331 lines
41 KiB
XML
331 lines
41 KiB
XML
<document id="EEA0FE8E8B9334CD4702A458F926C093" ID-CLB-Dataset="8781" ID-DOI="10.11646/zootaxa.4908.3.3" ID-GBIF-Dataset="de928cf2-80ea-4ede-ad12-3ac0300b1ae9" ID-ISSN="1175-5326" ID-Zenodo-Dep="4441906" ID-ZooBank="1A58171C-5BD8-4837-8708-3A1C7832DA8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610701278547" checkinUser="plazi" docAuthor="Bahder, Brian W., Zumbado Echavarria, Marco A., Barrantes Barrantes, Edwin A., Helmick, Ericka E. & Bartlett, Charles R." docDate="2021" docId="0393112EFFEAFFFE20A0BC9D3F0F3474" docLanguage="en" docName="zootaxa.4908.3.3.pdf" docOrigin="Zootaxa 4908 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tico Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett 2021, gen. n." docType="treatment" docVersion="7" lastPageNumber="374" masterDocId="FFAA6956FFEEFFFB2037B82E3B083179" masterDocTitle="A new genus and two new species of planthopper in the tribe Cenchreini (Hemiptera: Auchenorrhyncha: Derbidae) from lowland tropical rainforest in Costa Rica" masterLastPageNumber="392" masterPageNumber="369" pageNumber="373" updateTime="1698924501363" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="F57CCA9BCF55F984FEF966E29A11AA85">
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<mods:title id="376383EC19A92AF5C093A51AC4B58695">A new genus and two new species of planthopper in the tribe Cenchreini (Hemiptera: Auchenorrhyncha: Derbidae) from lowland tropical rainforest in Costa Rica</mods:title>
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<mods:name id="02660EF6BED26C6EC87DBF7CE94F97D0" type="personal">
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<mods:roleTerm id="654B0DA81FF9EEF608A32A99355B0DFB">Author</mods:roleTerm>
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<mods:namePart id="1CD6531B05E9BBF2CDB1656DEC9559FD">Bahder, Brian W.</mods:namePart>
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<mods:affiliation id="D708DEB3A9ACE5C7079919671900393B">University of Florida, Department of Entomology and Nematology-Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA.</mods:affiliation>
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<mods:role id="33AFDF537D00876D1F87E27D97BC3C67">
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<mods:roleTerm id="A3F5B724AE248D2879242C84AA807FAC">Author</mods:roleTerm>
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<mods:namePart id="A5119F52ABD4CFB3AAE96EE748A9ADF6">Zumbado Echavarria, Marco A.</mods:namePart>
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<mods:affiliation id="0B46BE7D764811FCA7FA5C875D1C464B">Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica.</mods:affiliation>
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<mods:name id="B2701D8EC9B350CE775884945D068AFE" type="personal">
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<mods:roleTerm id="EA6B649AC411DB47C1CACF2F8F492935">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2187108198C7B22B4DCAD3B242C4F68A">Barrantes Barrantes, Edwin A.</mods:namePart>
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<mods:affiliation id="1C1BD8867EDB35A70147085547D2F0FD">Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica.</mods:affiliation>
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</mods:name>
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<mods:namePart id="14880960C75606BB2B2C878F4DC25862">Helmick, Ericka E.</mods:namePart>
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<mods:affiliation id="AC3AFE2D9D9CA87ED16D2FAB392C76D5">University of Florida, Department of Entomology and Nematology-Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA.</mods:affiliation>
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</mods:role>
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<mods:namePart id="F4781AD6676ED07D9EA3DC4DEDBC367A">Bartlett, Charles R.</mods:namePart>
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<mods:affiliation id="CFCAA5E6F62221F0922BA7819476242A">University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716 - 2160, USA.</mods:affiliation>
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<mods:typeOfResource id="E5565FA0C3D90E20CECD21CB626F4AF2">text</mods:typeOfResource>
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<mods:title id="99369C096A4F964E78E0E1005D6C3EB1">Zootaxa</mods:title>
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<mods:part id="30C622DA334D02139CBEDB0017680439">
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<mods:date id="DFA8BB7ED58F4D53DB685DB9A4C1AE6D">2021</mods:date>
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<mods:number id="E660E51108FC3BD711FDC976C615209B">2021-01-15</mods:number>
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<mods:number id="1E76A7A8830C7A64078172896FE0764F">4908</mods:number>
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<mods:detail id="DB6AEB2B6D0FF857325E937B26C58480" type="issue">
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<mods:number id="8F35AC85F677FA8C26852572A80800A2">3</mods:number>
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<mods:classification id="684AECEFDDD5692A0DEE37942080F5ED">journal article</mods:classification>
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<mods:identifier id="A0E2E691F5A63C91209086D825D32C60" type="CLB-Dataset">8781</mods:identifier>
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<mods:identifier id="46282BC9269D229EAB542BE0522E4736" type="DOI">10.11646/zootaxa.4908.3.3</mods:identifier>
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<mods:identifier id="47DE9127660C053DB9C673EFFD155F45" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="054F2ECA2717AA1FCE6410C497D76EC4" type="Zenodo-Dep">4441906</mods:identifier>
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<treatment id="0393112EFFEAFFFE20A0BC9D3F0F3474" ID-DOI="http://doi.org/10.5281/zenodo.4450845" ID-GBIF-Taxon="177009916" ID-Zenodo-Dep="4450845" LSID="urn:lsid:plazi:treatment:0393112EFFEAFFFE20A0BC9D3F0F3474" httpUri="http://treatment.plazi.org/id/0393112EFFEAFFFE20A0BC9D3F0F3474" lastPageId="5" lastPageNumber="374" pageId="4" pageNumber="373">
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<subSubSection id="C320F3B3FFEAFFFF20A0BC9D391235B4" box="[151,538,1202,1229]" pageId="4" pageNumber="373" type="nomenclature">
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<paragraph id="8B85A038FFEAFFFF20A0BC9D391235B4" blockId="4.[151,538,1202,1229]" box="[151,538,1202,1229]" pageId="4" pageNumber="373">
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<heading id="D0CD1754FFEAFFFF20A0BC9D391235B4" bold="true" box="[151,538,1202,1229]" fontSize="11" level="1" pageId="4" pageNumber="373" reason="1">
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<emphasis id="B94E7C2AFFEAFFFF20A0BC9D391235B4" bold="true" box="[151,538,1202,1229]" pageId="4" pageNumber="373">
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||
<taxonomicName id="4C3ADBBBFFEAFFFF20A0BC9D3AB235B4" ID-CoL="9DNFB" authority="Bahder & Bartlett" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[151,442,1202,1229]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B94E7C2AFFEAFFFF20A0BC9D3BC435B4" bold="true" box="[151,204,1203,1229]" italics="true" pageId="4" pageNumber="373">Tico</emphasis>
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Bahder & Bartlett
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27DC151FFEAFFFF21F0BC9D391235B4" box="[455,538,1203,1229]" pageId="4" pageNumber="373" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C320F3B3FFEAFFFF20A0BCDC39DB3473" box="[151,723,1266,1290]" pageId="4" pageNumber="373" type="type_taxon">
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<paragraph id="8B85A038FFEAFFFF20A0BCDC39DB3473" blockId="4.[151,723,1266,1290]" box="[151,723,1266,1290]" pageId="4" pageNumber="373">
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<emphasis id="B94E7C2AFFEAFFFF20A0BCDC3A2B3473" bold="true" box="[151,291,1266,1290]" pageId="4" pageNumber="373">
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<typeStatus id="54811E9AFFEAFFFF20A0BCDC3BC43473" box="[151,204,1266,1290]" pageId="4" pageNumber="373">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C3ADBBBFFEAFFFF211DBCDD399B3473" ID-CoL="9FNPB" authority="Bahder & Bartlett" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[298,659,1266,1290]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="emmettcarri" status="sp. nov.">
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<emphasis id="B94E7C2AFFEAFFFF211DBCDD3AD03473" box="[298,472,1267,1290]" italics="true" pageId="4" pageNumber="373">Tico emmettcarri</emphasis>
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Bahder & Bartlett
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</taxonomicName>
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||
<emphasis id="B94E7C2AFFEAFFFF22ADBCDC39DB3473" bold="true" box="[666,723,1266,1290]" pageId="4" pageNumber="373">
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<taxonomicNameLabel id="A27DC151FFEAFFFF22ADBCDC39DB3473" box="[666,723,1266,1290]" pageId="4" pageNumber="373" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C320F3B3FFEAFFFF20A0BD1C390D3714" pageId="4" pageNumber="373" type="diagnosis">
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||
<paragraph id="8B85A038FFEAFFFF20A0BD1C390D3714" blockId="4.[151,1437,1330,2041]" pageId="4" pageNumber="373">
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<emphasis id="B94E7C2AFFEAFFFF20A0BD1C3A1A3435" bold="true" box="[151,274,1330,1356]" pageId="4" pageNumber="373">Diagnosis.</emphasis>
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Small (~
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<quantity id="4CC20DDDFFEAFFFF214BBD1D3AB73434" box="[380,447,1331,1357]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="373" unit="mm" value="3.0">3 mm</quantity>
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,
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<tableCitation id="C6B89583FFEAFFFF21FBBD1D39243434" box="[460,556,1331,1357]" captionStart="TABLE 2" captionStartId="12.[151,239,151,175]" captionText="TABLE 2. Biometric data for Tico emmettcarri gen. et. sp. n.; values = mm." pageId="4" pageNumber="373">Tables 2</tableCitation>
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,
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<tableCitation id="C6B89583FFEAFFFF220DBD1D39413434" box="[570,585,1331,1357]" captionStart="TABLE 3" captionStartId="17.[151,239,1316,1340]" captionText="TABLE 3. Biometric data for Tico pseudosororius gen. et. sp. n.; values = mm." pageId="4" pageNumber="373">3</tableCitation>
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), pale cenchreine derbids with variably marked, clear wings. Frons moderately compressed (length at midline about 4x dorsal width), median carina absent, with transverse carinae near fastigium separating frons from clypeus; lateral carinae of frons and clypeus keeled, bearing row of sensory pustules. Forewing with ScP branch preceding apex of clavus (resulting in long marginal cell). Pygofer with distinct expansion (but not projections) on lateral margins, ventral margin of opening without lobe. Gonostyli long and spatulate, apex rounded, with a medially directed dorsal lobe near midlength (lobes absent on ventral margin). Aedeagus nearly bilaterally symmetrical, lacking subapical projections on shaft, terminating in complex array of retrorse processes. Anal tube elongate, in lateral view ventral margin weakly concave, apex inflected downward, apex truncate; in dorsal view broadly spatulate.
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</paragraph>
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</subSubSection>
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<subSubSection id="C320F3B3FFEAFFFE20F0BE5838F43180" lastPageId="5" lastPageNumber="374" pageId="4" pageNumber="373" type="description">
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<paragraph id="8B85A038FFEAFFFE20F0BE5838F43180" blockId="4.[151,1437,1330,2041]" lastBlockId="5.[151,1437,151,1293]" lastPageId="5" lastPageNumber="374" pageId="4" pageNumber="373">
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<emphasis id="B94E7C2AFFEAFFFF20F0BE583A5137E9" bold="true" box="[199,345,1654,1680]" pageId="4" pageNumber="373">Description.</emphasis>
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Small, pale species (length with wings
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<quantity id="4CC20DDDFFEAFFFF231BBE5938DA37E8" box="[812,978,1655,1681]" metricMagnitude="-3" metricUnit="m" metricValue="3.04" metricValueMax="3.1" metricValueMin="2.98" pageId="4" pageNumber="373" unit="mm" value="3.04" valueMax="3.1" valueMin="2.98">2.98–3.10 mm</quantity>
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for included species), with clear wings bearing dark markings. Head with frons moderately compressed (median carina absent), narrowest between eyes, widest at frontoclypeal suture, bearing transverse carina separating frons and vertex (usually marked by small visible hump in lateral view). Lateral carinae of vertex and frons strongly keeled, bearing a row of pits. Vertex roughly triangular, narrowed distally, anterior margin concave between carinae, posterior margin strongly concave between eyes. Pronotum tricarinate on disc (median carina weak), elevated posteriorly; at midline subequal in length to vertex; anteriorly convex, following contours of head, posteriorly weakly concave; lateral paradiscal areas expansively developed into foliations surrounding (and broadly overtopping) the antennae posteriorly. Mesonotum tricarinate, wider than long. Post-tibia lacking lateral spines, apical spinulation (8-7)-(7-6)-6. Forewings broad, clavus apex near midlength; Sc branch just proximad of claval apex; forking of CuA proximad of fork of Sc+RA with RP. Terminalia bilaterally symmetrical or nearly so. Pygofer narrow in lateral view, opening caudally projected on lateral margins (lacking projection on ventral margin). Gonostyli spatulate with dorsal, medially directed projection near midlength, apex rounded. Aedeagus symmetrical, complex endosoma, shaft simple (lacking processes). Anal tube elongate in lateral view, apex truncate and arched downward, paraproct short.
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</paragraph>
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</subSubSection>
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<subSubSection id="C320F3B3FFEBFFFE20F0B92D39D335BC" pageId="5" pageNumber="374" type="discussion">
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<paragraph id="8B85A038FFEBFFFE20F0B92D3A3130D4" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="374">
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<emphasis id="B94E7C2AFFEBFFFE20F0B92D3A333064" bold="true" box="[199,315,259,285]" pageId="5" pageNumber="374">Remarks.</emphasis>
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<taxonomicName id="4C3ADBBBFFEBFFFE2174B92D3A7B3065" authority="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett, 2021" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[323,371,259,284]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B94E7C2AFFEBFFFE2174B92D3A7B3065" box="[323,371,259,284]" italics="true" pageId="5" pageNumber="374">Tico</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B94E7C2AFFEBFFFE214DB92D3AC23064" bold="true" box="[378,458,259,285]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="A27DC151FFEBFFFE214DB92D3AC23064" box="[378,458,259,285]" pageId="5" pageNumber="374" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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as comprised here is composed of three species
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<taxonomicName id="4C3ADBBBFFEBFFFE23DAB92D3FA63065" authority="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett, 2021" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[1005,1198,259,284]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="emmettcarri" status="sp. nov.">
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<emphasis id="B94E7C2AFFEBFFFE23DAB92D3FA63065" box="[1005,1198,259,284]" italics="true" pageId="5" pageNumber="374">Tico emmettcarri</emphasis>
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</taxonomicName>
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||
<emphasis id="B94E7C2AFFEBFFFE2482B92D3FF33064" bold="true" box="[1205,1275,259,285]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="A27DC151FFEBFFFE2482B92D3FFD3064" box="[1205,1269,259,285]" pageId="5" pageNumber="374" rank="species">sp. n.</taxonomicNameLabel>
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||
,
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||
</emphasis>
|
||
(the
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<typeStatus id="54811E9AFFEBFFFE2501B92D3E6E3064" box="[1334,1382,259,285]" pageId="5" pageNumber="374">type</typeStatus>
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||
species, described below),
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<taxonomicName id="4C3ADBBBFFEBFFFE21ACB90939743038" authority="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett, 2021" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[411,636,295,321]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="pseudosororius" status="sp. nov.">
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<emphasis id="B94E7C2AFFEBFFFE21ACB90939743038" box="[411,636,295,321]" italics="true" pageId="5" pageNumber="374">Tico pseudosororius</emphasis>
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</taxonomicName>
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||
<emphasis id="B94E7C2AFFEBFFFE22B5B90939C83038" bold="true" box="[642,704,295,321]" pageId="5" pageNumber="374">
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||
<taxonomicNameLabel id="A27DC151FFEBFFFE22B5B90939C83038" box="[642,704,295,321]" pageId="5" pageNumber="374" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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(described below), and
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<taxonomicName id="4C3ADBBBFFEBFFFE23FFB9093FCD303B" authority="(Fennah)" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" baseAuthorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" baseAuthorityYear="2021" box="[968,1221,295,322]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="sororius" status="comb. nov.">
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<emphasis id="B94E7C2AFFEBFFFE23FFB9093F513039" box="[968,1113,295,320]" italics="true" pageId="5" pageNumber="374">Tico sororius</emphasis>
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(Fennah)
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</taxonomicName>
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<emphasis id="B94E7C2AFFEBFFFE24FDB9093E943039" bold="true" box="[1226,1436,294,321]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="A27DC151FFEBFFFE24FDB9093E903039" box="[1226,1432,294,321]" pageId="5" pageNumber="374" rank="species">new combination</taxonomicNameLabel>
|
||
.
|
||
</emphasis>
|
||
Our molecular data supports
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE21E8B96539A8301D" authority="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett, 2021" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[479,672,331,356]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="emmettcarri" status="sp. nov.">
|
||
<emphasis id="B94E7C2AFFEBFFFE21E8B96539A8301D" box="[479,672,331,356]" italics="true" pageId="5" pageNumber="374">Tico emmettcarri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94E7C2AFFEBFFFE229FB96539E0301C" bold="true" box="[680,744,331,357]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="A27DC151FFEBFFFE229FB96539E0301C" box="[680,744,331,357]" pageId="5" pageNumber="374" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE2310B9653F03301C" authority="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett, 2021" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[807,1035,331,357]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="pseudosororius" status="sp. nov.">
|
||
<emphasis id="B94E7C2AFFEBFFFE2310B9653F03301C" box="[807,1035,331,357]" italics="true" pageId="5" pageNumber="374">Tico pseudosororius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94E7C2AFFEBFFFE2425B9653F5B301C" bold="true" box="[1042,1107,331,357]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="A27DC151FFEBFFFE2425B9653F5B301C" box="[1042,1107,331,357]" pageId="5" pageNumber="374" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
as a clade.
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE24E0B9653BE130F0" authority="Fennah" authorityName="Fennah. The" authorityYear="1952" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="sororia">
|
||
<emphasis id="B94E7C2AFFEBFFFE24E0B9653E94301D" box="[1239,1436,331,357]" italics="true" pageId="5" pageNumber="374">Cenchrea sororia</emphasis>
|
||
Fennah
|
||
</taxonomicName>
|
||
is here transferred into
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE21CBB941392430F1" authority="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett, 2021" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[508,556,367,392]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94E7C2AFFEBFFFE21CBB941392430F1" box="[508,556,367,392]" italics="true" pageId="5" pageNumber="374">Tico</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94E7C2AFFEBFFFE2202B941398E30F0" bold="true" box="[565,646,367,393]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="A27DC151FFEBFFFE2202B941398E30F0" box="[565,646,367,393]" pageId="5" pageNumber="374" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
to give the
|
||
<emphasis id="B94E7C2AFFEBFFFE2327B94138D630F1" bold="true" box="[784,990,366,393]" pageId="5" pageNumber="374">new combination</emphasis>
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE23D0B9413FE230F3" authority="(Fennah)" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" baseAuthorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" baseAuthorityYear="2021" box="[999,1258,367,394]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="sororius">
|
||
<emphasis id="B94E7C2AFFEBFFFE23D0B9413F7430F1" box="[999,1148,367,392]" italics="true" pageId="5" pageNumber="374">Tico sororius</emphasis>
|
||
(Fennah)
|
||
</taxonomicName>
|
||
for reasons described below.
|
||
</paragraph>
|
||
<paragraph id="8B85A038FFEBFFFE20F0B9993A4432DC" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="374">
|
||
The species described here in
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE2223B999394C30A9" authority="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett, 2021" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[532,580,439,464]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94E7C2AFFEBFFFE2223B999394C30A9" box="[532,580,439,464]" italics="true" pageId="5" pageNumber="374">Tico</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94E7C2AFFEBFFFE227DB999399030A8" bold="true" box="[586,664,439,465]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="A27DC151FFEBFFFE227DB999399030A8" box="[586,664,439,465]" pageId="5" pageNumber="374" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
initially appeared to belong to
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE23C6B9993F5530A8" authorityName="Westwood. The" authorityYear="1840" box="[1009,1117,439,465]" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94E7C2AFFEBFFFE23C6B9993F5530A8" box="[1009,1117,439,465]" italics="true" pageId="5" pageNumber="374">Cenchrea</emphasis>
|
||
</taxonomicName>
|
||
. Results of molecular analyses and consideration of some comments of
|
||
<bibRefCitation id="EFABDDC9FFEBFFFE22A4B9F53831308C" author="Fennah, R. G." box="[659,825,474,501]" pageId="5" pageNumber="374" pagination="109 - 170" refId="ref12306" refString="Fennah, R. G. (1952) On the generic classification of Derbidae (Fulgoroidea), with descriptions of new Neotropical species. Transactions of the Royal Entomological Society of London, 103 (4), 109 - 170. https: // doi. org / 10.1111 / j. 1365 - 2311.1952. tb 01063. x" type="journal article" year="1952">Fennah (1952)</bibRefCitation>
|
||
led us to describe these taxa in a new genus.
|
||
<bibRefCitation id="EFABDDC9FFEBFFFE257DB9F53BEA3360" author="Fennah, R. G." pageId="5" pageNumber="374" pagination="109 - 170" refId="ref12306" refString="Fennah, R. G. (1952) On the generic classification of Derbidae (Fulgoroidea), with descriptions of new Neotropical species. Transactions of the Royal Entomological Society of London, 103 (4), 109 - 170. https: // doi. org / 10.1111 / j. 1365 - 2311.1952. tb 01063. x" type="journal article" year="1952">Fennah (1952)</bibRefCitation>
|
||
emphasized the distinctness of
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE227FB9D139CB3360" authorityName="Westwood" authorityYear="1840" box="[584,707,511,537]" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="dorsalis">
|
||
<emphasis id="B94E7C2AFFEBFFFE227FB9D139CB3360" box="[584,707,511,537]" italics="true" pageId="5" pageNumber="374">C. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
(the
|
||
<typeStatus id="54811E9AFFEBFFFE22C9B9D138263360" box="[766,814,511,537]" pageId="5" pageNumber="374">type</typeStatus>
|
||
species) relative to the remaining species in
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE251EB9D13E9D3360" authorityName="Westwood. The" authorityYear="1840" box="[1321,1429,511,537]" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94E7C2AFFEBFFFE251EB9D13E9D3360" box="[1321,1429,511,537]" italics="true" pageId="5" pageNumber="374">Cenchrea</emphasis>
|
||
</taxonomicName>
|
||
, and suggested that
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE2145BA0D3AE73344" authorityName="Westwood" authorityYear="1840" box="[370,495,547,573]" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="dorsalis">
|
||
<emphasis id="B94E7C2AFFEBFFFE2145BA0D3AE73344" box="[370,495,547,573]" italics="true" pageId="5" pageNumber="374">C. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
and the remaining taxa might be placed in different subgenera. Study of available
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE20A0BA693A0B3318" authorityName="Westwood. The" authorityYear="1840" box="[151,259,583,609]" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94E7C2AFFEBFFFE20A0BA693A0B3318" box="[151,259,583,609]" italics="true" pageId="5" pageNumber="374">Cenchrea</emphasis>
|
||
</taxonomicName>
|
||
specimens, along with available molecular data seems to support
|
||
<bibRefCitation id="EFABDDC9FFEBFFFE23CDBA693FBB3318" author="Fennah, R. G." box="[1018,1203,582,609]" pageId="5" pageNumber="374" pagination="109 - 170" refId="ref12306" refString="Fennah, R. G. (1952) On the generic classification of Derbidae (Fulgoroidea), with descriptions of new Neotropical species. Transactions of the Royal Entomological Society of London, 103 (4), 109 - 170. https: // doi. org / 10.1111 / j. 1365 - 2311.1952. tb 01063. x" type="journal article" year="1952">Fennah’s (1952)</bibRefCitation>
|
||
assertion. However, because specimens (or photos) of
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE2222BA45385B33FF" authority="(Muir)" baseAuthorityName="Muir" baseAuthorityYear="1918" box="[533,851,619,646]" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="bipunctata">
|
||
<emphasis id="B94E7C2AFFEBFFFE2222BA45380933FC" box="[533,769,619,645]" italics="true" pageId="5" pageNumber="374">Cenchrea bipunctata</emphasis>
|
||
(Muir)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE23BDBA453F7F33FC" authority="Fennah" authorityName="Fennah" authorityYear="1952" box="[906,1143,619,645]" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="sexguttata">
|
||
<emphasis id="B94E7C2AFFEBFFFE23BDBA453F1633FD" box="[906,1054,619,645]" italics="true" pageId="5" pageNumber="374">C. sexguttata</emphasis>
|
||
Fennah
|
||
</taxonomicName>
|
||
(the latter described from females) are not available to us, we are reluctant to speculative on their generic placement. For
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE24FEBAA13E9133D0" authority="Uhler" authorityName="Uhler" authorityYear="1895" box="[1225,1433,655,681]" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="exquisita">
|
||
<emphasis id="B94E7C2AFFEBFFFE24FEBAA13E4733D1" box="[1225,1359,655,681]" italics="true" pageId="5" pageNumber="374">C. exquisita</emphasis>
|
||
Uhler
|
||
</taxonomicName>
|
||
, differences in morphology (viz. the shape of the face and the presence of large falciform processes on the lateral margins of the pygofer opening, absent in both species placed in
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE2355BAF9389A3389" authority="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett, 2021" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[866,914,727,752]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94E7C2AFFEBFFFE2355BAF9389A3389" box="[866,914,727,752]" italics="true" pageId="5" pageNumber="374">Tico</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94E7C2AFFEBFFFE23AEBAF938EE3388" bold="true" box="[921,998,727,753]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="A27DC151FFEBFFFE23AEBAF938EE3388" box="[921,998,727,753]" pageId="5" pageNumber="374" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) would require a much more expansive definition of
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE2110BAD53A5F326D" authority="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett, 2021" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[295,343,763,788]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94E7C2AFFEBFFFE2110BAD53A5F326D" box="[295,343,763,788]" italics="true" pageId="5" pageNumber="374">Tico</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94E7C2AFFEBFFFE216ABAD53AA2326C" bold="true" box="[349,426,763,789]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="A27DC151FFEBFFFE216ABAD53AA2326C" box="[349,426,763,789]" pageId="5" pageNumber="374" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
than we were comfortable making in the absence of supporting molecular data (sequencing available specimens was not successful). In contrast, we place
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE2362BB313F113241" authorityName="Fennah. The" authorityYear="1952" box="[853,1049,799,825]" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="sororia">
|
||
<emphasis id="B94E7C2AFFEBFFFE2362BB313F113241" box="[853,1049,799,825]" italics="true" pageId="5" pageNumber="374">Cenchrea sororia</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE2409BB313F663241" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[1086,1134,799,824]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94E7C2AFFEBFFFE2409BB313F663241" box="[1086,1134,799,824]" italics="true" pageId="5" pageNumber="374">Tico</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE24AABB313E953243" authority="(Fennah)" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" baseAuthorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" baseAuthorityYear="2021" box="[1181,1437,799,826]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="sororius">
|
||
<emphasis id="B94E7C2AFFEBFFFE24AABB313E383241" box="[1181,1328,799,824]" italics="true" pageId="5" pageNumber="374">Tico sororius</emphasis>
|
||
(Fennah)
|
||
</taxonomicName>
|
||
<emphasis id="B94E7C2AFFEBFFFE20A0BB6D3BD73224" bold="true" box="[151,223,835,861]" pageId="5" pageNumber="374">comb.</emphasis>
|
||
) because of its unmistakable similarity to
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE228CBB6D388A3224" authority="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett, 2021" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[699,898,835,861]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="pseudosororius" status="sp. nov.">
|
||
<emphasis id="B94E7C2AFFEBFFFE228CBB6D388A3224" box="[699,898,835,861]" italics="true" pageId="5" pageNumber="374">T. pseudosororius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94E7C2AFFEBFFFE23BEBB6D38C03224" bold="true" box="[905,968,835,861]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="A27DC151FFEBFFFE23BEBB6D38C03224" box="[905,968,835,861]" pageId="5" pageNumber="374" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(photos of
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE247FBB6D3FD43225" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" baseAuthorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" baseAuthorityYear="2021" box="[1096,1244,835,860]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="sororius">
|
||
<emphasis id="B94E7C2AFFEBFFFE247FBB6D3FD43225" box="[1096,1244,835,860]" italics="true" pageId="5" pageNumber="374">Tico sororius</emphasis>
|
||
</taxonomicName>
|
||
show only slight differences), and the placement removes the diagnostic exception in the key to genus by both
|
||
<bibRefCitation id="EFABDDC9FFEBFFFE24F1BB493E6332F8" author="Fennah, R. G." box="[1222,1387,870,897]" pageId="5" pageNumber="374" pagination="109 - 170" refId="ref12306" refString="Fennah, R. G. (1952) On the generic classification of Derbidae (Fulgoroidea), with descriptions of new Neotropical species. Transactions of the Royal Entomological Society of London, 103 (4), 109 - 170. https: // doi. org / 10.1111 / j. 1365 - 2311.1952. tb 01063. x" type="journal article" year="1952">Fennah (1952)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFABDDC9FFEBFFFE20A0BBA53A4032DC" author="O'Brien, L. B." box="[151,328,907,933]" pageId="5" pageNumber="374" pagination="306 - 321" refId="ref12799" refString="O'Brien, L. B. (1982) Two Neotropical derbid genera with observations on wing rolling (Fulgoroidea: Homoptera). Florida Entomologist, 65 (3), 306 - 321. https: // doi. org / 10.2307 / 3494303" type="journal article" year="1982">O’Brien (1982)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B85A038FFEBFFFE20F0BB813F633504" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="374">
|
||
Specifically, the diagnostic feature
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE227BBB8139CF32B0" authorityName="Muir" authorityYear="1917" box="[588,711,943,969]" class="Insecta" family="Derbidae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="tribe" tribe="Cenchreini">Cenchreini</taxonomicName>
|
||
keys by
|
||
<bibRefCitation id="EFABDDC9FFEBFFFE2310BB813F0832B0" author="Fennah, R. G." box="[807,1024,942,969]" pageId="5" pageNumber="374" pagination="109 - 170" refId="ref12306" refString="Fennah, R. G. (1952) On the generic classification of Derbidae (Fulgoroidea), with descriptions of new Neotropical species. Transactions of the Royal Entomological Society of London, 103 (4), 109 - 170. https: // doi. org / 10.1111 / j. 1365 - 2311.1952. tb 01063. x" type="journal article" year="1952">Fennah (1952: 128)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFABDDC9FFEBFFFE2403BB813E1032B0" author="O'Brien, L. B." box="[1076,1304,943,969]" pageId="5" pageNumber="374" pagination="306 - 321" refId="ref12799" refString="O'Brien, L. B. (1982) Two Neotropical derbid genera with observations on wing rolling (Fulgoroidea: Homoptera). Florida Entomologist, 65 (3), 306 - 321. https: // doi. org / 10.2307 / 3494303" type="journal article" year="1982">O’Brien (1982: 308)</bibRefCitation>
|
||
is “tegmina with subcostal cell short (not extending basad of claval apex)”, a feature found in both
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE2461BBFD3FCA3294" authorityName="Westwood. The" authorityYear="1840" box="[1110,1218,979,1005]" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94E7C2AFFEBFFFE2461BBFD3FCA3294" box="[1110,1218,979,1005]" italics="true" pageId="5" pageNumber="374">Cenchrea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE24C1BBFD3A6A3568" authority="Caldwell. This" authorityName="Caldwell. This" authorityYear="1944" class="Insecta" family="Derbidae" genus="Contigucephalus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94E7C2AFFEBFFFE24C1BBFD3BB03568" italics="true" pageId="5" pageNumber="374">Contigucephalus</emphasis>
|
||
Caldwell. This
|
||
</taxonomicName>
|
||
feature reflects that the Sc branches at the level of (or distal to) the apex of the clavus, making the marginal cell between the Sc and RA branches short. For species we place in
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE2432BC353F3D354D" authority="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett, 2021" authorityName="Bahder & Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett" authorityYear="2021" box="[1029,1077,1051,1076]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94E7C2AFFEBFFFE2432BC353F3D354D" box="[1029,1077,1051,1076]" italics="true" pageId="5" pageNumber="374">Tico</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94E7C2AFFEBFFFE240ABC353F9B354C" bold="true" box="[1085,1171,1051,1077]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="A27DC151FFEBFFFE240ABC353F86354C" box="[1085,1166,1051,1077]" pageId="5" pageNumber="374" rank="genus">gen. n.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
the Sc branches before the level of the claval apex, resulting in a longer marginal cell then the species we retain in
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE249CBC113E1F3520" authorityName="Westwood. The" authorityYear="1840" box="[1195,1303,1087,1113]" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94E7C2AFFEBFFFE249CBC113E1F3520" box="[1195,1303,1087,1113]" italics="true" pageId="5" pageNumber="374">Cenchrea</emphasis>
|
||
</taxonomicName>
|
||
. Additional diagnostic comments under ‘Remarks’ following the amended description of
|
||
<taxonomicName id="4C3ADBBBFFEBFFFE23CFBC4D3F6C3504" authorityName="Westwood. The" authorityYear="1840" box="[1016,1124,1123,1149]" class="Insecta" family="Derbidae" genus="Cenchrea" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94E7C2AFFEBFFFE23CFBC4D3F6C3504" box="[1016,1124,1123,1149]" italics="true" pageId="5" pageNumber="374">Cenchrea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B85A038FFEBFFFE20F0BCA939D335BC" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="374">There is some sexual dimorphism in the genus with males more slender and with narrower forewings than females and males tending to be more darkly marked.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C320F3B3FFEBFFFE20F0BCE13F0F3474" pageId="5" pageNumber="374" type="etymology">
|
||
<paragraph id="8B85A038FFEBFFFE20F0BCE13F0F3474" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="374">
|
||
<emphasis id="B94E7C2AFFEBFFFE20F0BCE13A463590" bold="true" box="[199,334,1231,1257]" pageId="5" pageNumber="374">Etymology.</emphasis>
|
||
The generic name is to be treated as an arbitrary combination of letters, alluding to a colloquial term referencing something from
|
||
<collectingCountry id="F32DE0A8FFEBFFFE2226BCDD39873474" box="[529,655,1267,1293]" name="Costa Rica" pageId="5" pageNumber="374">Costa Rica</collectingCountry>
|
||
. The term is masculine in gender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |