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<document id="BF1EAF955B084834B6C55F5BFAF5AC03" ID-DOI="10.5281/zenodo.274748" ID-GBIF-Dataset="e687202a-a625-4aab-94de-312277c7e3fb" ID-ISSN="1175-5326" ID-Zenodo-Dep="274748" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461047074832" checkinUser="plazi" docAuthor="Aarvik, Leif &amp; Karisch, Timm" docDate="2009" docId="039687B6FFA4FFF3FF6E5DFDFF684DDF" docLanguage="en" docName="zt02026p032.pdf" docOrigin="Zootaxa 2026" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Multiquaestia kingstoni Aarvik &amp; Karisch, new species" docType="treatment" docVersion="8" lastPageNumber="25" masterDocId="FFAFFFCEFFA2FFF4FFF95853FFB14F5D" masterDocTitle="Revision of Multiquaestia Karisch (Lepidoptera: Tortricidae: Grapholitini)" masterLastPageNumber="32" masterPageNumber="18" pageNumber="24" updateTime="1698577463540" updateUser="plazi">
<mods:mods id="4CDBCB01E4DCD319EEBF42A45BAEF815" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D0222916EC49DDABFED8DAAB7B915730">
<mods:title id="AA12ACF17442BC5C2663F64954C00C8F">Revision of Multiquaestia Karisch (Lepidoptera: Tortricidae: Grapholitini)</mods:title>
</mods:titleInfo>
<mods:name id="182A5C6FC05C1A11D326D647360D5FEF" type="personal">
<mods:role id="BA68CA524E1E017E879C561199DE417A">
<mods:roleTerm id="564C0DAA0129A4800F0BC48D632C1737">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FA7631E7C6F8409E73BF6CB016DD945A">Aarvik, Leif</mods:namePart>
</mods:name>
<mods:name id="B1C055361428742CAAF07B8059297305" type="personal">
<mods:role id="2806FD11228493B8112527928FD4A947">
<mods:roleTerm id="FDFE8316C45A13C3A10EBC3C0B5D3A6C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0CCFDE9AF6C2E6E546307107828515FB">Karisch, Timm</mods:namePart>
</mods:name>
<mods:typeOfResource id="8553798219DAC4D9CBE5C415FCB58DE0">text</mods:typeOfResource>
<mods:relatedItem id="AA223160DB36AB26B07964908129CDA7" type="host">
<mods:titleInfo id="DEE61FDD6BF48C4C936F9850020164CE">
<mods:title id="EA8947E391310EB30D3A6E4C037E6686">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9537CF1B7AE070BE7DF7FD877116D974">
<mods:date id="3E5DFDB5122F23CCC7E4668CEB36323E">2009</mods:date>
<mods:detail id="A209D8829E077400F9B44BAE99C05BFF" type="volume">
<mods:number id="C853CBD831499F0FA3F2FA9038B661F4">2026</mods:number>
</mods:detail>
<mods:extent id="04CA87D753DE108BF4108FC0A58E25E8" unit="page">
<mods:start id="FEF4E0F3BCB9164EB6E5F35CAD28AF3B">18</mods:start>
<mods:end id="F86841E8C059440113A1AC0A3B69869B">32</mods:end>
</mods:extent>
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<mods:classification id="E812FE744C77E8F227832BBF56E142B1">journal article</mods:classification>
<mods:identifier id="8D9EA07FE6EA6FA6A9DF681E840EE261" type="DOI">10.5281/zenodo.274748</mods:identifier>
<mods:identifier id="532B1B912AE53ADA67394670451FD16D" type="GBIF-Dataset">e687202a-a625-4aab-94de-312277c7e3fb</mods:identifier>
<mods:identifier id="AEB1ACB6AD1F97E7B182A16A6C4A7C89" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B7C50F33048FAABEDABE755CE0122A2F" type="Zenodo-Dep">274748</mods:identifier>
</mods:mods>
<treatment id="039687B6FFA4FFF3FF6E5DFDFF684DDF" ID-DOI="http://doi.org/10.5281/zenodo.6216215" ID-GBIF-Taxon="119611112" ID-Zenodo-Dep="6216215" LSID="urn:lsid:plazi:treatment:039687B6FFA4FFF3FF6E5DFDFF684DDF" httpUri="http://treatment.plazi.org/id/039687B6FFA4FFF3FF6E5DFDFF684DDF" lastPageId="7" lastPageNumber="25" pageId="6" pageNumber="24">
<subSubSection id="C325652BFFA4FFF2FF6E5DFDFED14AB1" pageId="6" pageNumber="24" type="nomenclature">
<paragraph id="8B8036A0FFA4FFF2FF6E5DFDFC8F4A95" blockId="6.[151,830,1454,1516]" box="[151,830,1454,1480]" pageId="6" pageNumber="24">
<heading id="D0C881CCFFA4FFF2FF6E5DFDFC8F4A95" bold="true" box="[151,830,1454,1480]" fontSize="11" level="1" pageId="6" pageNumber="24" reason="1">
<emphasis id="B94BEAB2FFA4FFF2FF6E5DFDFC8F4A95" bold="true" box="[151,830,1454,1480]" pageId="6" pageNumber="24">
<taxonomicName id="4C3F4D23FFA4FFF2FF6E5DFDFD104A95" ID-CoL="B3XJM" authority="Aarvik &amp; Karisch" authorityName="Aarvik &amp; Karisch" box="[151,673,1454,1480]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="24" phylum="Arthropoda" rank="species" species="kingstoni" status="sp. nov.">
<emphasis id="B94BEAB2FFA4FFF2FF6E5DFDFE064A95" bold="true" box="[151,439,1454,1480]" italics="true" pageId="6" pageNumber="24">Multiquaestia kingstoni</emphasis>
Aarvik &amp; Karisch
</taxonomicName>
,
<taxonomicNameLabel id="A27857C9FFA4FFF2FD575DFDFC8F4A95" box="[686,830,1454,1480]" pageId="6" pageNumber="24" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8036A0FFA4FFF2FF6E5D81FED14AB1" blockId="6.[151,830,1454,1516]" box="[151,352,1490,1516]" pageId="6" pageNumber="24">
(
<figureCitation id="13042A25FFA4FFF2FF665D81FEB04AB1" box="[159,257,1490,1516]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,269,1941,1965]" captionTargetBox="[151,1436,223,1920]" captionTargetId="figure@4.[151,1436,183,1922]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Adults of Multiquaestia. 7. M. skulei n. sp. Holotype. 8. M. agassizi n. sp. Holotype. 9. M. dallastai n. sp. Holotype. 10. M. kingstoni n. sp. Holotype. 11. M. purana n. sp. 12. M. iringana n. sp. 13. M. albimaculana Karisch. Holotype. 14. M. andersi n. sp. Holotype. Scale 10 mm." httpUri="https://zenodo.org/record/274750/files/figure.png" pageId="6" pageNumber="24">Figs. 10</figureCitation>
,
<figureCitation id="13042A25FFA4FFF2FEF75D81FEE84AB1" box="[270,345,1490,1516]" captionStart="FIGURES 23 – 28" captionStartId="9.[151,274,1557,1581]" captionTargetBox="[151,1436,194,1533]" captionTargetId="figure@9.[151,1436,194,1534]" captionTargetPageId="9" captionText="FIGURES 23 – 28. Male genitalia of Multiquaestia. 23. M. dallastai n. sp. 24. M. dallastai n. sp. aedeagus. 25. M. kingstoni n. sp. 26. M. kingstoni n. sp. aedeagus. 27. M. purana n. sp. 28. M. purana n. sp. aedeagus." httpUri="https://zenodo.org/record/274753/files/figure.png" pageId="6" pageNumber="24">25, 26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA4FFF2FF6E5E4DFC1B4902" pageId="6" pageNumber="24" type="materials_examined">
<paragraph id="8B8036A0FFA4FFF2FF6E5E4DFC1B4902" blockId="6.[151,1436,1566,1631]" pageId="6" pageNumber="24">
<emphasis id="B94BEAB2FFA4FFF2FF6E5E4DFEFD4965" bold="true" box="[151,332,1566,1592]" pageId="6" pageNumber="24">
<typeStatus id="54848802FFA4FFF2FF6E5E4DFF654965" box="[151,212,1566,1592]" pageId="6" pageNumber="24">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54848802FFA4FFF2FEAF5E4CFE774964" box="[342,454,1567,1593]" pageId="6" pageNumber="24" type="holotype">Holotype</typeStatus>
, 3,
<collectingCountry id="F3287630FFA4FFF2FE015E4CFDC44964" box="[504,629,1567,1593]" name="Malawi" pageId="6" pageNumber="24">MALAWI</collectingCountry>
, Central Region, Lilongwe District, Ntchisi Forest Reserve
<quantity id="4CC79B45FFA4FFF2FAC25E4CFA294964" box="[1339,1432,1567,1593]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="6" pageNumber="24" unit="m" value="1560.0">1560 m</quantity>
,
<date id="FF811060FFA4FFF2FF6E5E16FEA14902" box="[151,272,1605,1631]" pageId="6" pageNumber="24" value="2004-02-20">20.ii.2004</date>
, leg. L. Aarvik, genitalia slide 2743 L. Aarvik (
<collectionCode id="ED2EAE65FFA4FFF2FCC45E16FC2C4902" box="[829,925,1605,1631]" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="6" pageNumber="24">NHMO</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA4FFF3FF6E5EC1FED34E2E" lastPageId="7" lastPageNumber="25" pageId="6" pageNumber="24" type="description">
<paragraph id="8B8036A0FFA4FFF3FF6E5EC1FED34E2E" blockId="6.[151,1437,1682,2018]" lastBlockId="7.[151,1437,152,642]" lastPageId="7" lastPageNumber="25" pageId="6" pageNumber="24">
<emphasis id="B94BEAB2FFA4FFF2FF6E5EC1FE8149F1" bold="true" box="[151,304,1682,1708]" pageId="6" pageNumber="24">Description.</emphasis>
Adult (
<figureCitation id="13042A25FFA4FFF2FE755EC0FE5449F0" box="[396,485,1683,1709]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,269,1941,1965]" captionTargetBox="[151,1436,223,1920]" captionTargetId="figure@4.[151,1436,183,1922]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Adults of Multiquaestia. 7. M. skulei n. sp. Holotype. 8. M. agassizi n. sp. Holotype. 9. M. dallastai n. sp. Holotype. 10. M. kingstoni n. sp. Holotype. 11. M. purana n. sp. 12. M. iringana n. sp. 13. M. albimaculana Karisch. Holotype. 14. M. andersi n. sp. Holotype. Scale 10 mm." httpUri="https://zenodo.org/record/274750/files/figure.png" pageId="6" pageNumber="24">Fig. 10</figureCitation>
). Wingspan 14.0 mm.
<emphasis id="B94BEAB2FFA4FFF2FD0A5EC0FC8249F1" box="[755,819,1683,1708]" italics="true" pageId="6" pageNumber="24">Head</emphasis>
: Dirty white, mixed with grey around eye. Antenna brownish grey, scape dirty white. Labial palpus 1.5 times diameter of eye, externally dark grey, internally and on dorsal edge dirty white; due to scaling on second segment broadly triangular in lateral view.
<emphasis id="B94BEAB2FFA4FFF2FB035EB3FAFE49A4" box="[1274,1359,1760,1785]" italics="true" pageId="6" pageNumber="24">Thorax</emphasis>
: Dirty white in centre, anteriorly and posteriorly brownish grey; tegulae brown. Legs brownish grey, tarsi with pale rings. Forewing upperside with basal 2/3 blackish brown; terminal 1/3 ochreous; on dorsum two white marks; the inner mark reaching to about 2/5 from dorsum, the distal one reaching slightly above 1/4; a white dot present some distance beyond tip of distal white mark, and one between wing base and inner white mark; fringes with dark bar near apex, at middle of termen and at tornus. Forewing underside with patch of yellowish scales in cell below m-stem.
<collectingCountry id="F3287630FFA4FFF2FD955F9BFD1648BF" box="[620,679,1992,2018]" name="India" pageId="6" pageNumber="24">Hind</collectingCountry>
wing upperside dark brownish grey; ochreous hair-pencil from base of hind margin to scale cover on dorsum of abdomen.
<emphasis id="B94BEAB2FFA5FFF3FC7458CBFBB34FEC" box="[909,1026,152,177]" italics="true" pageId="7" pageNumber="25">Abdomen</emphasis>
: Grey, abdominal brush greyish ochreous. Male genitalia (
<figureCitation id="13042A25FFA5FFF3FE3F58EDFDAB4F85" box="[454,538,190,216]" captionStart="FIGURES 23 – 28" captionStartId="9.[151,274,1557,1581]" captionTargetBox="[151,1436,194,1533]" captionTargetId="figure@9.[151,1436,194,1534]" captionTargetPageId="9" captionText="FIGURES 23 – 28. Male genitalia of Multiquaestia. 23. M. dallastai n. sp. 24. M. dallastai n. sp. aedeagus. 25. M. kingstoni n. sp. 26. M. kingstoni n. sp. aedeagus. 27. M. purana n. sp. 28. M. purana n. sp. aedeagus." httpUri="https://zenodo.org/record/274753/files/figure.png" pageId="7" pageNumber="25">Fig. 25</figureCitation>
) with tegumen distally evenly curved, uncus not differentiated, socii present, but weak; lower edge of sacculus angled, cucullus large, roughly transversely semicircular, rounded on dorsal edge, ventral angle rounded; aedeagus (
<figureCitation id="13042A25FFA5FFF3FD82595FFD654E7B" box="[635,724,268,294]" captionStart="FIGURES 23 – 28" captionStartId="9.[151,274,1557,1581]" captionTargetBox="[151,1436,194,1533]" captionTargetId="figure@9.[151,1436,194,1534]" captionTargetPageId="9" captionText="FIGURES 23 – 28. Male genitalia of Multiquaestia. 23. M. dallastai n. sp. 24. M. dallastai n. sp. aedeagus. 25. M. kingstoni n. sp. 26. M. kingstoni n. sp. aedeagus. 27. M. purana n. sp. 28. M. purana n. sp. aedeagus." httpUri="https://zenodo.org/record/274753/files/figure.png" pageId="7" pageNumber="25">Fig. 26</figureCitation>
) angled at 1/5 basally broad, otherwise slender and nearly straight, slightly widening distally; with 9 cornuti of different length, from large spindle-shaped to thin and short; no carinae.
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA5FFF3FF3F592CFEB84D53" pageId="7" pageNumber="25" type="diagnosis">
<paragraph id="8B8036A0FFA5FFF3FF3F592CFEB84D53" blockId="7.[151,1437,152,642]" pageId="7" pageNumber="25">
<emphasis id="B94BEAB2FFA5FFF3FF3F592CFEF74EC4" bold="true" box="[198,326,383,409]" pageId="7" pageNumber="25">Diagnosis.</emphasis>
<taxonomicName id="4C3F4D23FFA5FFF3FEB459D3FE504EC4" box="[333,481,384,409]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="25" phylum="Arthropoda" rank="species" species="kingstoni">
<emphasis id="B94BEAB2FFA5FFF3FEB459D3FE504EC4" box="[333,481,384,409]" italics="true" pageId="7" pageNumber="25">M. kingstoni</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="4C3F4D23FFA5FFF3FCF959D3FCCD4EC4" box="[768,892,384,409]" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="25" phylum="Arthropoda" rank="species" species="purana">
<emphasis id="B94BEAB2FFA5FFF3FCF959D3FCCD4EC4" box="[768,892,384,409]" italics="true" pageId="7" pageNumber="25">M. purana</emphasis>
</taxonomicName>
in wing pattern and colouration. However,
<taxonomicName id="4C3F4D23FFA5FFF3FA8559D3FF5D4E9D" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="25" phylum="Arthropoda" rank="species" species="purana">
<emphasis id="B94BEAB2FFA5FFF3FA8559D3FF5D4E9D" italics="true" pageId="7" pageNumber="25">M. purana</emphasis>
</taxonomicName>
is larger, has a longer and more pointed distal white mark in the forewing, and has a distinct white spot posteriorly on the thorax. The roughly semicircular cucullus in the male genitalia is diagnostic for
<taxonomicName id="4C3F4D23FFA5FFF3FA82599EFEB34D50" class="Insecta" family="Tortricidae" genus="Multiquaestia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="25" phylum="Arthropoda" rank="species" species="kingstoni">
<emphasis id="B94BEAB2FFA5FFF3FA82599EFEB34D50" italics="true" pageId="7" pageNumber="25">M. kingstoni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA5FFF3FF3F5A49FE7E4D69" box="[198,463,538,564]" pageId="7" pageNumber="25" type="distribution">
<paragraph id="8B8036A0FFA5FFF3FF3F5A49FE7E4D69" blockId="7.[151,1437,152,642]" box="[198,463,538,564]" pageId="7" pageNumber="25">
<emphasis id="B94BEAB2FFA5FFF3FF3F5A49FED64D69" bold="true" box="[198,359,538,564]" pageId="7" pageNumber="25">Distribution.</emphasis>
<collectingCountry id="F3287630FFA5FFF3FE945A49FE7A4D69" box="[365,459,538,564]" name="Malawi" pageId="7" pageNumber="25">Malawi</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C325652BFFA5FFF3FF3F5A13FF684DDF" pageId="7" pageNumber="25" type="etymology">
<paragraph id="8B8036A0FFA5FFF3FF3F5A13FF684DDF" blockId="7.[151,1437,152,642]" pageId="7" pageNumber="25">
<emphasis id="B94BEAB2FFA5FFF3FF3F5A13FEE24D07" bold="true" box="[198,339,576,602]" pageId="7" pageNumber="25">Etymology.</emphasis>
The species is named after Anthony Kingston, who organized the collecting trip to
<collectingCountry id="F3287630FFA5FFF3FADC5A12FACE4D06" box="[1317,1407,577,603]" name="Malawi" pageId="7" pageNumber="25">Malawi</collectingCountry>
in 2004.
</paragraph>
</subSubSection>
</treatment>
</document>
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