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<document id="93E12B75F9E1FF7D716EA4766025EA34" ID-DOI="10.11646/zootaxa.5496.4.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="13617489" ID-ZooBank="EBC7D78A-A0AD-478A-BB59-2B81DB812A58" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725024750009" checkinUser="plazi" docAuthor="Ng, Heok Hee &amp; Tan, Heok Hui" docDate="2024" docId="039687C4B17DB40CDFF43593DF72FD81" docLanguage="en" docName="zootaxa.5496.4.4.pdf" docOrigin="Zootaxa 5496 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5496.4.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudomystus tuberosus Ng &amp; Tan, 2024, new species" docType="treatment" docUuid="55E60B17-6E2F-418F-A458-09CB35BB0089" docUuidSource="ZooBank" docVersion="2" lastPageNumber="549" masterDocId="FFAFFFBCB17CB40FDF633504DE3DFF98" masterDocTitle="Pseudomystus tuberosus, a new species of catfish from central Borneo (Actinopterygii: Bagridae)" masterLastPageNumber="554" masterPageNumber="546" pageNumber="547" updateTime="1725025148509" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="01D83BFD4214E33F04B783C5631380B4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AC4AE55E2FD32E84B11B6AC5AB59CAAF">
<mods:title id="5C2EBB0C337DD427DE1537A31536A38D">Pseudomystus tuberosus, a new species of catfish from central Borneo (Actinopterygii: Bagridae)</mods:title>
</mods:titleInfo>
<mods:name id="67D92C232BC617DB4344CCFCB7F5FDF1" type="personal">
<mods:role id="2E16F0468BDB12B517E252A886AC78E2">
<mods:roleTerm id="F0CA1FD600F88F2CD82BD5F42ECE4E43">Author</mods:roleTerm>
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<mods:namePart id="D20B647800A259A360D63CF4AFE12F6B">Ng, Heok Hee</mods:namePart>
<mods:affiliation id="62E8864DCF23EBDF50EA8BD071FDC1BC">c / o Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377.</mods:affiliation>
</mods:name>
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<mods:namePart id="1085E4C15DDF9FC0352B756859B8ADDA">Tan, Heok Hui</mods:namePart>
<mods:affiliation id="F222C4AE671F89637BF4EB2AA425690C">Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377.</mods:affiliation>
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<mods:title id="E4A1470F9AD7F0DF192AFDCDE092A048">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="64B482E686BC1384872AE4587904ACC0">
<mods:date id="DA281CB95E3225A1DAA99F916703F207">2024</mods:date>
<mods:detail id="54AC98878A1A1180127B3C6B8B9BDD6D" type="pubDate">
<mods:number id="F4093C89AF0780D773557DD3E9582F00">2024-08-21</mods:number>
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<mods:number id="56D38ED6B8CA2030ACE9133D69B83B2B">5496</mods:number>
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<mods:number id="C36BB6A0C133010737CDA68C6337CE65">4</mods:number>
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<mods:location id="F3671709D871625918DBD457F8C6EE85">
<mods:url id="5E254D646D196354563CFFBBCA804512">http://dx.doi.org/10.11646/zootaxa.5496.4.4</mods:url>
</mods:location>
<mods:classification id="077A047F3D1751E4379D9E2C59A8F44E">journal article</mods:classification>
<mods:identifier id="E46762FA2CE5C16EC855ED5E12744A6A" type="DOI">10.11646/zootaxa.5496.4.4</mods:identifier>
<mods:identifier id="32D6176EB269D4BBF78DFF0EB5910FCE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="84600E9E06C7B14D8AEEE424AB7371B6" type="Zenodo-Dep">13617489</mods:identifier>
<mods:identifier id="1B977E939DA1319E9EE298FE8105F2C2" type="ZooBank">EBC7D78A-A0AD-478A-BB59-2B81DB812A58</mods:identifier>
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<treatment id="039687C4B17DB40CDFF43593DF72FD81" LSID="urn:lsid:zoobank.org:act:55E60B17-6E2F-418F-A458-09CB35BB0089" httpUri="http://treatment.plazi.org/id/039687C4B17DB40CDFF43593DF72FD81" lastPageId="3" lastPageNumber="549" pageId="1" pageNumber="547">
<subSubSection id="C3256559B17DB40EDFF43593DC63FF29" box="[151,606,151,178]" pageId="1" pageNumber="547" type="nomenclature">
<paragraph id="8B8036D2B17DB40EDFF43593DC63FF29" blockId="1.[151,606,151,213]" box="[151,606,151,178]" pageId="1" pageNumber="547">
<heading id="D0C881BEB17DB40EDFF43593DC63FF29" bold="true" box="[151,606,151,178]" fontSize="11" level="1" pageId="1" pageNumber="547" reason="1">
<emphasis id="B94BEAC0B17DB40EDFF43593DC63FF29" bold="true" box="[151,606,151,178]" pageId="1" pageNumber="547">
<taxonomicName id="4C3F4D51B17DB40EDFF43593DF82FF2A" authorityName="Ng &amp; Tan" authorityYear="2024" box="[151,447,151,178]" family="Bagridae" genus="Pseudomystus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="547" phylum="Chordata" rank="species" species="tuberosus" status="sp. nov.">
<emphasis id="B94BEAC0B17DB40EDFF43593DF82FF2A" bold="true" box="[151,447,151,178]" italics="true" pageId="1" pageNumber="547">Pseudomystus tuberosus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27857BBB17DB40EDEAD3593DC63FF29" box="[462,606,151,177]" pageId="1" pageNumber="547" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3256559B17DB40EDFF435BFDEE4FF4D" box="[151,217,187,213]" pageId="1" pageNumber="547" type="description">
<paragraph id="8B8036D2B17DB40EDFF435BFDEE4FF4D" blockId="1.[151,606,151,213]" box="[151,217,187,213]" pageId="1" pageNumber="547">
<figureCitation id="13042A57B17DB40EDFF435BFDEE4FF4D" box="[151,217,187,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[151,1436,561,1860]" captionTargetId="figure-197@3.[151,1436,561,1860]" captionTargetPageId="3" captionText="FIGURE 1. Pseudomystus tuberosus, holotype, MZB 17244, 93.0 mm SL; Borneo: Kalimantan Tengah, Kahayan River drainage." figureDoi="http://doi.org/10.5281/zenodo.13617493" httpUri="https://zenodo.org/record/13617493/files/figure.png" pageId="1" pageNumber="547">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3256559B17DB40EDFF43407DDF9FEFD" pageId="1" pageNumber="547" type="materials_examined">
<paragraph id="8B8036D2B17DB40EDFF43407DD67FED9" blockId="1.[151,1437,259,1042]" pageId="1" pageNumber="547">
<materialsCitation id="3B573C8FB17DB40EDFF43407DD67FED9" collectingDate="2014-07-10" collectionCode="MZB" collectorName="H. H. Tan" location="Rungan River" municipality="Kahayan River" pageId="1" pageNumber="547" specimenCode="MZB 17244" specimenCount="1" stateProvince="Kalimantan Tengah" typeStatus="holotype">
<emphasis id="B94BEAC0B17DB40EDFF43407DF36FE85" bold="true" box="[151,267,259,285]" pageId="1" pageNumber="547">
<typeStatus id="54848870B17DB40EDFF43407DF3BFE85" box="[151,262,259,285]" pageId="1" pageNumber="547" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DB999EA9B17DB40EDE703400DF9CFE85" box="[275,417,259,286]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="547" type="Museum">MZB 17244</specimenCode>
, male, 93.0 mm SL; Borneo:
<collectingRegion id="49FBF830B17DB40EDD913407DDECFE85" box="[754,977,259,285]" country="Indonesia" name="Kalimantan Tengah" pageId="1" pageNumber="547">Kalimantan Tengah</collectingRegion>
,
<collectingMunicipality id="6BE4ACA8B17DB40EDCBE3407DABAFE85" box="[989,1159,259,285]" pageId="1" pageNumber="547">Kahayan River</collectingMunicipality>
drainage,
<location id="8EE06009B17DB40EDB9C3400DBA1FE85" LSID="urn:lsid:plazi:treatment:039687C4B17DB40CDFF43593DF72FD81:8EE06009B17DB40EDB9C3400DBA1FE85" box="[1279,1436,259,286]" municipality="Kahayan River" name="Rungan River" pageId="1" pageNumber="547" stateProvince="Kalimantan Tengah">Rungan River</location>
sub-drainage, stream near Tangkiling;
<collectorName id="26CA5304B17DB40EDD25342CDC8BFEDA" box="[582,694,296,322]" pageId="1" pageNumber="547">H. H. Tan</collectorName>
,
<date id="FF811012B17DB40EDDA23423DD6BFED9" box="[705,854,295,321]" pageId="1" pageNumber="547" value="2014-07-10">
<collectingDate id="EFC5E9FAB17DB40EDDA23423DD6BFED9" box="[705,854,295,321]" pageId="1" pageNumber="547" value="2014-07-10">10 July 2014</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8036D2B17DB40EDFA4344FDDF9FEFD" blockId="1.[151,1437,259,1042]" box="[199,964,331,358]" pageId="1" pageNumber="547">
<materialsCitation id="3B573C8FB17DB40EDFA4344FDDFCFEFD" box="[199,961,331,358]" collectingDate="2014-07-10" collectionCode="ZRC" collectorName="H. H. Tan" location="Rungan River" municipality="Kahayan River" pageId="1" pageNumber="547" specimenCode="ZRC 54541" specimenCount="1" stateProvince="Kalimantan Tengah" typeStatus="paratype">
<emphasis id="B94BEAC0B17DB40EDFA4344FDF7BFEFD" bold="true" box="[199,326,331,357]" pageId="1" pageNumber="547">
<typeStatus id="54848870B17DB40EDFA4344FDF7CFEFD" box="[199,321,331,357]" pageId="1" pageNumber="547" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="DB999EA9B17DB40EDE2E344FDFEDFEFE" box="[333,464,331,358]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="547">ZRC 54541</specimenCode>
(2),
<quantity id="4CC79B37B17DB40EDD65344FDC94FEFE" box="[518,681,331,358]" metricMagnitude="-2" metricUnit="m" metricValue="8.39" metricValueMax="8.809999999999999" metricValueMin="7.97" pageId="1" pageNumber="547" unit="mm" value="83.9" valueMax="88.1" valueMin="79.7">79.7–88.1 mm</quantity>
SL; data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256559B17DB40EDFA4346BDF69FD69" pageId="1" pageNumber="547" type="diagnosis">
<paragraph id="8B8036D2B17DB40EDFA4346BDF69FD69" blockId="1.[151,1437,259,1042]" pageId="1" pageNumber="547">
<emphasis id="B94BEAC0B17DB40EDFA4346BDF7FFE11" bold="true" box="[199,322,367,393]" pageId="1" pageNumber="547">Diagnosis.</emphasis>
<taxonomicName id="4C3F4D51B17DB40EDE2A346BDC61FE11" authorityName="Ng &amp; Tan" authorityYear="2024" box="[329,604,367,393]" family="Bagridae" genus="Pseudomystus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="547" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B94BEAC0B17DB40EDE2A346BDC61FE11" box="[329,604,367,393]" italics="true" pageId="1" pageNumber="547">Pseudomystus tuberosus</emphasis>
</taxonomicName>
can be distinguished from congeners in having a strongly humped dorsal profile (vs. weakly humped or without hump), and its head and flanks covered with rounded tubercles (vs. with smooth or with capillaceous epithelial projections). It additionally differs from congeners in having the following unique combination of characters: eye diameter 8% HL; head width 18.5–19.7% SL; head depth 15.8–16.0% SL; 12–14 rakers on first gill arch; posttemporal process indistinct, not as long as postcleithral process; dorsal margin of postcleithral process slightly concave; length of pectoral-fin spine 15.4–16.7% SL; 17–18 serrations on posterior margin of pectoral spine; length of dorsal-fin spine 18.5–22.4% SL; body depth at anus 14.2–16.0% SL; length of adipose-fin base 23.3–26.1% SL; posterior insertion of adipose fin separate from dorsal procurrent caudal-fin rays; caudal peduncle depth 5.5–6.0% SL; tubular extensions of laterosensory canals on body absent; dark-colored body with pale patches on flanks; caudal fin without dark submarginal longitudinal band running along both lobes; and 43–44 vertebrae.
</paragraph>
</subSubSection>
<subSubSection id="C3256559B17DB40CDFA437FFDCEDFE11" lastPageId="3" lastPageNumber="549" pageId="1" pageNumber="547" type="description">
<paragraph id="8B8036D2B17DB40EDFA437FFDA5CFB89" blockId="1.[151,1437,259,1042]" pageId="1" pageNumber="547">
<emphasis id="B94BEAC0B17DB40EDFA437FFDF64FC8D" bold="true" box="[199,345,763,789]" pageId="1" pageNumber="547">Description.</emphasis>
Morphometric data as in
<tableCitation id="C6BD0369B17DB40EDDE237FFDCEBFC8D" box="[641,726,763,789]" captionStart="TABLE 1" captionStartId="1.[152,249,1087,1113]" captionText="TABLE 1. Morphometric data for Pseudomystus tuberosus (n=3)." pageId="1" pageNumber="547">Table 1</tableCitation>
. Head depressed; dorsal profile slightly concave, with distinct nuchal hump. Ventral profile almost straight. Snout broadly rounded when viewed dorsally. Bony elements of dorsal surface of head covered with thin skin; bones visible, especially on posterior half of neurocranium, and ornamented with numerous fine, radial grooves. Midline of cranium with fontanelle extending from behind snout to beyond level of posterior orbital margin. Supraoccipital process broad, with gently converging sides and emarginate tip; not reaching nuchal plate. Eye ovoid, horizontal axis longest, subcutaneous; located entirely in dorsal half of head. Gill openings wide, extending from post-temporal to beyond isthmus. Gill membranes free from isthmus, with 8 (3) branchiostegal rays. First branchial arch with 4+8 (1), 4+9 (1) or 4+10* (1) gill rakers.
</paragraph>
<caption id="DF40665AB17DB40EDFFB313BDDBDFBC1" box="[152,896,1087,1113]" pageId="1" pageNumber="547" startId="1.[152,249,1087,1113]" targetType="table">
<paragraph id="8B8036D2B17DB40EDFFB313BDDBDFBC1" blockId="1.[152,896,1087,1113]" box="[152,896,1087,1113]" pageId="1" pageNumber="547">
<emphasis id="B94BEAC0B17DB40EDFFB313BDF28FBC1" bold="true" box="[152,277,1087,1113]" pageId="1" pageNumber="547">TABLE 1.</emphasis>
Morphometric data for
<taxonomicName id="4C3F4D51B17DB40EDD41313BDD09FBC1" authorityName="Ng &amp; Tan" authorityYear="2024" box="[546,820,1087,1113]" family="Bagridae" genus="Pseudomystus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="547" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B94BEAC0B17DB40EDD41313BDD09FBC1" box="[546,820,1087,1113]" italics="true" pageId="1" pageNumber="547">Pseudomystus tuberosus</emphasis>
</taxonomicName>
(n=3).
</paragraph>
</caption>
<paragraph id="8B8036D2B17DB40EDDF8316DDAFBF83B" pageId="1" pageNumber="547">
<table id="F93FC472B17D4BF0DFC0316DDB35F83C" box="[163,1288,1129,1956]" colsContinueIn="2.[163,1236,204,778]" gridcols="4" gridrows="20" pageId="1" pageNumber="547">
<tr id="350F3490B17D4BF0DFC0316DDB35FB1B" box="[163,1288,1129,1155]" gridrow="0" pageId="1" pageNumber="547" rowspan-0="1" rowspan-3="1">
<th id="76DE5DECB17D4BF0DDF8316DDD18FB1B" box="[667,805,1129,1155]" gridcol="1" gridrow="0" pageId="1" pageNumber="547">
<emphasis id="B94BEAC0B17DB40EDDF8316DDD35FB1B" bold="true" box="[667,776,1129,1155]" pageId="1" pageNumber="547">Holotype</emphasis>
</th>
<th id="76DE5DECB17D4BF0DCFC316DDA1EFB1B" box="[927,1059,1129,1155]" gridcol="2" gridrow="0" pageId="1" pageNumber="547">
<emphasis id="B94BEAC0B17DB40EDCFC316DDA2AFB1B" bold="true" box="[927,1047,1129,1155]" pageId="1" pageNumber="547">Paratypes</emphasis>
</th>
</tr>
<tr id="350F3490B17D4BF0DFC03197DB35FB36" box="[163,1288,1171,1198]" gridrow="1" pageId="1" pageNumber="547" rowspan-0="1" rowspan-3="1">
<td id="76DE5DECB17D4BF0DDF83197DD18FB36" box="[667,805,1171,1198]" gridcol="1" gridrow="1" pageId="1" pageNumber="547">MZB 17244</td>
<td id="76DE5DECB17D4BF0DCFC3197DA1EFB36" box="[927,1059,1171,1198]" gridcol="2" gridrow="1" pageId="1" pageNumber="547">ZRC 54541</td>
</tr>
<tr id="350F3490B17D4BF0DFC031B9DB35FB4F" box="[163,1288,1213,1239]" gridrow="2" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC031B9DFE1FB4F" box="[163,476,1213,1239]" gridcol="0" gridrow="2" pageId="1" pageNumber="547">Standard length (SL)</th>
<td id="76DE5DECB17D4BF0DDF831B9DD18FB4F" box="[667,805,1213,1239]" gridcol="1" gridrow="2" pageId="1" pageNumber="547">93.0</td>
<td id="76DE5DECB17D4BF0DCFC31B9DA1EFB4F" box="[927,1059,1213,1239]" gridcol="2" gridrow="2" pageId="1" pageNumber="547">79.7</td>
<td id="76DE5DECB17D4BF0DBC031B9DB35FB4F" box="[1187,1288,1213,1239]" gridcol="3" gridrow="2" pageId="1" pageNumber="547">88.4</td>
</tr>
<tr id="350F3490B17D4BF0DFC031ECDB35FA9A" box="[163,1288,1256,1282]" gridrow="3" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC031ECDB35FA9A" box="[163,1288,1256,1282]" colspan="4" colspanRight="3" gridcol="0" gridrow="3" pageId="1" pageNumber="547">%SL</th>
</tr>
<tr id="350F3490B17D4BF0DFC03015DB35FAB4" box="[163,1288,1297,1324]" gridrow="4" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC03015DFE1FAB4" box="[163,476,1297,1324]" gridcol="0" gridrow="4" pageId="1" pageNumber="547">Predorsal length</th>
<td id="76DE5DECB17D4BF0DDF83015DD18FAB4" box="[667,805,1297,1324]" gridcol="1" gridrow="4" pageId="1" pageNumber="547">40.9</td>
<td id="76DE5DECB17D4BF0DCFC3015DA1EFAB4" box="[927,1059,1297,1324]" gridcol="2" gridrow="4" pageId="1" pageNumber="547">40.7</td>
<td id="76DE5DECB17D4BF0DBC03015DB35FAB4" box="[1187,1288,1297,1324]" gridcol="3" gridrow="4" pageId="1" pageNumber="547">40.3</td>
</tr>
<tr id="350F3490B17D4BF0DFC0303FDB35FACE" box="[163,1288,1339,1366]" gridrow="5" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC0303FDFE1FACE" box="[163,476,1339,1366]" gridcol="0" gridrow="5" pageId="1" pageNumber="547">Preanal length</th>
<td id="76DE5DECB17D4BF0DDF8303FDD18FACE" box="[667,805,1339,1366]" gridcol="1" gridrow="5" pageId="1" pageNumber="547">63.9</td>
<td id="76DE5DECB17D4BF0DCFC303FDA1EFACE" box="[927,1059,1339,1366]" gridcol="2" gridrow="5" pageId="1" pageNumber="547">65.0</td>
<td id="76DE5DECB17D4BF0DBC0303FDB35FACE" box="[1187,1288,1339,1366]" gridcol="3" gridrow="5" pageId="1" pageNumber="547">62.9</td>
</tr>
<tr id="350F3490B17D4BF0DFC03062DB35FA19" box="[163,1288,1382,1409]" gridrow="6" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC03062DFE1FA19" box="[163,476,1382,1409]" gridcol="0" gridrow="6" pageId="1" pageNumber="547">Prepelvic length</th>
<td id="76DE5DECB17D4BF0DDF83062DD18FA19" box="[667,805,1382,1409]" gridcol="1" gridrow="6" pageId="1" pageNumber="547">53.0</td>
<td id="76DE5DECB17D4BF0DCFC3062DA1EFA19" box="[927,1059,1382,1409]" gridcol="2" gridrow="6" pageId="1" pageNumber="547">51.9</td>
<td id="76DE5DECB17D4BF0DBC03062DB35FA19" box="[1187,1288,1382,1409]" gridcol="3" gridrow="6" pageId="1" pageNumber="547">50.4</td>
</tr>
<tr id="350F3490B17D4BF0DFC03094DB35FA33" box="[163,1288,1424,1451]" gridrow="7" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC03094DFE1FA33" box="[163,476,1424,1451]" gridcol="0" gridrow="7" pageId="1" pageNumber="547">Prepectoral length</th>
<td id="76DE5DECB17D4BF0DDF83094DD18FA33" box="[667,805,1424,1451]" gridcol="1" gridrow="7" pageId="1" pageNumber="547">26.8</td>
<td id="76DE5DECB17D4BF0DCFC3094DA1EFA33" box="[927,1059,1424,1451]" gridcol="2" gridrow="7" pageId="1" pageNumber="547">25.5</td>
<td id="76DE5DECB17D4BF0DBC03094DB35FA33" box="[1187,1288,1424,1451]" gridcol="3" gridrow="7" pageId="1" pageNumber="547">25.5</td>
</tr>
<tr id="350F3490B17D4BF0DFC030BEDB35FA4D" box="[163,1288,1466,1493]" gridrow="8" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC030BEDFE1FA4D" box="[163,476,1466,1493]" gridcol="0" gridrow="8" pageId="1" pageNumber="547">Length of dorsal-fin base</th>
<td id="76DE5DECB17D4BF0DDF830BEDD18FA4D" box="[667,805,1466,1493]" gridcol="1" gridrow="8" pageId="1" pageNumber="547">12.5</td>
<td id="76DE5DECB17D4BF0DCFC30BEDA1EFA4D" box="[927,1059,1466,1493]" gridcol="2" gridrow="8" pageId="1" pageNumber="547">12.5</td>
<td id="76DE5DECB17D4BF0DBC030BEDB35FA4D" box="[1187,1288,1466,1493]" gridcol="3" gridrow="8" pageId="1" pageNumber="547">12.3</td>
</tr>
<tr id="350F3490B17D4BF0DFC030E0DB35FA67" box="[163,1288,1508,1535]" gridrow="9" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC030E0DFE1FA67" box="[163,476,1508,1535]" gridcol="0" gridrow="9" pageId="1" pageNumber="547">Length of dorsal-fin spine</th>
<td id="76DE5DECB17D4BF0DDF830E0DD18FA67" box="[667,805,1508,1535]" gridcol="1" gridrow="9" pageId="1" pageNumber="547">22.4</td>
<td id="76DE5DECB17D4BF0DCFC30E0DA1EFA67" box="[927,1059,1508,1535]" gridcol="2" gridrow="9" pageId="1" pageNumber="547">20.1</td>
<td id="76DE5DECB17D4BF0DBC030E0DB35FA67" box="[1187,1288,1508,1535]" gridcol="3" gridrow="9" pageId="1" pageNumber="547">18.5</td>
</tr>
<tr id="350F3490B17D4BF0DFC0330ADB35F9B0" box="[163,1288,1550,1576]" gridrow="10" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC0330ADFE1F9B0" box="[163,476,1550,1576]" gridcol="0" gridrow="10" pageId="1" pageNumber="547">Length of anal-fin base</th>
<td id="76DE5DECB17D4BF0DDF8330ADD18F9B0" box="[667,805,1550,1576]" gridcol="1" gridrow="10" pageId="1" pageNumber="547">17.4</td>
<td id="76DE5DECB17D4BF0DCFC330ADA1EF9B0" box="[927,1059,1550,1576]" gridcol="2" gridrow="10" pageId="1" pageNumber="547">18.6</td>
<td id="76DE5DECB17D4BF0DBC0330ADB35F9B0" box="[1187,1288,1550,1576]" gridcol="3" gridrow="10" pageId="1" pageNumber="547">18.8</td>
</tr>
<tr id="350F3490B17D4BF0DFC0333CDB35F9CB" box="[163,1288,1592,1619]" gridrow="11" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC0333CDFE1F9CB" box="[163,476,1592,1619]" gridcol="0" gridrow="11" pageId="1" pageNumber="547">Pelvic-fin length</th>
<td id="76DE5DECB17D4BF0DDF8333CDD18F9CB" box="[667,805,1592,1619]" gridcol="1" gridrow="11" pageId="1" pageNumber="547">14.8</td>
<td id="76DE5DECB17D4BF0DCFC333CDA1EF9CB" box="[927,1059,1592,1619]" gridcol="2" gridrow="11" pageId="1" pageNumber="547">15.3</td>
<td id="76DE5DECB17D4BF0DBC0333CDB35F9CB" box="[1187,1288,1592,1619]" gridcol="3" gridrow="11" pageId="1" pageNumber="547">14.5</td>
</tr>
<tr id="350F3490B17D4BF0DFC03366DB35F9E4" box="[163,1288,1634,1660]" gridrow="12" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC03366DFE1F9E4" box="[163,476,1634,1660]" gridcol="0" gridrow="12" pageId="1" pageNumber="547">Pectoral-fin length</th>
<td id="76DE5DECB17D4BF0DDF83366DD18F9E4" box="[667,805,1634,1660]" gridcol="1" gridrow="12" pageId="1" pageNumber="547">21.3</td>
<td id="76DE5DECB17D4BF0DCFC3366DA1EF9E4" box="[927,1059,1634,1660]" gridcol="2" gridrow="12" pageId="1" pageNumber="547">20.6</td>
<td id="76DE5DECB17D4BF0DBC03366DB35F9E4" box="[1187,1288,1634,1660]" gridcol="3" gridrow="12" pageId="1" pageNumber="547">20.2</td>
</tr>
<tr id="350F3490B17D4BF0DFC03388DB35F93F" box="[163,1288,1676,1703]" gridrow="13" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC03388DFE1F93F" box="[163,476,1676,1703]" gridcol="0" gridrow="13" pageId="1" pageNumber="547">Length of pectoral-fin spine</th>
<td id="76DE5DECB17D4BF0DDF83388DD18F93F" box="[667,805,1676,1703]" gridcol="1" gridrow="13" pageId="1" pageNumber="547">16.7</td>
<td id="76DE5DECB17D4BF0DCFC3388DA1EF93F" box="[927,1059,1676,1703]" gridcol="2" gridrow="13" pageId="1" pageNumber="547">15.4</td>
<td id="76DE5DECB17D4BF0DBC03388DB35F93F" box="[1187,1288,1676,1703]" gridcol="3" gridrow="13" pageId="1" pageNumber="547">15.8</td>
</tr>
<tr id="350F3490B17D4BF0DFC033B2DB35F949" box="[163,1288,1718,1745]" gridrow="14" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC033B2DFE1F949" box="[163,476,1718,1745]" gridcol="0" gridrow="14" pageId="1" pageNumber="547">Caudal-fin length</th>
<td id="76DE5DECB17D4BF0DDF833B2DD18F949" box="[667,805,1718,1745]" gridcol="1" gridrow="14" pageId="1" pageNumber="547">28.2</td>
<td id="76DE5DECB17D4BF0DCFC33B2DA1EF949" box="[927,1059,1718,1745]" gridcol="2" gridrow="14" pageId="1" pageNumber="547">damaged</td>
<td id="76DE5DECB17D4BF0DBC033B2DB35F949" box="[1187,1288,1718,1745]" gridcol="3" gridrow="14" pageId="1" pageNumber="547">damaged</td>
</tr>
<tr id="350F3490B17D4BF0DFC033E4DB35F963" box="[163,1288,1760,1787]" gridrow="15" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC033E4DFE1F963" box="[163,476,1760,1787]" gridcol="0" gridrow="15" pageId="1" pageNumber="547">Length of adipose-fin base</th>
<td id="76DE5DECB17D4BF0DDF833E4DD18F963" box="[667,805,1760,1787]" gridcol="1" gridrow="15" pageId="1" pageNumber="547">23.3</td>
<td id="76DE5DECB17D4BF0DCFC33E4DA1EF963" box="[927,1059,1760,1787]" gridcol="2" gridrow="15" pageId="1" pageNumber="547">26.1</td>
<td id="76DE5DECB17D4BF0DBC033E4DB35F963" box="[1187,1288,1760,1787]" gridcol="3" gridrow="15" pageId="1" pageNumber="547">24.0</td>
</tr>
<tr id="350F3490B17D4BF0DFC0320FDB35F8BD" box="[163,1288,1803,1829]" gridrow="16" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC0320FDFE1F8BD" box="[163,476,1803,1829]" gridcol="0" gridrow="16" pageId="1" pageNumber="547">Dorsal to adipose distance</th>
<td id="76DE5DECB17D4BF0DDF8320FDD18F8BD" box="[667,805,1803,1829]" gridcol="1" gridrow="16" pageId="1" pageNumber="547">11.2</td>
<td id="76DE5DECB17D4BF0DCFC320FDA1EF8BD" box="[927,1059,1803,1829]" gridcol="2" gridrow="16" pageId="1" pageNumber="547">9.8</td>
<td id="76DE5DECB17D4BF0DBC0320FDB35F8BD" box="[1187,1288,1803,1829]" gridcol="3" gridrow="16" pageId="1" pageNumber="547">9.6</td>
</tr>
<tr id="350F3490B17D4BF0DFC03231DB35F8C8" box="[163,1288,1845,1872]" gridrow="17" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC03231DFE1F8C8" box="[163,476,1845,1872]" gridcol="0" gridrow="17" pageId="1" pageNumber="547">Post-adipose distance</th>
<td id="76DE5DECB17D4BF0DDF83231DD18F8C8" box="[667,805,1845,1872]" gridcol="1" gridrow="17" pageId="1" pageNumber="547">15.3</td>
<td id="76DE5DECB17D4BF0DCFC3231DA1EF8C8" box="[927,1059,1845,1872]" gridcol="2" gridrow="17" pageId="1" pageNumber="547">19.1</td>
<td id="76DE5DECB17D4BF0DBC03231DB35F8C8" box="[1187,1288,1845,1872]" gridcol="3" gridrow="17" pageId="1" pageNumber="547">16.3</td>
</tr>
<tr id="350F3490B17D4BF0DFC0325BDB35F8E1" box="[163,1288,1887,1913]" gridrow="18" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC0325BDFE1F8E1" box="[163,476,1887,1913]" gridcol="0" gridrow="18" pageId="1" pageNumber="547">Caudal-peduncle length</th>
<td id="76DE5DECB17D4BF0DDF8325BDD18F8E1" box="[667,805,1887,1913]" gridcol="1" gridrow="18" pageId="1" pageNumber="547">17.0</td>
<td id="76DE5DECB17D4BF0DCFC325BDA1EF8E1" box="[927,1059,1887,1913]" gridcol="2" gridrow="18" pageId="1" pageNumber="547">17.3</td>
<td id="76DE5DECB17D4BF0DBC0325BDB35F8E1" box="[1187,1288,1887,1913]" gridcol="3" gridrow="18" pageId="1" pageNumber="547">17.9</td>
</tr>
<tr id="350F3490B17D4BF0DFC0328DDB35F83C" box="[163,1288,1929,1956]" gridrow="19" pageId="1" pageNumber="547">
<th id="76DE5DECB17D4BF0DFC0328DDFE1F83C" box="[163,476,1929,1956]" gridcol="0" gridrow="19" pageId="1" pageNumber="547">Caudal-peduncle depth</th>
<td id="76DE5DECB17D4BF0DDF8328DDD18F83C" box="[667,805,1929,1956]" gridcol="1" gridrow="19" pageId="1" pageNumber="547">5.6</td>
<td id="76DE5DECB17D4BF0DCFC328DDA1EF83C" box="[927,1059,1929,1956]" gridcol="2" gridrow="19" pageId="1" pageNumber="547">5.5</td>
<td id="76DE5DECB17D4BF0DBC0328DDB35F83C" box="[1187,1288,1929,1956]" gridcol="3" gridrow="19" pageId="1" pageNumber="547">6.0</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8036D2B17DB40EDB5A32B0DBADF856" blockId="1.[1081,1424,1972,1998]" box="[1081,1424,1972,1998]" pageId="1" pageNumber="547">
<emphasis id="B94BEAC0B17DB40EDB5A32B0DBADF856" box="[1081,1424,1972,1998]" italics="true" pageId="1" pageNumber="547">......continued on the next page</emphasis>
</paragraph>
<paragraph id="8B8036D2B17EB40DDBC03424DAE9FE20" box="[1187,1236,288,440]" pageId="2" pageNumber="548">
<table id="F93FC472B17E4BF0DFC035C8DAE9FC92" box="[163,1236,204,778]" colsContinueFrom="1.[163,1288,1129,1956]" gridcols="4" gridrows="14" pageId="2" pageNumber="548">
<td id="76DE5DECB17E4BF0DBC03424DAE9FEA3" box="[1187,1236,288,315]" gridcol="3" gridrow="2" pageId="2" pageNumber="548">16.0</td>
<td id="76DE5DECB17E4BF0DBC0344EDAE9FEFC" box="[1187,1236,330,356]" gridcol="3" gridrow="3" pageId="2" pageNumber="548">28.6</td>
<td id="76DE5DECB17E4BF0DBC03470DAE9FE17" box="[1187,1236,372,399]" gridcol="3" gridrow="4" pageId="2" pageNumber="548">18.5</td>
<td id="76DE5DECB17E4BF0DBC0349ADAE9FE21" box="[1187,1236,414,441]" gridcol="3" gridrow="5" pageId="2" pageNumber="548">16.0</td>
</table>
</paragraph>
<paragraph id="8B8036D2B17EB40DDFA43659DB4DFB9F" blockId="2.[151,1437,861,2039]" pageId="2" pageNumber="548">Mouth subterminal. Oral teeth small, villiform, in irregular rows on all tooth-bearing surfaces. Premaxillary tooth band rounded, of uniform width throughout. Dentary tooth band much narrower than premaxillary tooth band at symphysis, tapering laterally. Vomerine tooth band unpaired, continuous across midline; smoothly arched along anterior margin, tapering laterally to point extending posteriorly past level of premaxillary band; band width narrower than premaxillary band at midline, widening laterally, then tapering to a sharp point postero-laterally.</paragraph>
<paragraph id="8B8036D2B17EB40DDFA43115DED8FB23" blockId="2.[151,1437,861,2039]" pageId="2" pageNumber="548">Barbels in four pairs. Maxillary barbel slender, extending for four-fifths of head length. Nasal barbel slender, extending to one third of distance between posterior orbital margin and dorsalmost extent of gill opening. Inner mandibular-barbel origin close to midline, thicker and longer than nasal barbel, extending for half of head length. Outer mandibular barbel originating postero-lateral of inner mandibular barbel, extending for three quarters of head length.</paragraph>
<paragraph id="8B8036D2B17EB40DDFA431C1DAC3FAF7" blockId="2.[151,1437,861,2039]" pageId="2" pageNumber="548">Body slightly compressed, becoming more so toward caudal peduncle. Dorsal profile gently concave from tip of snout to nuchal region, distinctly humped from nuchal region to origin of dorsal fin, sloping gently ventrad from origin of dorsal fin to end of caudal peduncle. Ventral profile slightly convex to anal-fin base, then sloping slightly dorsally to end of caudal peduncle. Skin smooth, with rounded tubercles; projections especially prominent on head and flanks. Lateral line complete, mid-lateral in position. Vertebrae 17+26=43 (1) or 19+25=44* (2).</paragraph>
<paragraph id="8B8036D2B17EB40DDFA4307DDAECFA43" blockId="2.[151,1437,861,2039]" pageId="2" pageNumber="548">Dorsal fin with spinelet, spine, and 7 (3) rays. Origin of dorsal fin anterior to mid-body, about two-fifths of body. Dorsal fin margin convex, anterior branch of fin rays usually longer than other branches. Dorsal fin spine of moderate length, straight, slender, posterior edge without serrations. Nuchal plate broadly ovoid.</paragraph>
<paragraph id="8B8036D2B17EB40DDFA430E1DED8F9F3" blockId="2.[151,1437,861,2039]" pageId="2" pageNumber="548">Pectoral fin with stout spine, sharply pointed at tip, and 8 (3) rays. Anterior spine margin smooth; posterior spine margin with 17–18 large serrations along entire length. Pectoral fin margin straight anteriorly, convex posteriorly. Cleithral process moderately broad, with slightly concave dorsal margin, extending for two thirds of pectoral-spine length.</paragraph>
<paragraph id="8B8036D2B17EB40DDFA43371DA47F94F" blockId="2.[151,1437,861,2039]" pageId="2" pageNumber="548">Pelvic fin with origin at vertical through posterior end of dorsal-fin base, with i,5 (3) rays, its margin slightly convex; tip of adpressed fin reaching slightly beyond anal-fin origin. Anus and urogenital openings located at vertical through middle of adpressed pelvic fin. Males with short, conical genital papilla.</paragraph>
<paragraph id="8B8036D2B17EB40DDFA433E5DB58F8DB" blockId="2.[151,1437,861,2039]" pageId="2" pageNumber="548">Adipose fin with convex margin for entire length, with deeply incised posterior portion and origin, separate from base of last dorsal-fin ray; fin base of moderate length, spanning half of postdorsal distance. Anal fin originating just posterior to vertical through origin of adipose fin, with iv,11,i* (1) or v,10,1 (2) rays, posterior margin curved.</paragraph>
<paragraph id="8B8036D2B17EB40DDFA43249DD63F813" blockId="2.[151,1437,861,2039]" pageId="2" pageNumber="548">Caudal peduncle slender. Caudal fin deeply forked, with i,7,8,i (3) principal rays; upper lobe slender, pointed, lower lobe rounded. Procurrent rays extend anterior to fin base.</paragraph>
<paragraph id="8B8036D2B17EB40CDFA43290DCEDFE11" blockId="2.[151,1437,861,2039]" lastBlockId="3.[151,1437,151,537]" lastPageId="3" lastPageNumber="549" pageId="2" pageNumber="548">
<emphasis id="B94BEAC0B17EB40DDFA43290DF72F836" bold="true" box="[199,335,1940,1966]" pageId="2" pageNumber="548">Coloration.</emphasis>
In 70% ethanol: head and body dark brownish-gray above, fading on sides of head and abdomen; cream on ventral surfaces. Caudal fin hyaline with diffuse chocolate-brown color on proximal half of fin rays on each lobe. Irregular, wavy yellowish bar over nape. Series of yellowish patches on body: first patch ovoid, behind tympanic region and below dorsal fin; second patch ovoid, on dorsal surface of body at base of last dorsal-fin ray; third patch ovoid, at pelvic-fin base; fourth patch consisting of two coalescent ovoid patches on flank posterior to dorsal-fin base and between pelvic- and anal-fin origins; fifth patch narrowly ellipsoid on dorsal surface of adipose fin at its anterior origin; sixth patch consisting of two coalescent ovoid patches on caudal peduncle, extending into posteroventral corner of adipose fin. Dorsal and anal fins hyaline with chocolate-brown basal, and diffuse brown submarginal bars. Pelvic fin hyaline with diffuse chocolate-brown color on proximal half of fin rays. Posterior edge of adipose fin hyaline. Coloration in life unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3256559B17FB40CDFA43496DA9AFE35" box="[199,1191,402,429]" pageId="3" pageNumber="549" type="distribution">
<paragraph id="8B8036D2B17FB40CDFA43496DA9AFE35" blockId="3.[151,1437,151,537]" box="[199,1191,402,429]" pageId="3" pageNumber="549">
<emphasis id="B94BEAC0B17FB40CDFA43496DF5CFE34" bold="true" box="[199,353,402,428]" pageId="3" pageNumber="549">Distribution.</emphasis>
Known from the Kahayan River drainage in south central Borneo (
<figureCitation id="13042A57B17FB40CDB373497DAA4FE35" box="[1108,1177,403,429]" captionStart="FIGURE 2" captionStartId="4.[151,250,1615,1639]" captionTargetBox="[153,1419,181,1572]" captionTargetId="figure-17@4.[151,1436,181,1589]" captionTargetPageId="4" captionText="FIGURE 2. Map showing collecting locality of Pseudomystus tuberosus." figureDoi="http://doi.org/10.5281/zenodo.13617495" httpUri="https://zenodo.org/record/13617495/files/figure.png" pageId="3" pageNumber="549">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3256559B17FB40CDFA434B3DF72FD81" pageId="3" pageNumber="549" type="etymology">
<paragraph id="8B8036D2B17FB40CDFA434B3DF72FD81" blockId="3.[151,1437,151,537]" pageId="3" pageNumber="549">
<emphasis id="B94BEAC0B17FB40CDFA434B3DF73FE49" bold="true" box="[199,334,439,465]" pageId="3" pageNumber="549">Etymology.</emphasis>
The Latin adjective
<taxonomicName id="4C3F4D51B17FB40CDD2134B3DC93FE49" authorityName="Ng &amp; Tan" authorityYear="2024" box="[578,686,439,465]" family="Bagridae" genus="Pseudomystus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="549" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B94BEAC0B17FB40CDD2134B3DC93FE49" box="[578,686,439,465]" italics="true" pageId="3" pageNumber="549">tuberosus</emphasis>
</taxonomicName>
(
<emphasis id="B94BEAC0B17FB40CDDA234B3DCE5FE48" box="[705,728,439,464]" italics="true" pageId="3" pageNumber="549">-a</emphasis>
,
<emphasis id="B94BEAC0B17FB40CDD8934B3DD28FE48" box="[746,789,439,464]" italics="true" pageId="3" pageNumber="549">-um</emphasis>
) means “full of humps, lumps and protuberances”. The name is used in reference to both the distinctively humpbacked appearance of this species, as well as its tuberculate head and flanks.
</paragraph>
</subSubSection>
</treatment>
</document>
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