C314D59FFD3FF9295844D8DFDFD91F5" box="[482,609,485,509]" class="Actinopterygii" family="Loricariidae" genus="Aphanotorulus" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="526" phylum="Chordata" rank="species" species="unicolor">
<emphasis id="B945EAC8FFD3FF9295844D8DFDFD91F5" box="[482,609,485,509]" italics="true" pageId="25" pageNumber="526">A. unicolor</emphasis>
</taxonomicName>
. The urohyal in
<taxonomicName id="4C314D59FFD3FF9297424D8DFC3891F5" box="[804,932,485,509]" class="Actinopterygii" family="Loricariidae" genus="Aphanotorulus" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="526" phylum="Chordata" rank="species" species="unicolor">
<emphasis id="B945EAC8FFD3FF9297424D8DFC3891F5" box="[804,932,485,509]" italics="true" pageId="25" pageNumber="526">A. unicolor</emphasis>
</taxonomicName>
has arms, which are thin and elongate, that overlap with the anterohyal (vs. short lateral arms of the urohyal that do not overlap the anterohyal in most other Loricariids) (
<bibRefCitation id="EFA04B2BFFD3FF92954F4E46FE73924E" author="Armbruster" box="[297,495,557,582]" pageId="25" pageNumber="526" refString="Armbruster, J. W. (1998) Review of the loricariid catfish genus Aphanotorulus and redescription of A. unicolor (Teleostei: Siluriformes). Ichthyological Exploration of Freshwaters, 8, 253 - 262." type="journal article" year="1998">Armbruster 1998</bibRefCitation>
). In specimens larger than
<quantity id="4CC99B3FFFD3FF92977E4E46FCFC924E" box="[792,864,557,582]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="25" pageNumber="526" unit="mm" value="75.0">75mm</quantity>
SL,
<taxonomicName id="4C314D59FFD3FF9297F34E45FB8F924D" box="[917,1043,557,581]" class="Actinopterygii" family="Loricariidae" genus="Aphanotorulus" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="526" phylum="Chordata" rank="species" species="unicolor">
<emphasis id="B945EAC8FFD3FF9297F34E45FB8F924D" box="[917,1043,557,581]" italics="true" pageId="25" pageNumber="526">A. unicolor</emphasis>
</taxonomicName>
has a larger orbit diameter to snout length ratio (28.042.2% vs. 18.527.9%), a larger caudal depth to pectoral spine length ratio (25.635.1% vs. 19.825.5%), and a smaller pectoral spine length to thorax length ratio (99.3116.9% vs. 117.0136.3%) than
<taxonomicName id="4C314D59FFD3FF9291E24E1DFE6092BA" authority="Armbruster 1998" authorityName="Armbruster" authorityYear="1998" class="Actinopterygii" family="Loricariidae" genus="Aphanotorulus" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="526" phylum="Chordata" rank="species" species="ammophilus">
<emphasis id="B945EAC8FFD3FF9291E24E1DFEBC92B9" italics="true" pageId="25" pageNumber="526">A. ammophilus</emphasis>
(
<bibRefCitation id="EFA04B2BFFD3FF9295484EF2FE6892BA" author="Armbruster" box="[302,500,665,690]" pageId="25" pageNumber="526" refString="Armbruster, J. W. (1998) Review of the loricariid catfish genus Aphanotorulus and redescription of A. unicolor (Teleostei: Siluriformes). Ichthyological Exploration of Freshwaters, 8, 253 - 262." type="journal article" year="1998">Armbruster 1998</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B6551FFD3FF9294A14ED6FD039556" pageId="25" pageNumber="526" type="description">
<paragraph id="8B8E36DAFFD3FF9294A14ED6FCD09316" blockId="25.[151,1437,341,1446]" pageId="25" pageNumber="526">
<emphasis id="B945EAC8FFD3FF9294A14ED6FEC592DE" bold="true" box="[199,345,701,726]" pageId="25" pageNumber="526">Description.</emphasis>
Morphometric data given in
<tableCitation id="C6B30361FFD3FF9296C04ED6FD6192DE" box="[678,765,701,726]" captionStart="TABLE 2" captionStartId="8.[151,239,152,175]" captionTargetBox="[157,1389,202,1261]" captionTargetPageId="8" captionText="TABLE 2. Selected morphometrics of Aphanotorulus ammophilus and A. unicolor." httpUri="http://table.plazi.org/id/DF4E6652FFC2FF8394F14CF3FB9390A7" pageId="25" pageNumber="526" tableUuid="DF4E6652FFC2FF8394F14CF3FB9390A7">Table 2</tableCitation>
. Head slightly compressed with raised supraoccipital crest; crest ending abruptly with posterior edge perpendicular to head. Large flap covering posterior opening of nare. Interorbital surface flat. Nape increasing in height posteriorly.
</paragraph>
<paragraph id="8B8E36DAFFD3FF9294A14F42FDBC9382" blockId="25.[151,1437,341,1446]" pageId="25" pageNumber="526">Dorsal-fin spine slightly shorter than or equal in length to first dorsal-fin ray. Pectoral-fin spine even with point of insertion on pelvic fin. Pelvic fin spine comes even to insertion of the anal fin. Adipose-fin membrane extending to end of fourth adipose base plate.</paragraph>
<paragraph id="8B8E36DAFFD3FF9294A14FFEFDE8946A" blockId="25.[151,1437,341,1446]" pageId="25" pageNumber="526">Lateral line plates 2832 (mode 29; n = 307); dorsal fin base plates six to nine (mode seven; n = 306); folded dorsal plates 1013 (mode 12; n = 300); plates between dorsal and adipose fins six to 10 (mode eight; n = 305); adipose fin plates two to four (mode three; n =305); anal-fin base plates two to three (mode three; n =307); plates from anal fin insertion to last plate on caudal peduncle 1316 (mode =14; n =307); plates in folded pectoral fin four to seven (mode five; n =302); number of teeth on dentary four to 18 (mode =12; n =302); number of teeth on premaxilla four to 17 (mode =12; n =304).</paragraph>
<paragraph id="8B8E36DAFFD3FF9294A14806FF4694C6" blockId="25.[151,1437,341,1446]" pageId="25" pageNumber="526">
<emphasis id="B945EAC8FFD3FF9294A14806FE22948E" bold="true" box="[199,446,1133,1158]" pageId="25" pageNumber="526">Sexual Dimorphism.</emphasis>
Breeding males have elongate, unicuspid teeth. Hypertrophied odontodes also present on the posterior dorsal and lateral plates. Odontodes also present on the adipose- fin spine and upper caudal- fin spine.
</paragraph>
<paragraph id="8B8E36DAFFD3FF9294A148B2FD039556" blockId="25.[151,1437,341,1446]" pageId="25" pageNumber="526">
<emphasis id="B945EAC8FFD3FF9294A148B2FE1294FA" bold="true" box="[199,398,1241,1266]" pageId="25" pageNumber="526">Color in alcohol.</emphasis>
Light tan to white background. Ventral surface white. Dorsal and lateral surfaces with dark, round spots. Spots small in size, less than half the width of the pupil. Spots form longitudinal rows in some specimens, more random in other specimens. Spots on caudal fin create vertical bars when fin is compressed. Spots on dorsal fin lie anterior to each dorsal fin ray.
</paragraph>
</subSubSection>
<subSubSection id="C32B6551FFD3FF9294A14902FD0895AE" pageId="25" pageNumber="526" type="distribution">
<paragraph id="8B8E36DAFFD3FF9294A14902FD0895AE" blockId="25.[151,1437,341,1446]" pageId="25" pageNumber="526">
<emphasis id="B945EAC8FFD3FF9294A14902FEFC958A" bold="true" box="[199,352,1385,1410]" pageId="25" pageNumber="526">Distribution.</emphasis>
A wide-ranging species, with populations known from upper Amazon tributaries of
<collectingCountry id="F326764AFFD3FF9291254902FA04958A" box="[1347,1432,1385,1410]" name="Bolivia" pageId="25" pageNumber="526">Bolivia</collectingCountry>
,
<collectingCountry id="F326764AFFD3FF9294F149E6FF4295AE" box="[151,222,1421,1446]" name="Brazil" pageId="25" pageNumber="526">Brazil</collectingCountry>
,
<collectingCountry id="F326764AFFD3FF92948F49E6FEC695AE" box="[233,346,1421,1446]" name="Colombia" pageId="25" pageNumber="526">Colombia</collectingCountry>
,
<collectingCountry id="F326764AFFD3FF92950049E6FE5995AE" box="[358,453,1421,1446]" name="Ecuador" pageId="25" pageNumber="526">Ecuador</collectingCountry>
, and
<collectingCountry id="F326764AFFD3FF92959949E5FDAE95AD" box="[511,562,1422,1445]" name="Peru" pageId="25" pageNumber="526">Peru</collectingCountry>
(
<figureCitation id="130A2A5FFFD3FF92962449E6FD1A95AE" box="[578,646,1421,1446]" captionStart="FIGURE 6" captionStartId="10.[151,250,1736,1758]" captionTargetBox="[332,1254,1058,1714]" captionTargetId="figure@10.[325,1262,1048,1715]" captionTargetPageId="10" captionText="FIGURE 6. Distribution of Aphanotorulus ammophilus and A. unicolor. Circles represent A. ammophilus, squares represent A. unicolor. Open shapes represent type localities. From Armbruster (1998)." httpUri="https://zenodo.org/record/263026/files/figure.png" pageId="25" pageNumber="526">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>