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<document id="95C835579237FF94F45B11F439814AF0" ID-DOI="10.5281/zenodo.178539" ID-GBIF-Dataset="dda53384-9974-4b83-b780-a9cd51fffacf" ID-ISSN="1175-5326" ID-Zenodo-Dep="178539" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459899930198" checkinUser="plazi" docAuthor="Freitag, Hendrik &amp; Jäch, Manfred A." docDate="2007" docId="03939707FFB52548FF2CFD52E510F81B" docLanguage="en" docName="zt01590p059.pdf" docOrigin="Zootaxa 1590" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Ancyronyx pseudopatrolus Freitag &amp; Jach, sp. n." docType="treatment" docVersion="10" lastPageNumber="46" masterDocId="FFAAEF7FFFBC2541FFBBFFBDE027FF97" masterDocTitle="The genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae) in Palawan and Busuanga, (Philippines) with descriptions of six new species" masterLastPageNumber="59" masterPageNumber="37" pageNumber="46" updateTime="1698228715028" updateUser="plazi">
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<mods:titleInfo id="4FF7DA8DAFB5E1B9564E86853E94526A">
<mods:title id="D98F6313E82594E05822077A8FA417CF">The genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae) in Palawan and Busuanga, (Philippines) with descriptions of six new species</mods:title>
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<mods:namePart id="C649106FAF2C25ED25F7F2A3311B82FE">Freitag, Hendrik</mods:namePart>
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<mods:namePart id="5C28707C967834369DF877A6BE397656">Jäch, Manfred A.</mods:namePart>
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<mods:typeOfResource id="7396B91086C638D24BC711477C571F4B">text</mods:typeOfResource>
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<mods:title id="B96ADB26F6D1B9CBC730F5F67504D2BA">Zootaxa</mods:title>
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<mods:date id="C2D7A5575FB2CEBC648056B538271116">2007</mods:date>
<mods:detail id="BA70D8FDCD13DFD143DA2230CFFC4A8B" type="volume">
<mods:number id="D65B6FD3811DBB89804CA485C256935E">1590</mods:number>
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<mods:classification id="6228FA3457CA5738100FB1C347E95C07">journal article</mods:classification>
<mods:identifier id="9BC3346A92D14C010FFED4B79D89DF6C" type="DOI">10.5281/zenodo.178539</mods:identifier>
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<mods:identifier id="020B69085BD91355020DF765E1BA87F4" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03939707FFB52548FF2CFD52E510F81B" ID-DOI="http://doi.org/10.5281/zenodo.6243451" ID-GBIF-Taxon="119372906" ID-Zenodo-Dep="6243451" LSID="urn:lsid:zoobank.org:act:86C5107A-A81E-4D58-95FB-88AA498FF81C" httpUri="http://treatment.plazi.org/id/03939707FFB52548FF2CFD52E510F81B" lastPageNumber="46" pageId="9" pageNumber="46">
<subSubSection id="C320759AFFB52548FF2CFD52E177FCBB" pageId="9" pageNumber="46" type="nomenclature">
<paragraph id="8B852611FFB52548FF2CFD52E2C0FC9E" blockId="9.[151,743,751,812]" box="[151,743,751,777]" pageId="9" pageNumber="46">
<heading id="D0CD917DFFB52548FF2CFD52E2C0FC9E" bold="true" box="[151,743,751,777]" fontSize="11" level="1" pageId="9" pageNumber="46" reason="1">
<emphasis id="B94EFA03FFB52548FF2CFD52E2C0FC9E" bold="true" box="[151,743,751,777]" pageId="9" pageNumber="46">
<taxonomicName id="4C3A5D92FFB52548FF2CFD52E2B9FC9E" ID-CoL="DPSX" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[151,670,751,777]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="pseudopatrolus" status="sp. nov.">
<emphasis id="B94EFA03FFB52548FF2CFD52E1FDFC9E" bold="true" box="[151,474,751,777]" italics="true" pageId="9" pageNumber="46">Ancyronyx pseudopatrolus</emphasis>
Freitag &amp; Jäch
</taxonomicName>
<taxonomicNameLabel id="A27D4778FFB52548FD1EFD52E2C0FC9E" box="[677,743,751,777]" pageId="9" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B852611FFB52548FF2CFCAFE177FCBB" blockId="9.[151,743,751,812]" box="[151,336,786,812]" pageId="9" pageNumber="46">
(
<figureCitation id="13013A94FFB52548FF24FCAFE0D5FCBB" box="[159,242,786,812]" captionStart="FIGURES 3 – 6" captionStartId="5.[151,269,1843,1867]" captionTargetBox="[153,1436,208,1819]" captionTargetId="figure@5.[151,1436,194,1819]" captionTargetPageId="5" captionText="FIGURES 3 – 6. Habitus of 3) Ancyronyx patrolus sp. n., paratype ♂, usual colouration; 4) A. patrolus sp. n., paratype Ψ, dark coloured individual, partly completed; 5) A. pseudopatrolus sp. n., holotype ♂, partly completed; 6) A. punkti sp. n., paratype ♂, partly completed." httpUri="https://zenodo.org/record/178542/files/figure.png" pageId="9" pageNumber="46">Figs. 5</figureCitation>
,
<figureCitation id="13013A94FFB52548FF44FCAFE13CFCBB" box="[255,283,786,812]" captionStart="FIGURE 12" captionStartId="10.[151,255,1319,1343]" captionTargetBox="[162,1427,157,1263]" captionTargetId="figure@10.[162,1427,157,1265]" captionTargetPageId="10" captionText="FIGURE 12. Ancyronyx pseudopatrolus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male ventrite 5 (left half)." httpUri="https://zenodo.org/record/178545/files/figure.png" pageId="9" pageNumber="46">12</figureCitation>
a–d)
</paragraph>
</subSubSection>
<subSubSection id="C320759AFFB52548FF2CFCDFE1BEF8AB" pageId="9" pageNumber="46" type="materials_examined">
<paragraph id="8B852611FFB52548FF2CFCDFE104FC33" blockId="9.[151,1437,866,1932]" pageId="9" pageNumber="46">
<emphasis id="B94EFA03FFB52548FF2CFCDFE119FCEB" bold="true" box="[151,318,866,892]" pageId="9" pageNumber="46">
<typeStatus id="548198B3FFB52548FF2CFCDFE0F2FCEB" box="[151,213,866,892]" pageId="9" pageNumber="46">Type</typeStatus>
locality.
</emphasis>
Panaguman River, St. Paul National Park, Marofinas District, Puerto Princesa City, Palawan,
<collectingCountry id="F32D6681FFB52548FF2CFC37E138FC33" box="[151,287,906,932]" name="Philippines" pageId="9" pageNumber="46">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8B852611FFB52548FF7CFC0FE499FB8B" blockId="9.[151,1437,866,1932]" pageId="9" pageNumber="46">
<emphasis id="B94EFA03FFB52548FF7CFC0FE22AFC5B" bold="true" box="[199,525,946,972]" pageId="9" pageNumber="46">
<typeStatus id="548198B3FFB52548FF7CFC0FE125FC5B" box="[199,258,946,972]" pageId="9" pageNumber="46">Type</typeStatus>
material.
<typeStatus id="548198B3FFB52548FE20FC0FE22AFC5B" box="[411,525,946,972]" pageId="9" pageNumber="46" type="holotype">Holotype</typeStatus>
</emphasis>
ď (
<collectionCode id="ED2BBED4FFB52548FDF5FC0FE286FC5B" box="[590,673,946,972]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="9" pageNumber="46">NMW</collectionCode>
): “
<collectionCode id="ED2BBED4FFB52548FD71FC0FE332FC5B" box="[714,789,946,972]" pageId="9" pageNumber="46">PHIL</collectionCode>
.: Palawan, P. Princesa Panaguman R.
<geoCoordinate id="EE0E40D6FFB52548FAA8FC0FE5BBFC5B" box="[1299,1436,946,972]" direction="north" orientation="latitude" pageId="9" pageNumber="46" precision="15" value="10.2525">10°15'09''N</geoCoordinate>
<geoCoordinate id="EE0E40D6FFB52548FF2CFC67E10DFC63" box="[151,298,986,1012]" direction="east" orientation="longitude" pageId="9" pageNumber="46" precision="15" value="118.9675">
118°58'03''
<geoCoordinate id="EE0E40D6FFB52548FEADFC67E10DFC63" box="[278,298,986,1012]" direction="east" orientation="longitude" pageId="9" pageNumber="46" precision="5555" value="17.5">E</geoCoordinate>
</geoCoordinate>
<date id="FF8400D1FFB52548FE8AFC67E18EFC63" box="[305,425,986,1012]" pageId="9" pageNumber="46" value="2001-05-17">17.5.2001</date>
, leg. Freitag (PR1)D”, dissected (terminal parts of abdomen, aedeagus, one labial palpus, and parts of legs glued separately), two legs missing, other legs broken or incomplete.
</paragraph>
<paragraph id="8B852611FFB52548FF7CFB97E5BBFB2B" blockId="9.[151,1437,866,1932]" pageId="9" pageNumber="46">
<emphasis id="B94EFA03FFB52548FF7CFB97E1FAFBD3" bold="true" box="[199,477,1066,1092]" pageId="9" pageNumber="46">Differential diagnosis.</emphasis>
In general habitus and colour (
<figureCitation id="13013A94FFB52548FCE5FB97E380FBD3" box="[862,935,1066,1092]" captionStart="FIGURES 3 – 6" captionStartId="5.[151,269,1843,1867]" captionTargetBox="[153,1436,208,1819]" captionTargetId="figure@5.[151,1436,194,1819]" captionTargetPageId="5" captionText="FIGURES 3 – 6. Habitus of 3) Ancyronyx patrolus sp. n., paratype ♂, usual colouration; 4) A. patrolus sp. n., paratype Ψ, dark coloured individual, partly completed; 5) A. pseudopatrolus sp. n., holotype ♂, partly completed; 6) A. punkti sp. n., paratype ♂, partly completed." httpUri="https://zenodo.org/record/178542/files/figure.png" pageId="9" pageNumber="46">Fig. 5</figureCitation>
) this species conspicuously resembles
<taxonomicName id="4C3A5D92FFB52548FA38FB97E0DFFBFB" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="patrolus">
<emphasis id="B94EFA03FFB52548FA38FB97E0DFFBFB" italics="true" pageId="9" pageNumber="46">A. patrolus</emphasis>
</taxonomicName>
. It can be distinguished from the latter by the colour pattern of the legs and the stout aedeagus. In size and proportions, this specimen (CL:
<quantity id="4CC28BF4FFB52548FDFAFBC7E28BFB03" box="[577,684,1146,1172]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="9" pageNumber="46" unit="mm" value="1.42">1.42 mm</quantity>
, HW:
<quantity id="4CC28BF4FFB52548FD4DFBC7E347FB03" box="[758,864,1146,1172]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="9" pageNumber="46" unit="mm" value="0.33">0.33 mm</quantity>
, ID:
<quantity id="4CC28BF4FFB52548FC22FBC7E423FB03" box="[921,1028,1146,1172]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="46" unit="mm" value="0.21">0.21 mm</quantity>
, PL:
<quantity id="4CC28BF4FFB52548FBFAFBC7E48CFB03" box="[1089,1195,1146,1172]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="9" pageNumber="46" unit="mm" value="0.41">0.41 mm</quantity>
, MW:
<quantity id="4CC28BF4FFB52548FB41FBC7E542FB03" box="[1274,1381,1146,1172]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="46" unit="mm" value="0.45">0.45 mm</quantity>
, EL:
<quantity id="4CC28BF4FFB52548FF2CFB1FE127FB2B" box="[151,256,1186,1212]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="9" pageNumber="46" unit="mm" value="0.96">0.96 mm</quantity>
) agrees quite well with small specimens of
<taxonomicName id="4C3A5D92FFB52548FD46FB1EE358FB2B" box="[765,895,1187,1212]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="patrolus">
<emphasis id="B94EFA03FFB52548FD46FB1EE358FB2B" box="[765,895,1187,1212]" italics="true" pageId="9" pageNumber="46">A. patrolus</emphasis>
</taxonomicName>
, except for a slightly smaller EW of
<quantity id="4CC28BF4FFB52548FA96FB1FE5BFFB2B" box="[1325,1432,1186,1212]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="9" pageNumber="46" unit="mm" value="0.59">0.59 mm</quantity>
.
</paragraph>
<paragraph id="8B852611FFB52548FF7CFB77E5ADFA9B" blockId="9.[151,1437,866,1932]" pageId="9" pageNumber="46">
Ventral side dark brown, darker than in
<taxonomicName id="4C3A5D92FFB52548FD11FB76E309FB73" box="[682,814,1227,1252]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="patrolus">
<emphasis id="B94EFA03FFB52548FD11FB76E309FB73" box="[682,814,1227,1252]" italics="true" pageId="9" pageNumber="46">A. patrolus</emphasis>
</taxonomicName>
(except for dark coloured individuals of the latter); legs brown to dark brown, distal half of tibia (except tip) yellowish; antennae pale brown, distally darkened.
</paragraph>
<paragraph id="8B852611FFB52548FF7CFAA7E19CFACB" blockId="9.[151,1437,866,1932]" pageId="9" pageNumber="46">Ventral (inner) side of femora with conspicuous setae. Claw with two teeth, distal one much larger than inconspicuous basal one.</paragraph>
<paragraph id="8B852611FFB52548FF7CFAD7E150FA3B" blockId="9.[151,1437,866,1932]" pageId="9" pageNumber="46">
Prosternum punctate to granulose; punctures of metaventrite and ventrites 1–5 more deeply impressed than in
<taxonomicName id="4C3A5D92FFB52548FF54FA2EE148FA3B" box="[239,367,1427,1452]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="patrolus">
<emphasis id="B94EFA03FFB52548FF54FA2EE148FA3B" box="[239,367,1427,1452]" italics="true" pageId="9" pageNumber="46">A. patrolus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B852611FFB52548FF7DFA07E298FA6B" blockId="9.[151,1437,866,1932]" pageId="9" pageNumber="46">
Sternite IX (
<figureCitation id="13013A94FFB52548FEDCFA07E1E4FA43" box="[359,451,1466,1492]" captionStart="FIGURE 12" captionStartId="10.[151,255,1319,1343]" captionTargetBox="[162,1427,157,1263]" captionTargetId="figure@10.[162,1427,157,1265]" captionTargetPageId="10" captionText="FIGURE 12. Ancyronyx pseudopatrolus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male ventrite 5 (left half)." httpUri="https://zenodo.org/record/178545/files/figure.png" pageId="9" pageNumber="46">Fig. 12</figureCitation>
c) apically slightly rounded, not emarginate, lateroapically without setae, with micropores; paraprocts reaching apical margin.
</paragraph>
<paragraph id="8B852611FFB52548FF7CF9B7E380F97B" blockId="9.[151,1437,866,1932]" pageId="9" pageNumber="46">
Aedeagus (
<figureCitation id="13013A94FFB52548FEF0F9B7E1B8F9B3" box="[331,415,1546,1572]" captionStart="FIGURE 12" captionStartId="10.[151,255,1319,1343]" captionTargetBox="[162,1427,157,1263]" captionTargetId="figure@10.[162,1427,157,1265]" captionTargetPageId="10" captionText="FIGURE 12. Ancyronyx pseudopatrolus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male ventrite 5 (left half)." httpUri="https://zenodo.org/record/178545/files/figure.png" pageId="9" pageNumber="46">Fig. 12</figureCitation>
a, b) ca. 350 µm long. Median lobe broader and stouter than in
<taxonomicName id="4C3A5D92FFB52548FB3EF9B6E520F9B3" box="[1157,1287,1547,1572]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="patrolus">
<emphasis id="B94EFA03FFB52548FB3EF9B6E520F9B3" box="[1157,1287,1547,1572]" italics="true" pageId="9" pageNumber="46">A. patrolus</emphasis>
</taxonomicName>
; subapically distinctly widened (ca. 110 µm broad), with numerous distinct microtube-like structures ending sublateroapically; ventral sac plicate; fibula weekly sclerotized, inconspicuous; corona well developed. Phallobase subsymmetrical, basally somewhat tapered and with conspicuous, strongly sclerotized margin; basolateral (penile) apophyses short. Parameres subtriangular (lateral view), reaching about basal 0.65 of aedeagus; basal margin slightly emarginate (lateral view), not contiguous ventrally.
</paragraph>
<paragraph id="8B852611FFB52548FF7CF947E3C7F883" blockId="9.[151,1437,866,1932]" box="[199,992,1786,1812]" pageId="9" pageNumber="46">
Male ventrite 5 (
<figureCitation id="13013A94FFB52548FE31F947E1CEF883" box="[394,489,1786,1812]" captionStart="FIGURE 12" captionStartId="10.[151,255,1319,1343]" captionTargetBox="[162,1427,157,1263]" captionTargetId="figure@10.[162,1427,157,1265]" captionTargetPageId="10" captionText="FIGURE 12. Ancyronyx pseudopatrolus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male ventrite 5 (left half)." httpUri="https://zenodo.org/record/178545/files/figure.png" pageId="9" pageNumber="46">Figs. 12</figureCitation>
d) subtrapezoidal, apically microreticulate.
</paragraph>
<paragraph id="8B852611FFB52548FF7CF89FE1BEF8AB" blockId="9.[151,1437,866,1932]" box="[199,409,1826,1852]" pageId="9" pageNumber="46">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C320759AFFB52548FF7CF8F7E35FF8F3" box="[199,888,1866,1892]" pageId="9" pageNumber="46" type="distribution">
<paragraph id="8B852611FFB52548FF7CF8F7E35FF8F3" blockId="9.[151,1437,866,1932]" box="[199,888,1866,1892]" pageId="9" pageNumber="46">
<emphasis id="B94EFA03FFB52548FF7CF8F7E147F8F3" bold="true" box="[199,352,1866,1892]" pageId="9" pageNumber="46">Distribution</emphasis>
(Fig. 17). Only known from the
<typeStatus id="548198B3FFB52548FD5AF8F7E334F8F3" box="[737,787,1866,1892]" pageId="9" pageNumber="46">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C320759AFFB52548FF7CF8CFE510F81B" box="[199,1335,1906,1932]" pageId="9" pageNumber="46" type="etymology">
<paragraph id="8B852611FFB52548FF7CF8CFE510F81B" blockId="9.[151,1437,866,1932]" box="[199,1335,1906,1932]" pageId="9" pageNumber="46">
<emphasis id="B94EFA03FFB52548FF7CF8CFE173F81B" bold="true" box="[199,340,1906,1932]" pageId="9" pageNumber="46">Etymology.</emphasis>
The species is named in reference to its close relationships with
<taxonomicName id="4C3A5D92FFB52548FBF7F8CEE517F81B" box="[1100,1328,1907,1932]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="patrolus">
<emphasis id="B94EFA03FFB52548FBF7F8CEE517F81B" box="[1100,1328,1907,1932]" italics="true" pageId="9" pageNumber="46">Ancyronyx patrolus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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