546 lines
63 KiB
XML
546 lines
63 KiB
XML
<document id="C82DDCF6CF7A3750D6BC2B22B240C77B" ID-DOI="10.5281/zenodo.186138" ID-GBIF-Dataset="8cf8a401-bc91-4a30-99b8-27492f4f254e" ID-ISSN="1175-5326" ID-Zenodo-Dep="186138" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461046376021" checkinUser="plazi" docAuthor="De, Jeferson L., Lindquist, Evert E. & De, Gilberto J." docDate="2009" docId="039687DDFFC28D59FF76B0CEFC827AD0" docLanguage="en" docName="zt02024p032.pdf" docOrigin="Zootaxa 2024" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Proctolaelaps paulista De, Lindquist & De, 2009, sp. nov." docType="treatment" docVersion="10" lastPageNumber="25" masterDocId="FFAFFFA5FFD78D41FFE1B44BFFF27D22" masterDocTitle="Edaphic ascid mites (Acari: Mesostigmata: Ascidae) from the state of São Paulo, Brazil, with description of five new species" masterLastPageNumber="32" masterPageNumber="1" pageNumber="22" updateTime="1698577253990" updateUser="plazi">
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<mods:mods id="65D2B74D4F16F7168709BEEB642141BC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3146ED79FB8E91CFD16E5A164FABE43B">
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<mods:title id="BE90ACDA7995D8D163C7C757F5C2F919">Edaphic ascid mites (Acari: Mesostigmata: Ascidae) from the state of São Paulo, Brazil, with description of five new species</mods:title>
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</mods:titleInfo>
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<mods:name id="45457AB26E3C3CAC64D37467F7BA0EAA" type="personal">
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<mods:role id="07436BAE94526AEE68CA2EF5EE40486E">
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<mods:roleTerm id="29C5BB15C6CB2E18171A2FD95E216BAE">Author</mods:roleTerm>
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<mods:namePart id="862233C8F04FEA612E7B67C11445C39C">De, Jeferson L.</mods:namePart>
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</mods:name>
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<mods:name id="E0CC51CE511B00590290379F5052CBEC" type="personal">
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<mods:roleTerm id="4524884DF5F4A8EA0555B6B9C13E8C9B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B566A99762F2B3B4B105416D34083877">Lindquist, Evert E.</mods:namePart>
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</mods:name>
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<mods:name id="6DD6223EFC534BE27BA8415BCDF3DFB9" type="personal">
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<mods:role id="6AD900509656BDD69BC42F36CADD46BC">
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<mods:roleTerm id="74E853E928CB6AAF5AA0D394377F964B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F326BA2DA191CBC18381C611E9D80CB2">De, Gilberto J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4656CA9B660C7CE0C8E6B312B3BDB9F6">text</mods:typeOfResource>
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<mods:titleInfo id="9FA5C69AC57B3E91F3B4622D0C162B6A">
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<mods:title id="7BE5B68FF1B458624754CA46759BF671">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="438EC4D076ED232EDABDDC50A9F65372">
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<mods:date id="06FD08FDF0616B5C2BC01139248E3C3A">2009</mods:date>
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<mods:detail id="E895CB3D7028C0E2AC79DBA588B2C0AF" type="volume">
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<mods:number id="F7FC8102E5ADC8CC7DFCC5424D9E9047">2024</mods:number>
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<mods:start id="A4986F2DDB261E34A9ADB0F65BA46B8B">1</mods:start>
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<mods:classification id="8A6BA077AE80306E0C28FD26945520A8">journal article</mods:classification>
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<mods:identifier id="781258A1DD08450CCFEA733EDDD9358E" type="DOI">10.5281/zenodo.186138</mods:identifier>
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<mods:identifier id="BF3D3835FF857A581743CA9F60C90964" type="GBIF-Dataset">8cf8a401-bc91-4a30-99b8-27492f4f254e</mods:identifier>
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<mods:identifier id="064879F66CD857C1473325A7499C085C" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8C84623F98B623BDE2D9E1BBF4479D7A" type="Zenodo-Dep">186138</mods:identifier>
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<treatment id="039687DDFFC28D59FF76B0CEFC827AD0" ID-DOI="http://doi.org/10.5281/zenodo.6222962" ID-GBIF-Taxon="119610879" ID-Zenodo-Dep="6222962" LSID="urn:lsid:plazi:treatment:039687DDFFC28D59FF76B0CEFC827AD0" httpUri="http://treatment.plazi.org/id/039687DDFFC28D59FF76B0CEFC827AD0" lastPageId="24" lastPageNumber="25" pageId="21" pageNumber="22">
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<subSubSection id="C3256540FFC28D54FF76B0CEFEDF79E0" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="8B8036CBFFC28D54FF76B0CEFDF579BD" blockId="21.[151,519,1157,1218]" box="[151,519,1157,1183]" pageId="21" pageNumber="22">
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<heading id="D0C881A7FFC28D54FF76B0CEFDF579BD" bold="true" box="[151,519,1157,1183]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
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<emphasis id="B94BEAD9FFC28D54FF76B0CEFDF579BD" bold="true" box="[151,519,1157,1183]" pageId="21" pageNumber="22">
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<taxonomicName id="4C3F4D48FFC28D54FF76B0CEFE5179BD" box="[151,419,1157,1183]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="paulista" status="sp. nov.">
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<emphasis id="B94BEAD9FFC28D54FF76B0CEFE5179BD" bold="true" box="[151,419,1157,1183]" italics="true" pageId="21" pageNumber="22">Proctolaelaps paulista</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27857A2FFC28D54FE4BB0CEFDF579BD" box="[426,519,1157,1183]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8036CBFFC28D54FF76B0E3FEDF79E0" blockId="21.[151,519,1157,1218]" box="[151,301,1192,1218]" pageId="21" pageNumber="22">
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(
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<figureCitation id="13042A4EFFC28D54FF7EB0E3FED779E0" box="[159,293,1192,1218]" captionStart="FIGURES 51 – 59" captionStartId="22.[151,273,1926,1950]" captionTargetBox="[152,1428,433,1925]" captionTargetId="figure@22.[151,1436,385,1981]" captionTargetPageId="22" captionText="FIGURES 51 – 59. Proctolaelaps spp.: P. paulista n. sp. Adult female, dorsal idiosoma (51), ventral idiosoma (52), spermathecal apparatus (53), tectum (54), chelicera (55), ventral gnathosoma (56) and tarsus IV (57). P. intermedius Athias – Henriot. Chelicera (58). P. wenkochingi (Samsinak). Chelicera, holotype female (59)." httpUri="https://zenodo.org/record/186146/files/figure.png" pageId="21" pageNumber="22">Figs 51–57</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3256540FFC28D57FF76B0BEFE5E7C51" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="diagnosis">
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<paragraph id="8B8036CBFFC28D54FF76B0BEFBB17A71" blockId="21.[151,1437,1269,2030]" pageId="21" pageNumber="22">
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<emphasis id="B94BEAD9FFC28D54FF76B0BEFEE9782D" bold="true" box="[151,283,1269,1295]" pageId="21" pageNumber="22">Diagnosis:</emphasis>
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Adult females of this species resemble those of
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<taxonomicName id="4C3F4D48FFC28D54FC82B0BEFA6F782D" authority="Athias-Henriot 1959" authorityName="Athias-Henriot" authorityYear="1959" box="[867,1437,1269,1295]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="intermedius">
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<emphasis id="B94BEAD9FFC28D54FC82B0BEFB6F782C" box="[867,1181,1269,1294]" italics="true" pageId="21" pageNumber="22">Proctolaelaps intermedius</emphasis>
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<bibRefCitation id="EFAE4B3AFFC28D54FB47B0BEFA6F782D" author="Athias-Henriot" box="[1190,1437,1269,1295]" pageId="21" pageNumber="30" refString="Athias-Henriot, C. (1959) Phytoseiidae et Aceosejidae (Acarina, Gamasina) d'Algerie. III. Contribution aux Aceosejinae. Bulletin de la Societe d'Historie Naturalle de L'Afrique du Nord, 50, 158 - 195." type="journal article" year="1959">Athias-Henriot 1959</bibRefCitation>
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</taxonomicName>
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and similar forms (see remarks below) in having usually one or rarely two pairs of opisthogastric setae (
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<emphasis id="B94BEAD9FFC28D54FA86B157FA667817" box="[1383,1428,1308,1333]" italics="true" pageId="21" pageNumber="22">JV2</emphasis>
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, sometimes
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<emphasis id="B94BEAD9FFC28D54FEC3B108FEA3787E" box="[290,337,1347,1372]" italics="true" pageId="21" pageNumber="22">JV1</emphasis>
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or
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<emphasis id="B94BEAD9FFC28D54FE9AB108FE59787E" box="[379,427,1347,1372]" italics="true" pageId="21" pageNumber="22">JV4</emphasis>
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) in addition to the circum-anal setae on a ventri-anal shield that is widest near its anterior margin, a triramous tectum, the anterior hypostomatic setae thickened, and the dorsal shield ornamented with few diagonal lateral striae on the anterior region. The dorsal shield in
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<taxonomicName id="4C3F4D48FFC28D54FB4BB1DBFAD5788B" box="[1194,1319,1424,1450]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="paulista">
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<emphasis id="B94BEAD9FFC28D54FB4BB1DBFB4B788B" box="[1194,1209,1424,1449]" italics="true" pageId="21" pageNumber="22">P</emphasis>
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.
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<emphasis id="B94BEAD9FFC28D54FB29B1DBFAD5788B" box="[1224,1319,1424,1449]" italics="true" pageId="21" pageNumber="22">paulista</emphasis>
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</taxonomicName>
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differs in having a pair of deep podonotal incisions between setae
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<emphasis id="B94BEAD9FFC28D54FCD2B1FCFCBE78F2" box="[819,844,1463,1488]" italics="true" pageId="21" pageNumber="22">s1</emphasis>
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and
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<emphasis id="B94BEAD9FFC28D54FC66B1FCFC5378F2" box="[903,929,1463,1488]" italics="true" pageId="21" pageNumber="22">s2</emphasis>
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that extend nearly to the bases of setae
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<emphasis id="B94BEAD9FFC28D54FA9AB1FCFA6678F2" box="[1403,1428,1463,1488]" italics="true" pageId="21" pageNumber="22">z2</emphasis>
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, the opisthonotal region fully ornamented with transverse striae, with 20 pairs of opisthonotal setae including
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<emphasis id="B94BEAD9FFC28D54FF76B24FFF167B3F" box="[151,228,1540,1566]" italics="true" pageId="21" pageNumber="22">R1-R5</emphasis>
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, setae
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<emphasis id="B94BEAD9FFC28D54FED8B24FFEAB7B3F" box="[313,345,1540,1565]" italics="true" pageId="21" pageNumber="22">Z5</emphasis>
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distinctly stouter than neighboring opisthonotal setae, the sternal, genital and ventri-anal shields fully reticulated, the palptarsal apotele 2-tined, the corniculus tricuspidate apically, and the chelicerae with a moderately multidenticulate fixed digit with 12 or 13 teeth, and a tridentate movable digit that lacks a proximal ridge of denticles. Those of
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<taxonomicName id="4C3F4D48FFC28D54FDAFB233FD047BB3" box="[590,758,1656,1682]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="intermedius">
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<emphasis id="B94BEAD9FFC28D54FDAFB233FDAF7BB3" box="[590,605,1656,1681]" italics="true" pageId="21" pageNumber="22">P</emphasis>
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.
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<emphasis id="B94BEAD9FFC28D54FD8AB233FD047BB3" box="[619,758,1656,1681]" italics="true" pageId="21" pageNumber="22">intermedius</emphasis>
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</taxonomicName>
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and similar forms lack the deep podonotal incisions, have the posterior region of the dorsal shield unornamented, with 19 pairs of opisthonotal setae including
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<emphasis id="B94BEAD9FFC28D54FAA8B2D4FA667B9A" box="[1353,1428,1694,1720]" italics="true" pageId="21" pageNumber="22">R1-R4</emphasis>
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, setae
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<emphasis id="B94BEAD9FFC28D54FF38B28EFF0A7BFC" box="[217,248,1733,1758]" italics="true" pageId="21" pageNumber="22">Z5</emphasis>
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similar in form to
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<emphasis id="B94BEAD9FFC28D54FE35B28EFE017BFC" box="[468,499,1733,1758]" italics="true" pageId="21" pageNumber="22">Z4</emphasis>
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and
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<emphasis id="B94BEAD9FFC28D54FDCDB28EFDBB7BFC" box="[556,585,1733,1758]" italics="true" pageId="21" pageNumber="22">S5</emphasis>
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, the ventral shields unornamented, the palptarsal apotele clearly 3-tined, the corniculus bicuspidate apically, and the chelicerae with a more finely multidenticulate fixed digit with 18–20 teeth, and a tridentate movable digit that is also provided more basally and paraxially with a ridge of 8 –10 fine denticles that appose similar denticles on the fixed digit (see Remarks).
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</paragraph>
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<paragraph id="8B8036CBFFC28D57FF27B32BFE5E7C51" blockId="21.[151,1437,1269,2030]" lastBlockId="22.[151,1437,152,371]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
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<emphasis id="B94BEAD9FFC28D54FF27B32BFDC67A5B" box="[198,564,1888,1913]" italics="true" pageId="21" pageNumber="22">Adult female. Dorsal idiosoma</emphasis>
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(
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<figureCitation id="13042A4EFFC28D54FDA4B32BFD6C7A58" box="[581,670,1888,1914]" captionStart="FIGURES 51 – 59" captionStartId="22.[151,273,1926,1950]" captionTargetBox="[152,1428,433,1925]" captionTargetId="figure@22.[151,1436,385,1981]" captionTargetPageId="22" captionText="FIGURES 51 – 59. Proctolaelaps spp.: P. paulista n. sp. Adult female, dorsal idiosoma (51), ventral idiosoma (52), spermathecal apparatus (53), tectum (54), chelicera (55), ventral gnathosoma (56) and tarsus IV (57). P. intermedius Athias – Henriot. Chelicera (58). P. wenkochingi (Samsinak). Chelicera, holotype female (59)." httpUri="https://zenodo.org/record/186146/files/figure.png" pageId="21" pageNumber="22">Fig. 51</figureCitation>
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): Dorsal shield 305–315 µm long, 182–190 µm wide at level of setae
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<emphasis id="B94BEAD9FFC28D54FF3BB3CCFF007A82" box="[218,242,1927,1952]" italics="true" pageId="21" pageNumber="22">j6</emphasis>
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, with few diagonal striae laterally on anterior region but fully covered with light, mostly transverse striae on posterior region; antero-lateral margins of shield with pair of deep clefts between setae
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<emphasis id="B94BEAD9FFC28D54FB1DB3E6FAE47AE4" box="[1276,1302,1965,1990]" italics="true" pageId="21" pageNumber="22">s1</emphasis>
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and
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<emphasis id="B94BEAD9FFC28D54FAAEB3E6FA9A7AE4" box="[1359,1384,1965,1990]" italics="true" pageId="21" pageNumber="22">s2</emphasis>
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that extend nearly to bases of setae
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<emphasis id="B94BEAD9FFC28D54FDE6B39FFDD37ACF" box="[519,545,2004,2029]" italics="true" pageId="21" pageNumber="22">z2</emphasis>
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. Dorsal shield with 43 pairs of setae, 23 pairs including marginals
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<emphasis id="B94BEAD9FFC28D54FAD8B39FFA857ACF" box="[1337,1399,2004,2030]" italics="true" pageId="21" pageNumber="22">r2-r6</emphasis>
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on anterior region, 20 pairs including marginals
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<emphasis id="B94BEAD9FFC18D57FD27B4D3FCE67D93" box="[710,788,152,178]" italics="true" pageId="22" pageNumber="23">R1-R5</emphasis>
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on posterior region; marginal
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<emphasis id="B94BEAD9FFC18D57FB6CB4D3FB5D7D93" box="[1165,1199,152,177]" italics="true" pageId="22" pageNumber="23">R6</emphasis>
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inserted somewhat ventrolaterally on posterior soft cuticle; submarginal
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<emphasis id="B94BEAD9FFC18D57FCF8B4F4FCB37DFA" box="[793,833,191,216]" italics="true" pageId="22" pageNumber="23">UR</emphasis>
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setae absent. Most dorsal shield setae moderately short (15–22 µm, except
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<emphasis id="B94BEAD9FFC18D57FE37B4AEFE007DDC" box="[470,498,229,254]" italics="true" pageId="22" pageNumber="23">z1</emphasis>
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,
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<emphasis id="B94BEAD9FFC18D57FDE5B4AEFDED7DDC" box="[516,543,229,254]" italics="true" pageId="22" pageNumber="23">s1</emphasis>
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,
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<emphasis id="B94BEAD9FFC18D57FDD0B4AEFDBC7DDC" box="[561,590,229,254]" italics="true" pageId="22" pageNumber="23">J5</emphasis>
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and all
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<emphasis id="B94BEAD9FFC18D57FD57B4AEFD3A7DDC" box="[694,712,229,254]" italics="true" pageId="22" pageNumber="23">R</emphasis>
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-setae slightly shorter, 11–14 µm), generally shorter than successive distances between their bases,
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<emphasis id="B94BEAD9FFC18D57FD42B547FD327C07" box="[675,704,268,293]" italics="true" pageId="22" pageNumber="23">J3</emphasis>
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and
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<emphasis id="B94BEAD9FFC18D57FD1FB547FCED7C07" box="[766,799,268,293]" italics="true" pageId="22" pageNumber="23">Z3</emphasis>
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reaching 0.6–0.7 of distance to bases of
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<emphasis id="B94BEAD9FFC18D57FAC0B547FACC7C07" box="[1313,1342,268,293]" italics="true" pageId="22" pageNumber="23">J4</emphasis>
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and
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<emphasis id="B94BEAD9FFC18D57FA9DB547FA6E7C07" box="[1404,1436,268,293]" italics="true" pageId="22" pageNumber="23">Z4</emphasis>
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respectively,
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<emphasis id="B94BEAD9FFC18D57FED6B579FE8B7C69" box="[311,377,306,331]" italics="true" pageId="22" pageNumber="23">S3 ca</emphasis>
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. 0.5 as long as interval
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<emphasis id="B94BEAD9FFC18D57FD7FB579FD167C69" box="[670,740,306,332]" italics="true" pageId="22" pageNumber="23">S3-S4</emphasis>
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; all setae smooth, slender, except
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<emphasis id="B94BEAD9FFC18D57FB66B579FB557C69" box="[1159,1191,306,331]" italics="true" pageId="22" pageNumber="23">Z5</emphasis>
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longer (34–35 µm), thicker, slightly barbed.
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</paragraph>
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</subSubSection>
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<caption id="DF406643FFC18D57FF76B3CDFB8A7AC1" httpUri="https://zenodo.org/record/186146/files/figure.png" pageId="22" pageNumber="23" targetBox="[152,1428,433,1925]" targetPageId="22">
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<paragraph id="8B8036CBFFC18D57FF76B3CDFB8A7AC1" blockId="22.[151,1437,1926,2019]" pageId="22" pageNumber="23">
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<emphasis id="B94BEAD9FFC18D57FF76B3CDFE967ABC" bold="true" box="[151,356,1926,1950]" pageId="22" pageNumber="23">FIGURES 51–59.</emphasis>
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<taxonomicName id="4C3F4D48FFC18D57FE8DB3CDFDF47ABF" box="[364,518,1926,1949]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B94BEAD9FFC18D57FE8DB3CDFDF47ABF" box="[364,518,1926,1949]" italics="true" pageId="22" pageNumber="23">Proctolaelaps</emphasis>
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</taxonomicName>
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spp.:
|
||
<taxonomicName id="4C3F4D48FFC18D57FDAAB3CDFD327ABF" box="[587,704,1926,1949]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="paulista" status="sp. nov.">
|
||
<emphasis id="B94BEAD9FFC18D57FDAAB3CDFDAB7ABF" box="[587,601,1926,1949]" italics="true" pageId="22" pageNumber="23">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFC18D57FD86B3CDFD327ABF" box="[615,704,1926,1949]" italics="true" pageId="22" pageNumber="23">paulista</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94BEAD9FFC18D57FD29B3CDFCFA7ABC" bold="true" box="[712,776,1926,1950]" pageId="22" pageNumber="23">
|
||
<taxonomicNameLabel id="A27857A2FFC18D57FD29B3CDFCFA7ABC" box="[712,776,1926,1950]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Adult female, dorsal idiosoma (51), ventral idiosoma (52), spermathecal apparatus (53), tectum (54), chelicera (55), ventral
|
||
<taxonomicName id="4C3F4D48FFC18D57FC87B3E2FC187AE2" box="[870,1002,1961,1984]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="22" pageNumber="28" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
|
||
(56) and tarsus IV (57).
|
||
<taxonomicName id="4C3F4D48FFC18D57FB1FB3E2FF2F7AC1" authority="Athias" authorityName="Athias" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="B94BEAD9FFC18D57FB1FB3E2FAFE7AE2" box="[1278,1292,1961,1984]" italics="true" pageId="22" pageNumber="23">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFC18D57FAFAB3E2FA6E7AE2" box="[1307,1436,1961,1984]" italics="true" pageId="22" pageNumber="23">intermedius</emphasis>
|
||
Athias
|
||
</taxonomicName>
|
||
–Henriot. Chelicera (58).
|
||
<taxonomicName id="4C3F4D48FFC18D57FE0EB387FCE17AC1" authority="(Samsinak)" authorityName="Samsinak" baseAuthorityName="Samsinak" box="[495,787,1996,2019]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="wenkochingi">
|
||
<emphasis id="B94BEAD9FFC18D57FE0EB387FE0E7AC1" box="[495,508,1996,2019]" italics="true" pageId="22" pageNumber="23">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFC18D57FDEBB387FD637AC1" box="[522,657,1996,2019]" italics="true" pageId="22" pageNumber="23">wenkochingi</emphasis>
|
||
(Samsinak)
|
||
</taxonomicName>
|
||
. Chelicera, holotype female (59).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3256540FFC08D59FF24B4D3FE227DDD" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
|
||
<paragraph id="8B8036CBFFC08D56FF24B4D3FF0A7909" blockId="23.[151,1437,152,2034]" pageId="23" pageNumber="24">
|
||
<emphasis id="B94BEAD9FFC08D56FF24B4D3FE7F7D93" box="[197,397,152,177]" italics="true" pageId="23" pageNumber="24">Ven t ra l idiosoma</emphasis>
|
||
(
|
||
<figureCitation id="13042A4EFFC08D56FE7CB4D3FE057D90" box="[413,503,152,178]" captionStart="FIGURES 51 – 59" captionStartId="22.[151,273,1926,1950]" captionTargetBox="[152,1428,433,1925]" captionTargetId="figure@22.[151,1436,385,1981]" captionTargetPageId="22" captionText="FIGURES 51 – 59. Proctolaelaps spp.: P. paulista n. sp. Adult female, dorsal idiosoma (51), ventral idiosoma (52), spermathecal apparatus (53), tectum (54), chelicera (55), ventral gnathosoma (56) and tarsus IV (57). P. intermedius Athias – Henriot. Chelicera (58). P. wenkochingi (Samsinak). Chelicera, holotype female (59)." httpUri="https://zenodo.org/record/186146/files/figure.png" pageId="23" pageNumber="24">Fig. 52</figureCitation>
|
||
): Tritosternum with slender base (length 10 µm) and laciniae fused along basal 15 µm of their length (50–55 µm). Pre-sternal area transversely lineate, without evident platelets. Sternal shield 80–84 µm in median length by 72–75 µm wide at narrowest width between coxae II, with 3 pairs of setae, 2 pairs of poroids, and nearly straight posterior margin; shield reticulated over entire surface, with prominent endopodal projections between coxae I and II, and fully connected with more densely sclerotized endopodal strips between coxae II and III. Third pair of sternal poroids with sternal setae
|
||
<emphasis id="B94BEAD9FFC08D56FAE5B512FADB7C50" box="[1284,1321,345,370]" italics="true" pageId="23" pageNumber="24">st4</emphasis>
|
||
on small metasternal plates. Sternal setae similar in form,
|
||
<emphasis id="B94BEAD9FFC08D56FD2FB5CBFD1D7CBB" box="[718,751,384,409]" italics="true" pageId="23" pageNumber="24">st1</emphasis>
|
||
(29–30 µm) slightly longer and
|
||
<emphasis id="B94BEAD9FFC08D56FB88B5CBFB797CBB" box="[1129,1163,384,409]" italics="true" pageId="23" pageNumber="24">st3</emphasis>
|
||
slightly shorter (24–25 µm) than
|
||
<emphasis id="B94BEAD9FFC08D56FEEAB5ECFEDF7CE2" box="[267,301,423,448]" italics="true" pageId="23" pageNumber="24">st2</emphasis>
|
||
and
|
||
<emphasis id="B94BEAD9FFC08D56FE89B5ECFE797CE2" box="[360,395,423,448]" italics="true" pageId="23" pageNumber="24">st4</emphasis>
|
||
(26–27 µm). Pair of thin endopodal strips alongside coxae III–IV connecting anteriorly with those of sternal shield projecting between coxae II–III. Genital shield reticulated over nearly entire surface, well widened behind level of insertions of genital setae such that greatest width (75–80 µm) at level of slightly convex posterior margin whose corners may touch openings of paragenital gland pores; hyaline anterior margin of shield broadly rounded, overlapping posterior margin of sternal shield. Post-genital furrow without evident platelets. Metapodal plates divided into smaller inner pair (length 5–7 µm) and larger outer pair (15–18 µm). Ventri-anal shield cup-shaped, leaving only narrow transverse strip of soft cuticle bearing setae
|
||
<emphasis id="B94BEAD9FFC08D56FF3DB6FEFEF97FEC" box="[220,267,693,718]" italics="true" pageId="23" pageNumber="24">JV1</emphasis>
|
||
and
|
||
<emphasis id="B94BEAD9FFC08D56FEA9B6FEFE887FEC" box="[328,378,693,718]" italics="true" pageId="23" pageNumber="24">ZV1</emphasis>
|
||
between it and genital shield, its greatest width (81–85 µm) at nearly straight anterior margin
|
||
<emphasis id="B94BEAD9FFC08D56FF11B697FEFE7FD7" box="[240,268,732,757]" italics="true" pageId="23" pageNumber="24">ca</emphasis>
|
||
. 0.9 as long as its length (90 µm); shield transversely striated anteriorly, reticulated posteriorly, with enlarged anal opening (length
|
||
<emphasis id="B94BEAD9FFC08D56FE18B749FDE67E39" box="[505,532,770,795]" italics="true" pageId="23" pageNumber="24">ca</emphasis>
|
||
. 30 µm); shield with one (
|
||
<emphasis id="B94BEAD9FFC08D56FCA8B749FC857E39" box="[841,887,770,795]" italics="true" pageId="23" pageNumber="24">JV2</emphasis>
|
||
) or sometimes asymmetrically 2 (
|
||
<emphasis id="B94BEAD9FFC08D56FAE4B749FAC17E39" box="[1285,1331,770,795]" italics="true" pageId="23" pageNumber="24">JV4</emphasis>
|
||
) pairs of opisthogastric setae plus 3 circum-anal setae, all simple,
|
||
<emphasis id="B94BEAD9FFC08D56FCDAB762FC9B7E60" box="[827,873,809,834]" italics="true" pageId="23" pageNumber="24">JV2</emphasis>
|
||
slightly the longest (26–27 µm); para-anal seta subequally as long as post-anal seta (21–23 µm). Eight or 9 pairs of opisthogastric setae on soft cuticle around ventri-anal shield include
|
||
<emphasis id="B94BEAD9FFC08D56FE2CB73CFE097EB2" box="[461,507,887,912]" italics="true" pageId="23" pageNumber="24">JV1</emphasis>
|
||
,
|
||
<emphasis id="B94BEAD9FFC08D56FDEBB73CFDCA7EB2" box="[522,568,887,912]" italics="true" pageId="23" pageNumber="24">JV3</emphasis>
|
||
,
|
||
<emphasis id="B94BEAD9FFC08D56FDA9B73CFD877EB2" box="[584,629,887,912]" italics="true" pageId="23" pageNumber="24">JV5</emphasis>
|
||
,
|
||
<emphasis id="B94BEAD9FFC08D56FD64B73CFD037EB2" box="[645,753,886,912]" italics="true" pageId="23" pageNumber="24">ZV1-ZV5</emphasis>
|
||
and usually
|
||
<emphasis id="B94BEAD9FFC08D56FC6BB73CFC457EB2" box="[906,951,887,912]" italics="true" pageId="23" pageNumber="24">JV4</emphasis>
|
||
, these similar in size (12–18 µm, except
|
||
<emphasis id="B94BEAD9FFC08D56FF76B7D6FF377E94" box="[151,197,925,950]" italics="true" pageId="23" pageNumber="24">JV1</emphasis>
|
||
and
|
||
<emphasis id="B94BEAD9FFC08D56FF1FB7D6FED97E94" box="[254,299,925,950]" italics="true" pageId="23" pageNumber="24">JV5</emphasis>
|
||
slightly longer, 19–21 µm) and flanked laterally by marginal pair
|
||
<emphasis id="B94BEAD9FFC08D56FBDAB7D6FBA97E94" box="[1083,1115,925,950]" italics="true" pageId="23" pageNumber="24">R6</emphasis>
|
||
(resembling a submarginal
|
||
<emphasis id="B94BEAD9FFC08D56FF76B78FFF4C7EFF" box="[151,190,964,989]" italics="true" pageId="23" pageNumber="24">UR</emphasis>
|
||
-seta); setae
|
||
<emphasis id="B94BEAD9FFC08D56FEB4B78FFE777EFF" box="[341,389,964,989]" italics="true" pageId="23" pageNumber="24">SV1</emphasis>
|
||
absent. Exopodal plate a continuous strip alongside coxae II–IV, not consolidated with peritrematal shield alongside coxae II–III, but moderately broadly connected with latter beside and behind coxa IV.
|
||
</paragraph>
|
||
<paragraph id="8B8036CBFFC08D56FF24B073FC9A795A" blockId="23.[151,1437,152,2034]" pageId="23" pageNumber="24">
|
||
<emphasis id="B94BEAD9FFC08D56FF24B073FDA27973" box="[197,592,1080,1105]" italics="true" pageId="23" pageNumber="24">Peritrematal shield and peritreme</emphasis>
|
||
(
|
||
<figureCitation id="13042A4EFFC08D56FDBEB073FD1A7970" box="[607,744,1080,1106]" captionStart="FIGURES 51 – 59" captionStartId="22.[151,273,1926,1950]" captionTargetBox="[152,1428,433,1925]" captionTargetId="figure@22.[151,1436,385,1981]" captionTargetPageId="22" captionText="FIGURES 51 – 59. Proctolaelaps spp.: P. paulista n. sp. Adult female, dorsal idiosoma (51), ventral idiosoma (52), spermathecal apparatus (53), tectum (54), chelicera (55), ventral gnathosoma (56) and tarsus IV (57). P. intermedius Athias – Henriot. Chelicera (58). P. wenkochingi (Samsinak). Chelicera, holotype female (59)." httpUri="https://zenodo.org/record/186146/files/figure.png" pageId="23" pageNumber="24">Figs 51–52</figureCitation>
|
||
): Peritrematal shields fused to dorsal shield at level slightly anterior to setae
|
||
<emphasis id="B94BEAD9FFC08D56FEB8B014FE81795A" box="[345,371,1119,1144]" italics="true" pageId="23" pageNumber="24">s1</emphasis>
|
||
; peritremes extending to bases of setae
|
||
<emphasis id="B94BEAD9FFC08D56FCA6B014FC90795A" box="[839,866,1119,1144]" italics="true" pageId="23" pageNumber="24">z1</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8036CBFFC08D56FF24B0CEFCFC79CF" blockId="23.[151,1437,152,2034]" pageId="23" pageNumber="24">
|
||
<emphasis id="B94BEAD9FFC08D56FF24B0CEFE1C79BC" box="[197,494,1157,1182]" italics="true" pageId="23" pageNumber="24">Spermathecal apparatus</emphasis>
|
||
(
|
||
<figureCitation id="13042A4EFFC08D56FE1EB0CEFDAE79BD" box="[511,604,1157,1183]" captionStart="FIGURES 51 – 59" captionStartId="22.[151,273,1926,1950]" captionTargetBox="[152,1428,433,1925]" captionTargetId="figure@22.[151,1436,385,1981]" captionTargetPageId="22" captionText="FIGURES 51 – 59. Proctolaelaps spp.: P. paulista n. sp. Adult female, dorsal idiosoma (51), ventral idiosoma (52), spermathecal apparatus (53), tectum (54), chelicera (55), ventral gnathosoma (56) and tarsus IV (57). P. intermedius Athias – Henriot. Chelicera (58). P. wenkochingi (Samsinak). Chelicera, holotype female (59)." httpUri="https://zenodo.org/record/186146/files/figure.png" pageId="23" pageNumber="24">Fig. 53</figureCitation>
|
||
): Not clearly discernible in specimens at hand; a short unsclerotized major duct leading from solenostome between coxae III and IV to a weakly sclerotized tubular portion (length
|
||
<emphasis id="B94BEAD9FFC08D56FF76B098FF4179CE" box="[151,179,1235,1260]" italics="true" pageId="23" pageNumber="24">ca</emphasis>
|
||
. 10 µm, distal width
|
||
<emphasis id="B94BEAD9FFC08D56FE4CB098FE3B79CE" box="[429,457,1235,1260]" italics="true" pageId="23" pageNumber="24">ca</emphasis>
|
||
. 8 µm) evident in
|
||
<typeStatus id="54848869FFC08D56FD43B098FCF979CF" box="[674,779,1235,1261]" pageId="23" pageNumber="24" type="paratype">paratype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8036CBFFC08D56FF24B0B2FC8D7BB4" blockId="23.[151,1437,152,2034]" pageId="23" pageNumber="24">
|
||
<taxonomicName id="4C3F4D48FFC08D56FF24B0B2FEAC7830" box="[197,350,1273,1298]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="23" pageNumber="24" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B94BEAD9FFC08D56FF24B0B2FEAC7830" box="[197,350,1273,1298]" italics="true" pageId="23" pageNumber="24">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13042A4EFFC08D56FE8EB0B2FE0C7831" box="[367,510,1273,1299]" captionStart="FIGURES 51 – 59" captionStartId="22.[151,273,1926,1950]" captionTargetBox="[152,1428,433,1925]" captionTargetId="figure@22.[151,1436,385,1981]" captionTargetPageId="22" captionText="FIGURES 51 – 59. Proctolaelaps spp.: P. paulista n. sp. Adult female, dorsal idiosoma (51), ventral idiosoma (52), spermathecal apparatus (53), tectum (54), chelicera (55), ventral gnathosoma (56) and tarsus IV (57). P. intermedius Athias – Henriot. Chelicera (58). P. wenkochingi (Samsinak). Chelicera, holotype female (59)." httpUri="https://zenodo.org/record/186146/files/figure.png" pageId="23" pageNumber="24">Figs 54–56</figureCitation>
|
||
): Anterior margin of tectum triramous, lateral tines slender, slightly serrated apically, medial tine triangular, broader basally, simply pointed apically. Fixed digit of chelicera with row of 12–13 teeth, the more distal ones slightly larger and more spaced than the proximal ones, with hyaline-lobed
|
||
<emphasis id="B94BEAD9FFC08D56FF76B126FEC178A4" box="[151,307,1389,1414]" italics="true" pageId="23" pageNumber="24">pilus dentilis</emphasis>
|
||
, and with hyaline weakly serrated rim near base on paraxial face; movable digit 20 µm long, tridentate, with basal paraxial ridge lacking teeth in apposition to proximal teeth of fixed digit, and with single prominent ventral mucro near base. Deutosternum with 6 rows of denticles, rows 1–5 connected; anterior 4 rows each with 2–5 denticles, fifth row widened with 10–15 denticles, 6th row widened with
|
||
<emphasis id="B94BEAD9FFC08D56FAC2B1AAFAB278D8" box="[1315,1344,1505,1530]" italics="true" pageId="23" pageNumber="24">ca</emphasis>
|
||
. 20–25 denticles, 7th row vestigial, indicated by concave line without denticles; corniculi slender, nearly parallel to each other, with trifid tips; internal mala fringed laterally,
|
||
<emphasis id="B94BEAD9FFC08D56FCB4B264FC837B6A" box="[853,881,1583,1608]" italics="true" pageId="23" pageNumber="24">ca</emphasis>
|
||
. as long as corniculus. Anterior hypostomatic setae very thick, longer (24–25 µm) than slender pairs
|
||
<emphasis id="B94BEAD9FFC08D56FCC7B21EFC827B4C" box="[806,880,1621,1647]" italics="true" pageId="23" pageNumber="24">h2–h3</emphasis>
|
||
which are similar in length (17–18 µm); basal palpcoxal pair much longer (25 µm). Palptarsal apotele 2-tined.
|
||
</paragraph>
|
||
<paragraph id="8B8036CBFFC08D59FF24B2E8FC387DFA" blockId="23.[151,1437,152,2034]" lastBlockId="24.[151,1438,152,2034]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
|
||
<emphasis id="B94BEAD9FFC08D56FF24B2E8FF0E7B9E" box="[197,252,1699,1724]" italics="true" pageId="23" pageNumber="24">Legs</emphasis>
|
||
(
|
||
<figureCitation id="13042A4EFFC08D56FEEDB2E9FE977B9E" box="[268,357,1698,1724]" captionStart="FIGURES 51 – 59" captionStartId="22.[151,273,1926,1950]" captionTargetBox="[152,1428,433,1925]" captionTargetId="figure@22.[151,1436,385,1981]" captionTargetPageId="22" captionText="FIGURES 51 – 59. Proctolaelaps spp.: P. paulista n. sp. Adult female, dorsal idiosoma (51), ventral idiosoma (52), spermathecal apparatus (53), tectum (54), chelicera (55), ventral gnathosoma (56) and tarsus IV (57). P. intermedius Athias – Henriot. Chelicera (58). P. wenkochingi (Samsinak). Chelicera, holotype female (59)." httpUri="https://zenodo.org/record/186146/files/figure.png" pageId="23" pageNumber="24">Fig. 57</figureCitation>
|
||
): Legs I (325–337 µm) and IV (390 µm) longer, and legs II (215–230 µm) and III (230–240 µm) clearly shorter than dorsal shield. Pretarsi of legs I–III of similar length (15–17 µm), that of IV longer (25 µm), with normally developed claws; tarsi II–IV with apical setal processes
|
||
<emphasis id="B94BEAD9FFC08D56FBF9B2BBFBBC7A2B" box="[1048,1102,1776,1802]" italics="true" pageId="23" pageNumber="24">ad -1</emphasis>
|
||
,
|
||
<emphasis id="B94BEAD9FFC08D56FBBDB2BBFB607A2B" box="[1116,1170,1776,1802]" italics="true" pageId="23" pageNumber="24">pd -1</emphasis>
|
||
inconspicuous (12–13 µm), and with ventral and subapical setae not thickened or spinelike. Coxae of all legs lineate ventrally; anterodorsal edge of coxa I with ridge of 6–8 serrations. Setation of genua of legs I-II-III-IV, respectively,
|
||
<date id="FF81100BFFC08D56FA94B376FF1A7A5C" pageId="23" pageNumber="24" value="1913-11-09" valueMax="1913-11-09" valueMin="1913-11-09">13- 11-9-9</date>
|
||
; that of tibiae,
|
||
<date id="FF81100BFFC08D56FE7CB32FFDD47A5C" box="[413,550,1892,1918]" pageId="23" pageNumber="24" value="1913-08-10" valueMax="1913-10-10" valueMin="1913-08-10">13-10-8-10</date>
|
||
; leg chaetotactic formulae holotrichous for genus and tribe Melicharini as presented by
|
||
<bibRefCitation id="EFAE4B3AFFC08D56FED4B3C1FD9D7A86" author="Lindquist" box="[309,623,1930,1956]" pageId="23" pageNumber="32" refString="Lindquist, E. E. & Evans, G. O. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata). Memoirs of the Entomological Society of Canada, 47, 1 - 64." type="journal article" year="1965">Lindquist & Evans (1965)</bibRefCitation>
|
||
; leg setae slender, smooth, of generally similar size, without evident macrosetae (but see below). Leg IV with tarsal seta
|
||
<emphasis id="B94BEAD9FFC08D56FD1AB3FAFCED7AE8" box="[763,799,1969,1994]" italics="true" pageId="23" pageNumber="24">md</emphasis>
|
||
as long as length of tibia (60 µm), and slightly longer than distance from its insertion to base of pretarsus (50 µm); alveoli of setae
|
||
<emphasis id="B94BEAD9FFC08D56FBC0B393FBA57AD3" box="[1057,1111,2008,2034]" italics="true" pageId="23" pageNumber="24">pd -2</emphasis>
|
||
and
|
||
<emphasis id="B94BEAD9FFC08D56FB70B393FB357AD3" box="[1169,1223,2008,2034]" italics="true" pageId="23" pageNumber="24">pd -3</emphasis>
|
||
on elevated bases and with diameters slightly larger (3.0–3.5 µm) than those of other setae (2.0–2.5 µm) on tarsus IV, possibly indicating bases of macrosetae (shafts missing on specimens at hand).
|
||
</paragraph>
|
||
<paragraph id="8B8036CBFFCF8D59FF24B4AEFE227DDD" blockId="24.[151,1438,152,2034]" box="[197,464,229,255]" pageId="24" pageNumber="25">
|
||
<emphasis id="B94BEAD9FFCF8D59FF24B4AEFEB77DDC" box="[197,325,229,254]" italics="true" pageId="24" pageNumber="25">Adult male</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3256540FFCF8D59FF24B540FE1C7C51" pageId="24" pageNumber="25" type="materials_examined">
|
||
<paragraph id="8B8036CBFFCF8D59FF24B540FE1C7C51" blockId="24.[151,1438,152,2034]" pageId="24" pageNumber="25">
|
||
<emphasis id="B94BEAD9FFCF8D59FF24B540FE497C07" bold="true" box="[197,443,267,293]" pageId="24" pageNumber="25">Material examined:</emphasis>
|
||
<typeStatus id="54848869FFCF8D59FE22B547FDC27C04" box="[451,560,268,294]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
|
||
female,
|
||
<date id="FF81100BFFCF8D59FD74B547FD0C7C04" box="[661,766,268,294]" pageId="24" pageNumber="25" value="1998-10-02">2.x.1998</date>
|
||
, from soil of a corn field, ESALQ-USP, Piracicaba, State of São Paulo, col. J.L. de C. Mineiro, deposited at ESALQ-USP. One
|
||
<typeStatus id="54848869FFCF8D59FC24B579FBD87C6E" box="[965,1066,306,332]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
|
||
female, same collection data as
|
||
<typeStatus id="54848869FFCF8D59FF76B512FEF37C51" box="[151,257,345,371]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
, deposited at
|
||
<collectionCode id="ED2EAE0EFFCF8D59FE41B512FE187C51" box="[416,490,345,371]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="24" pageNumber="25">CNCI</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3256540FFCF8D59FF27B534FED77A86" pageId="24" pageNumber="25" type="discussion">
|
||
<paragraph id="8B8036CBFFCF8D59FF27B534FDB97F79" blockId="24.[151,1438,152,2034]" pageId="24" pageNumber="25">
|
||
<emphasis id="B94BEAD9FFCF8D59FF27B534FEB07CBB" bold="true" box="[198,322,383,409]" pageId="24" pageNumber="25">Remarks:</emphasis>
|
||
<taxonomicName id="4C3F4D48FFCF8D59FEB0B5CBFE267CBB" box="[337,468,384,410]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="paulista">
|
||
<emphasis id="B94BEAD9FFCF8D59FEB0B5CBFE927CBB" box="[337,352,384,409]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FE97B5CBFE267CBB" box="[374,468,384,409]" italics="true" pageId="24" pageNumber="25">paulista</emphasis>
|
||
</taxonomicName>
|
||
is at least facultatively fungivorous, as indicated by the presence of fungal fragments in the gut cavity of the
|
||
<typeStatus id="54848869FFCF8D59FDDFB5EDFD5B7CE2" box="[574,681,422,448]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
specimen. The apically toothed corniculi, thickened anterior hypostomatic setae and enlarged anal opening are also indicative of fungivory. Our anticipation that setae
|
||
<emphasis id="B94BEAD9FFCF8D59FA94B586FF547F2F" italics="true" pageId="24" pageNumber="25">pd - 2</emphasis>
|
||
and
|
||
<emphasis id="B94BEAD9FFCF8D59FF01B5BFFEE47F2F" box="[224,278,500,526]" italics="true" pageId="24" pageNumber="25">
|
||
pd -
|
||
<date id="FF81100BFFCF8D59FEE6B5BFFEE47F2F" box="[263,278,500,525]" pageId="24" pageNumber="25">3</date>
|
||
</emphasis>
|
||
may be macrosetae on tarsus IV in
|
||
<taxonomicName id="4C3F4D48FFCF8D59FD20B5BFFCCD7F2F" box="[705,831,500,526]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="paulista">
|
||
<emphasis id="B94BEAD9FFCF8D59FD20B5BFFD227F2F" box="[705,720,500,525]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FD01B5BFFCCD7F2F" box="[736,831,500,525]" italics="true" pageId="24" pageNumber="25">paulista</emphasis>
|
||
</taxonomicName>
|
||
is supported by our observations of the same setae on leg IV in
|
||
<taxonomicName id="4C3F4D48FFCF8D59FECDB651FE267F11" box="[300,468,538,564]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="B94BEAD9FFCF8D59FECDB651FEC97F11" box="[300,315,538,563]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FEABB651FE267F11" box="[330,468,538,563]" italics="true" pageId="24" pageNumber="25">intermedius</emphasis>
|
||
</taxonomicName>
|
||
, in which
|
||
<emphasis id="B94BEAD9FFCF8D59FDB0B651FD757F11" box="[593,647,538,564]" italics="true" pageId="24" pageNumber="25">pd -2</emphasis>
|
||
(
|
||
<emphasis id="B94BEAD9FFCF8D59FD79B651FD467F11" box="[664,692,538,563]" italics="true" pageId="24" pageNumber="25">ca</emphasis>
|
||
. 40 µm) and
|
||
<emphasis id="B94BEAD9FFCF8D59FCB2B651FC7B7F11" box="[851,905,538,564]" italics="true" pageId="24" pageNumber="25">pd -3</emphasis>
|
||
(
|
||
<emphasis id="B94BEAD9FFCF8D59FC7BB651FC477F11" box="[922,949,538,563]" italics="true" pageId="24" pageNumber="25">ca</emphasis>
|
||
. 35 µm) are shorter than the tibia (length
|
||
<emphasis id="B94BEAD9FFCF8D59FF76B60AFF417F78" box="[151,179,577,602]" italics="true" pageId="24" pageNumber="25">ca</emphasis>
|
||
. 50 µm) and lack enlarged alveoli.
|
||
</paragraph>
|
||
<paragraph id="8B8036CBFFCF8D59FF24B623FD55795A" blockId="24.[151,1438,152,2034]" pageId="24" pageNumber="25">
|
||
One of us (EEL) has examined the
|
||
<typeStatus id="54848869FFCF8D59FD89B623FD697FA0" box="[616,667,616,642]" pageId="24" pageNumber="25">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C3F4D48FFCF8D59FCCDB623FC277FA3" box="[812,981,616,642]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="B94BEAD9FFCF8D59FCCDB623FCC97FA3" box="[812,827,616,641]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FCABB623FC277FA3" box="[842,981,616,641]" italics="true" pageId="24" pageNumber="25">intermedius</emphasis>
|
||
</taxonomicName>
|
||
as well as that of two other species of
|
||
<taxonomicName id="4C3F4D48FFCF8D59FF76B6C4FE387F8A" box="[151,458,655,680]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="originally">
|
||
<emphasis id="B94BEAD9FFCF8D59FF76B6C4FE387F8A" box="[151,458,655,680]" italics="true" pageId="24" pageNumber="25">Proctolaelaps originally</emphasis>
|
||
</taxonomicName>
|
||
named
|
||
<taxonomicName id="4C3F4D48FFCF8D59FDCEB6C4FBE27F8A" authority="Chant, 1963" authorityName="Chant" authorityYear="1963" box="[559,1040,654,680]" class="Arachnida" family="Ascidae" genus="Neojordensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="B94BEAD9FFCF8D59FDCEB6C4FCAA7F8A" box="[559,856,655,680]" italics="true" pageId="24" pageNumber="25">Neojordensia orientalis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAE4B3AFFCF8D59FC8BB6C5FBF57F8A" author="Chant" box="[874,1031,654,680]" pageId="24" pageNumber="30" refString="Chant, D. A. (1963) The subfamily Blattisocinae Garman (= Aceosejinae Evans) (Acarina: Blattisocidae Garman) (= Aceosejidae Baker and Wharton) in North America, with descriptions of new species. Canadian Journal of Zoology, 41, 243 - 305." type="journal article" year="1963">Chant, 1963</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3F4D48FFCF8D59FBAEB6C4FEAE7FED" authority="Samsinak, 1964" authorityName="Samsinak" authorityYear="1964" class="Arachnida" family="Ascidae" genus="Neojordensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="wenkochingi">
|
||
<emphasis id="B94BEAD9FFCF8D59FBAEB6C4FA6F7F8A" box="[1103,1437,655,680]" italics="true" pageId="24" pageNumber="25">Neojordensia wenkochingi</emphasis>
|
||
<bibRefCitation id="EFAE4B3AFFCF8D59FF76B6FEFEAE7FED" author="Samsinak" box="[151,348,693,719]" pageId="24" pageNumber="32" refString="Samsinak, K. (1964) Termitophile Milben aus der VR China. 1. Mesostigmata. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden, 32, 33 - 52." type="journal article" year="1964">Samšinák, 1964</bibRefCitation>
|
||
</taxonomicName>
|
||
. Adult females of these species are alike in all descriptive aspects, including the peculiar dentition of the movable cheliceral digit, which was not noted in the original descriptions for these forms (
|
||
<figureCitation id="13042A4EFFCF8D59FF7EB749FED57E3E" box="[159,295,770,796]" captionStart="FIGURES 51 – 59" captionStartId="22.[151,273,1926,1950]" captionTargetBox="[152,1428,433,1925]" captionTargetId="figure@22.[151,1436,385,1981]" captionTargetPageId="22" captionText="FIGURES 51 – 59. Proctolaelaps spp.: P. paulista n. sp. Adult female, dorsal idiosoma (51), ventral idiosoma (52), spermathecal apparatus (53), tectum (54), chelicera (55), ventral gnathosoma (56) and tarsus IV (57). P. intermedius Athias – Henriot. Chelicera (58). P. wenkochingi (Samsinak). Chelicera, holotype female (59)." httpUri="https://zenodo.org/record/186146/files/figure.png" pageId="24" pageNumber="25">Figs 58, 59</figureCitation>
|
||
) (see below). The form of the ventri-anal shield among these species varies considerably among females of the same population and in any case does not differ visually between the named species as much as shown by
|
||
<bibRefCitation id="EFAE4B3AFFCF8D59FEF2B71BFDFB7E48" author="Karg" box="[275,521,848,874]" pageId="24" pageNumber="31" refString="Karg, W. (1985) Die Raubmilbengattung Proctolaelaps Berlese, 1923. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 112, 185 - 206." type="journal article" year="1985">Karg (1985, fig. 10)</bibRefCitation>
|
||
. Among the
|
||
<typeStatus id="54848869FFCF8D59FD44B71BFD2B7E48" box="[677,729,848,874]" pageId="24" pageNumber="25">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="4C3F4D48FFCF8D59FC66B71BFBC67E4B" box="[903,1076,848,874]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="B94BEAD9FFCF8D59FC66B71BFC647E4B" box="[903,918,848,873]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FC47B71BFBC67E4B" box="[934,1076,848,873]" italics="true" pageId="24" pageNumber="25">intermedius</emphasis>
|
||
</taxonomicName>
|
||
as well as the
|
||
<typeStatus id="54848869FFCF8D59FB06B71BFAA97E48" box="[1255,1371,848,874]" pageId="24" pageNumber="25" type="holotype">holotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C3F4D48FFCF8D59FA64B71BFEF97EB2" class="Arachnida" family="Ascidae" genus="Neojordensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="B94BEAD9FFCF8D59FA64B71BFA667E4B" box="[1413,1428,848,873]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FF76B73CFEF97EB2" box="[151,267,887,912]" italics="true" pageId="24" pageNumber="25">orientalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3F4D48FFCF8D59FEA7B73CFE0F7EB2" box="[326,509,886,912]" class="Arachnida" family="Ascidae" genus="Neojordensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="wenkochingi">
|
||
<emphasis id="B94BEAD9FFCF8D59FEA7B73CFEA77EB2" box="[326,341,887,912]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FE84B73CFE0F7EB2" box="[357,509,887,912]" italics="true" pageId="24" pageNumber="25">wenkochingi</emphasis>
|
||
</taxonomicName>
|
||
, the ventri-anal shield may carry one pair, or asymmetrically a third seta, or symmetrically two pairs of “pre-anal” setae, due to considerable variation in the position of seta
|
||
<emphasis id="B94BEAD9FFCF8D59FAE1B7D6FADF7E94" box="[1280,1325,925,950]" italics="true" pageId="24" pageNumber="25">JV1</emphasis>
|
||
either on the anterior margin, or touching that margin, or clearly within the margin of the shield. As a result, these species are here placed in synonymy, with
|
||
<taxonomicName id="4C3F4D48FFCF8D59FD44B7A1FCC87921" box="[677,826,1002,1028]" class="Arachnida" family="Ascidae" genus="Neojordensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="B94BEAD9FFCF8D59FD44B7A1FD467921" box="[677,692,1002,1027]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FD24B7A1FCC87921" box="[709,826,1002,1027]" italics="true" pageId="24" pageNumber="25">orientalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A27857A2FFCF8D59FCAAB7A1FC067926" box="[843,1012,1002,1028]" pageId="24" pageNumber="25">new synonym</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C3F4D48FFCF8D59FBD0B7A1FB197921" box="[1073,1259,1002,1028]" class="Arachnida" family="Ascidae" genus="Neojordensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="wenkochingi">
|
||
<emphasis id="B94BEAD9FFCF8D59FBD0B7A1FBB27921" box="[1073,1088,1002,1027]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FBB0B7A1FB197921" box="[1105,1259,1002,1027]" italics="true" pageId="24" pageNumber="25">wenkochingi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27857A2FFCF8D59FB13B7A1FA6E7926" box="[1266,1436,1002,1028]" pageId="24" pageNumber="25">new synonym</taxonomicNameLabel>
|
||
treated as junior subjective synonyms of
|
||
<taxonomicName id="4C3F4D48FFCF8D59FD94B05AFCEE7908" box="[629,796,1041,1067]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="B94BEAD9FFCF8D59FD94B05AFD767908" box="[629,644,1041,1066]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FD73B05AFCEE7908" box="[658,796,1041,1066]" italics="true" pageId="24" pageNumber="25">intermedius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C3F4D48FFCF8D59FCCBB05AFB197909" authority="Karg, 1971" authorityName="Karg" authorityYear="1971" box="[810,1259,1041,1067]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="ventrianalis">
|
||
<emphasis id="B94BEAD9FFCF8D59FCCBB05AFB937908" box="[810,1121,1041,1066]" italics="true" pageId="24" pageNumber="25">Proctolaelaps ventrianalis</emphasis>
|
||
<bibRefCitation id="EFAE4B3AFFCF8D59FB89B05AFB197909" author="Karg" box="[1128,1259,1041,1067]" pageId="24" pageNumber="31" refString="Karg, W. (1971) Acari (Acarina), Milben. Unterordnung Anactinochaeta (Parasitiformes). Die freilebenden Gamasina (Gamasides), Raubmilben. Die Tierwelt Deutschlands, 1 - 475." type="book chapter" year="1971">Karg, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
is also thought to be conspecific and a junior synonym of
|
||
<taxonomicName id="4C3F4D48FFCF8D59FD55B073FC9A7973" box="[692,872,1080,1106]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="B94BEAD9FFCF8D59FD55B073FD317973" box="[692,707,1080,1105]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FD35B073FC9A7973" box="[724,872,1080,1105]" italics="true" pageId="24" pageNumber="25">intermedius</emphasis>
|
||
</taxonomicName>
|
||
, but we have not studied its
|
||
<typeStatus id="54848869FFCF8D59FB37B073FAFE7970" box="[1238,1292,1080,1106]" pageId="24" pageNumber="25">type</typeStatus>
|
||
material to determine the attributes of the movable digit.
|
||
</paragraph>
|
||
<paragraph id="8B8036CBFFCF8D59FF24B0CEFCD37831" blockId="24.[151,1438,152,2034]" pageId="24" pageNumber="25">
|
||
The peculiar nature of the dentition of the movable digit in
|
||
<taxonomicName id="4C3F4D48FFCF8D59FC99B0CEFBED79BC" box="[888,1055,1157,1183]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="B94BEAD9FFCF8D59FC99B0CEFC7579BC" box="[888,903,1157,1182]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FC74B0CEFBED79BC" box="[917,1055,1157,1182]" italics="true" pageId="24" pageNumber="25">intermedius</emphasis>
|
||
</taxonomicName>
|
||
and related forms was not given attention in the original or subsequent descriptions of those species. The secondarily dentate ridge on the basal paraxial surface of the movable digit in apposition to the similarly fine dentition proximally on the fixed digit (
|
||
<figureCitation id="13042A4EFFCF8D59FF7EB0B2FED77831" box="[159,293,1273,1299]" captionStart="FIGURES 51 – 59" captionStartId="22.[151,273,1926,1950]" captionTargetBox="[152,1428,433,1925]" captionTargetId="figure@22.[151,1436,385,1981]" captionTargetPageId="22" captionText="FIGURES 51 – 59. Proctolaelaps spp.: P. paulista n. sp. Adult female, dorsal idiosoma (51), ventral idiosoma (52), spermathecal apparatus (53), tectum (54), chelicera (55), ventral gnathosoma (56) and tarsus IV (57). P. intermedius Athias – Henriot. Chelicera (58). P. wenkochingi (Samsinak). Chelicera, holotype female (59)." httpUri="https://zenodo.org/record/186146/files/figure.png" pageId="24" pageNumber="25">Figs 58–59</figureCitation>
|
||
) may be a further adaptation for fungivory.
|
||
</paragraph>
|
||
<paragraph id="8B8036CBFFCF8D59FF24B16BFC587B4D" blockId="24.[151,1438,152,2034]" pageId="24" pageNumber="25">
|
||
Some additional females identified as
|
||
<taxonomicName id="4C3F4D48FFCF8D59FD75B16BFCB2781B" box="[660,832,1312,1338]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="B94BEAD9FFCF8D59FD75B16BFD51781B" box="[660,675,1312,1337]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FD52B16BFCB2781B" box="[691,832,1312,1337]" italics="true" pageId="24" pageNumber="25">intermedius</emphasis>
|
||
</taxonomicName>
|
||
by Athias-Henriot apparently subsequently to her original description of that species, and deposited as reference specimens with the
|
||
<typeStatus id="54848869FFCF8D59FB7EB10DFB267842" box="[1183,1236,1350,1376]" pageId="24" pageNumber="25">type</typeStatus>
|
||
material of that species, are clearly of a different, and apparently undescribed, species that is more similar to the one newly described here. Those on slides labeled W383 and V331 have the dorsal, sternal, genital and ventri-anal shields fully reticulated; the dorsal shield setae are collectively long, much longer than the intervals between their bases; the ventri-anal shield is widest at the anus, tapered anteriorly, and bears only pre-anal seta
|
||
<emphasis id="B94BEAD9FFCF8D59FAABB1AAFA8478D8" box="[1354,1398,1505,1530]" italics="true" pageId="24" pageNumber="25">JV2</emphasis>
|
||
on its antero-lateral corners; the corniculus is apically tricuspidate; the palpal apotele 2-tined; the fixed cheliceral dentition begins with 2 large coarse teeth basally; and leg IV tarsal setae
|
||
<emphasis id="B94BEAD9FFCF8D59FC19B264FBDC7B6A" box="[1016,1070,1582,1608]" italics="true" pageId="24" pageNumber="25">pd -2</emphasis>
|
||
(58 µm) and
|
||
<emphasis id="B94BEAD9FFCF8D59FB30B264FAFA7B6A" box="[1233,1288,1582,1608]" italics="true" pageId="24" pageNumber="25">pd -3</emphasis>
|
||
(44 µm) are not conspicuous as macrosetae relative to the tibial length (55 µm).
|
||
</paragraph>
|
||
<paragraph id="8B8036CBFFCF8D59FF24B237FED77A86" blockId="24.[151,1438,152,2034]" pageId="24" pageNumber="25">
|
||
<bibRefCitation id="EFAE4B3AFFCF8D59FF24B237FEA97BB4" author="Karg" box="[197,347,1660,1686]" pageId="24" pageNumber="31" refString="Karg, W. (1979) Zur Kenntnis der Raubmilbengattungen Cheiroseius Berlese, 1916, Asca v. Heyden, 1826 und Halolaelaps Berlese et Trouessart, 1889 (Acarina, Parasitiformes). Mitteilungen aus dem Zoologischen Museum in Berlin, 55, 251 - 267." type="journal article" year="1979">Karg (1979</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAE4B3AFFCF8D59FE88B237FE5E7BB4" author="Karg" box="[361,428,1660,1686]" pageId="24" pageNumber="31" refString="Karg, W. (1985) Die Raubmilbengattung Proctolaelaps Berlese, 1923. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 112, 185 - 206." type="journal article" year="1985">1985</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAE4B3AFFCF8D59FE5AB237FE0E7BB4" author="Karg" box="[443,508,1660,1686]" pageId="24" pageNumber="31" refString="Karg, W. (1988) Zur Kenntnis der Gattung Proctolaelaps Berlese, 1923 (Acarina, Parasitiformes). Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 115, 441 - 454." type="journal article" year="1988">1988</bibRefCitation>
|
||
) recognized 3 subgenera of
|
||
<taxonomicName id="4C3F4D48FFCF8D59FC8CB237FBD27BB7" box="[877,1056,1660,1685]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94BEAD9FFCF8D59FC8CB237FBD27BB7" box="[877,1056,1660,1685]" italics="true" pageId="24" pageNumber="25">Proctolaelaps</emphasis>
|
||
</taxonomicName>
|
||
, of which
|
||
<taxonomicName id="4C3F4D48FFCF8D59FB4CB237FE947B9F" authority="Bregetova (1977)" authorityName="Bregetova" authorityYear="1977" class="Arachnida" family="Ascidae" genus="Paraproctolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94BEAD9FFCF8D59FB4CB237FA6E7BB7" box="[1197,1436,1660,1685]" italics="true" pageId="24" pageNumber="25">Paraproctolaelaps</emphasis>
|
||
<bibRefCitation id="EFAE4B3AFFCF8D59FF76B2E8FE947B9F" author="Bregetova" box="[151,358,1699,1725]" pageId="24" pageNumber="30" refString="Bregetova, N. G. (1977) Family Aceosejidae Baker and Wharton, 1952 (sensu Evans, 1958). In ' Key to the Soilinhabiting Mites. Mesostigmata' (Ed. M. S. Ghilyarov & N. G. Bregetova) pp. 169 - 226. Nauka: Leningrad. In Russian." type="journal article" year="1977">Bregetova (1977)</bibRefCitation>
|
||
</taxonomicName>
|
||
was defined by the female having a ventri-anal shield, and
|
||
<taxonomicName id="4C3F4D48FFCF8D59FBCBB2E8FAA17B9F" authority="Karg (1979)" authorityName="Karg" authorityYear="1979" box="[1066,1363,1699,1725]" class="Arachnida" family="Ascidae" genus="Proctofissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94BEAD9FFCF8D59FBCBB2E8FB487B9E" box="[1066,1210,1699,1724]" italics="true" pageId="24" pageNumber="25">Proctofissus</emphasis>
|
||
<bibRefCitation id="EFAE4B3AFFCF8D59FB23B2E8FAA17B9F" author="Karg" box="[1218,1363,1699,1725]" pageId="24" pageNumber="31" refString="Karg, W. (1979) Zur Kenntnis der Raubmilbengattungen Cheiroseius Berlese, 1916, Asca v. Heyden, 1826 und Halolaelaps Berlese et Trouessart, 1889 (Acarina, Parasitiformes). Mitteilungen aus dem Zoologischen Museum in Berlin, 55, 251 - 267." type="journal article" year="1979">Karg (1979)</bibRefCitation>
|
||
</taxonomicName>
|
||
by its adults having the corniculus divided, or multicuspidate, distally. The phylogenetic value of these distinctions is doubtful and they are not followed here, in part because the nominate subgenus is not defined apomorphically, but also because some species, including
|
||
<taxonomicName id="4C3F4D48FFCF8D59FCB4B35CFBF37A12" box="[853,1025,1814,1840]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="B94BEAD9FFCF8D59FCB4B35CFC967A12" box="[853,868,1815,1840]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FC95B35CFBF37A12" box="[884,1025,1815,1840]" italics="true" pageId="24" pageNumber="25">intermedius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3F4D48FFCF8D59FBF0B35CFB357A12" box="[1041,1223,1814,1840]" class="Arachnida" family="Ascidae" genus="Neojordensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="wenkochingi">
|
||
<emphasis id="B94BEAD9FFCF8D59FBF0B35CFBD27A12" box="[1041,1056,1815,1840]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FBD1B35CFB357A12" box="[1072,1223,1815,1840]" italics="true" pageId="24" pageNumber="25">wenkochingi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3F4D48FFCF8D59FAE2B35CFA677A12" box="[1283,1429,1814,1840]" class="Arachnida" family="Ascidae" genus="Neojordensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="B94BEAD9FFCF8D59FAE2B35CFAE07A12" box="[1283,1298,1815,1840]" italics="true" pageId="24" pageNumber="25">P</emphasis>
|
||
.
|
||
<emphasis id="B94BEAD9FFCF8D59FAC3B35CFA677A12" box="[1314,1429,1815,1840]" italics="true" pageId="24" pageNumber="25">orientalis</emphasis>
|
||
</taxonomicName>
|
||
, have the apomorphic attributes of both of the other subgroups. Also,
|
||
<taxonomicName id="4C3F4D48FFCF8D59FC00B376FB877A74" box="[993,1141,1853,1878]" class="Arachnida" family="Ascidae" genus="Proctofissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94BEAD9FFCF8D59FC00B376FB877A74" box="[993,1141,1853,1878]" italics="true" pageId="24" pageNumber="25">Proctofissus</emphasis>
|
||
</taxonomicName>
|
||
is an unavailable name, according to the International Code of Zoological Nomenclature, as no
|
||
<typeStatus id="54848869FFCF8D59FC3CB32FFBE27A5C" box="[989,1040,1892,1918]" pageId="24" pageNumber="25">type</typeStatus>
|
||
species was designated (
|
||
<bibRefCitation id="EFAE4B3AFFCF8D59FAD4B32FFEEA7A86" author="Halliday" pageId="24" pageNumber="31" refString="Halliday, R. B., Walter, D. E. & Lindquist, E. E. (1998) Revision of the Australian Ascidae (Acarina: Mesostigmata). Invertebrate Taxonomy, 12, 1 - 54." type="journal article" year="1998">
|
||
Halliday
|
||
<emphasis id="B94BEAD9FFCF8D59FF76B3C0FF387A86" box="[151,202,1931,1956]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
|
||
. 1998
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3256540FFCF8D59FF27B3FBFC827AD0" pageId="24" pageNumber="25" type="etymology">
|
||
<paragraph id="8B8036CBFFCF8D59FF27B3FBFC827AD0" blockId="24.[151,1438,152,2034]" pageId="24" pageNumber="25">
|
||
<emphasis id="B94BEAD9FFCF8D59FF27B3FBFEA57AE8" bold="true" box="[198,343,1968,1994]" pageId="24" pageNumber="25">Etymology:</emphasis>
|
||
The name
|
||
<taxonomicName id="4C3F4D48FFCF8D59FE03B3FAFDB27AE8" box="[482,576,1969,1994]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="paulista">
|
||
<emphasis id="B94BEAD9FFCF8D59FE03B3FAFDB27AE8" box="[482,576,1969,1994]" italics="true" pageId="24" pageNumber="25">paulista</emphasis>
|
||
</taxonomicName>
|
||
corresponds to “from the State of São Paulo,
|
||
<collectingCountry id="F328765BFFCF8D59FB90B3FAFB4C7AE9" box="[1137,1214,1969,1995]" name="Brazil" pageId="24" pageNumber="25">Brazil</collectingCountry>
|
||
”, and refers to the place where the
|
||
<typeStatus id="54848869FFCF8D59FEB7B393FE7A7AD0" box="[342,392,2008,2034]" pageId="24" pageNumber="25">type</typeStatus>
|
||
specimens of this species were collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |