391 lines
60 KiB
XML
391 lines
60 KiB
XML
<document id="96829222945A0A99F4C47A7A3DCA610E" ID-DOI="10.11646/zootaxa.5016.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="5222106" ID-ZooBank="4CD56237-D83B-4604-9915-1C82AE4836AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629378235035" checkinUser="plazi" docAuthor="Matalin, Andrey V., Perkovsky, Evgeny E. & Vasilenko, Dmitry V." docDate="2021" docId="0394ED212C64FF84FF727DC4FC48FE94" docLanguage="en" docName="zootaxa.5016.2.5.pdf" docOrigin="Zootaxa 5016 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Goriresina Matalin & Perkovsky & Vasilenko 2021, new genus" docType="treatment" docVersion="3" lastPageNumber="249" masterDocId="FFAD95592C66FF82FFE5780FFFD8FFF2" masterDocTitle="First record of tiger beetles (Coleoptera, Cicindelidae) from Rovno amber, with the description of a new genus and species" masterLastPageNumber="256" masterPageNumber="243" pageNumber="245" updateTime="1699070361325" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="5D239D6ECE52E1A6B2AA5DC1A3812690">
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<mods:title id="98ECCB3ACD5B45892C1D4085F975D393">First record of tiger beetles (Coleoptera, Cicindelidae) from Rovno amber, with the description of a new genus and species</mods:title>
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<mods:name id="57EF0720CB8C472E2F92658419A76027" type="personal">
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<mods:roleTerm id="1B554A1DD65100154D41A34D9AC75179">Author</mods:roleTerm>
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<mods:namePart id="6F337ADCCD5AF5B9ADB69AD95F062C05">Matalin, Andrey V.</mods:namePart>
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<mods:affiliation id="146C5E6067E046854374FF99584FBCF2">Moscow State Pedagogical University, Education-Scientific Centre Ecology & Biodiversity, Kibalchicha str. 6, build. 3, Moscow 129164, Russia & Pirogov National Research Medical University, Department of Biology, Ostrovitianova str. 1, Moscow 117997, Russia</mods:affiliation>
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<mods:namePart id="9A168E1C79C8BEB26AF7D7A6C88A1652">Perkovsky, Evgeny E.</mods:namePart>
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<mods:affiliation id="ABAFDBC9727DFDEFA183701FFCDE6A02">Department of Entomology and Collection Management, I. I. Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmel’nyts’koho 15, Kiev 01601, Ukraine</mods:affiliation>
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<mods:namePart id="E34FCCA953D9A22BED3D6C528AF48647">Vasilenko, Dmitry V.</mods:namePart>
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<mods:affiliation id="B688EEEC0BC9C7332CB461671106CBEC">A. A. Borissiak Paleontological Institute of the Russian Academy of Sciences, Moscow, Russia & Cherepovets State University, Lunacharsky Avenue, 5, Cherepovets, 162600 Russia</mods:affiliation>
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<mods:title id="608B35C0EEA16038D55C8FC0DFFA0E36">Zootaxa</mods:title>
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<mods:part id="FB6A2809E46C986D0980512E42BF2802">
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<mods:date id="6DA25803EA4C269E676D4967FBE6C486">2021</mods:date>
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<mods:number id="DD8AD8E4C4002D4D3DF873C4725F6A3D">2021-08-05</mods:number>
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<mods:identifier id="15BC6EA0F92E514FE5737FD04268143B" type="DOI">10.11646/zootaxa.5016.2.5</mods:identifier>
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<subSubSection id="C3270FBC2C64FF80FF727DC4FE72FA17" box="[151,426,1483,1509]" pageId="2" pageNumber="245" type="nomenclature">
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<paragraph id="8B825C372C64FF80FF727DC4FE72FA17" blockId="2.[151,426,1483,1545]" box="[151,426,1483,1509]" pageId="2" pageNumber="245">
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<heading id="D0CAEB5B2C64FF80FF727DC4FE72FA17" bold="true" box="[151,426,1483,1509]" fontSize="11" level="1" pageId="2" pageNumber="245" reason="1">
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<emphasis id="B94980252C64FF80FF727DC4FE72FA17" bold="true" box="[151,426,1483,1509]" pageId="2" pageNumber="245">
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<taxonomicName id="4C3D27B42C64FF80FF727DC4FEC1FA17" authority="Matalin & Perkovsky & Vasilenko, 2021" authorityName="Matalin & Perkovsky & Vasilenko" authorityYear="2021" box="[151,281,1483,1509]" class="Insecta" family="Carabidae" genus="Goriresina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B94980252C64FF80FF727DC4FEC1FA17" bold="true" box="[151,281,1483,1509]" italics="true" pageId="2" pageNumber="245">Goriresina</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27A3D5E2C64FF80FECD7DC4FE72FA17" box="[296,426,1483,1509]" pageId="2" pageNumber="245" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3270FBC2C64FF80FF727DE0FEC1F9FB" box="[151,281,1519,1545]" pageId="2" pageNumber="245" type="description">
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<paragraph id="8B825C372C64FF80FF727DE0FEC1F9FB" blockId="2.[151,426,1483,1545]" box="[151,281,1519,1545]" pageId="2" pageNumber="245">
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(
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<figureCitation id="130640B22C64FF80FF7A7DE0FEC9F9FB" box="[159,273,1519,1545]" captionStart-0="FIGURES 3–4" captionStart-1="FIGURES 5–7" captionStart-2="FIGURES 8–10" captionStart-3="FIGURES 11–16" captionStart-4="FIGURE 17" captionStartId-0="4.[151,264,1971,1995]" captionStartId-1="5.[151,264,1867,1891]" captionStartId-2="6.[151,264,1860,1884]" captionStartId-3="7.[151,264,1899,1923]" captionStartId-4="8.[151,250,1569,1593]" captionTargetBox-0="[178,1409,181,1948]" captionTargetBox-1="[257,1330,181,1843]" captionTargetBox-2="[167,1418,383,1836]" captionTargetBox-3="[211,1379,187,1874]" captionTargetBox-4="[568,1020,207,1528]" captionTargetId-0="figure-24@4.[177,1410,181,1948]" captionTargetId-1="figure-17@5.[257,1330,181,1843]" captionTargetId-2="figure-108@6.[166,1421,381,1837]" captionTargetId-3="figure-17@7.[209,1379,181,1875]" captionTargetId-4="figure-24@8.[548,1040,181,1545]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURES 3–4. Goriresina fungifora gen. n., sp. n., holotype: 3—left lateral view; 4—right dorso-lateral view." captionText-1="FIGURES 5–7. Goriresina fungifora gen. n., sp. n., holotype, left lateral view: 5—head and pronotum; 6—labrum; 7—mouth parts. Abbreviations: lss—left scape seta; lcs—left clypeal seta; fts—left fore-trochanter seta; llb—left latero-basal seta; lla— left latero-apical seta; la—left apical seta; ra—right apical seta; rla—right latero-apical seta; lb—labrum; lmn—left mandible; mxp—maxillary palps; lbp—labial palps; cas—cardo setae; sts—stipes setae." captionText-2="FIGURES 8–10. Goriresina fungifora gen. n., sp. n., holotype: 8—head and labrum, right dorso-lateral view (arrow points to the synnema of Ophiocordyceps sp.); 9—labrum, right dorso-lateral view; 10—reconstructed contour of the labrum, dorsal view. Abbreviations: raso—right anterior supraorbital seta; rlb—right latero-basal labial seta; rla—right latero-apical labial seta; rmn—right mandible; rmxp—right maxillary palp." captionText-3="FIGURES 11–16. Goriresina fungifora gen. n., sp. n., holotype: 11—elytra; 12—punctation and pubescence of shoulders area of right elytron; 13—lateral edge of right elytron; 14—right thoracic and abdominal segments; 15—legs; 16—middle left tarsus. Abbreviations: cv—costal vein of right wing; mte—metepisternum; as1—first abdominal sternite; hc—hind coxa; fr—fore right leg; mr—middle right leg; fl—fore left leg; ml—middle left leg; hl—hind left leg; I–V—middle tarsomeres; cl—claws." captionText-4="FIGURE 17. Simplified reconstruction of the habitus of Goriresina fungifora gen. n., sp. n., holotype, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.5222110" figureDoi-1="http://doi.org/10.5281/zenodo.5222112" figureDoi-2="http://doi.org/10.5281/zenodo.5222114" figureDoi-3="http://doi.org/10.5281/zenodo.5222118" figureDoi-4="http://doi.org/10.5281/zenodo.5222120" httpUri-0="https://zenodo.org/record/5222110/files/figure.png" httpUri-1="https://zenodo.org/record/5222112/files/figure.png" httpUri-2="https://zenodo.org/record/5222114/files/figure.png" httpUri-3="https://zenodo.org/record/5222118/files/figure.png" httpUri-4="https://zenodo.org/record/5222120/files/figure.png" pageId="2" pageNumber="245">Figs 3–17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3270FBC2C64FF80FF727E3CFD5EF9B9" box="[151,646,1587,1611]" pageId="2" pageNumber="245" type="type_taxon">
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<paragraph id="8B825C372C64FF80FF727E3CFD5EF9B9" blockId="2.[151,646,1587,1611]" box="[151,646,1587,1611]" pageId="2" pageNumber="245">
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<emphasis id="B94980252C64FF80FF727E3CFEFBF9B9" bold="true" box="[151,291,1587,1611]" pageId="2" pageNumber="245">
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<typeStatus id="5486E2952C64FF80FF727E3CFF14F9B9" box="[151,204,1587,1611]" pageId="2" pageNumber="245">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C3D27B42C64FF80FECF7E3BFE24F9B9" authority="Matalin & Perkovsky & Vasilenko, 2021" authorityName="Matalin & Perkovsky & Vasilenko" authorityYear="2021" box="[298,508,1587,1611]" class="Insecta" family="Carabidae" genus="Goriresina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="fungifora" status="sp. nov.">
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<emphasis id="B94980252C64FF80FECF7E3BFE24F9B9" box="[298,508,1587,1611]" italics="true" pageId="2" pageNumber="245">Goriresina fungifora</emphasis>
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</taxonomicName>
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,
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<emphasis id="B94980252C64FF80FDEC7E3CFD5EF9B9" bold="true" box="[521,646,1587,1611]" pageId="2" pageNumber="245">
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<taxonomicNameLabel id="A27A3D5E2C64FF80FDEC7E3CFD5EF9B9" box="[521,646,1587,1611]" pageId="2" pageNumber="245" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="C3270FBC2C64FF81FF727E76FB9AFCCB" lastPageId="3" lastPageNumber="246" pageId="2" pageNumber="245" type="description">
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<paragraph id="8B825C372C64FF80FF727E76FD50F961" blockId="2.[151,1437,1657,2008]" box="[151,648,1657,1684]" pageId="2" pageNumber="245">
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<emphasis id="B94980252C64FF80FF727E76FEF1F961" bold="true" box="[151,297,1657,1683]" pageId="2" pageNumber="245">Description.</emphasis>
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TL =
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<quantity id="4CC5F1D22C64FF80FE8B7E76FE0AF966" box="[366,466,1657,1684]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="2" pageNumber="245" unit="mm" value="10.7">10.7 mm</quantity>
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in the
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<typeStatus id="5486E2952C64FF80FDFA7E76FD5DF961" box="[543,645,1657,1683]" pageId="2" pageNumber="245" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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<paragraph id="8B825C372C64FF81FF227E92FE7BFEEF" blockId="2.[151,1437,1657,2008]" lastBlockId="3.[151,1437,150,1977]" lastPageId="3" lastPageNumber="246" pageId="2" pageNumber="245">
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Head with large, protruding, hemispherical eyes (right eye damaged by an
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<taxonomicName id="4C3D27B42C64FF80FBF07E92FB2BF94A" box="[1045,1267,1693,1720]" class="Sordariomycetes" family="Ophiocordycipitaceae" genus="Ophiocordyceps" kingdom="Fungi" order="Hypocreales" pageId="2" pageNumber="245" phylum="Ascomycota" rank="species" species="undetermined">
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<emphasis id="B94980252C64FF80FBF07E92FB13F945" box="[1045,1227,1693,1719]" italics="true" pageId="2" pageNumber="245">Ophiocordyceps</emphasis>
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sp.
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</taxonomicName>
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synnema); left posterior and right anterior supra-orbital setae, as well as left clypeal seta clearly visible, while right posterior and left anterior supra-orbital, as well as right clypeal setae could not be found (
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<figureCitation id="130640B22C64FF80FBA87EEAFB42F8F2" box="[1101,1178,1765,1792]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[257,1330,181,1843]" captionTargetId="figure-17@5.[257,1330,181,1843]" captionTargetPageId="5" captionText="FIGURES 5–7. Goriresina fungifora gen. n., sp. n., holotype, left lateral view: 5—head and pronotum; 6—labrum; 7—mouth parts. Abbreviations: lss—left scape seta; lcs—left clypeal seta; fts—left fore-trochanter seta; llb—left latero-basal seta; lla— left latero-apical seta; la—left apical seta; ra—right apical seta; rla—right latero-apical seta; lb—labrum; lmn—left mandible; mxp—maxillary palps; lbp—labial palps; cas—cardo setae; sts—stipes setae." figureDoi="http://doi.org/10.5281/zenodo.5222112" httpUri="https://zenodo.org/record/5222112/files/figure.png" pageId="2" pageNumber="245">Figs 5</figureCitation>
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,
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<figureCitation id="130640B22C64FF80FB4F7EEAFB61F90D" box="[1194,1209,1765,1791]" captionStart="FIGURES 8–10" captionStartId="6.[151,264,1860,1884]" captionTargetBox="[167,1418,383,1836]" captionTargetId="figure-108@6.[166,1421,381,1837]" captionTargetPageId="6" captionText="FIGURES 8–10. Goriresina fungifora gen. n., sp. n., holotype: 8—head and labrum, right dorso-lateral view (arrow points to the synnema of Ophiocordyceps sp.); 9—labrum, right dorso-lateral view; 10—reconstructed contour of the labrum, dorsal view. Abbreviations: raso—right anterior supraorbital seta; rlb—right latero-basal labial seta; rla—right latero-apical labial seta; rmn—right mandible; rmxp—right maxillary palp." figureDoi="http://doi.org/10.5281/zenodo.5222114" httpUri="https://zenodo.org/record/5222114/files/figure.png" pageId="2" pageNumber="245">8</figureCitation>
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); orbital plates and lateral portions of frons with at least four longitudinal furrows (
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<figureCitation id="130640B22C64FF80FC8B7F06FC6DF8D1" box="[878,949,1801,1827]" captionStart="FIGURES 3–4" captionStartId="4.[151,264,1971,1995]" captionTargetBox="[178,1409,181,1948]" captionTargetId="figure-24@4.[177,1410,181,1948]" captionTargetPageId="4" captionText="FIGURES 3–4. Goriresina fungifora gen. n., sp. n., holotype: 3—left lateral view; 4—right dorso-lateral view." figureDoi="http://doi.org/10.5281/zenodo.5222110" httpUri="https://zenodo.org/record/5222110/files/figure.png" pageId="2" pageNumber="245">Fig. 4</figureCitation>
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). Labrum most likely pale, relatively long, moderately convex, covering the mandibles to 4/5 of their length, dilated in the base with straight lateral margins, six submarginal setae (latero-basal setae very long), and two apical teeth with notch between them (
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<figureCitation id="130640B22C64FF80FB037F5EFA91F899" box="[1254,1353,1873,1900]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[257,1330,181,1843]" captionTargetId="figure-17@5.[257,1330,181,1843]" captionTargetPageId="5" captionText="FIGURES 5–7. Goriresina fungifora gen. n., sp. n., holotype, left lateral view: 5—head and pronotum; 6—labrum; 7—mouth parts. Abbreviations: lss—left scape seta; lcs—left clypeal seta; fts—left fore-trochanter seta; llb—left latero-basal seta; lla— left latero-apical seta; la—left apical seta; ra—right apical seta; rla—right latero-apical seta; lb—labrum; lmn—left mandible; mxp—maxillary palps; lbp—labial palps; cas—cardo setae; sts—stipes setae." figureDoi="http://doi.org/10.5281/zenodo.5222112" httpUri="https://zenodo.org/record/5222112/files/figure.png" pageId="2" pageNumber="245">Figs 5, 6</figureCitation>
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,
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<figureCitation id="130640B22C64FF80FAB17F5EFA57F899" box="[1364,1423,1873,1899]" captionStart="FIGURES 8–10" captionStartId="6.[151,264,1860,1884]" captionTargetBox="[167,1418,383,1836]" captionTargetId="figure-108@6.[166,1421,381,1837]" captionTargetPageId="6" captionText="FIGURES 8–10. Goriresina fungifora gen. n., sp. n., holotype: 8—head and labrum, right dorso-lateral view (arrow points to the synnema of Ophiocordyceps sp.); 9—labrum, right dorso-lateral view; 10—reconstructed contour of the labrum, dorsal view. Abbreviations: raso—right anterior supraorbital seta; rlb—right latero-basal labial seta; rla—right latero-apical labial seta; rmn—right mandible; rmxp—right maxillary palp." figureDoi="http://doi.org/10.5281/zenodo.5222114" httpUri="https://zenodo.org/record/5222114/files/figure.png" pageId="2" pageNumber="245">8–10</figureCitation>
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). Antennae filiform, apparently extending behind shoulders, scape with a single long apical seta (
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<figureCitation id="130640B22C64FF80FB2A7F7AFAC0F87D" box="[1231,1304,1909,1935]" captionStart="FIGURES 3–4" captionStartId="4.[151,264,1971,1995]" captionTargetBox="[178,1409,181,1948]" captionTargetId="figure-24@4.[177,1410,181,1948]" captionTargetPageId="4" captionText="FIGURES 3–4. Goriresina fungifora gen. n., sp. n., holotype: 3—left lateral view; 4—right dorso-lateral view." figureDoi="http://doi.org/10.5281/zenodo.5222110" httpUri="https://zenodo.org/record/5222110/files/figure.png" pageId="2" pageNumber="245">Figs 3</figureCitation>
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,
|
||
<figureCitation id="130640B22C64FF80FAC07F7AFAECF862" box="[1317,1332,1909,1936]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[257,1330,181,1843]" captionTargetId="figure-17@5.[257,1330,181,1843]" captionTargetPageId="5" captionText="FIGURES 5–7. Goriresina fungifora gen. n., sp. n., holotype, left lateral view: 5—head and pronotum; 6—labrum; 7—mouth parts. Abbreviations: lss—left scape seta; lcs—left clypeal seta; fts—left fore-trochanter seta; llb—left latero-basal seta; lla— left latero-apical seta; la—left apical seta; ra—right apical seta; rla—right latero-apical seta; lb—labrum; lmn—left mandible; mxp—maxillary palps; lbp—labial palps; cas—cardo setae; sts—stipes setae." figureDoi="http://doi.org/10.5281/zenodo.5222112" httpUri="https://zenodo.org/record/5222112/files/figure.png" pageId="2" pageNumber="245">5</figureCitation>
|
||
), pedicel glabrous, 3
|
||
<superScript id="7C48F17F2C64FF80FEF07F96FEFBF855" attach="left" box="[277,291,1945,1959]" fontSize="6" pageId="2" pageNumber="245">rd</superScript>
|
||
antennomere the longest. Maxillary palpus with well-visible palpomeres 2–4; 2
|
||
<superScript id="7C48F17F2C64FF80FB547F96FB19F855" attach="left" box="[1201,1217,1945,1959]" fontSize="6" pageId="2" pageNumber="245">nd</superScript>
|
||
palpomere slightly fusiform, with five visible stout setae; 3
|
||
<superScript id="7C48F17F2C64FF80FDB07FB2FDBBF839" attach="left" box="[597,611,1981,1995]" fontSize="6" pageId="2" pageNumber="245">rd</superScript>
|
||
palpomere with three visible stout setae; 4
|
||
<superScript id="7C48F17F2C64FF80FBA67FB2FB88F839" attach="left" box="[1091,1104,1981,1995]" fontSize="6" pageId="2" pageNumber="245">th</superScript>
|
||
palpomere flattened, asetose, with a dilated and truncate apex; cardo and stipes each with two visible, long and stout setae (
|
||
<figureCitation id="130640B22C65FF81FB317898FAE5FF43" box="[1236,1341,150,177]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[257,1330,181,1843]" captionTargetId="figure-17@5.[257,1330,181,1843]" captionTargetPageId="5" captionText="FIGURES 5–7. Goriresina fungifora gen. n., sp. n., holotype, left lateral view: 5—head and pronotum; 6—labrum; 7—mouth parts. Abbreviations: lss—left scape seta; lcs—left clypeal seta; fts—left fore-trochanter seta; llb—left latero-basal seta; lla— left latero-apical seta; la—left apical seta; ra—right apical seta; rla—right latero-apical seta; lb—labrum; lmn—left mandible; mxp—maxillary palps; lbp—labial palps; cas—cardo setae; sts—stipes setae." figureDoi="http://doi.org/10.5281/zenodo.5222112" httpUri="https://zenodo.org/record/5222112/files/figure.png" pageId="3" pageNumber="246">Figs 5, 7</figureCitation>
|
||
). Labial palpus with distinctly visible palpomeres 2–4, 3
|
||
<superScript id="7C48F17F2C65FF81FD5578B5FD66FF3A" attach="left" box="[688,702,186,200]" fontSize="6" pageId="3" pageNumber="246">rd</superScript>
|
||
palpomere with at least 18 long stout setae along anterior margin and four setae (one apical, one medial and two basal) along posterior margin, 4
|
||
<superScript id="7C48F17F2C65FF81FBFF78D1FBFFFF1E" attach="left" box="[1050,1063,222,236]" fontSize="6" pageId="3" pageNumber="246">th</superScript>
|
||
palpomere asetose, elongate and drop-shaped (
|
||
<figureCitation id="130640B22C65FF81FED5790CFE4DFEEF" box="[304,405,258,285]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[257,1330,181,1843]" captionTargetId="figure-17@5.[257,1330,181,1843]" captionTargetPageId="5" captionText="FIGURES 5–7. Goriresina fungifora gen. n., sp. n., holotype, left lateral view: 5—head and pronotum; 6—labrum; 7—mouth parts. Abbreviations: lss—left scape seta; lcs—left clypeal seta; fts—left fore-trochanter seta; llb—left latero-basal seta; lla— left latero-apical seta; la—left apical seta; ra—right apical seta; rla—right latero-apical seta; lb—labrum; lmn—left mandible; mxp—maxillary palps; lbp—labial palps; cas—cardo setae; sts—stipes setae." figureDoi="http://doi.org/10.5281/zenodo.5222112" httpUri="https://zenodo.org/record/5222112/files/figure.png" pageId="3" pageNumber="246">Figs 5, 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B825C372C65FF81FF227928FC5FFE5F" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="246">
|
||
Pronotum with well-developed apical and basal lobes, deep anterior and posterior grooves and a smooth virtually globular disc (
|
||
<figureCitation id="130640B22C65FF81FEDA7944FE53FE97" box="[319,395,331,357]" captionStart="FIGURES 3–4" captionStartId="4.[151,264,1971,1995]" captionTargetBox="[178,1409,181,1948]" captionTargetId="figure-24@4.[177,1410,181,1948]" captionTargetPageId="4" captionText="FIGURES 3–4. Goriresina fungifora gen. n., sp. n., holotype: 3—left lateral view; 4—right dorso-lateral view." figureDoi="http://doi.org/10.5281/zenodo.5222110" httpUri="https://zenodo.org/record/5222110/files/figure.png" pageId="3" pageNumber="246">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="130640B22C65FF81FE797945FE73FE97" box="[412,427,330,357]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[257,1330,181,1843]" captionTargetId="figure-17@5.[257,1330,181,1843]" captionTargetPageId="5" captionText="FIGURES 5–7. Goriresina fungifora gen. n., sp. n., holotype, left lateral view: 5—head and pronotum; 6—labrum; 7—mouth parts. Abbreviations: lss—left scape seta; lcs—left clypeal seta; fts—left fore-trochanter seta; llb—left latero-basal seta; lla— left latero-apical seta; la—left apical seta; ra—right apical seta; rla—right latero-apical seta; lb—labrum; lmn—left mandible; mxp—maxillary palps; lbp—labial palps; cas—cardo setae; sts—stipes setae." figureDoi="http://doi.org/10.5281/zenodo.5222112" httpUri="https://zenodo.org/record/5222112/files/figure.png" pageId="3" pageNumber="246">5</figureCitation>
|
||
), anterior margin of apical lobe and posterior margin of basal lobe narrowly bordered (
|
||
<figureCitation id="130640B22C65FF81FF457960FF3FFE7B" box="[160,231,367,393]" captionStart="FIGURES 3–4" captionStartId="4.[151,264,1971,1995]" captionTargetBox="[178,1409,181,1948]" captionTargetId="figure-24@4.[177,1410,181,1948]" captionTargetPageId="4" captionText="FIGURES 3–4. Goriresina fungifora gen. n., sp. n., holotype: 3—left lateral view; 4—right dorso-lateral view." figureDoi="http://doi.org/10.5281/zenodo.5222110" httpUri="https://zenodo.org/record/5222110/files/figure.png" pageId="3" pageNumber="246">Fig. 4</figureCitation>
|
||
). Right metepisternum and metepimeron, as well as visible portions of abdominal sternites without setae; metepisternum approximately 2.8 times longer than wide (
|
||
<figureCitation id="130640B22C65FF81FCC2799CFCA2FE5F" box="[807,890,403,429]" captionStart="FIGURES 11–16" captionStartId="7.[151,264,1899,1923]" captionTargetBox="[211,1379,187,1874]" captionTargetId="figure-17@7.[209,1379,181,1875]" captionTargetPageId="7" captionText="FIGURES 11–16. Goriresina fungifora gen. n., sp. n., holotype: 11—elytra; 12—punctation and pubescence of shoulders area of right elytron; 13—lateral edge of right elytron; 14—right thoracic and abdominal segments; 15—legs; 16—middle left tarsus. Abbreviations: cv—costal vein of right wing; mte—metepisternum; as1—first abdominal sternite; hc—hind coxa; fr—fore right leg; mr—middle right leg; fl—fore left leg; ml—middle left leg; hl—hind left leg; I–V—middle tarsomeres; cl—claws." figureDoi="http://doi.org/10.5281/zenodo.5222118" httpUri="https://zenodo.org/record/5222118/files/figure.png" pageId="3" pageNumber="246">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B825C372C65FF81FF2279B8FCCBFD93" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="246">
|
||
Elytra elongate, converging to the apex, indistinctly dilated in basal third, moderately convex, with rounded obtuse-angulated shoulders, narrowly bordered from shoulders to suture; humeral area with dense punctuation and sparse thin setae, as well as a short groove between shoulder and an indistinct basal hump (
|
||
<figureCitation id="130640B22C65FF81FB9A79F0FB09FDEB" box="[1151,1233,511,537]" captionStart="FIGURES 11–16" captionStartId="7.[151,264,1899,1923]" captionTargetBox="[211,1379,187,1874]" captionTargetId="figure-17@7.[209,1379,181,1875]" captionTargetPageId="7" captionText="FIGURES 11–16. Goriresina fungifora gen. n., sp. n., holotype: 11—elytra; 12—punctation and pubescence of shoulders area of right elytron; 13—lateral edge of right elytron; 14—right thoracic and abdominal segments; 15—legs; 16—middle left tarsus. Abbreviations: cv—costal vein of right wing; mte—metepisternum; as1—first abdominal sternite; hc—hind coxa; fr—fore right leg; mr—middle right leg; fl—fore left leg; ml—middle left leg; hl—hind left leg; I–V—middle tarsomeres; cl—claws." figureDoi="http://doi.org/10.5281/zenodo.5222118" httpUri="https://zenodo.org/record/5222118/files/figure.png" pageId="3" pageNumber="246">Fig. 12</figureCitation>
|
||
); scutellum small; each elytron clearly narrowing in apical portion and angularly, but smoothly rounded to the suture; sutural spine small and sharp; epipleura narrow (
|
||
<figureCitation id="130640B22C65FF81FDC67A48FD50FD93" box="[547,648,583,609]" captionStart="FIGURES 3–4" captionStartId="4.[151,264,1971,1995]" captionTargetBox="[178,1409,181,1948]" captionTargetId="figure-24@4.[177,1410,181,1948]" captionTargetPageId="4" captionText="FIGURES 3–4. Goriresina fungifora gen. n., sp. n., holotype: 3—left lateral view; 4—right dorso-lateral view." figureDoi="http://doi.org/10.5281/zenodo.5222110" httpUri="https://zenodo.org/record/5222110/files/figure.png" pageId="3" pageNumber="246">Figs 3, 4</figureCitation>
|
||
,
|
||
<figureCitation id="130640B22C65FF81FD717A48FD03FD93" box="[660,731,583,609]" captionStart="FIGURES 11–16" captionStartId="7.[151,264,1899,1923]" captionTargetBox="[211,1379,187,1874]" captionTargetId="figure-17@7.[209,1379,181,1875]" captionTargetPageId="7" captionText="FIGURES 11–16. Goriresina fungifora gen. n., sp. n., holotype: 11—elytra; 12—punctation and pubescence of shoulders area of right elytron; 13—lateral edge of right elytron; 14—right thoracic and abdominal segments; 15—legs; 16—middle left tarsus. Abbreviations: cv—costal vein of right wing; mte—metepisternum; as1—first abdominal sternite; hc—hind coxa; fr—fore right leg; mr—middle right leg; fl—fore left leg; ml—middle left leg; hl—hind left leg; I–V—middle tarsomeres; cl—claws." figureDoi="http://doi.org/10.5281/zenodo.5222118" httpUri="https://zenodo.org/record/5222118/files/figure.png" pageId="3" pageNumber="246">11, 13</figureCitation>
|
||
,
|
||
<figureCitation id="130640B22C65FF81FD027A48FCDEFD93" box="[743,774,583,609]" captionStart="FIGURE 17" captionStartId="8.[151,250,1569,1593]" captionTargetBox="[568,1020,207,1528]" captionTargetId="figure-24@8.[548,1040,181,1545]" captionTargetPageId="8" captionText="FIGURE 17. Simplified reconstruction of the habitus of Goriresina fungifora gen. n., sp. n., holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5222120" httpUri="https://zenodo.org/record/5222120/files/figure.png" pageId="3" pageNumber="246">17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B825C372C65FF81FF227A64FB61FCE4" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="246">
|
||
Legs long and slender (
|
||
<figureCitation id="130640B22C65FF81FE327A64FDF3FD77" box="[471,555,618,645]" captionStart="FIGURES 11–16" captionStartId="7.[151,264,1899,1923]" captionTargetBox="[211,1379,187,1874]" captionTargetId="figure-17@7.[209,1379,181,1875]" captionTargetPageId="7" captionText="FIGURES 11–16. Goriresina fungifora gen. n., sp. n., holotype: 11—elytra; 12—punctation and pubescence of shoulders area of right elytron; 13—lateral edge of right elytron; 14—right thoracic and abdominal segments; 15—legs; 16—middle left tarsus. Abbreviations: cv—costal vein of right wing; mte—metepisternum; as1—first abdominal sternite; hc—hind coxa; fr—fore right leg; mr—middle right leg; fl—fore left leg; ml—middle left leg; hl—hind left leg; I–V—middle tarsomeres; cl—claws." figureDoi="http://doi.org/10.5281/zenodo.5222118" httpUri="https://zenodo.org/record/5222118/files/figure.png" pageId="3" pageNumber="246">Fig. 15</figureCitation>
|
||
), fore- and mesotrochanters each with a single long seta (
|
||
<figureCitation id="130640B22C65FF81FB2D7A64FAD7FD77" box="[1224,1295,618,645]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[257,1330,181,1843]" captionTargetId="figure-17@5.[257,1330,181,1843]" captionTargetPageId="5" captionText="FIGURES 5–7. Goriresina fungifora gen. n., sp. n., holotype, left lateral view: 5—head and pronotum; 6—labrum; 7—mouth parts. Abbreviations: lss—left scape seta; lcs—left clypeal seta; fts—left fore-trochanter seta; llb—left latero-basal seta; lla— left latero-apical seta; la—left apical seta; ra—right apical seta; rla—right latero-apical seta; lb—labrum; lmn—left mandible; mxp—maxillary palps; lbp—labial palps; cas—cardo setae; sts—stipes setae." figureDoi="http://doi.org/10.5281/zenodo.5222112" httpUri="https://zenodo.org/record/5222112/files/figure.png" pageId="3" pageNumber="246">Fig. 5</figureCitation>
|
||
); profemora 1.19 times longer than protibia, meso- and metafemora virtually as long as meso and metatibia, respectively; all tarsomeres with two parallel rows of short stout setae on bottom surface and two rows of thin setae on upper surface; 1
|
||
<superScript id="7C48F17F2C65FF81FEE17AD8FED7FD17" attach="left" box="[260,271,727,741]" fontSize="6" pageId="3" pageNumber="246">st</superScript>
|
||
tarsomere the longest; 4
|
||
<superScript id="7C48F17F2C65FF81FDC47AD9FDF6FD16" attach="left" box="[545,558,726,740]" fontSize="6" pageId="3" pageNumber="246">th</superScript>
|
||
tarsomere the shortest, with a group of long apical setae on bottom surface; 5
|
||
<superScript id="7C48F17F2C65FF81FA757AD9FA45FD16" attach="none" box="[1424,1437,726,740]" fontSize="6" pageId="3" pageNumber="246">th</superScript>
|
||
tarsomere inserted towards upper apical side of 4
|
||
<superScript id="7C48F17F2C65FF81FD587AF5FD12FCFA" attach="left" box="[701,714,762,776]" fontSize="6" pageId="3" pageNumber="246">th</superScript>
|
||
one; claws paired, equal in length (
|
||
<figureCitation id="130640B22C65FF81FBBD7AF4FB74FCE4" box="[1112,1196,763,790]" captionStart="FIGURES 11–16" captionStartId="7.[151,264,1899,1923]" captionTargetBox="[211,1379,187,1874]" captionTargetId="figure-17@7.[209,1379,181,1875]" captionTargetPageId="7" captionText="FIGURES 11–16. Goriresina fungifora gen. n., sp. n., holotype: 11—elytra; 12—punctation and pubescence of shoulders area of right elytron; 13—lateral edge of right elytron; 14—right thoracic and abdominal segments; 15—legs; 16—middle left tarsus. Abbreviations: cv—costal vein of right wing; mte—metepisternum; as1—first abdominal sternite; hc—hind coxa; fr—fore right leg; mr—middle right leg; fl—fore left leg; ml—middle left leg; hl—hind left leg; I–V—middle tarsomeres; cl—claws." figureDoi="http://doi.org/10.5281/zenodo.5222118" httpUri="https://zenodo.org/record/5222118/files/figure.png" pageId="3" pageNumber="246">Fig. 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B825C372C65FF81FF227B10FB9AFCCB" blockId="3.[151,1437,150,1977]" box="[199,1090,799,825]" pageId="3" pageNumber="246">Posterior margin of last abdominal sternite with at least 30 clearly visible setae.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3270FBC2C65FF81FF227B4CFC74FC73" pageId="3" pageNumber="246" type="etymology">
|
||
<paragraph id="8B825C372C65FF81FF227B4CFC74FC73" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="246">
|
||
<emphasis id="B94980252C65FF81FF227B4CFE97FCAF" bold="true" box="[199,335,835,861]" pageId="3" pageNumber="246">Etymology.</emphasis>
|
||
The name is composed of the first part from Ukrainian hydronym “
|
||
<emphasis id="B94980252C65FF81FBB67B4CFB7EFCAF" box="[1107,1190,835,861]" italics="true" pageId="3" pageNumber="246">ГОРинь</emphasis>
|
||
” (tributary of Pripyat River) as the main regional river and the name of the subtribe
|
||
<taxonomicName id="4C3D27B42C65FF81FCB57B68FC70FC73" authorityName="Rivalier" authorityYear="1971" box="[848,936,871,897]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="subTribe" subTribe="Iresiina">Iresiina</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3270FBC2C65FF84FF227B85FC48FE94" lastPageId="6" lastPageNumber="249" pageId="3" pageNumber="246" type="diagnosis">
|
||
<paragraph id="8B825C372C65FF81FF227B85FACDFB8F" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="246">
|
||
<emphasis id="B94980252C65FF81FF227B85FE9AFC56" bold="true" box="[199,322,906,932]" pageId="3" pageNumber="246">Diagnosis.</emphasis>
|
||
The glabrous head with two clypeal and four supra-orbital setae, the labrum with six submarginal setae, the glabrous and globular pronotum, the lack of setae from the metepisternum and metepimeron, as well as from the visible portions of abdominal sternites are combined the main reasons for considering the fossil as a new genus in the subtribe
|
||
<taxonomicName id="4C3D27B42C65FF81FE717BF8FD40FBE3" authority="Rivalier, 1971" authorityName="Rivalier" authorityYear="1971" box="[404,664,1015,1041]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="subTribe" subTribe="Iresiina">
|
||
Iresiina
|
||
<bibRefCitation id="EFAC21C62C65FF81FE177BF8FD40FBE3" author="Rivalier, E." box="[498,664,1015,1041]" pageId="3" pageNumber="246" pagination="135 - 143" refId="ref8314" refString="Rivalier, E. (1971) Remarques sur la Tribu des Cicindelini (Col. Cicindelidae) et sa subdivision en sous-tribus. Nouvelle Revue d'Entomologie, 1, 135 - 143." type="journal article" year="1971">Rivalier, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
. At present, nine extant (
|
||
<taxonomicName id="4C3D27B42C65FF81FC257BF8FB7DFBE3" authority="Dejean, 1831" authorityName="Dejean" authorityYear="1831" box="[960,1189,1015,1041]" class="Insecta" family="Carabidae" genus="Iresia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FC257BF8FBD8FBE2" box="[960,1024,1015,1040]" italics="true" pageId="3" pageNumber="246">Iresia</emphasis>
|
||
<bibRefCitation id="EFAC21C62C65FF81FBEF7BF8FB7DFBE3" author="Dejean, P. F. M. A." box="[1034,1189,1015,1041]" pageId="3" pageNumber="246" refId="ref5775" refString="Dejean, P. F. M. A. (1831) Species General des Coleopteres de la Collection de M. le Comte Dejean. Vol. 5. Crevot, Paris, 883 pp. [https: // www. biodiversitylibrary. org / item / 36755 # page / 7 / mode / 1 up]" type="book" year="1831">Dejean, 1831</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D27B42C65FF81FB517BF8FA4FFBE3" authority="Horn, 1901" authorityName="Horn" authorityYear="1901" box="[1204,1431,1015,1041]" class="Insecta" family="Carabidae" genus="Langea" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FB517BF8FAD0FBE2" box="[1204,1288,1015,1040]" italics="true" pageId="3" pageNumber="246">Langea</emphasis>
|
||
<bibRefCitation id="EFAC21C62C65FF81FAF77BF8FA4FFBE3" author="Horn, W." box="[1298,1431,1015,1041]" pageId="3" pageNumber="246" pagination="353 - 358" refId="ref6069" refString="Horn, W. (1901) De 7 novis Cicindelidis. Deutsche Entomologische Zeitschrift, 1901, 353 - 358. [https: // www. biodiversitylibrary. org / item / 103314 % 2523 page / 364 / mode / 1 up]" type="journal article" year="1901">Horn, 1901</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D27B42C65FF81FF727C14FE1BFBC7" authority="Dejean, 1825" authorityName="Dejean" authorityYear="1825" box="[151,451,1050,1077]" class="Insecta" family="Carabidae" genus="Euprosopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FF727C14FEC4FBC6" box="[151,284,1051,1076]" italics="true" pageId="3" pageNumber="246">Euprosopus</emphasis>
|
||
<bibRefCitation id="EFAC21C62C65FF81FEC27C14FE1BFBC7" author="Dejean, P. F. M. A." box="[295,451,1050,1077]" pageId="3" pageNumber="246" refId="ref5700" refString="Dejean, P. F. M. A. (1825) Species General des Coleopteres de la Collection de M. le Comte Dejean. Vol. 1. Crevot, Paris, 463 pp. [https: // www. biodiversitylibrary. org / item / 18355 # page / 11 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 8863" type="book" year="1825">Dejean, 1825</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D27B42C65FF81FE347C14FC88FBC7" authority="Guerin-Meneville, 1844" authorityName="Guerin-Meneville" authorityYear="1844" box="[465,848,1051,1077]" class="Insecta" family="Carabidae" genus="Eucallia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FE347C14FDE8FBC7" box="[465,560,1051,1077]" italics="true" pageId="3" pageNumber="246">Eucallia</emphasis>
|
||
<bibRefCitation id="EFAC21C62C65FF81FDDE7C14FC88FBC7" author="Guerin-Meneville, F. E." box="[571,848,1051,1077]" pageId="3" pageNumber="246" refId="ref5953" refString="Guerin-Meneville, F. E. (1844) Iconographie de regne animal de G. Cuvier, ou representation d'apres nature de l'une des especes les plus remarquables, et souvent non encore figurees, de chaque d'animaux. Vol. 3. Insectes. J. B. Bailliere, Paris, 576 pp." type="book" year="1844">Guérin-Méneville, 1844</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D27B42C65FF81FCBA7C14FB4EFBC7" authority="Bates 1891" authorityName="Bates" authorityYear="1891" box="[863,1174,1051,1077]" class="Insecta" family="Carabidae" genus="Rhytidophaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FCBA7C14FBD1FBC7" box="[863,1033,1051,1077]" italics="true" pageId="3" pageNumber="246">Rhytidophaena</emphasis>
|
||
<bibRefCitation id="EFAC21C62C65FF81FBF67C14FB4EFBC7" author="Bates, H. W." box="[1043,1174,1051,1077]" pageId="3" pageNumber="246" pagination="7 - 23" refId="ref5183" refString="Bates, H. W. (1891) Coleoptera from Kulu in N. W. India. The Entomologist, Supplement 24, 7 - 23." type="journal article" year="1891">Bates 1891</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D27B42C65FF81FB407C14FF0AFBAB" authority="Horn, 1929" authorityName="Horn" authorityYear="1929" class="Insecta" family="Carabidae" genus="Pronyssiformia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FB407C14FA89FBC7" box="[1189,1361,1051,1077]" italics="true" pageId="3" pageNumber="246">Pronyssiformia</emphasis>
|
||
<bibRefCitation id="EFAC21C62C65FF81FAB97C14FF0AFBAB" author="Horn, W." pageId="3" pageNumber="246" pagination="4 - 9" refId="ref6257" refString="Horn, W. (1929) Ueber einige orientalische und aethiopische Cicindelinae-Genera. Entomologische Nachrichtsblatt, 3 (1), 4 - 9." type="journal article" year="1929">Horn, 1929</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D27B42C65FF81FF047C30FDEFFBAB" authority="Westwood, 1837" authorityName="Westwood" authorityYear="1837" box="[225,567,1087,1113]" class="Insecta" family="Carabidae" genus="Distipsidera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FF047C30FEB3FBAB" box="[225,363,1087,1113]" italics="true" pageId="3" pageNumber="246">Distipsidera</emphasis>
|
||
<bibRefCitation id="EFAC21C62C65FF81FE907C30FDEFFBAB" author="Westwood, J. O." box="[373,567,1087,1113]" pageId="3" pageNumber="246" pagination="251 - 257" refId="ref8716" refString="Westwood, J. O. (1837) Descriptions of some new species of exotic coleopterous insects from the collection of Sir Patrick Walker. Magazine of Zoology and Botany, 1, 251 - 257. [https: // www. biodiversitylibrary. org / item / 123712 # page / 265 / mode / 1 up]" type="journal article" year="1837">Westwood, 1837</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D27B42C65FF81FDA37C30FC9BFBAB" authority="Horn, 1899" authorityName="Horn" authorityYear="1899" box="[582,835,1087,1113]" class="Insecta" family="Carabidae" genus="Nickerlea" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FDA37C30FD6BFBAB" box="[582,691,1087,1113]" italics="true" pageId="3" pageNumber="246">Nickerlea</emphasis>
|
||
<bibRefCitation id="EFAC21C62C65FF81FD587C30FC9BFBAB" author="Horn, W." box="[701,835,1087,1113]" pageId="3" pageNumber="246" pagination="135 - 136" refId="ref6010" refString="Horn, W. (1899) Nickerlea nov. gen. Euryodinorum. Deutsche Entomologische Zeitschrift, 1899, 135 - 136. [https: // www. biodiversitylibrary. org / item / 103426 # page / 141 / mode / 1 up] https: // doi. org / 10.1002 / mmnd. 48018990128" type="journal article" year="1899">Horn, 1899</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D27B42C65FF81FCB77C30FB59FBA8" authority="Sloane, 1906" authorityName="Sloane" authorityYear="1906" box="[850,1153,1087,1114]" class="Insecta" family="Carabidae" genus="Rhysopleura" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FCB77C30FC38FBAB" box="[850,992,1087,1113]" italics="true" pageId="3" pageNumber="246">Rhysopleura</emphasis>
|
||
<bibRefCitation id="EFAC21C62C65FF81FC0F7C30FB59FBA8" author="Sloane, T. G." box="[1002,1153,1087,1114]" pageId="3" pageNumber="246" pagination="309 - 360" refId="ref8528" refString="Sloane, T. G. (1906) Revision of the Cicindelidae of Australia. Proceedings of the Linnean Society of New South Wales, 31 (2), 309 - 360." type="journal article" year="1906">Sloane, 1906</bibRefCitation>
|
||
</taxonomicName>
|
||
) and two extinct genera (
|
||
<taxonomicName id="4C3D27B42C65FF81FF447C6CFE41FB8F" box="[161,409,1123,1149]" class="Insecta" family="Carabidae" genus="Palaeopronyssiformia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FF447C6CFE41FB8F" box="[161,409,1123,1149]" italics="true" pageId="3" pageNumber="246">Palaeopronyssiformia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3D27B42C65FF81FE2A7C6CFDB3FB8F" box="[463,619,1123,1149]" class="Insecta" family="Carabidae" genus="Palaeoiresina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FE2A7C6CFDB3FB8F" box="[463,619,1123,1149]" italics="true" pageId="3" pageNumber="246">Palaeoiresina</emphasis>
|
||
</taxonomicName>
|
||
) are considered as members of this subtribe (
|
||
<bibRefCitation id="EFAC21C62C65FF81FB827C6CFAD1FB8F" author="Wiesner, J." box="[1127,1289,1123,1149]" pageId="3" pageNumber="246" refId="ref8777" refString="Wiesner, J. (2020) Checklist of the Tiger Beetles of the World. 2 nd Edition. Borsdorf, Winterwork, 540 pp." type="book" year="2020">Wiesner 2020</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B825C372C65FF81FF227C88FA86FAC3" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="246">
|
||
Among them, by the shape of the labrum, by the elytral pattern and by the 5
|
||
<superScript id="7C48F17F2C65FF81FBF37C89FBFBFB66" attach="left" box="[1046,1059,1158,1172]" fontSize="6" pageId="3" pageNumber="246">th</superScript>
|
||
tarsomere being inserted towards the upper apical side of the 4
|
||
<superScript id="7C48F17F2C65FF81FE0F7CA5FE2FFB4A" attach="left" box="[490,503,1194,1208]" fontSize="6" pageId="3" pageNumber="246">th</superScript>
|
||
one, the new genus resembles some species of
|
||
<taxonomicName id="4C3D27B42C65FF81FBC47CA4FB73FB37" authorityName="Westwood" authorityYear="1837" box="[1057,1195,1195,1221]" class="Insecta" family="Carabidae" genus="Distipsidera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FBC47CA4FB73FB37" box="[1057,1195,1195,1221]" italics="true" pageId="3" pageNumber="246">Distipsidera</emphasis>
|
||
</taxonomicName>
|
||
. However, all extant
|
||
<taxonomicName id="4C3D27B42C65FF81FF727CC0FEA0FB1B" box="[151,376,1231,1257]" class="Insecta" family="Carabidae" genus="Distipsidera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94980252C65FF81FF727CC0FEF9FB1B" box="[151,289,1231,1257]" italics="true" pageId="3" pageNumber="246">Distipsidera</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
are characterised by a very broad preapical labial palpomere, by the apical setae starting directly from the anterior labial margin, by a deep horseshoe-shaped groove on the metepisternum, and by the punctateundulate elytral sculpture. Thus, these features make these two genera reliably distinguished from each other.
|
||
</paragraph>
|
||
<paragraph id="8B825C372C65FF81FF227D34FA57FA8B" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="246">
|
||
From
|
||
<taxonomicName id="4C3D27B42C65FF81FEE97D34FEA1FAA7" authorityName="Horn" authorityYear="1899" box="[268,377,1339,1365]" class="Insecta" family="Carabidae" genus="Nickerlea" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FEE97D34FEA1FAA7" box="[268,377,1339,1365]" italics="true" pageId="3" pageNumber="246">Nickerlea</emphasis>
|
||
</taxonomicName>
|
||
, the new genus differs by the thin preapical labial palpomere, the submarginal apical setae of the labrum, the 5
|
||
<superScript id="7C48F17F2C65FF81FEB07D51FEBAFA9E" attach="left" box="[341,354,1374,1388]" fontSize="6" pageId="3" pageNumber="246">th</superScript>
|
||
tarsomere inserted towards the upper apical side of the 4
|
||
<superScript id="7C48F17F2C65FF81FC047D51FC36FA9E" attach="left" box="[993,1006,1374,1388]" fontSize="6" pageId="3" pageNumber="246">th</superScript>
|
||
one, as well as by the elytral pattern.
|
||
</paragraph>
|
||
<paragraph id="8B825C372C65FF81FF227D8CFF35F9FB" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="246">
|
||
From
|
||
<taxonomicName id="4C3D27B42C65FF81FEEC7D8CFE4FFA6F" authorityName="Sloane" authorityYear="1906" box="[265,407,1411,1437]" class="Insecta" family="Carabidae" genus="Rhysopleura" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FEEC7D8CFE4FFA6F" box="[265,407,1411,1437]" italics="true" pageId="3" pageNumber="246">Rhysopleura</emphasis>
|
||
</taxonomicName>
|
||
, the new genus is easily distinct by the globular, not hump-shaped (in lateral view) pronotum, the very much shorter preapical labial palpomere, the apical setae starting in the submarginal area of the labrum, the smooth basal third of the elytra, the angularly rounded elytral apex with small sharp sutural spine, and the elytral pattern.
|
||
</paragraph>
|
||
<paragraph id="8B825C372C65FF81FF227E1CFE8CF9A3" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="246">
|
||
<taxonomicName id="4C3D27B42C65FF81FF227E1CFE94F9DE" authorityName="Dejean" authorityYear="1825" box="[199,332,1555,1580]" class="Insecta" family="Carabidae" genus="Euprosopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FF227E1CFE94F9DE" box="[199,332,1555,1580]" italics="true" pageId="3" pageNumber="246">Euprosopus</emphasis>
|
||
</taxonomicName>
|
||
clearly differs from the new genus by the very long, narrow and parallel-sided elytra, as well as the elytral sculpture.
|
||
</paragraph>
|
||
<paragraph id="8B825C372C65FF81FF227E54FF0CF96B" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="246">
|
||
<taxonomicName id="4C3D27B42C65FF81FF227E54FEC3F986" authorityName="Horn" authorityYear="1901" box="[199,283,1627,1652]" class="Insecta" family="Carabidae" genus="Langea" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FF227E54FEC3F986" box="[199,283,1627,1652]" italics="true" pageId="3" pageNumber="246">Langea</emphasis>
|
||
</taxonomicName>
|
||
, unlike the new genus, is characterised by the rough elytral sculpture and the smoothly rounded elytral apex.
|
||
</paragraph>
|
||
<paragraph id="8B825C372C65FF81FF227EACFB6DF913" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="246">
|
||
The monotypic
|
||
<taxonomicName id="4C3D27B42C65FF81FE937EACFE0DF94F" authorityName="Guerin-Meneville" authorityYear="1844" box="[374,469,1699,1725]" class="Insecta" family="Carabidae" genus="Eucallia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FE937EACFE0DF94F" box="[374,469,1699,1725]" italics="true" pageId="3" pageNumber="246">Eucallia</emphasis>
|
||
</taxonomicName>
|
||
is reliably distinguished by the very short labrum, the deep and rugose elytral sculpture and narrow apices of each elytron without sutural spine, both latter never convergent together.
|
||
</paragraph>
|
||
<paragraph id="8B825C372C65FF81FF227EE4FB43F8BF" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="246">
|
||
<taxonomicName id="4C3D27B42C65FF81FF227EE4FEDFF8F6" authorityName="Dejean" authorityYear="1831" box="[199,263,1771,1796]" class="Insecta" family="Carabidae" genus="Iresia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FF227EE4FEDFF8F6" box="[199,263,1771,1796]" italics="true" pageId="3" pageNumber="246">Iresia</emphasis>
|
||
</taxonomicName>
|
||
, in contrast to the new genus, shows a very long and thin apical joint, and the preapical joint of the labial palpomere slightly curved along the posterior margin, a stripe-shaped impression along the bottom margin of the metepisternum, as well as the elytra with punctate-undulate sculpture devoid of maculation.
|
||
</paragraph>
|
||
<paragraph id="8B825C372C65FF81FF227F58FCA7F84B" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="246">
|
||
Unlike the new genus, the monotypic
|
||
<taxonomicName id="4C3D27B42C65FF81FD937F58FCFAF883" authorityName="Horn" authorityYear="1929" box="[630,802,1879,1905]" class="Insecta" family="Carabidae" genus="Pronyssiformia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C65FF81FD937F58FCFAF883" box="[630,802,1879,1905]" italics="true" pageId="3" pageNumber="246">Pronyssiformia</emphasis>
|
||
</taxonomicName>
|
||
is distinct in the larger size and a robust habitus, by the presence of an irregular row of large setigerous pores along the suture, as well as in only two setae located at the posterior margin of each abdominal sternite including the last one.
|
||
</paragraph>
|
||
<caption id="DF420CBF2C62FF86FF727FBCFADDF83E" ID-DOI="http://doi.org/10.5281/zenodo.5222110" ID-Zenodo-Dep="5222110" box="[151,1285,1971,1996]" httpUri="https://zenodo.org/record/5222110/files/figure.png" pageId="4" pageNumber="247" startId="4.[151,264,1971,1995]" targetBox="[178,1409,181,1948]" targetPageId="4">
|
||
<paragraph id="8B825C372C62FF86FF727FBCFADDF83E" blockId="4.[151,1285,1971,1996]" box="[151,1285,1971,1996]" pageId="4" pageNumber="247">
|
||
<emphasis id="B94980252C62FF86FF727FBCFEE3F83E" bold="true" box="[151,315,1971,1996]" pageId="4" pageNumber="247">FIGURES 3–4.</emphasis>
|
||
<taxonomicName id="4C3D27B42C62FF86FEA47FBBFDCBF83E" authority="Matalin & Perkovsky & Vasilenko, 2021" authorityName="Matalin & Perkovsky & Vasilenko" authorityYear="2021" box="[321,531,1972,1996]" class="Insecta" family="Carabidae" genus="Goriresina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="fungifora" status="gen. n., sp. n.">
|
||
<emphasis id="B94980252C62FF86FEA47FBBFDCBF83E" box="[321,531,1972,1996]" italics="true" pageId="4" pageNumber="247">Goriresina fungifora</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94980252C62FF86FDFF7FBBFD7FF83E" bold="true" box="[538,679,1972,1996]" pageId="4" pageNumber="247">
|
||
<taxonomicNameLabel id="A27A3D5E2C62FF86FDFF7FBBFD7FF83E" box="[538,679,1972,1996]" pageId="4" pageNumber="247" rank="species">gen. n., sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype: 3—left lateral view; 4—right dorso-lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF420CBF2C63FF87FF727F44FC70F83C" ID-DOI="http://doi.org/10.5281/zenodo.5222112" ID-Zenodo-Dep="5222112" httpUri="https://zenodo.org/record/5222112/files/figure.png" pageId="5" pageNumber="248" startId="5.[151,264,1867,1891]" targetBox="[257,1330,181,1843]" targetPageId="5">
|
||
<paragraph id="8B825C372C63FF87FF727F44FC70F83C" blockId="5.[151,1437,1867,1999]" pageId="5" pageNumber="248">
|
||
<emphasis id="B94980252C63FF87FF727F44FEE3F891" bold="true" box="[151,315,1867,1891]" pageId="5" pageNumber="248">FIGURES 5–7.</emphasis>
|
||
<taxonomicName id="4C3D27B42C63FF87FEA47F44FDCBF891" authority="Matalin & Perkovsky & Vasilenko, 2021" authorityName="Matalin & Perkovsky & Vasilenko" authorityYear="2021" box="[321,531,1867,1891]" class="Insecta" family="Carabidae" genus="Goriresina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="248" phylum="Arthropoda" rank="species" species="fungifora" status="gen. n., sp. n.">
|
||
<emphasis id="B94980252C63FF87FEA47F44FDCBF891" box="[321,531,1867,1891]" italics="true" pageId="5" pageNumber="248">Goriresina fungifora</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94980252C63FF87FDFF7F44FD7EF891" bold="true" box="[538,678,1867,1891]" pageId="5" pageNumber="248">
|
||
<taxonomicNameLabel id="A27A3D5E2C63FF87FDFF7F44FD7EF891" box="[538,678,1867,1891]" pageId="5" pageNumber="248" rank="species">gen. n., sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, left lateral view: 5—head and pronotum; 6—labrum; 7—mouth parts. Abbreviations:
|
||
<emphasis id="B94980252C63FF87FE977F60FE55F875" box="[370,397,1903,1927]" italics="true" pageId="5" pageNumber="248">lss</emphasis>
|
||
—left scape seta;
|
||
<emphasis id="B94980252C63FF87FDA07F60FDB9F875" box="[581,609,1903,1927]" italics="true" pageId="5" pageNumber="248">lcs</emphasis>
|
||
—left clypeal seta;
|
||
<emphasis id="B94980252C63FF87FCCC7F60FC99F875" box="[809,833,1903,1927]" italics="true" pageId="5" pageNumber="248">fts</emphasis>
|
||
—left fore-trochanter seta;
|
||
<emphasis id="B94980252C63FF87FBBD7F60FBABF875" box="[1112,1139,1903,1927]" italics="true" pageId="5" pageNumber="248">llb</emphasis>
|
||
—left latero-basal seta;
|
||
<emphasis id="B94980252C63FF87FA8D7F60FA5BF875" box="[1384,1411,1903,1927]" italics="true" pageId="5" pageNumber="248">lla</emphasis>
|
||
— left latero-apical seta;
|
||
<emphasis id="B94980252C63FF87FE9F7F9CFE56F859" box="[378,398,1939,1963]" italics="true" pageId="5" pageNumber="248">la</emphasis>
|
||
—left apical seta;
|
||
<emphasis id="B94980252C63FF87FDA27F9BFD86F859" box="[583,606,1940,1963]" italics="true" pageId="5" pageNumber="248">ra</emphasis>
|
||
—right apical seta;
|
||
<emphasis id="B94980252C63FF87FCC07F9CFC9BF859" box="[805,835,1939,1963]" italics="true" pageId="5" pageNumber="248">rla</emphasis>
|
||
—right latero-apical seta;
|
||
<emphasis id="B94980252C63FF87FBA87F9CFBB9F859" box="[1101,1121,1939,1963]" italics="true" pageId="5" pageNumber="248">lb</emphasis>
|
||
—labrum;
|
||
<emphasis id="B94980252C63FF87FB2A7F9CFB2DF859" box="[1231,1269,1939,1963]" italics="true" pageId="5" pageNumber="248">lmn</emphasis>
|
||
—left mandible;
|
||
<emphasis id="B94980252C63FF87FF727FB7FF19F83D" box="[151,193,1976,1999]" italics="true" pageId="5" pageNumber="248">mxp</emphasis>
|
||
—maxillary palps;
|
||
<emphasis id="B94980252C63FF87FE607FB8FE7DF83D" box="[389,421,1975,1999]" italics="true" pageId="5" pageNumber="248">lbp</emphasis>
|
||
—labial palps;
|
||
<emphasis id="B94980252C63FF87FDA57FB7FDBAF83D" box="[576,610,1976,1999]" italics="true" pageId="5" pageNumber="248">cas</emphasis>
|
||
—cardo setae;
|
||
<emphasis id="B94980252C63FF87FD1F7FB7FCCDF83D" box="[762,789,1976,1999]" italics="true" pageId="5" pageNumber="248">sts</emphasis>
|
||
—stipes setae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B825C372C60FF84FF227898FE1EFF0B" blockId="6.[151,1437,151,358]" pageId="6" pageNumber="249">
|
||
<taxonomicName id="4C3D27B42C60FF84FF227898FEA9FF43" authorityName="Bates" authorityYear="1891" box="[199,369,151,177]" class="Insecta" family="Carabidae" genus="Rhytidophaena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="249" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C60FF84FF227898FEA9FF43" box="[199,369,151,177]" italics="true" pageId="6" pageNumber="249">Rhytidophaena</emphasis>
|
||
</taxonomicName>
|
||
differs well from the new genus by the shorter labrum, short apical labial palpomere, asetose clypeus, flattened nonglobular pronotum, numerous stout setae on the femora and the hairless posterior margin of the last abdominal sternite.
|
||
</paragraph>
|
||
<paragraph id="8B825C372C60FF84FF22790CFC48FE94" blockId="6.[151,1437,151,358]" pageId="6" pageNumber="249">
|
||
From both fossil genera,
|
||
<taxonomicName id="4C3D27B42C60FF84FE3B790CFD0EFEEF" box="[478,726,259,285]" class="Insecta" family="Carabidae" genus="Palaeopronyssiformia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="249" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C60FF84FE3B790CFD0EFEEF" box="[478,726,259,285]" italics="true" pageId="6" pageNumber="249">Palaeopronyssiformia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3D27B42C60FF84FCE9790CFC70FEEF" box="[780,936,259,285]" class="Insecta" family="Carabidae" genus="Palaeoiresina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="249" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94980252C60FF84FCE9790CFC70FEEF" box="[780,936,259,285]" italics="true" pageId="6" pageNumber="249">Palaeoiresina</emphasis>
|
||
</taxonomicName>
|
||
, the new genus is distinguished by the shape of a bidentate labrum, the apicad converging elytra and the shape of elytral apex with a sharp sutural spine (for more details, see the description and diagnosis of the new species below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |