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<document id="94ED78714EB1B81116239E5E601047C9" ID-CLB-Dataset="8781" ID-DOI="10.11646/zootaxa.4908.3.3" ID-GBIF-Dataset="de928cf2-80ea-4ede-ad12-3ac0300b1ae9" ID-ISSN="1175-5326" ID-Zenodo-Dep="4441906" ID-ZooBank="1A58171C-5BD8-4837-8708-3A1C7832DA8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610701278547" checkinUser="plazi" docAuthor="Bahder, Brian W., Zumbado Echavarria, Marco A., Barrantes Barrantes, Edwin A., Helmick, Ericka E. &amp; Bartlett, Charles R." docDate="2021" docId="0393112EFFEBFFF120A0BED93AD23690" docLanguage="en" docName="zootaxa.4908.3.3.pdf" docOrigin="Zootaxa 4908 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tico emmettcarri Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett 2021, sp. n." docType="treatment" docVersion="7" lastPageNumber="379" masterDocId="FFAA6956FFEEFFFB2037B82E3B083179" masterDocTitle="A new genus and two new species of planthopper in the tribe Cenchreini (Hemiptera: Auchenorrhyncha: Derbidae) from lowland tropical rainforest in Costa Rica" masterLastPageNumber="392" masterPageNumber="369" pageNumber="374" updateTime="1698924501363" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="587057CA6F22021EAE4B923251A7716F">A new genus and two new species of planthopper in the tribe Cenchreini (Hemiptera: Auchenorrhyncha: Derbidae) from lowland tropical rainforest in Costa Rica</mods:title>
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<mods:namePart id="470559AF5C83D6DA2CB409E435F121D6">Bahder, Brian W.</mods:namePart>
<mods:affiliation id="9EA638BFF0F021B13745E60ADE6B442A">University of Florida, Department of Entomology and Nematology-Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA.</mods:affiliation>
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<mods:namePart id="ABDAE7E7AC86F67F105F310996F8B533">Zumbado Echavarria, Marco A.</mods:namePart>
<mods:affiliation id="F1C54FFB0F8EF90629F4337C9130D5FC">Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica.</mods:affiliation>
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<mods:namePart id="78CB8452999A0AC491696E549F41D7B3">Barrantes Barrantes, Edwin A.</mods:namePart>
<mods:affiliation id="5FCD53AE1128A00BBB5D77481AFE3FD1">Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica.</mods:affiliation>
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<mods:namePart id="A0C18E7FC19AFE0019C9DF5DCE2A155A">Helmick, Ericka E.</mods:namePart>
<mods:affiliation id="2612782BA4B4E23E3B556E61A4B9666F">University of Florida, Department of Entomology and Nematology-Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA.</mods:affiliation>
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<mods:namePart id="A734107A2681E1C92EDA86C7E7D5FDE0">Bartlett, Charles R.</mods:namePart>
<mods:affiliation id="CB90F3A384738710AFD2C10C5320BA7B">University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716 - 2160, USA.</mods:affiliation>
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<treatment id="0393112EFFEBFFF120A0BED93AD23690" ID-DOI="http://doi.org/10.5281/zenodo.4450853" ID-GBIF-Taxon="177009917" ID-Zenodo-Dep="4450853" LSID="urn:lsid:plazi:treatment:0393112EFFEBFFF120A0BED93AD23690" httpUri="http://treatment.plazi.org/id/0393112EFFEBFFF120A0BED93AD23690" lastPageId="10" lastPageNumber="379" pageId="5" pageNumber="374">
<subSubSection id="C320F3B3FFEBFFFE20A0BED939943668" box="[151,668,1782,1809]" pageId="5" pageNumber="374" type="nomenclature">
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<heading id="D0CD1754FFEBFFFE20A0BED939943668" bold="true" box="[151,668,1782,1809]" fontSize="11" level="1" pageId="5" pageNumber="374" reason="1">
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<taxonomicName id="4C3ADBBBFFEBFFFE20A0BED9395B3668" ID-CoL="9FNPB" authority="Bahder &amp; Bartlett" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[151,595,1782,1809]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="emmettcarri" status="sp. nov.">
<emphasis id="B94E7C2AFFEBFFFE20A0BED93A6E3668" bold="true" box="[151,358,1783,1809]" italics="true" pageId="5" pageNumber="374">Tico emmettcarri</emphasis>
Bahder &amp; Bartlett
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<taxonomicNameLabel id="A27DC151FFEBFFFE226DBED939943668" box="[602,668,1783,1809]" pageId="5" pageNumber="374" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="8B85A038FFEBFFFE20A0BF353A10364C" blockId="5.[151,668,1782,1845]" box="[151,280,1819,1845]" pageId="5" pageNumber="374">
(
<figureCitation id="1301BCBDFFEBFFFE20A8BF353A18364C" box="[159,272,1819,1845]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStart-4="FIGURE 8" captionStart-5="FIGURE 9" captionStart-6="FIGURE 10" captionStart-7="FIGURE 11" captionStartId-0="6.[151,250,1320,1344]" captionStartId-1="6.[151,250,1981,2005]" captionStartId-2="7.[151,250,1543,1567]" captionStartId-3="8.[151,250,1928,1952]" captionStartId-4="9.[151,250,1481,1505]" captionStartId-5="10.[151,250,1352,1376]" captionStartId-6="11.[151,250,1096,1120]" captionStartId-7="11.[151,250,1847,1871]" captionTargetBox-0="[228,1360,597,1297]" captionTargetBox-1="[211,1376,1412,1953]" captionTargetBox-2="[151,1430,418,1520]" captionTargetBox-3="[152,1435,814,1904]" captionTargetBox-4="[157,1430,181,1458]" captionTargetBox-5="[188,1372,197,1328]" captionTargetBox-6="[161,1433,188,1065]" captionTargetBox-7="[155,1432,1180,1822]" captionTargetId-0="figure-239@6.[228,1360,597,1297]" captionTargetId-1="figure-278@6.[207,1380,1404,1957]" captionTargetId-2="figure-133@7.[150,1437,417,1520]" captionTargetId-3="figure-336@8.[152,1435,813,1905]" captionTargetId-4="figure-18@9.[157,1430,181,1458]" captionTargetId-5="figure-23@10.[188,1400,181,1328]" captionTargetId-6="figure-17@11.[154,1434,181,1072]" captionTargetId-7="figure-51@11.[155,1432,1180,1822]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="11" captionText-0="FIGURE 4. Adult habitus Tico gen. n. emmettcarri sp. n.; A) female body lateral view, B) dorsal view, C) male body lateral view, and D) dorsal view, scale = 1 mm." captionText-1="FIGURE 5. Adult Tico gen. n. emmettcarri sp. n.; A) head and pronotum frontal view, B) head, pronotum, and mesonotum dorsal view, and C) head, pronotum, and mesonotum lateral view, scale = 1 mm." captionText-2="FIGURE 6. Male Tico gen. n. emmettcarri sp. n. wing venation; black = vein, green = cell." captionText-3="FIGURE 7. Male Tico gen. n. emmettcarri sp. n. terminalia; A) lateral view, B) ventral view, and C) dorsal view." captionText-4="FIGURE 8. Gonostylus of Tico gen. n. emmettcarri sp. n.; A) lateral view demonstrating ventral-angular projection and dorsal process and B) inner surface demonstrating sclerotized ridge with tubercles and spinose apex." captionText-5="FIGURE 9. Aedeagus of adult male Tico gen. n. emmettcarri sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." captionText-6="FIGURE 10. Illustration of aedeagus of adult male Tico gen. n. emmettcarri sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." captionText-7="FIGURE 11. Female genitalia of Tico gen. n. emmettcarri sp. n.; A) pregenital plate ventral view and B) first gonapophysis of ovipositor." figureDoi-0="http://doi.org/10.5281/zenodo.4441914" figureDoi-1="http://doi.org/10.5281/zenodo.4441916" figureDoi-2="http://doi.org/10.5281/zenodo.4441920" figureDoi-3="http://doi.org/10.5281/zenodo.4441922" figureDoi-4="http://doi.org/10.5281/zenodo.4441924" figureDoi-5="http://doi.org/10.5281/zenodo.4441926" figureDoi-6="http://doi.org/10.5281/zenodo.4441928" figureDoi-7="http://doi.org/10.5281/zenodo.4441930" httpUri-0="https://zenodo.org/record/4441914/files/figure.png" httpUri-1="https://zenodo.org/record/4441916/files/figure.png" httpUri-2="https://zenodo.org/record/4441920/files/figure.png" httpUri-3="https://zenodo.org/record/4441922/files/figure.png" httpUri-4="https://zenodo.org/record/4441924/files/figure.png" httpUri-5="https://zenodo.org/record/4441926/files/figure.png" httpUri-6="https://zenodo.org/record/4441928/files/figure.png" httpUri-7="https://zenodo.org/record/4441930/files/figure.png" pageId="5" pageNumber="374">Figs 411</figureCitation>
)
</paragraph>
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<paragraph id="8B85A038FFEBFFFE20A0BF4D386F3604" blockId="5.[151,1437,1890,2025]" box="[151,871,1890,1917]" pageId="5" pageNumber="374">
<materialsCitation id="3B52AA65FFEBFFFE20A0BF4D386C3604" ID-GBIF-Occurrence="3019651305" box="[151,868,1890,1917]" country="Costa Rica" location="La Selva Biological Station" pageId="5" pageNumber="374" specimenCount="1" stateProvince="Heredia" typeStatus="holotype">
<emphasis id="B94E7C2AFFEBFFFE20A0BF4D3A3E3605" bold="true" box="[151,310,1890,1917]" pageId="5" pageNumber="374">
<typeStatus id="54811E9AFFEBFFFE20A0BF4D3BDA3604" box="[151,210,1891,1917]" pageId="5" pageNumber="374">Type</typeStatus>
locality.
</emphasis>
<location id="8EE5F6E3FFEBFFFE210BBF4D397C3604" LSID="urn:lsid:plazi:treatment:0393112EFFEBFFF120A0BED93AD23690:8EE5F6E3FFEBFFFE210BBF4D397C3604" box="[316,628,1891,1917]" country="Costa Rica" name="La Selva Biological Station" pageId="5" pageNumber="374" stateProvince="Heredia">La Selva Biological Station</location>
,
<collectingRegion id="49FE6EDAFFEBFFFE2248BF4D39D33604" box="[639,731,1891,1917]" country="Costa Rica" name="Heredia" pageId="5" pageNumber="374">Heredia</collectingRegion>
,
<collectingCountry id="F32DE0A8FFEBFFFE22D1BF4D386C3604" box="[742,868,1891,1917]" name="Costa Rica" pageId="5" pageNumber="374">Costa Rica</collectingCountry>
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.
</paragraph>
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<subSubSection id="C320F3B3FFEBFFFD20F0BFA838A13064" lastPageId="6" lastPageNumber="375" pageId="5" pageNumber="374" type="diagnosis">
<paragraph id="8B85A038FFEBFFFD20F0BFA838A13064" blockId="5.[151,1437,1890,2025]" lastBlockId="6.[151,1437,151,574]" lastPageId="6" lastPageNumber="375" pageId="5" pageNumber="374">
<emphasis id="B94E7C2AFFEBFFFE20F0BFA83A4A36D9" bold="true" box="[199,322,1926,1952]" pageId="5" pageNumber="374">Diagnosis.</emphasis>
A pale species without distinct body markings bearing clear wings with limited, diffuse markings, including a distinct spot near the fork of the CuA and a more diffuse marking at the claval apex among other markings. Sc branching from R proximad of claval apex. RP and CuA collectively have 8 branches reaching wing margin. The gonostyli have a rounded apex and a dorsal process near midlength including a sclerotized ridge and a large, curved spine angled distad.Aedeagus compact and narrow, shaft arched upward near midlength, devoid of subapical processes, with complex array of apical retrorse processes associated with endosoma. Pygofer with lateral margins of opening convex with a rounded lobe near the anal tube articulation.
</paragraph>
</subSubSection>
<subSubSection id="C320F3B3FFE8FFF220F0B90839F737A6" lastPageId="9" lastPageNumber="378" pageId="6" pageNumber="375" type="description">
<paragraph id="8B85A038FFE8FFFD20F0B9083E8E3347" blockId="6.[151,1437,151,574]" pageId="6" pageNumber="375">
<emphasis id="B94E7C2AFFE8FFFD20F0B9083A513039" bold="true" box="[199,345,294,320]" pageId="6" pageNumber="375">Description.</emphasis>
<emphasis id="B94E7C2AFFE8FFFD2168B9093AA83038" box="[351,416,295,321]" italics="true" pageId="6" pageNumber="375">Color</emphasis>
. In life, uniform ivory-white, males with diffuse orange markings on genae (diagonally below antennae), and laterad of lateral carinae on pro- and mesonotum and on median carina of mesonotum. (
<figureCitation id="1301BCBDFFE8FFFD250DB9653E88301C" box="[1338,1408,331,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1320,1344]" captionTargetBox="[228,1360,597,1297]" captionTargetId="figure-239@6.[228,1360,597,1297]" captionTargetPageId="6" captionText="FIGURE 4. Adult habitus Tico gen. n. emmettcarri sp. n.; A) female body lateral view, B) dorsal view, C) male body lateral view, and D) dorsal view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4441914" httpUri="https://zenodo.org/record/4441914/files/figure.png" pageId="6" pageNumber="375">Figs 4</figureCitation>
&amp;
<figureCitation id="1301BCBDFFE8FFFD20A0B9403BAF30F0" box="[151,167,366,393]" captionStart="FIGURE 5" captionStartId="6.[151,250,1981,2005]" captionTargetBox="[211,1376,1412,1953]" captionTargetId="figure-278@6.[207,1380,1404,1957]" captionTargetPageId="6" captionText="FIGURE 5. Adult Tico gen. n. emmettcarri sp. n.; A) head and pronotum frontal view, B) head, pronotum, and mesonotum dorsal view, and C) head, pronotum, and mesonotum lateral view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4441916" httpUri="https://zenodo.org/record/4441916/files/figure.png" pageId="6" pageNumber="375">5</figureCitation>
). Lateral carinae of head embrowned (
<figureCitation id="1301BCBDFFE8FFFD2260B941399330F0" box="[599,667,366,393]" captionStart="FIGURE 5" captionStartId="6.[151,250,1981,2005]" captionTargetBox="[211,1376,1412,1953]" captionTargetId="figure-278@6.[207,1380,1404,1957]" captionTargetPageId="6" captionText="FIGURE 5. Adult Tico gen. n. emmettcarri sp. n.; A) head and pronotum frontal view, B) head, pronotum, and mesonotum dorsal view, and C) head, pronotum, and mesonotum lateral view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4441916" httpUri="https://zenodo.org/record/4441916/files/figure.png" pageId="6" pageNumber="375">Fig. 5</figureCitation>
). Forewings clear, male wings with three prominent black spots; one near midlength in costal cell, second near fork of CuA, third between branches MP
<subScript id="17BEA27DFFE8FFFD241CB98B3F3B30CA" attach="left" box="[1067,1075,421,435]" fontSize="6" pageId="6" pageNumber="375">3</subScript>
and MP
<subScript id="17BEA27DFFE8FFFD24A7B98B3F9030CA" attach="left" box="[1168,1176,421,435]" fontSize="6" pageId="6" pageNumber="375">4</subScript>
proximad to crossvein; additional less prominent markings (some reddish) include large diffuse patch on Pcu; a patch cell between apex of clavus and CuA; weak markings at apical vein in each apical cell, and two poorly defined bands, one arising about at marginal apex of Sc, the second just beyond marginal apex of RA (
<figureCitation id="1301BCBDFFE8FFFD234AB9D138C93363" box="[893,961,511,538]" captionStart="FIGURE 6" captionStartId="7.[151,250,1543,1567]" captionTargetBox="[151,1430,418,1520]" captionTargetId="figure-133@7.[150,1437,417,1520]" captionTargetPageId="7" captionText="FIGURE 6. Male Tico gen. n. emmettcarri sp. n. wing venation; black = vein, green = cell." figureDoi="http://doi.org/10.5281/zenodo.4441920" httpUri="https://zenodo.org/record/4441920/files/figure.png" pageId="6" pageNumber="375">Fig. 6</figureCitation>
). Females with similar markings, differing in size and definition, except lacking spots in subcostal cell and Pcu and nearly all markings are fuscous (
<figureCitation id="1301BCBDFFE8FFFD2504BA0D3E703347" box="[1331,1400,547,574]" captionStart="FIGURE 6" captionStartId="7.[151,250,1543,1567]" captionTargetBox="[151,1430,418,1520]" captionTargetId="figure-133@7.[150,1437,417,1520]" captionTargetPageId="7" captionText="FIGURE 6. Male Tico gen. n. emmettcarri sp. n. wing venation; black = vein, green = cell." figureDoi="http://doi.org/10.5281/zenodo.4441920" httpUri="https://zenodo.org/record/4441920/files/figure.png" pageId="6" pageNumber="375">Fig. 6</figureCitation>
).
</paragraph>
<caption id="DF45F0B0FFE8FFFD20A0BD063927341D" ID-DOI="http://doi.org/10.5281/zenodo.4441914" ID-Zenodo-Dep="4441914" httpUri="https://zenodo.org/record/4441914/files/figure.png" pageId="6" pageNumber="375" startId="6.[151,250,1320,1344]" targetBox="[228,1360,597,1297]" targetPageId="6">
<paragraph id="8B85A038FFE8FFFD20A0BD063927341D" blockId="6.[151,1436,1320,1380]" pageId="6" pageNumber="375">
<emphasis id="B94E7C2AFFE8FFFD20A0BD063A1C3439" bold="true" box="[151,276,1320,1344]" pageId="6" pageNumber="375">FIGURE 4.</emphasis>
Adult habitus
<taxonomicName id="4C3ADBBBFFE8FFFD219BBD073AD03439" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[428,472,1321,1344]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E7C2AFFE8FFFD219BBD073AD03439" box="[428,472,1321,1344]" italics="true" pageId="6" pageNumber="375">Tico</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE8FFFD21E8BD06392F3439" bold="true" box="[479,551,1320,1344]" pageId="6" pageNumber="375">
<taxonomicNameLabel id="A27DC151FFE8FFFD21E8BD06392F3439" box="[479,551,1320,1344]" pageId="6" pageNumber="375" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C3ADBBBFFE8FFFD2218BD0739A33439" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[559,683,1321,1344]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="emmettcarri" status="sp. nov.">
<emphasis id="B94E7C2AFFE8FFFD2218BD0739A33439" box="[559,683,1321,1344]" italics="true" pageId="6" pageNumber="375">emmettcarri</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE8FFFD2285BD0639E43439" bold="true" box="[690,748,1320,1344]" pageId="6" pageNumber="375">
<taxonomicNameLabel id="A27DC151FFE8FFFD2285BD0639E43439" box="[690,748,1320,1344]" pageId="6" pageNumber="375" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; A) female body lateral view, B) dorsal view, C) male body lateral view, and D) dorsal view, scale = 1 mm.
</paragraph>
</caption>
<caption id="DF45F0B0FFE8FFFD20A0BF9338CF3680" ID-DOI="http://doi.org/10.5281/zenodo.4441916" ID-Zenodo-Dep="4441916" httpUri="https://zenodo.org/record/4441916/files/figure.png" pageId="6" pageNumber="375" startId="6.[151,250,1981,2005]" targetBox="[211,1376,1412,1953]" targetPageId="6">
<paragraph id="8B85A038FFE8FFFD20A0BF9338CF3680" blockId="6.[151,1436,1981,2042]" pageId="6" pageNumber="375">
<emphasis id="B94E7C2AFFE8FFFD20A0BF933A1D36AF" bold="true" box="[151,277,1981,2006]" pageId="6" pageNumber="375">FIGURE 5.</emphasis>
Adult
<taxonomicName id="4C3ADBBBFFE8FFFD216ABF903A8136AC" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[349,393,1982,2005]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E7C2AFFE8FFFD216ABF903A8136AC" box="[349,393,1982,2005]" italics="true" pageId="6" pageNumber="375">Tico</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE8FFFD21A6BF903AD236AF" bold="true" box="[401,474,1982,2006]" pageId="6" pageNumber="375">
<taxonomicNameLabel id="A27DC151FFE8FFFD21A6BF903AD236AF" box="[401,474,1982,2006]" pageId="6" pageNumber="375" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C3ADBBBFFE8FFFD21D5BF90395636AC" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[482,606,1982,2005]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="emmettcarri" status="sp. nov.">
<emphasis id="B94E7C2AFFE8FFFD21D5BF90395636AC" box="[482,606,1982,2005]" italics="true" pageId="6" pageNumber="375">emmettcarri</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE8FFFD2252BF9039A836AF" bold="true" box="[613,672,1982,2006]" pageId="6" pageNumber="375">
<taxonomicNameLabel id="A27DC151FFE8FFFD2252BF9039A836AF" box="[613,672,1982,2006]" pageId="6" pageNumber="375" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; A) head and pronotum frontal view, B) head, pronotum, and mesonotum dorsal view, and C) head, pronotum, and mesonotum lateral view, scale = 1 mm.
</paragraph>
</caption>
<paragraph id="8B85A038FFE9FFFC20F0B8B93E1530F0" blockId="7.[151,1436,151,393]" pageId="7" pageNumber="376">
<emphasis id="B94E7C2AFFE9FFFC20F0B8B93A3E31C8" box="[199,310,151,177]" italics="true" pageId="7" pageNumber="376">Structure.</emphasis>
Body length 3.00 mm (males),
<quantity id="4CC20DDDFFE9FFFC22A5B8B939FC31C8" box="[658,756,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.02" pageId="7" pageNumber="376" unit="mm" value="2.02">2.02 mm</quantity>
(females) (
<tableCitation id="C6B89583FFE9FFFC2347B8B938CF31C8" box="[880,967,151,177]" captionStart="TABLE 2" captionStartId="12.[151,239,151,175]" captionText="TABLE 2. Biometric data for Tico emmettcarri gen. et. sp. n.; values = mm." pageId="7" pageNumber="376">Table 2</tableCitation>
). Head. In lateral view, generally rounded, slightly humped at transverse carina between vertex and frons (
<figureCitation id="1301BCBDFFE9FFFC2360B895389131AC" box="[855,921,186,213]" captionStart="FIGURE 5" captionStartId="6.[151,250,1981,2005]" captionTargetBox="[211,1376,1412,1953]" captionTargetId="figure-278@6.[207,1380,1404,1957]" captionTargetPageId="6" captionText="FIGURE 5. Adult Tico gen. n. emmettcarri sp. n.; A) head and pronotum frontal view, B) head, pronotum, and mesonotum dorsal view, and C) head, pronotum, and mesonotum lateral view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4441916" httpUri="https://zenodo.org/record/4441916/files/figure.png" pageId="7" pageNumber="376">Fig. 5</figureCitation>
), anterior margin somewhat flattened (
<figureCitation id="1301BCBDFFE9FFFC257DB8953E8631AC" box="[1354,1422,186,213]" captionStart="FIGURE 5" captionStartId="6.[151,250,1981,2005]" captionTargetBox="[211,1376,1412,1953]" captionTargetId="figure-278@6.[207,1380,1404,1957]" captionTargetPageId="6" captionText="FIGURE 5. Adult Tico gen. n. emmettcarri sp. n.; A) head and pronotum frontal view, B) head, pronotum, and mesonotum dorsal view, and C) head, pronotum, and mesonotum lateral view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4441916" httpUri="https://zenodo.org/record/4441916/files/figure.png" pageId="7" pageNumber="376">Fig. 5</figureCitation>
). Frons and vertex with lateral margins strongly keeled, bearing sensory pits, median carinae absent (
<figureCitation id="1301BCBDFFE9FFFC24C1B8F13E323180" box="[1270,1338,222,249]" captionStart="FIGURE 5" captionStartId="6.[151,250,1981,2005]" captionTargetBox="[211,1376,1412,1953]" captionTargetId="figure-278@6.[207,1380,1404,1957]" captionTargetPageId="6" captionText="FIGURE 5. Adult Tico gen. n. emmettcarri sp. n.; A) head and pronotum frontal view, B) head, pronotum, and mesonotum dorsal view, and C) head, pronotum, and mesonotum lateral view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4441916" httpUri="https://zenodo.org/record/4441916/files/figure.png" pageId="7" pageNumber="376">Fig. 5</figureCitation>
). Antennae short and bulbous. In dorsal view, vertex roughly triangular, widest posteriorly, narrowed anteriorly to transverse carina; anterior margin concave between carinae, posterior margin strongly concave between eyes (
<figureCitation id="1301BCBDFFE9FFFC251BB9093E783038" box="[1324,1392,294,321]" captionStart="FIGURE 5" captionStartId="6.[151,250,1981,2005]" captionTargetBox="[211,1376,1412,1953]" captionTargetId="figure-278@6.[207,1380,1404,1957]" captionTargetPageId="6" captionText="FIGURE 5. Adult Tico gen. n. emmettcarri sp. n.; A) head and pronotum frontal view, B) head, pronotum, and mesonotum dorsal view, and C) head, pronotum, and mesonotum lateral view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4441916" httpUri="https://zenodo.org/record/4441916/files/figure.png" pageId="7" pageNumber="376">Fig. 5</figureCitation>
). In frontal view, frons relatively broad, lateral margins parallel between eyes, expanding below eyes to frontoclypeal suture; transverse suture between frons and clypeus distinct. Clypeus triangular, bearing median carina.
</paragraph>
<caption id="DF45F0B0FFE9FFFC20A0BE293F333766" ID-DOI="http://doi.org/10.5281/zenodo.4441920" ID-Zenodo-Dep="4441920" box="[151,1083,1543,1568]" httpUri="https://zenodo.org/record/4441920/files/figure.png" pageId="7" pageNumber="376" startId="7.[151,250,1543,1567]" targetBox="[151,1430,418,1520]" targetPageId="7">
<paragraph id="8B85A038FFE9FFFC20A0BE293F333766" blockId="7.[151,1083,1543,1568]" box="[151,1083,1543,1568]" pageId="7" pageNumber="376">
<emphasis id="B94E7C2AFFE9FFFC20A0BE293A1B3766" bold="true" box="[151,275,1543,1567]" pageId="7" pageNumber="376">FIGURE 6.</emphasis>
Male
<taxonomicName id="4C3ADBBBFFE9FFFC2163BE263A883766" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[340,384,1544,1567]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E7C2AFFE9FFFC2163BE263A883766" box="[340,384,1544,1567]" italics="true" pageId="7" pageNumber="376">Tico</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE9FFFC21B1BE293AC53766" bold="true" box="[390,461,1543,1567]" pageId="7" pageNumber="376">
<taxonomicNameLabel id="A27DC151FFE9FFFC21B1BE293AC53766" box="[390,461,1543,1567]" pageId="7" pageNumber="376" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C3ADBBBFFE9FFFC21E4BE2639473766" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[467,591,1544,1567]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="emmettcarri" status="sp. nov.">
<emphasis id="B94E7C2AFFE9FFFC21E4BE2639473766" box="[467,591,1544,1567]" italics="true" pageId="7" pageNumber="376">emmettcarri</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE9FFFC2262BE2939863766" bold="true" box="[597,654,1543,1567]" pageId="7" pageNumber="376">
<taxonomicNameLabel id="A27DC151FFE9FFFC2262BE2939863766" box="[597,654,1543,1567]" pageId="7" pageNumber="376" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
wing venation; black = vein, green = cell.
</paragraph>
</caption>
<paragraph id="8B85A038FFE9FFFC20F0BE633A4D36A9" blockId="7.[151,1437,1613,2000]" pageId="7" pageNumber="376">
Thorax. Pronotum relatively long at midline (subequal to vertex length), weakly tricarinate; anterior margin strongly convex, following contours of head; posterior margin concave; pronotal paradiscal region broadly foliate, forming a cup-like fossa surrounding and strongly overtopping the antennae, foliations appearing semi-quadrate in frontal view (
<figureCitation id="1301BCBDFFE9FFFC2118BE973A8137AD" box="[303,393,1721,1748]" captionStart="FIGURE 9" captionStartId="10.[151,250,1352,1376]" captionTargetBox="[188,1372,197,1328]" captionTargetId="figure-23@10.[188,1400,181,1328]" captionTargetPageId="10" captionText="FIGURE 9. Aedeagus of adult male Tico gen. n. emmettcarri sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441926" httpUri="https://zenodo.org/record/4441926/files/figure.png" pageId="7" pageNumber="376">Fig. 9A</figureCitation>
). Mesonotum slightly wider than long, tricarinate, lateral carinae subparallel, slightly convex (
<figureCitation id="1301BCBDFFE9FFFC2097BEF33BEB3781" box="[160,227,1757,1784]" captionStart="FIGURE 5" captionStartId="6.[151,250,1981,2005]" captionTargetBox="[211,1376,1412,1953]" captionTargetId="figure-278@6.[207,1380,1404,1957]" captionTargetPageId="6" captionText="FIGURE 5. Adult Tico gen. n. emmettcarri sp. n.; A) head and pronotum frontal view, B) head, pronotum, and mesonotum dorsal view, and C) head, pronotum, and mesonotum lateral view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4441916" httpUri="https://zenodo.org/record/4441916/files/figure.png" pageId="7" pageNumber="376">Fig. 5</figureCitation>
). Rostrum just exceeding hind coxae, apical segment about as long as wide. Hind tibia lacking lateral spines, flared at apex; spinulation of hind leg 7-6-6. Forewing (
<figureCitation id="1301BCBDFFE9FFFC2332BF2F38403665" box="[773,840,1793,1820]" captionStart="FIGURE 6" captionStartId="7.[151,250,1543,1567]" captionTargetBox="[151,1430,418,1520]" captionTargetId="figure-133@7.[150,1437,417,1520]" captionTargetPageId="7" captionText="FIGURE 6. Male Tico gen. n. emmettcarri sp. n. wing venation; black = vein, green = cell." figureDoi="http://doi.org/10.5281/zenodo.4441920" httpUri="https://zenodo.org/record/4441920/files/figure.png" pageId="7" pageNumber="376">Fig. 6</figureCitation>
) with sensory pits along costal, subcostal+radial, and postcubital veins. Sc branching proximad of claval apex and fork of RA and RP. Branching pattern; RA 1-branched, RP 2-branched, MP 4-branched (fork of MP
<subScript id="17BEA27DFFE9FFFC22A3BF7239A63613" attach="left" box="[660,686,1884,1898]" fontSize="6" pageId="7" pageNumber="376">3+4</subScript>
from MP
<subScript id="17BEA27DFFE9FFFC2328BF7238313613" attach="left" box="[799,825,1884,1898]" fontSize="6" pageId="7" pageNumber="376">1+2</subScript>
and MP
<subScript id="17BEA27DFFE9FFFC23AABF7238AD3613" attach="left" box="[925,933,1884,1898]" fontSize="6" pageId="7" pageNumber="376">1</subScript>
from MP
<subScript id="17BEA27DFFE9FFFC2424BF723F133613" attach="left" box="[1043,1051,1884,1898]" fontSize="6" pageId="7" pageNumber="376">2</subScript>
nearly coincident; MP may be 5- branched if RP
<subScript id="17BEA27DFFE9FFFC2108BFAE3A5136F7" attach="left" box="[319,345,1920,1934]" fontSize="6" pageId="7" pageNumber="376">3+4</subScript>
is assumed to fork), CuA 2-branched (distally fused if RP
<subScript id="17BEA27DFFE9FFFC23D6BFAE38F336F7" attach="left" box="[993,1019,1920,1934]" fontSize="6" pageId="7" pageNumber="376">3+4</subScript>
forks); Sc branch just preceding fork of RA and RP, and proximad of claval apex; CuA fork preceding fork of RA and RP (latter about level of claval apex) (
<figureCitation id="1301BCBDFFE9FFFC20D2BF9B3A3036A9" box="[229,312,1973,2000]" captionStart="FIGURE 10" captionStartId="11.[151,250,1096,1120]" captionTargetBox="[161,1433,188,1065]" captionTargetId="figure-17@11.[154,1434,181,1072]" captionTargetPageId="11" captionText="FIGURE 10. Illustration of aedeagus of adult male Tico gen. n. emmettcarri sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441928" httpUri="https://zenodo.org/record/4441928/files/figure.png" pageId="7" pageNumber="376">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8B85A038FFE6FFF320F0B8B938AC326C" blockId="8.[151,1437,151,789]" pageId="8" pageNumber="377">
Terminalia. Pygofer in lateral view narrow, irregularly sinuate on both posterior and anterior margin, widest ventrally, lateral margin of opening distinctly projecting posteriorly; dorsal margin with ovate lobe near articulation of anal tube (
<figureCitation id="1301BCBDFFE6FFF3211EB8F13A653180" box="[297,365,223,249]" captionStart="FIGURE 7" captionStartId="8.[151,250,1928,1952]" captionTargetBox="[152,1435,814,1904]" captionTargetId="figure-336@8.[152,1435,813,1905]" captionTargetPageId="8" captionText="FIGURE 7. Male Tico gen. n. emmettcarri sp. n. terminalia; A) lateral view, B) ventral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441922" httpUri="https://zenodo.org/record/4441922/files/figure.png" pageId="8" pageNumber="377">Fig. 7</figureCitation>
); in ventral view, pygofer lacking medioventral process (
<figureCitation id="1301BCBDFFE6FFF323DDB8F13F263180" box="[1002,1070,223,249]" captionStart="FIGURE 7" captionStartId="8.[151,250,1928,1952]" captionTargetBox="[152,1435,814,1904]" captionTargetId="figure-336@8.[152,1435,813,1905]" captionTargetPageId="8" captionText="FIGURE 7. Male Tico gen. n. emmettcarri sp. n. terminalia; A) lateral view, B) ventral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441922" httpUri="https://zenodo.org/record/4441922/files/figure.png" pageId="8" pageNumber="377">Fig. 7</figureCitation>
); in dorsal view, ovate lobes distinct (
<figureCitation id="1301BCBDFFE6FFF320EEB92D3A163064" box="[217,286,259,285]" captionStart="FIGURE 7" captionStartId="8.[151,250,1928,1952]" captionTargetBox="[152,1435,814,1904]" captionTargetId="figure-336@8.[152,1435,813,1905]" captionTargetPageId="8" captionText="FIGURE 7. Male Tico gen. n. emmettcarri sp. n. terminalia; A) lateral view, B) ventral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441922" httpUri="https://zenodo.org/record/4441922/files/figure.png" pageId="8" pageNumber="377">Fig. 7</figureCitation>
). Gonostyli in lateral view spatulate, apex rounded, with proximal angular projection ventrally, dorsal margin near midlength bearing basal, sclerotized hook (apex directed laterad) and distal sclerotized ridge (
<figureCitation id="1301BCBDFFE6FFF32579B9093E943038" box="[1358,1436,295,321]" captionStart="FIGURE 7" captionStartId="8.[151,250,1928,1952]" captionTargetBox="[152,1435,814,1904]" captionTargetId="figure-336@8.[152,1435,813,1905]" captionTargetPageId="8" captionText="FIGURE 7. Male Tico gen. n. emmettcarri sp. n. terminalia; A) lateral view, B) ventral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441922" httpUri="https://zenodo.org/record/4441922/files/figure.png" pageId="8" pageNumber="377">Figs. 7</figureCitation>
&amp;
<figureCitation id="1301BCBDFFE6FFF32082B9653BCC301C" box="[181,196,331,357]" captionStart="FIGURE 8" captionStartId="9.[151,250,1481,1505]" captionTargetBox="[157,1430,181,1458]" captionTargetId="figure-18@9.[157,1430,181,1458]" captionTargetPageId="9" captionText="FIGURE 8. Gonostylus of Tico gen. n. emmettcarri sp. n.; A) lateral view demonstrating ventral-angular projection and dorsal process and B) inner surface demonstrating sclerotized ridge with tubercles and spinose apex." figureDoi="http://doi.org/10.5281/zenodo.4441924" httpUri="https://zenodo.org/record/4441924/files/figure.png" pageId="8" pageNumber="377">8</figureCitation>
); in ventral view, lateral margin irregularly sinuate, inner margins strongly concave, greatly expanded at base forming medial angular projections (
<figureCitation id="1301BCBDFFE6FFF32206B941397D30F0" box="[561,629,367,393]" captionStart="FIGURE 7" captionStartId="8.[151,250,1928,1952]" captionTargetBox="[152,1435,814,1904]" captionTargetId="figure-336@8.[152,1435,813,1905]" captionTargetPageId="8" captionText="FIGURE 7. Male Tico gen. n. emmettcarri sp. n. terminalia; A) lateral view, B) ventral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441922" httpUri="https://zenodo.org/record/4441922/files/figure.png" pageId="8" pageNumber="377">Fig. 7</figureCitation>
); in dorsal view, inner margins with serrulate sclerotized ridge terminating with small spine, approximately ¾ the length of gonostyli, basal portion with dark tubercles (
<figureCitation id="1301BCBDFFE6FFF324E1B9BD3E1230D4" box="[1238,1306,403,429]" captionStart="FIGURE 8" captionStartId="9.[151,250,1481,1505]" captionTargetBox="[157,1430,181,1458]" captionTargetId="figure-18@9.[157,1430,181,1458]" captionTargetPageId="9" captionText="FIGURE 8. Gonostylus of Tico gen. n. emmettcarri sp. n.; A) lateral view demonstrating ventral-angular projection and dorsal process and B) inner surface demonstrating sclerotized ridge with tubercles and spinose apex." figureDoi="http://doi.org/10.5281/zenodo.4441924" httpUri="https://zenodo.org/record/4441924/files/figure.png" pageId="8" pageNumber="377">Fig. 8</figureCitation>
). Aedeagus nearly bilaterally symmetrical, upcurved near midlength (
<figureCitation id="1301BCBDFFE6FFF32321B999385130A8" box="[790,857,439,465]" captionStart="FIGURE 9" captionStartId="10.[151,250,1352,1376]" captionTargetBox="[188,1372,197,1328]" captionTargetId="figure-23@10.[188,1400,181,1328]" captionTargetPageId="10" captionText="FIGURE 9. Aedeagus of adult male Tico gen. n. emmettcarri sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441926" httpUri="https://zenodo.org/record/4441926/files/figure.png" pageId="8" pageNumber="377">Fig. 9</figureCitation>
), shaft without subapical processes, but bearing pair median dorsal flanges, dorsally serrate in in left lateral view (dorsum partly occluded by endosoma); aedeagus with single pair of large retrorse processes arising from apex (
<figureCitation id="1301BCBDFFE6FFF3232FB9D138633360" box="[792,875,511,537]" captionStart="FIGURE 10" captionStartId="11.[151,250,1096,1120]" captionTargetBox="[161,1433,188,1065]" captionTargetId="figure-17@11.[154,1434,181,1072]" captionTargetPageId="11" captionText="FIGURE 10. Illustration of aedeagus of adult male Tico gen. n. emmettcarri sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441928" httpUri="https://zenodo.org/record/4441928/files/figure.png" pageId="8" pageNumber="377">Fig. 10</figureCitation>
, A
<figureCitation id="1301BCBDFFE6FFF323B9B9D138903360" box="[910,920,511,537]" captionStart="FIGURE 1" captionStartId="2.[151,250,1013,1037]" captionTargetBox="[524,1063,181,989]" captionTargetId="figure-23@2.[524,1063,181,989]" captionTargetPageId="2" captionText="FIGURE 1. Host plant and general habitat where specimens were collected; all collected from underside of leaves." figureDoi="http://doi.org/10.5281/zenodo.4441908" httpUri="https://zenodo.org/record/4441908/files/figure.png" pageId="8" pageNumber="377">1</figureCitation>
&amp; A
<figureCitation id="1301BCBDFFE6FFF323E6B9D138D73360" box="[977,991,511,537]" captionStart="FIGURE 2" captionStartId="3.[151,250,1956,1980]" captionTargetBox="[155,1419,1355,1931]" captionTargetId="figure-404@3.[155,1432,1354,1931]" captionTargetPageId="3" captionText="FIGURE 2. Cenchrea dorsalis; A) lateral view, B) gonostylus lateral view, C) aedeagus right lateral view, D) aedeagus dorsal view, E) male terminalia lateral view, and F) male terminalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.4441910" httpUri="https://zenodo.org/record/4441910/files/figure.png" pageId="8" pageNumber="377">2</figureCitation>
, resting above endosoma) and complex endosoma; endosoma symmetrical, compact and complex, bearing large pair of robust elongate ventral processes (E1 &amp; E2) bearing 2 pair of small processes at their base (E3 &amp; E5 on right ride, E4 &amp; E6 on left side) (
<figureCitation id="1301BCBDFFE6FFF3250DBA693E863318" box="[1338,1422,583,609]" captionStart="FIGURE 10" captionStartId="11.[151,250,1096,1120]" captionTargetBox="[161,1433,188,1065]" captionTargetId="figure-17@11.[154,1434,181,1072]" captionTargetPageId="11" captionText="FIGURE 10. Illustration of aedeagus of adult male Tico gen. n. emmettcarri sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441928" httpUri="https://zenodo.org/record/4441928/files/figure.png" pageId="8" pageNumber="377">Fig. 10</figureCitation>
); two processes arise near aedeagal apex (medial, in dorsal view,
<figureCitation id="1301BCBDFFE6FFF32353BA4538BE33FC" box="[868,950,619,645]" captionStart="FIGURE 10" captionStartId="11.[151,250,1096,1120]" captionTargetBox="[161,1433,188,1065]" captionTargetId="figure-17@11.[154,1434,181,1072]" captionTargetPageId="11" captionText="FIGURE 10. Illustration of aedeagus of adult male Tico gen. n. emmettcarri sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441928" httpUri="https://zenodo.org/record/4441928/files/figure.png" pageId="8" pageNumber="377">Fig. 10</figureCitation>
, E
<figureCitation id="1301BCBDFFE6FFF323E2BA4538EB33FC" box="[981,995,619,645]" captionStart="FIGURE 7" captionStartId="8.[151,250,1928,1952]" captionTargetBox="[152,1435,814,1904]" captionTargetId="figure-336@8.[152,1435,813,1905]" captionTargetPageId="8" captionText="FIGURE 7. Male Tico gen. n. emmettcarri sp. n. terminalia; A) lateral view, B) ventral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441922" httpUri="https://zenodo.org/record/4441922/files/figure.png" pageId="8" pageNumber="377">7</figureCitation>
, E
<figureCitation id="1301BCBDFFE6FFF32433BA453F1933FC" box="[1028,1041,619,645]" captionStart="FIGURE 8" captionStartId="9.[151,250,1481,1505]" captionTargetBox="[157,1430,181,1458]" captionTargetId="figure-18@9.[157,1430,181,1458]" captionTargetPageId="9" captionText="FIGURE 8. Gonostylus of Tico gen. n. emmettcarri sp. n.; A) lateral view demonstrating ventral-angular projection and dorsal process and B) inner surface demonstrating sclerotized ridge with tubercles and spinose apex." figureDoi="http://doi.org/10.5281/zenodo.4441924" httpUri="https://zenodo.org/record/4441924/files/figure.png" pageId="8" pageNumber="377">8</figureCitation>
), and two pair of apical endosomal processes, 1 pair of long, narrow processes arising from inner margin of endosoma
<figureCitation id="1301BCBDFFE6FFF32409BAA13F9833D0" box="[1086,1168,655,681]" captionStart="FIGURE 10" captionStartId="11.[151,250,1096,1120]" captionTargetBox="[161,1433,188,1065]" captionTargetId="figure-17@11.[154,1434,181,1072]" captionTargetPageId="11" captionText="FIGURE 10. Illustration of aedeagus of adult male Tico gen. n. emmettcarri sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441928" httpUri="https://zenodo.org/record/4441928/files/figure.png" pageId="8" pageNumber="377">Fig. 10</figureCitation>
, E
<figureCitation id="1301BCBDFFE6FFF32499BAA13FB333D0" box="[1198,1211,655,681]" captionStart="FIGURE 9" captionStartId="10.[151,250,1352,1376]" captionTargetBox="[188,1372,197,1328]" captionTargetId="figure-23@10.[188,1400,181,1328]" captionTargetPageId="10" captionText="FIGURE 9. Aedeagus of adult male Tico gen. n. emmettcarri sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441926" httpUri="https://zenodo.org/record/4441926/files/figure.png" pageId="8" pageNumber="377">9</figureCitation>
&amp; E
<figureCitation id="1301BCBDFFE6FFF324C6BAA13E0533D0" box="[1265,1293,655,681]" captionStart="FIGURE 10" captionStartId="11.[151,250,1096,1120]" captionTargetBox="[161,1433,188,1065]" captionTargetId="figure-17@11.[154,1434,181,1072]" captionTargetPageId="11" captionText="FIGURE 10. Illustration of aedeagus of adult male Tico gen. n. emmettcarri sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441928" httpUri="https://zenodo.org/record/4441928/files/figure.png" pageId="8" pageNumber="377">10</figureCitation>
) and a small median pair of processes visible in dorsal view (
<figureCitation id="1301BCBDFFE6FFF322F7BA9D381C33B4" box="[704,788,691,717]" captionStart="FIGURE 10" captionStartId="11.[151,250,1096,1120]" captionTargetBox="[161,1433,188,1065]" captionTargetId="figure-17@11.[154,1434,181,1072]" captionTargetPageId="11" captionText="FIGURE 10. Illustration of aedeagus of adult male Tico gen. n. emmettcarri sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441928" httpUri="https://zenodo.org/record/4441928/files/figure.png" pageId="8" pageNumber="377">Fig. 10</figureCitation>
, E
<figureCitation id="1301BCBDFFE6FFF32302BA9D384633B4" box="[821,846,691,717]" captionStart="FIGURE 11" captionStartId="11.[151,250,1847,1871]" captionTargetBox="[155,1432,1180,1822]" captionTargetId="figure-51@11.[155,1432,1180,1822]" captionTargetPageId="11" captionText="FIGURE 11. Female genitalia of Tico gen. n. emmettcarri sp. n.; A) pregenital plate ventral view and B) first gonapophysis of ovipositor." figureDoi="http://doi.org/10.5281/zenodo.4441930" httpUri="https://zenodo.org/record/4441930/files/figure.png" pageId="8" pageNumber="377">11</figureCitation>
&amp; E
<figureCitation id="1301BCBDFFE6FFF323BEBA9D38AE33B4" box="[905,934,691,717]" captionStart="FIGURE 12" captionStartId="13.[151,250,1426,1450]" captionTargetBox="[160,1428,309,1403]" captionTargetId="figure-93@13.[160,1428,309,1403]" captionTargetPageId="13" captionText="FIGURE 12. Adult habitus Tico gen. n. pseudosororius sp. n.; A) lateral view and B) dorsal view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4441932" httpUri="https://zenodo.org/record/4441932/files/figure.png" pageId="8" pageNumber="377">12</figureCitation>
). Anal tube elongate in lateral view, distally downcurved (ventrally convex), apex truncate with ventrally pointed projection dorsal margin irregularly sinuate; in dorsal view anal tube broadly spatulate; paraproct short (
<figureCitation id="1301BCBDFFE6FFF32306BAD53890326C" box="[817,920,763,789]" captionStart="FIGURE 10" captionStartId="11.[151,250,1096,1120]" captionTargetBox="[161,1433,188,1065]" captionTargetId="figure-17@11.[154,1434,181,1072]" captionTargetPageId="11" captionText="FIGURE 10. Illustration of aedeagus of adult male Tico gen. n. emmettcarri sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4441928" httpUri="https://zenodo.org/record/4441928/files/figure.png" pageId="8" pageNumber="377">Fig. 10C</figureCitation>
).
</paragraph>
<caption id="DF45F0B0FFE6FFF320A0BFA63E1136D9" ID-DOI="http://doi.org/10.5281/zenodo.4441922" ID-Zenodo-Dep="4441922" box="[151,1305,1928,1952]" httpUri="https://zenodo.org/record/4441922/files/figure.png" pageId="8" pageNumber="377" startId="8.[151,250,1928,1952]" targetBox="[152,1435,814,1904]" targetPageId="8">
<paragraph id="8B85A038FFE6FFF320A0BFA63E1136D9" blockId="8.[151,1305,1928,1952]" box="[151,1305,1928,1952]" pageId="8" pageNumber="377">
<emphasis id="B94E7C2AFFE6FFF320A0BFA63A1B36D9" bold="true" box="[151,275,1928,1952]" pageId="8" pageNumber="377">FIGURE 7.</emphasis>
Male
<taxonomicName id="4C3ADBBBFFE6FFF32163BFA73A8836D9" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[340,384,1929,1952]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E7C2AFFE6FFF32163BFA73A8836D9" box="[340,384,1929,1952]" italics="true" pageId="8" pageNumber="377">Tico</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE6FFF321B1BFA63AC536D9" bold="true" box="[390,461,1928,1952]" pageId="8" pageNumber="377">
<taxonomicNameLabel id="A27DC151FFE6FFF321B1BFA63AC536D9" box="[390,461,1928,1952]" pageId="8" pageNumber="377" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C3ADBBBFFE6FFF321E4BFA7394736D9" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[467,591,1929,1952]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="emmettcarri" status="sp. nov.">
<emphasis id="B94E7C2AFFE6FFF321E4BFA7394736D9" box="[467,591,1929,1952]" italics="true" pageId="8" pageNumber="377">emmettcarri</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE6FFF32262BFA6398636D9" bold="true" box="[597,654,1928,1952]" pageId="8" pageNumber="377">
<taxonomicNameLabel id="A27DC151FFE6FFF32262BFA6398636D9" box="[597,654,1928,1952]" pageId="8" pageNumber="377" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
terminalia; A) lateral view, B) ventral view, and C) dorsal view.
</paragraph>
</caption>
<caption id="DF45F0B0FFE7FFF220A0BDE73F45377C" ID-DOI="http://doi.org/10.5281/zenodo.4441924" ID-Zenodo-Dep="4441924" httpUri="https://zenodo.org/record/4441924/files/figure.png" pageId="9" pageNumber="378" startId="9.[151,250,1481,1505]" targetBox="[157,1430,181,1458]" targetPageId="9">
<paragraph id="8B85A038FFE7FFF220A0BDE73F45377C" blockId="9.[151,1436,1481,1542]" pageId="9" pageNumber="378">
<emphasis id="B94E7C2AFFE7FFF220A0BDE73A1A349B" bold="true" box="[151,274,1481,1506]" pageId="9" pageNumber="378">FIGURE 8.</emphasis>
Gonostylus of
<taxonomicName id="4C3ADBBBFFE7FFF2219ABDE43AD13498" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[429,473,1482,1505]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="378" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E7C2AFFE7FFF2219ABDE43AD13498" box="[429,473,1482,1505]" italics="true" pageId="9" pageNumber="378">Tico</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE7FFF221E8BDE4392E349B" bold="true" box="[479,550,1482,1506]" pageId="9" pageNumber="378">
<taxonomicNameLabel id="A27DC151FFE7FFF221E8BDE4392E349B" box="[479,550,1482,1506]" pageId="9" pageNumber="378" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C3ADBBBFFE7FFF2221BBDE439A03498" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[556,680,1482,1505]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="emmettcarri" status="sp. nov.">
<emphasis id="B94E7C2AFFE7FFF2221BBDE439A03498" box="[556,680,1482,1505]" italics="true" pageId="9" pageNumber="378">emmettcarri</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE7FFF22299BDE439EE349B" bold="true" box="[686,742,1482,1506]" pageId="9" pageNumber="378">
<taxonomicNameLabel id="A27DC151FFE7FFF22299BDE439EE349B" box="[686,742,1482,1506]" pageId="9" pageNumber="378" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; A) lateral view demonstrating ventral-angular projection and dorsal process and B) inner surface demonstrating sclerotized ridge with tubercles and spinose apex.
</paragraph>
</caption>
<paragraph id="8B85A038FFE7FFF220F0BE1A3A7137C3" blockId="9.[151,1437,1588,1974]" pageId="9" pageNumber="378">
Female terminalia: Subgenital plate subtriangular, widest at base, lateral margins sinuate, converging slightly basally, distal margin broad, sinuate (
<figureCitation id="1301BCBDFFE7FFF2220CBE7639AB370B" box="[571,675,1624,1650]" captionStart="FIGURE 11" captionStartId="11.[151,250,1847,1871]" captionTargetBox="[155,1432,1180,1822]" captionTargetId="figure-51@11.[155,1432,1180,1822]" captionTargetPageId="11" captionText="FIGURE 11. Female genitalia of Tico gen. n. emmettcarri sp. n.; A) pregenital plate ventral view and B) first gonapophysis of ovipositor." figureDoi="http://doi.org/10.5281/zenodo.4441930" httpUri="https://zenodo.org/record/4441930/files/figure.png" pageId="9" pageNumber="378">Fig. 11A</figureCitation>
). First gonapophyses of ovipositor bifid at apex, ventral process robust, curved dorsad, dorsal process slightly curved upward, strongly serrate on dorsal margin near apex, serrations irregular (
<figureCitation id="1301BCBDFFE7FFF22130BE8E3A6437C3" box="[263,364,1696,1722]" captionStart="FIGURE 11" captionStartId="11.[151,250,1847,1871]" captionTargetBox="[155,1432,1180,1822]" captionTargetId="figure-51@11.[155,1432,1180,1822]" captionTargetPageId="11" captionText="FIGURE 11. Female genitalia of Tico gen. n. emmettcarri sp. n.; A) pregenital plate ventral view and B) first gonapophysis of ovipositor." figureDoi="http://doi.org/10.5281/zenodo.4441930" httpUri="https://zenodo.org/record/4441930/files/figure.png" pageId="9" pageNumber="378">Fig. 11B</figureCitation>
).
</paragraph>
<paragraph id="8B85A038FFE7FFF220F0BEEA39F737A6" blockId="9.[151,1437,1588,1974]" box="[199,767,1732,1759]" pageId="9" pageNumber="378">
<emphasis id="B94E7C2AFFE7FFF220F0BEEA3AAB37A7" bold="true" box="[199,419,1732,1758]" pageId="9" pageNumber="378">Plant associations.</emphasis>
<taxonomicName id="4C3ADBBBFFE7FFF2219DBEEA391437A7" authorityName="Harling" authorityYear="1954" box="[426,540,1732,1758]" class="Liliopsida" family="Cyclanthaceae" genus="Asplundia" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="378" phylum="Tracheophyta" rank="genus">
<emphasis id="B94E7C2AFFE7FFF2219DBEEA391437A7" box="[426,540,1732,1758]" italics="true" pageId="9" pageNumber="378">Asplundia</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C3ADBBBFFE7FFF22267BEEA39FF37A6" box="[592,759,1732,1759]" class="Liliopsida" family="Cyclanthaceae" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="378" phylum="Tracheophyta" rank="family">Cyclanthaceae</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection id="C320F3B3FFE7FFF220F0BEC63953367A" box="[199,603,1768,1795]" pageId="9" pageNumber="378" type="distribution">
<paragraph id="8B85A038FFE7FFF220F0BEC63953367A" blockId="9.[151,1437,1588,1974]" box="[199,603,1768,1795]" pageId="9" pageNumber="378">
<emphasis id="B94E7C2AFFE7FFF220F0BEC63A69367B" bold="true" box="[199,353,1768,1794]" pageId="9" pageNumber="378">Distribution.</emphasis>
<collectingCountry id="F32DE0A8FFE7FFF22150BEC63AE9367B" box="[359,481,1768,1794]" name="Costa Rica" pageId="9" pageNumber="378">Costa Rica</collectingCountry>
(
<collectingRegion id="49FE6EDAFFE7FFF221C7BEC63947367A" box="[496,591,1768,1795]" country="Costa Rica" name="Heredia" pageId="9" pageNumber="378">Heredia</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C320F3B3FFE7FFF220F0BF2239DD3633" pageId="9" pageNumber="378" type="etymology">
<paragraph id="8B85A038FFE7FFF220F0BF2239DD3633" blockId="9.[151,1437,1588,1974]" pageId="9" pageNumber="378">
<emphasis id="B94E7C2AFFE7FFF220F0BF223A46365F" bold="true" box="[199,334,1804,1830]" pageId="9" pageNumber="378">Etymology.</emphasis>
The specific name is given in honor of the young entomologist Emmett Carr who has a natural passion for palms, their insect pests and pathogens.
</paragraph>
</subSubSection>
<subSubSection id="C320F3B3FFE7FFF220F0BF7A3AB336CF" pageId="9" pageNumber="378" type="materials_examined">
<paragraph id="8B85A038FFE7FFF220F0BF7A3AB336CF" blockId="9.[151,1437,1588,1974]" pageId="9" pageNumber="378">
<emphasis id="B94E7C2AFFE7FFF220F0BF7A3ABF3617" bold="true" box="[199,439,1876,1902]" pageId="9" pageNumber="378">Material examined.</emphasis>
<materialsCitation id="3B52AA65FFE7FFF221F3BF7A38F336EB" ID-GBIF-Occurrence="3019651302" collectingDate="2019-06-22" collectionCode="FLREC" collectorName="B. W. Bahder" country="Costa Rica" location="La Selva Biological Station" pageId="9" pageNumber="378" specimenCount="2" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="54811E9AFFE7FFF221F3BF7A39243617" box="[452,556,1876,1902]" pageId="9" pageNumber="378" type="holotype">Holotype</typeStatus>
male, “
<collectingCountry id="F32DE0A8FFE7FFF222B8BF7A381D3617" box="[655,789,1876,1902]" name="Costa Rica" pageId="9" pageNumber="378">Costa Rica</collectingCountry>
,
<collectingRegion id="49FE6EDAFFE7FFF22310BF7A38883617" box="[807,896,1876,1902]" country="Costa Rica" name="Heredia" pageId="9" pageNumber="378">Heredia</collectingRegion>
/
<location id="8EE5F6E3FFE7FFF22395BF7A3FE23617" LSID="urn:lsid:plazi:treatment:0393112EFFEBFFF120A0BED93AD23690:8EE5F6E3FFE7FFF22395BF7A3FE23617" box="[930,1258,1876,1902]" country="Costa Rica" name="La Selva Biological Station" pageId="9" pageNumber="378" stateProvince="Heredia">La Selva Biological Station</location>
/
<collectorName id="26CFC5EEFFE7FFF2253ABF7A3E903617" box="[1293,1432,1876,1902]" pageId="9" pageNumber="378">B.W.Bahder</collectorName>
,
<date id="FF8486F8FFE7FFF220A0BF563A1F36EB" box="[151,279,1912,1938]" pageId="9" pageNumber="378" value="2019-06-22">
<collectingDate id="EFC07F10FFE7FFF220A0BF563A1F36EB" box="[151,279,1912,1938]" pageId="9" pageNumber="378" value="2019-06-22">22.VI.2019</collectingDate>
</date>
/ Host:
<taxonomicName id="4C3ADBBBFFE7FFF22145BF563AEC36EB" authorityName="Harling" authorityYear="1954" box="[370,484,1912,1938]" class="Liliopsida" family="Cyclanthaceae" genus="Asplundia" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="378" phylum="Tracheophyta" rank="genus">
<emphasis id="B94E7C2AFFE7FFF22145BF563AEC36EB" box="[370,484,1912,1938]" italics="true" pageId="9" pageNumber="378">Asplundia</emphasis>
</taxonomicName>
sp. //
<typeStatus id="54811E9AFFE7FFF2221DBF56399A36EB" box="[554,658,1912,1938]" pageId="9" pageNumber="378" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4C3ADBBBFFE7FFF2229EBF5739D136EB" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[681,729,1913,1938]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="378" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7C2AFFE7FFF2229EBF5739D136EB" box="[681,729,1913,1938]" italics="true" pageId="9" pageNumber="378">Tico</emphasis>
</taxonomicName>
/
<taxonomicName id="4C3ADBBBFFE7FFF222C6BF57387236EB" authority="(FLREC)" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" baseAuthorityName="FLREC" box="[753,890,1913,1938]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="emmettcarri">
<emphasis id="B94E7C2AFFE7FFF222C6BF57387236EB" box="[753,890,1913,1938]" italics="true" pageId="9" pageNumber="378">emmettcarri</emphasis>
</taxonomicName>
” (
<collectionCode id="ED2B38FDFFE7FFF223A1BF5638FC36EB" box="[918,1012,1912,1938]" pageId="9" pageNumber="378">FLREC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B52AA65FFE7FFF22431BF563ABF36CF" ID-GBIF-Occurrence="3019651306" collectingDate="2019-06-22" collectionCode="FSCA" collectorName="B. W. Bahder" country="Costa Rica" location="La Selva Biological Station" pageId="9" pageNumber="378" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Heredia" typeStatus="paratype">
<typeStatus id="54811E9AFFE7FFF22431BF563F7E36EB" box="[1030,1142,1912,1938]" pageId="9" pageNumber="378" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D3C6BB1FFE7FFF224B5BF563FD336EB" box="[1154,1243,1912,1938]" pageId="9" pageNumber="378" type="male">2 males</specimenCount>
,
<specimenCount id="9D3C6BB1FFE7FFF224D0BF563E5F36EB" box="[1255,1367,1912,1938]" pageId="9" pageNumber="378" type="female">3 females</specimenCount>
, same data as holotype, (
<collectionCode id="ED2B38FDFFE7FFF22154BFB23AA736CF" box="[355,431,1948,1974]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="9" pageNumber="378" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF45F0B0FFE4FFF120A0BD663BC634FD" ID-DOI="http://doi.org/10.5281/zenodo.4441926" ID-Zenodo-Dep="4441926" httpUri="https://zenodo.org/record/4441926/files/figure.png" pageId="10" pageNumber="379" startId="10.[151,250,1352,1376]" targetBox="[188,1372,197,1328]" targetPageId="10">
<paragraph id="8B85A038FFE4FFF120A0BD663BC634FD" blockId="10.[151,1436,1352,1412]" pageId="10" pageNumber="379">
<emphasis id="B94E7C2AFFE4FFF120A0BD663A1C3419" bold="true" box="[151,276,1352,1376]" pageId="10" pageNumber="379">FIGURE 9.</emphasis>
Aedeagus of adult male
<taxonomicName id="4C3ADBBBFFE4FFF12221BD67394A3419" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[534,578,1353,1376]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E7C2AFFE4FFF12221BD67394A3419" box="[534,578,1353,1376]" italics="true" pageId="10" pageNumber="379">Tico</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE4FFF1227EBD6639993419" bold="true" box="[585,657,1352,1376]" pageId="10" pageNumber="379">
<taxonomicNameLabel id="A27DC151FFE4FFF1227EBD6639993419" box="[585,657,1352,1376]" pageId="10" pageNumber="379" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C3ADBBBFFE4FFF122AEBD67381D3419" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[665,789,1353,1376]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="emmettcarri" status="sp. nov.">
<emphasis id="B94E7C2AFFE4FFF122AEBD67381D3419" box="[665,789,1353,1376]" italics="true" pageId="10" pageNumber="379">emmettcarri</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE4FFF1232ABD66385F3419" bold="true" box="[797,855,1352,1376]" pageId="10" pageNumber="379">
<taxonomicNameLabel id="A27DC151FFE4FFF1232ABD66385F3419" box="[797,855,1352,1376]" pageId="10" pageNumber="379" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; A) right lateral view, B) left lateral view, and C) dorsal view.
</paragraph>
</caption>
<subSubSection id="C320F3B3FFE4FFF120F0BD9D3AD23690" pageId="10" pageNumber="379" type="discussion">
<paragraph id="8B85A038FFE4FFF120F0BD9D39033605" blockId="10.[151,1437,1458,2025]" pageId="10" pageNumber="379">
<emphasis id="B94E7C2AFFE4FFF120F0BD9D3A3334B4" bold="true" box="[199,315,1459,1485]" pageId="10" pageNumber="379">Remarks.</emphasis>
<taxonomicName id="4C3ADBBBFFE4FFF12174BD9D390D34B5" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[323,517,1459,1484]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="emmettcarri" status="sp. nov.">
<emphasis id="B94E7C2AFFE4FFF12174BD9D390D34B5" box="[323,517,1459,1484]" italics="true" pageId="10" pageNumber="379">Tico emmettcarri</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE4FFF1223ABD9D394634B4" bold="true" box="[525,590,1459,1485]" pageId="10" pageNumber="379">
<taxonomicNameLabel id="A27DC151FFE4FFF1223ABD9D394634B4" box="[525,590,1459,1485]" pageId="10" pageNumber="379" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be readily separated from the other two species in
<taxonomicName id="4C3ADBBBFFE4FFF124F2BD9D3FFD34B5" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[1221,1269,1459,1484]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E7C2AFFE4FFF124F2BD9D3FFD34B5" box="[1221,1269,1459,1484]" italics="true" pageId="10" pageNumber="379">Tico</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE4FFF124CABD9D3E4634B4" bold="true" box="[1277,1358,1459,1485]" pageId="10" pageNumber="379">
<taxonomicNameLabel id="A27DC151FFE4FFF124CABD9D3E4634B4" box="[1277,1358,1459,1485]" pageId="10" pageNumber="379" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
by the nature of the markings on the wings and the lack of a black marking in front of the eyes. The collective number of branches of RP and CuA reaching the wing margin is 8, such that an additional cell (relative to other included species) is defined (here identified as C3a,
<figureCitation id="1301BCBDFFE4FFF1224CBE3039B73740" box="[635,703,1566,1593]" captionStart="FIGURE 6" captionStartId="7.[151,250,1543,1567]" captionTargetBox="[151,1430,418,1520]" captionTargetId="figure-133@7.[150,1437,417,1520]" captionTargetPageId="7" captionText="FIGURE 6. Male Tico gen. n. emmettcarri sp. n. wing venation; black = vein, green = cell." figureDoi="http://doi.org/10.5281/zenodo.4441920" httpUri="https://zenodo.org/record/4441920/files/figure.png" pageId="10" pageNumber="379">Fig. 6</figureCitation>
). The species is also distinctive in possessing proximal lobes and a dorsal sclerotized ridge on the gonostyli, the aedeagal shaft is arched upward near midlength and devoid of processes (although dorsal flange visible below retrorse flagellum, flange dorsally serrate from left view). The anal tube is elongate, distally downcurved and apically truncate. In contrast, both
<taxonomicName id="4C3ADBBBFFE4FFF123F5BEA53FAE37DD" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[962,1190,1674,1700]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="pseudosororius" status="sp. nov.">
<emphasis id="B94E7C2AFFE4FFF123F5BEA53FAE37DD" box="[962,1190,1674,1700]" italics="true" pageId="10" pageNumber="379">Tico pseudosororius</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE4FFF1249ABEA53FE537DC" bold="true" box="[1197,1261,1675,1701]" pageId="10" pageNumber="379">
<taxonomicNameLabel id="A27DC151FFE4FFF1249ABEA53FE537DC" box="[1197,1261,1675,1701]" pageId="10" pageNumber="379" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C3ADBBBFFE4FFF12513BEA53BF537B0" authority="(Fennah)" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" baseAuthorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" baseAuthorityYear="2021" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="sororius" status="comb. nov.">
<emphasis id="B94E7C2AFFE4FFF12513BEA53E9437DD" box="[1316,1436,1675,1700]" italics="true" pageId="10" pageNumber="379">T. sororius</emphasis>
(Fennah)
</taxonomicName>
<emphasis id="B94E7C2AFFE4FFF12134BE813A6237B0" bold="true" box="[259,362,1711,1737]" pageId="10" pageNumber="379">
<taxonomicNameLabel id="A27DC151FFE4FFF12134BE813A6237B0" box="[259,362,1711,1737]" pageId="10" pageNumber="379" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
bear a row of 4 spots on the wing and have a dark marking in front of the eyes. Also, in both of these species, the cumulative number of veins reaching the wing margin from RP and CuA is 7, such that cell C3a is not formed (compare
<figureCitation id="1301BCBDFFE4FFF1219CBED83AE73669" box="[427,495,1782,1808]" captionStart="FIGURE 6" captionStartId="7.[151,250,1543,1567]" captionTargetBox="[151,1430,418,1520]" captionTargetId="figure-133@7.[150,1437,417,1520]" captionTargetPageId="7" captionText="FIGURE 6. Male Tico gen. n. emmettcarri sp. n. wing venation; black = vein, green = cell." figureDoi="http://doi.org/10.5281/zenodo.4441920" httpUri="https://zenodo.org/record/4441920/files/figure.png" pageId="10" pageNumber="379">Fig. 6</figureCitation>
with
<figureCitation id="1301BCBDFFE4FFF12205BED8398E3668" box="[562,646,1782,1809]" captionStart="FIGURE 14" captionStartId="14.[151,250,1173,1197]" captionTargetBox="[163,1421,183,1145]" captionTargetId="figure-25@14.[162,1426,181,1150]" captionTargetPageId="14" captionText="FIGURE 14. Male Tico gen. n. pseudosororius sp. n. wing venation; black = vein, green = cell." figureDoi="http://doi.org/10.5281/zenodo.4441938" httpUri="https://zenodo.org/record/4441938/files/figure.png" pageId="10" pageNumber="379">Fig. 14</figureCitation>
).
<taxonomicName id="4C3ADBBBFFE4FFF122ABBED938893669" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[668,897,1782,1808]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="pseudosororius" status="sp. nov.">
<emphasis id="B94E7C2AFFE4FFF122ABBED938893669" box="[668,897,1782,1808]" italics="true" pageId="10" pageNumber="379">Tico pseudosororius</emphasis>
</taxonomicName>
<emphasis id="B94E7C2AFFE4FFF123BDBED938C33668" bold="true" box="[906,971,1783,1809]" pageId="10" pageNumber="379">
<taxonomicNameLabel id="A27DC151FFE4FFF123BDBED938C33668" box="[906,971,1783,1809]" pageId="10" pageNumber="379" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has evident differences in the terminalia including the dorsal lobe of the gonostyli is thumb-like (dorsal flange absent); also, the longest retrorse process on the aedeagus in dorsal view is a large flattened flange bearing a few large teeth. The anal tube is elongate, distally downcurved into rounded apices.
</paragraph>
<paragraph id="8B85A038FFE4FFF120F0BFA83AD23690" blockId="10.[151,1437,1458,2025]" pageId="10" pageNumber="379">
The male terminalia of
<taxonomicName id="4C3ADBBBFFE4FFF121FEBFA9395336D9" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" baseAuthorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" baseAuthorityYear="2021" box="[457,603,1927,1952]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="sororius">
<emphasis id="B94E7C2AFFE4FFF121FEBFA9395336D9" box="[457,603,1927,1952]" italics="true" pageId="10" pageNumber="379">Tico sororius</emphasis>
</taxonomicName>
are not known. When discovered, we anticipate that they will have features in common with described species in
<taxonomicName id="4C3ADBBBFFE4FFF12200BF85396F36BD" authority="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett, 2021" authorityName="Bahder &amp; Zumbado Echavarria &amp; Barrantes Barrantes &amp; Helmick &amp; Bartlett" authorityYear="2021" box="[567,615,1963,1988]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E7C2AFFE4FFF12200BF85396F36BD" box="[567,615,1963,1988]" italics="true" pageId="10" pageNumber="379">Tico</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DC151FFE4FFF1225ABF8539B236BC" box="[621,698,1963,1989]" pageId="10" pageNumber="379" rank="genus">
<emphasis id="B94E7C2AFFE4FFF1225ABF8539BC36BC" bold="true" box="[621,692,1963,1989]" pageId="10" pageNumber="379">gen. n</emphasis>
.
</taxonomicNameLabel>
, and their description would further elaborate genus-level features genera allied with
<taxonomicName id="4C3ADBBBFFE4FFF12150BFE03ADB3691" authorityName="Westwood. The" authorityYear="1840" box="[359,467,1998,2024]" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7C2AFFE4FFF12150BFE03ADB3691" box="[359,467,1998,2024]" italics="true" pageId="10" pageNumber="379">Cenchrea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>