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<document id="67FD7AC2E4D172C5F6E8148D29547D19" ID-CLB-Dataset="3021" ID-DOI="10.11646/zootaxa.5081.1.1" ID-GBIF-Dataset="b96aa772-0701-4b75-a308-f3ff755bb749" ID-ISSN="1175-5326" ID-Zenodo-Dep="5769323" ID-ZooBank="F4BE101A-30D3-43BA-B468-CF4A6ED59496" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639038973822" checkinUser="plazi" docAuthor="Zeidler, Wolfgang" docDate="2021" docId="038F1944587AFFB9829D1926FBD1FD47" docLanguage="en" docName="zootaxa.5081.1.1.pdf" docOrigin="Zootaxa 5081 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lycaea lilia Volkov 1982" docType="treatment" docVersion="8" lastPageNumber="21" masterDocId="FFB6613C586BFFAD820A1A39FFCDFFB6" masterDocTitle="Review of the hyperiidean amphipod family Lycaeidae Claus, 1879 (Crustacea: Amphipoda: Hyperiidea)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="18" updateTime="1699472199594" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C7B6FAEFB7DB707F56B5974D627E905B">Review of the hyperiidean amphipod family Lycaeidae Claus, 1879 (Crustacea: Amphipoda: Hyperiidea)</mods:title>
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<mods:namePart id="928BFF40B23E8EB06707AA3530B10356">Zeidler, Wolfgang</mods:namePart>
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<mods:date id="8CA4AB2FDCA433DC2478624751E5121D">2021</mods:date>
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<treatment id="038F1944587AFFB9829D1926FBD1FD47" ID-DOI="http://doi.org/10.5281/zenodo.5770319" ID-GBIF-Taxon="191261836" ID-Zenodo-Dep="5770319" LSID="urn:lsid:plazi:treatment:038F1944587AFFB9829D1926FBD1FD47" httpUri="http://treatment.plazi.org/id/038F1944587AFFB9829D1926FBD1FD47" lastPageId="20" lastPageNumber="21" pageId="17" pageNumber="18">
<subSubSection id="C33CFBD9587AFFBC829D1926FD0BFC8E" box="[151,710,798,825]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8B99A852587AFFBC829D1926FD0BFC8E" blockId="17.[151,710,798,862]" box="[151,710,798,825]" pageId="17" pageNumber="18">
<heading id="D0D11F3E587AFFBC829D1926FD0BFC8E" bold="true" box="[151,710,798,825]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="B9527440587AFFBC829D1926FD0BFC8E" bold="true" box="[151,710,798,825]" pageId="17" pageNumber="18">
<taxonomicName id="4C26D3D1587AFFBC829D1926FE4CFC8F" ID-CoL="3WHSR" authority="Volkov" authorityName="Volkov" authorityYear="1982" box="[151,385,798,825]" class="Malacostraca" family="Lycaeidae" genus="Lycaea" kingdom="Animalia" order="Amphipoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lilia">
<emphasis id="B9527440587AFFBC829D1926FEEEFC8F" bold="true" box="[151,291,798,825]" italics="true" pageId="17" pageNumber="18">Lycaea lilia</emphasis>
Volkov
</taxonomicName>
in
<bibRefCitation id="EFB7D5A3587AFFBC83AD1927FD0BFC8E" author="Vinogradov, M. E. &amp; Volkov, A. F. &amp; Semenova, T. N." box="[423,710,798,825]" pageId="17" pageNumber="18" refId="ref44917" refString="Vinogradov, M. E., Volkov, A. F. &amp; Semenova, T. N. (1982) Amfipody-Giperiidy (Amphipoda: Hyperiidea) Mirovogo Okeanea. Akademiya Nauk SSSR, Opredeliteli po Faune SSSR No. 132. Akademiya Nauk SSSR, Leningrad, 492 pp. [in Russian]" type="book" year="1982">
Vinogradov
<emphasis id="B9527440587AFFBC80481926FDB0FC8F" bold="true" box="[578,637,798,825]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
, 1982
</bibRefCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33CFBD9587AFFBC829D197AFEC6FCE8" box="[151,267,835,862]" pageId="17" pageNumber="18" type="description">
<paragraph id="8B99A852587AFFBC829D197AFEC6FCE8" blockId="17.[151,710,798,862]" box="[151,267,835,862]" pageId="17" pageNumber="18">
(
<figureCitation id="131DB4D7587AFFBC8295197AFECEFCE8" box="[159,259,835,862]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="18.[151,250,1859,1883]" captionStartId-1="19.[151,250,1882,1906]" captionTargetBox-0="[159,1424,181,1835]" captionTargetBox-1="[175,1404,196,1842]" captionTargetId-0="figure-25@18.[151,1436,181,1835]" captionTargetId-1="figure-15@19.[151,1436,181,1859]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURE 6. Lycaea lilia Volkov, 1982. Female, 6.3 mm, USNM 1242788. Scale bars = 0.5 mm (habitus); 0.2 mm (remainder)." captionText-1="FIGURE 7. Lycaea lilia Volkov, 1982. Male, 8.5 mm (recently moulted), USNM 1242788. A2 from male 7.8 mm, NHMD- 228129. Scale bars = 0.2 mm (A1); 0.5 mm (remainder)." figureDoi-0="http://doi.org/10.5281/zenodo.5769338" figureDoi-1="http://doi.org/10.5281/zenodo.5769342" httpUri-0="https://zenodo.org/record/5769338/files/figure.png" httpUri-1="https://zenodo.org/record/5769342/files/figure.png" pageId="17" pageNumber="18">Figs 6–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33CFBD9587AFFBC829D19B1FCEAFC6A" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8B99A852587AFFBC829D19B1FCEAFC6A" blockId="17.[151,1437,903,989]" pageId="17" pageNumber="18">
<taxonomicName id="4C26D3D1587AFFBC829D19B1FE90FC29" ID-CoL="3WHSR" authority="Volkov" authorityName="Volkov" authorityYear="1982" box="[151,349,903,927]" class="Malacostraca" family="Lycaeidae" genus="Lycaea" kingdom="Animalia" order="Amphipoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lilia">
<emphasis id="B9527440587AFFBC829D19B1FEC2FC29" box="[151,271,903,927]" italics="true" pageId="17" pageNumber="18">Lycaea lilia</emphasis>
Volkov
</taxonomicName>
in
<bibRefCitation id="EFB7D5A3587AFFBC837619BEFD5AFC29" author="Vinogradov, M. E. &amp; Volkov, A. F. &amp; Semenova, T. N." box="[380,663,903,927]" pageId="17" pageNumber="18" refId="ref44917" refString="Vinogradov, M. E., Volkov, A. F. &amp; Semenova, T. N. (1982) Amfipody-Giperiidy (Amphipoda: Hyperiidea) Mirovogo Okeanea. Akademiya Nauk SSSR, Opredeliteli po Faune SSSR No. 132. Akademiya Nauk SSSR, Leningrad, 492 pp. [in Russian]" type="book" year="1982">
Vinogradov
<emphasis id="B9527440587AFFBC83F319B1FDE1FC29" box="[505,556,903,927]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
1982: 382
</bibRefCitation>
(key), 389–390, fig. 210.—Vinogradov
<emphasis id="B9527440587AFFBC862519B1FB96FC29" box="[1071,1115,903,927]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. 1996: 472 (key), 480–482, fig. 210.—
<bibRefCitation id="EFB7D5A3587AFFBC8301199FFD85FC08" author="Lavaniegos, B. E. &amp; Hereu, C. M." box="[267,584,934,958]" pageId="17" pageNumber="18" pagination="137 - 152" refId="ref41571" refString="Lavaniegos, B. E. &amp; Hereu, C. M. (2009) Seasonal variation in hyperiid amphipod abundance and diversity and influence of mesoscale structures off Baja California. Marine Ecology Progress Series, 394, 137 - 152. https: // doi. org / 10.3354 / meps 08285" type="journal article" year="2009">Lavaniegos &amp; Hereu 2009: 152</bibRefCitation>
(tab.).—
<bibRefCitation id="EFB7D5A3587AFFBC8094199FFC4EFC08" author="Gasca, R. &amp; Suarez-Morales, E. &amp; Franco-Gordo, L." box="[670,899,934,958]" pageId="17" pageNumber="18" pagination="927 - 940" refId="ref40207" refString="Gasca, R., Suarez-Morales, E. &amp; Franco-Gordo, L. (2010) New records of hyperiids (Amphipoda, Hyperiidea) from surface waters of the Central Mexican Pacific. Crustaceana, 83 (8), 927 - 940. https: // doi. org / 10.1163 / 001121610 X 504298" type="journal article" year="2010">
Gasca
<emphasis id="B9527440587AFFBC80EC199EFCD5FC08" box="[742,792,934,958]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2010: 934
</bibRefCitation>
, 938.—Gasca
<emphasis id="B9527440587AFFBC861C199EFB85FC08" box="[1046,1096,934,958]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2012: passim.—Gasca &amp; Franco- Gordo 2014: 75 (list).—
<bibRefCitation id="EFB7D5A3587AFFBC83B319FDFD1DFC6B" author="Espinosa-Leal, L. &amp; Escribano, R. &amp; Riquelme-Bugueno, R. &amp; Corredor-Acosta, A." box="[441,720,964,989]" pageId="17" pageNumber="18" pagination="1 - 17" refId="ref38909" refString="Espinosa-Leal, L., Escribano, R., Riquelme-Bugueno, R. &amp; Corredor-Acosta, A. (2021 a) Distribution and biodiversity patterns of hyperiid amphipods across the coastal-offshore gradient of the sub-tropical Southeastern Pacific. Marine Biodiversity, 51 (13), 1 - 17. https: // doi. org / 10.1007 / s 12526 - 020 - 01152 - x" type="journal article" year="2021">
Espinosa-Leal
<emphasis id="B9527440587AFFBC805F19FCFD4CFC6A" box="[597,641,964,988]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. 2021a
</bibRefCitation>
: passim.
</paragraph>
</subSubSection>
<subSubSection id="C33CFBD9587AFFBC829D1E32FD18FA97" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8B99A852587AFFBC829D1E32FE97FBDB" blockId="17.[151,1437,1034,2033]" pageId="17" pageNumber="18">
<emphasis id="B9527440587AFFBC829D1E32FE8EFB92" bold="true" box="[151,323,1034,1061]" pageId="17" pageNumber="18">Type material.</emphasis>
<materialsCitation id="3B4EA20F587AFFBC83431E32FE97FBDB" ID-GBIF-Occurrence="3417039349" collectionCode="TINTRO" collectorName="The &amp; S. E. Pacific" county="Oceanography" elevation="50" latitude="-13.0" location="The" longLatPrecision="41492" longitude="-86.0" municipality="Pacific Ocean Scientific Research Institution of Fisheries" pageId="17" pageNumber="18" specimenCount="2" typeStatus="holotype">
<location id="8EF9FE89587AFFBC83431E32FEB8FB93" LSID="urn:lsid:plazi:treatment:038F1944587AFFB9829D1926FBD1FD47:8EF9FE89587AFFBC83431E32FEB8FB93" box="[329,373,1035,1061]" county="Oceanography" latitude="-13.0" longLatPrecision="41492" longitude="-86.0" municipality="Pacific Ocean Scientific Research Institution of Fisheries" name="The" pageId="17" pageNumber="18">The</location>
<typeStatus id="549D16F0587AFFBC83701E32FE11FB93" box="[378,476,1035,1061]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
female (9.0 mm) and some
<typeStatus id="549D16F0587AFFBC81071E32FCB5FB93" box="[781,888,1035,1061]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
are held by the
<collectingMunicipality id="6BFD3228587AFFBC862F1E32FE57FBFF" pageId="17" pageNumber="18">Pacific Ocean Scientific Research Institution of Fisheries</collectingMunicipality>
and
<collectingCounty id="62F8D0DE587AFFBC83DC1E16FDB4FBFF" box="[470,633,1071,1097]" pageId="17" pageNumber="18">Oceanography</collectingCounty>
(
<collectionCode id="ED373097587AFFBC80811E16FD35FBFF" box="[651,760,1071,1097]" pageId="17" pageNumber="18">TINTRO</collectionCode>
).
<collectorName id="26D3CD84587AFFBC81061E16FCF5FBFF" box="[780,824,1071,1097]" pageId="17" pageNumber="18">The</collectorName>
type locality is the
<collectorName id="26D3CD84587AFFBC862B1E16FB65FBFF" box="[1057,1192,1071,1097]" pageId="17" pageNumber="18">S.E. Pacific</collectorName>
[
<geoCoordinate id="EE12CE95587AFFBC86B21E16FB3CFBFF" box="[1208,1265,1071,1097]" degrees="08" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="55555" value="-8.0">08°S</geoCoordinate>
<geoCoordinate id="EE12CE95587AFFBC86F11E16FAF1FBFF" box="[1275,1340,1071,1097]" degrees="90" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="55555" value="-90.0">90°W</geoCoordinate>
&amp; 13°S 86°W],
<quantity id="4CDE05B7587AFFBC82FA1E6AFE9BFBDB" box="[240,342,1107,1133]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="17" pageNumber="18" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">
<elevation id="000B4F61587AFFBC82FA1E6AFE9BFBDB" box="[240,342,1107,1133]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="17" pageNumber="18" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
<paragraph id="8B99A852587AFFBC82CD1E4FFD18FA97" blockId="17.[151,1437,1034,2033]" pageId="17" pageNumber="18">
<emphasis id="B9527440587AFFBC82CD1E4FFE7CFB26" bold="true" box="[199,433,1142,1168]" pageId="17" pageNumber="18">Material examined.</emphasis>
<materialsCitation id="3B4EA20F587AFFBC83B31E4EFD19FB03" ID-GBIF-Occurrence="3417039378" collectingDate="1934-02-15" collectionCode="NHMD, USNM" country="China" county="Chaco" location="Colombia" municipality="Port Utria" pageId="17" pageNumber="18" specimenCount="14" specimenCount-female="9" specimenCount-male="5">
<collectorName id="26D3CD84587AFFBC83B31E4EFE1DFB27" box="[441,464,1143,1169]" pageId="17" pageNumber="18">In</collectorName>
<collectionCode id="ED373097587AFFBC83D21E4EFDFFFB27" box="[472,562,1143,1169]" pageId="17" pageNumber="18">NHMD</collectionCode>
:
<emphasis id="B9527440587AFFBC80371E4FFD78FB26" bold="true" box="[573,693,1142,1168]" pageId="17" pageNumber="18">
<collectorName id="26D3CD84587AFFBC80371E4FFD7DFB26" box="[573,688,1142,1168]" pageId="17" pageNumber="18">S. Pacific</collectorName>
:
</emphasis>
near
<collectorName id="26D3CD84587AFFBC80FF1E4EFC89FB27" box="[757,836,1143,1169]" pageId="17" pageNumber="18">Samoa</collectorName>
,
<collectorName id="26D3CD84587AFFBC815A1E4EFC46FB27" box="[848,907,1143,1169]" pageId="17" pageNumber="18">Dana</collectorName>
stns 3558 ii (228124), 3584 vi (228126), 3584 vii (228129), 3588 iii (228130),
<specimenCount id="9D2063DB587AFFBC83F41EA2FDA0FB03" box="[510,621,1179,1205]" pageId="17" pageNumber="18" type="female">3 females</specimenCount>
,
<specimenCount id="9D2063DB587AFFBC807D1EA2FD1DFB03" box="[631,720,1179,1205]" pageId="17" pageNumber="18" type="male">2 males</specimenCount>
.
</materialsCitation>
<materialsCitation id="3B4EA20F587AFFBC80D01EA3FA50FB03" ID-GBIF-Occurrence="3417039363" box="[730,1437,1178,1205]" collectingDate="1934-02-15" collectionCode="NHMD, USNM" country="China" county="Chaco" location="Colombia" municipality="Port Utria" pageId="17" pageNumber="18" specimenCount="14" specimenCount-female="9" specimenCount-male="5">
<emphasis id="B9527440587AFFBC80D01EA3FC91FB02" bold="true" box="[730,860,1178,1204]" pageId="17" pageNumber="18">
<collectingCountry id="F331E8C2587AFFBC80D01EA3FCEEFB02" box="[730,803,1178,1204]" name="China" pageId="17" pageNumber="18">China</collectingCountry>
Sea:
</emphasis>
N. of Luzon, Dana stn. 3729 iii (228131),
<specimenCount id="9D2063DB587AFFBC873F1EA2FA54FB03" box="[1333,1433,1179,1205]" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
.
</materialsCitation>
<materialsCitation id="3B4EA20F587AFFBC829D1E87FC14FB6F" ID-GBIF-Occurrence="3417039431" box="[151,985,1214,1241]" collectingDate="1934-02-15" collectionCode="NHMD, USNM" country="China" county="Chaco" location="Colombia" municipality="Port Utria" pageId="17" pageNumber="18" specimenCount="14" specimenCount-female="9" specimenCount-male="5">
<emphasis id="B9527440587AFFBC829D1E87FEB7FB6E" bold="true" box="[151,378,1214,1240]" pageId="17" pageNumber="18">
<collectorName id="26D3CD84587AFFBC829D1E87FEB9FB6E" box="[151,372,1214,1240]" pageId="17" pageNumber="18">S.E. Indian Ocean</collectorName>
:
</emphasis>
S. of Sumatra, Dana stn. 3856 iv (228133),
<specimenCount id="9D2063DB587AFFBC816C1E86FC1BFB6F" box="[870,982,1215,1241]" pageId="17" pageNumber="18" type="female">2 females</specimenCount>
.
</materialsCitation>
<materialsCitation id="3B4EA20F587AFFBC81D51E87FD6CFB4B" ID-GBIF-Occurrence="3417039442" collectingDate="1934-02-15" collectionCode="NHMD, USNM" country="China" county="Chaco" location="Colombia" municipality="Port Utria" pageId="17" pageNumber="18" specimenCount="14" specimenCount-female="9" specimenCount-male="5">
<emphasis id="B9527440587AFFBC81D51E87FB05FB6E" bold="true" box="[991,1224,1214,1240]" pageId="17" pageNumber="18">
<collectorName id="26D3CD84587AFFBC81D51E87FB0EFB6E" box="[991,1219,1214,1240]" pageId="17" pageNumber="18">S.W. Indian Ocean</collectorName>
:
</emphasis>
N. of Madagascar, Dana stn. 3931 (228135),
<specimenCount id="9D2063DB587AFFBC83B11EDAFDEDFB4B" box="[443,544,1251,1277]" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
. In
<collectionCode id="ED373097587AFFBC80421EDAFD50FB4B" box="[584,669,1251,1277]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18" type="Museum">USNM</collectionCode>
:
</materialsCitation>
<materialsCitation id="3B4EA20F587AFFBC80AD1EDAFD1CFA97" ID-GBIF-Occurrence="3417039387" collectingDate="1934-02-15" collectionCode="NHMD, USNM" country="China" county="Chaco" location="Colombia" municipality="Port Utria" pageId="17" pageNumber="18" specimenCount="14" specimenCount-female="9" specimenCount-male="5">
<emphasis id="B9527440587AFFBC80AD1EDAFCF3FB4A" bold="true" box="[679,830,1250,1277]" pageId="17" pageNumber="18">
<collectorName id="26D3CD84587AFFBC80AD1EDAFCF5FB4A" box="[679,824,1250,1277]" pageId="17" pageNumber="18">N.E. Pacific</collectorName>
:
</emphasis>
<specimenCount id="9D2063DB587AFFBC814E1EDAFC7EFB4B" box="[836,947,1251,1277]" pageId="17" pageNumber="18" type="female">2 females</specimenCount>
,
<specimenCount id="9D2063DB587AFFBC81B41EDAFBD9FB4B" box="[958,1044,1251,1277]" pageId="17" pageNumber="18" type="male">3 males</specimenCount>
(1242788), “
<location id="8EF9FE89587AFFBC86AD1EDAFAD6FB4B" LSID="urn:lsid:plazi:treatment:038F1944587AFFB9829D1926FBD1FD47:8EF9FE89587AFFBC86AD1EDAFAD6FB4B" box="[1191,1307,1251,1277]" country="China" county="Chaco" municipality="Port Utria" name="Colombia" pageId="17" pageNumber="18">Colombia</location>
,
<collectingMunicipality id="6BFD3228587AFFBC872C1EDAFA55FB4B" box="[1318,1432,1251,1277]" pageId="17" pageNumber="18">Port Utria</collectingMunicipality>
,
<collectingCounty id="62F8D0DE587AFFBC829D1F3EFF29FA97" box="[151,228,1287,1313]" pageId="17" pageNumber="18">Chaco</collectingCounty>
”, R/
<collectorName id="26D3CD84587AFFBC831E1F3EFEBAFA97" box="[276,375,1287,1313]" pageId="17" pageNumber="18">V Velero</collectorName>
III, surface,
<date id="FF988E92587AFFBC800D1F3FFD1CFA97" box="[519,721,1286,1313]" pageId="17" pageNumber="18" value="1934-02-15">
<collectingDate id="EFDC777A587AFFBC800D1F3FFD1CFA97" box="[519,721,1286,1313]" pageId="17" pageNumber="18" value="1934-02-15">15 February 1934</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33CFBD9587AFFB982CD1F13FAADFF4F" lastPageId="20" lastPageNumber="21" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8B99A852587AFFB982CD1F13FAADFF4F" blockId="17.[151,1437,1034,2033]" lastBlockId="20.[151,1437,151,753]" lastPageId="20" lastPageNumber="21" pageId="17" pageNumber="18">
<emphasis id="B9527440587AFFBC82CD1F13FE8FFAF2" bold="true" box="[199,322,1322,1348]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
Body length up to about 9.0 mm. Head of females relatively large, rounded, much deeper than long (about 1.5 x), as long as first 3–4 pereonites combined. Head of males almost as long as first 3 pereonites combined, usually with small, anterior knob. Buccal mass protruded well below head. Pereonite 2 of females sometimes with slight dorsal hump. Callynophore of A1 of males without antero-distal corner; postero-distal corner small, rounded, separated from following article by small notch. G1 and G2 sub-chelate, G2 slightly longer than G1. G1 basis slightly shorter than for G2; carpus rectangular with small, rounded postero-distal tooth, reaching just past base of dactylus; propodus with small postero-distal corner produced very slightly over dactylus; carpus and propodus with smooth margins; dactylus very short, stubby, length less than 0.3 x propodus. G2 carpus similar to G1 but distal margin only extends to about the middle of the propodus; propodus of males with few small serrations distally on posterior margin, otherwise similar to G1. P3–6 with very short, stubby dactylus, those of P3 and P4 about 0.2 x as long as propodus, or slightly less. P3 and P4 morphologically similar, P4 slightly longer than P3; merus marginally inflated anteriorly, slightly shorter than propodus, about 0.4–0.5 x basis; carpus length about 0.7 x propodus. P5 length about 1.4 x P4 and about 1.3 x P
<quantity id="4CDE05B7587AFFBC83CD1CE2FE3FF943" box="[455,498,1755,1781]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="17" pageNumber="18" unit="in" value="6.0">6 in</quantity>
both sexes; basis rectangular, length about 2 x maximum width; merus marginally inflated, slightly longer than propodus, about 0.6 x basis; carpus length about 0.7 x propodus. P6 basis oval-shaped, slightly wider distally in males, length about 2 x maximum width, slightly shorter than basis of P5; merus distinctly inflated, maximum width slightly more than 0.6 x length, sub-equal in length to propodus, about 0.5 x basis; carpus also slightly inflated, length almost 0.7 x propodus; anterior margin of carpus and propodus, and antero-distal corner of merus, slightly serrated. P7 basis with bulging posterior margin, wider in females, length about 1.5 x maximum width, in males length about 1.7 x maximum width, about 0.7 x basis of P6; remaining articles together relatively long, almost 0.7 x basis in males and almost as long as basis in females; propodus with antero-distal corner produced into small, rounded lobe; dactylus sharp, hook-like. U1 and U2; endopod not fused with peduncle. U1; peduncle relatively short, about 2.5 x length of exopod or only slightly longer; rami relatively slender, equal in length. Telson length about equal to width at base, relatively shorter and more rounded than for other species.
</paragraph>
</subSubSection>
<caption id="DF59F8DA5879FFBF829D1D7AFEDBF8C9" ID-DOI="http://doi.org/10.5281/zenodo.5769338" ID-Zenodo-Dep="5769338" httpUri="https://zenodo.org/record/5769338/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,250,1859,1883]" targetBox="[159,1424,181,1835]" targetPageId="18">
<paragraph id="8B99A8525879FFBF829D1D7AFEDBF8C9" blockId="18.[151,1436,1859,1919]" pageId="18" pageNumber="19">
<emphasis id="B95274405879FFBF829D1D7AFED7F8ED" bold="true" box="[151,282,1859,1883]" pageId="18" pageNumber="19">FIGURE 6.</emphasis>
<taxonomicName id="4C26D3D15879FFBF83221D7DFD89F8ED" authority="Volkov, 1982" authorityName="Volkov" authorityYear="1982" box="[296,580,1859,1883]" class="Malacostraca" family="Lycaeidae" genus="Lycaea" kingdom="Animalia" order="Amphipoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lilia">
<emphasis id="B95274405879FFBF83221D7DFE6AF8ED" box="[296,423,1859,1883]" italics="true" pageId="18" pageNumber="19">Lycaea lilia</emphasis>
Volkov, 1982
</taxonomicName>
. Female, 6.3 mm, USNM 1242788. Scale bars = 0.5 mm (habitus); 0.2 mm (remainder).
</paragraph>
</caption>
<caption id="DF59F8DA5878FFBE829D1D63FD1BF821" ID-DOI="http://doi.org/10.5281/zenodo.5769342" ID-Zenodo-Dep="5769342" httpUri="https://zenodo.org/record/5769342/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1882,1906]" targetBox="[175,1404,196,1842]" targetPageId="19">
<paragraph id="8B99A8525878FFBE829D1D63FD1BF821" blockId="19.[151,1436,1882,1943]" pageId="19" pageNumber="20">
<emphasis id="B95274405878FFBE829D1D63FED8F8C5" bold="true" box="[151,277,1882,1907]" pageId="19" pageNumber="20">FIGURE 7.</emphasis>
<taxonomicName id="4C26D3D15878FFBE83171D62FDE4F8C5" authority="Volkov, 1982" authorityName="Volkov" authorityYear="1982" box="[285,553,1883,1907]" class="Malacostraca" family="Lycaeidae" genus="Lycaea" kingdom="Animalia" order="Amphipoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lilia">
<emphasis id="B95274405878FFBE83171D62FE5BF8C5" box="[285,406,1883,1907]" italics="true" pageId="19" pageNumber="20">Lycaea lilia</emphasis>
Volkov, 1982
</taxonomicName>
. Male, 8.5 mm (recently moulted), USNM 1242788. A2 from male 7.8 mm, NHMD- 228129. Scale bars = 0.2 mm (A1); 0.5 mm (remainder).
</paragraph>
</caption>
<subSubSection id="C33CFBD9587FFFB982CD1B3AFCE0FD33" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8B99A852587FFFB982CD1B3AFA89FE43" blockId="20.[151,1437,151,753]" pageId="20" pageNumber="21">
<emphasis id="B9527440587FFFB982CD1B3AFEF6FEAB" bold="true" box="[199,315,259,285]" pageId="20" pageNumber="21">Remarks.</emphasis>
<taxonomicName id="4C26D3D1587FFFB9834E1B3AFE06FEAB" authorityName="Volkov" authorityYear="1982" box="[324,459,259,285]" class="Malacostraca" family="Lycaeidae" genus="Lycaea" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="lilia">
<emphasis id="B9527440587FFFB9834E1B3AFE06FEAB" box="[324,459,259,285]" italics="true" pageId="20" pageNumber="21">Lycaea lilia</emphasis>
</taxonomicName>
is readily distinguished from all its congeners, except
<taxonomicName id="4C26D3D1587FFFB986491B3AFB63FEAA" authorityName="Claus" authorityYear="1879" box="[1091,1198,259,284]" class="Malacostraca" family="Lycaeidae" genus="Lycaea" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9527440587FFFB986491B3AFB63FEAA" box="[1091,1198,259,284]" italics="true" pageId="20" pageNumber="21">L. nasuta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C26D3D1587FFFB986E31B3AFABCFEAB" authority="Zeidler, 2021" authorityName="Zeidler" authorityYear="2021" box="[1257,1393,259,285]" class="Malacostraca" family="Lycaeidae" genus="Lycaea" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="osbornae" status="sp. nov.">
<emphasis id="B9527440587FFFB986E31B3AFABCFEAB" box="[1257,1393,259,285]" italics="true" pageId="20" pageNumber="21">L. osbornae</emphasis>
</taxonomicName>
<emphasis id="B9527440587FFFB987711B3AFF05FEF7" bold="true" pageId="20" pageNumber="21">
<taxonomicNameLabel id="A261C93B587FFFB987711B3AFF05FEF7" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the morphology of G
<quantity id="4CDE05B7587FFFB983EB1B1EFDC1FEF7" box="[481,524,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="20" pageNumber="21" unit="in" value="2.0">2 in</quantity>
that the carpus is relatively narrow so that the propodus extends well beyond the postero-distal corner of the carpus. It also shares two other distinctive characters with
<taxonomicName id="4C26D3D1587FFFB986651B72FB14FED2" authorityName="Claus" authorityYear="1879" box="[1135,1241,331,356]" class="Malacostraca" family="Lycaeidae" genus="Lycaea" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9527440587FFFB986651B72FB14FED2" box="[1135,1241,331,356]" italics="true" pageId="20" pageNumber="21">L. nasuta</emphasis>
</taxonomicName>
: the morphology of the head of males is similar and, in some females, the second pereonite has a small dorsal hump. Clearly the two species are closely related but
<taxonomicName id="4C26D3D1587FFFB983E61BAAFDFAFE1B" authorityName="Volkov" authorityYear="1982" box="[492,567,403,429]" class="Malacostraca" family="Lycaeidae" genus="Lycaea" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="lilia">
<emphasis id="B9527440587FFFB983E61BAAFDFAFE1B" box="[492,567,403,429]" italics="true" pageId="20" pageNumber="21">L. lilia</emphasis>
</taxonomicName>
is readily distinguished by the morphology of U2 (peduncle and endopod not fused) and the morphology of G1 and G
<quantity id="4CDE05B7587FFFB980691B8EFD42FE67" box="[611,655,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="20" pageNumber="21" unit="in" value="2.0">2 in</quantity>
that the postero-distal corner of the carpus is relatively rounded and does not end in a distinct tooth. It is distinguished from
<taxonomicName id="4C26D3D1587FFFB981021BE2FC43FE43" authority="Zeidler, 2021" authorityName="Zeidler" authorityYear="2021" box="[776,910,475,501]" class="Malacostraca" family="Lycaeidae" genus="Lycaea" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="osbornae" status="sp. nov.">
<emphasis id="B9527440587FFFB981021BE2FC43FE43" box="[776,910,475,501]" italics="true" pageId="20" pageNumber="21">L. osbornae</emphasis>
</taxonomicName>
<emphasis id="B9527440587FFFB9819F1BE2FC22FE43" bold="true" box="[917,1007,475,501]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="A261C93B587FFFB9819F1BE2FC22FE43" box="[917,1007,475,501]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
as detailed under that species.
</paragraph>
<paragraph id="8B99A852587FFFB982CD1BC6FA91FDD7" blockId="20.[151,1437,151,753]" pageId="20" pageNumber="21">
The male of
<taxonomicName id="4C26D3D1587FFFB983531BC6FE13FDAF" authorityName="Volkov" authorityYear="1982" box="[345,478,511,537]" class="Malacostraca" family="Lycaeidae" genus="Lycaea" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="lilia">
<emphasis id="B9527440587FFFB983531BC6FE13FDAF" box="[345,478,511,537]" italics="true" pageId="20" pageNumber="21">Lycaea lilia</emphasis>
</taxonomicName>
has not been illustrated previously in the literature and is unusual in that the head is produced into a small, rounded knob, anteriorly, similar to males of
<taxonomicName id="4C26D3D1587FFFB9817E181AFC17FD8A" authorityName="Claus" authorityYear="1879" box="[884,986,547,572]" class="Malacostraca" family="Lycaeidae" genus="Lycaea" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9527440587FFFB9817E181AFC17FD8A" box="[884,986,547,572]" italics="true" pageId="20" pageNumber="21">L. nasuta</emphasis>
</taxonomicName>
. Unfortunately, of the
<specimenCount id="9D2063DB587FFFB986C4181AFA8FFD8B" box="[1230,1346,547,573]" pageId="20" pageNumber="21" type="male">four males</specimenCount>
at hand, two are immature and the other two are recent moults so it is impossible to adequately describe the antennae.
</paragraph>
<paragraph id="8B99A852587FFFB982CD1852FCE0FD33" blockId="20.[151,1437,151,753]" box="[199,813,619,645]" pageId="20" pageNumber="21">A salp associate has not been recorded for this species.</paragraph>
</subSubSection>
<subSubSection id="C33CFBD9587FFFB982CD18B7FBD1FD47" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8B99A852587FFFB982CD18B7FBD1FD47" blockId="20.[151,1437,151,753]" pageId="20" pageNumber="21">
<emphasis id="B9527440587FFFB982CD18B7FEACFD1E" bold="true" box="[199,353,654,680]" pageId="20" pageNumber="21">Distribution.</emphasis>
A relatively rare species known only from the tropical E. Pacific, the tropical S.E. and S.W. Indian Ocean and the
<collectingCountry id="F331E8C2587FFFB9839F188AFE15FD7B" box="[405,472,691,717]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
Sea as detailed above, and from off central
<collectingCountry id="F331E8C2587FFFB981DC188AFBE1FD7B" box="[982,1068,691,717]" name="Mexico" pageId="20" pageNumber="21">Mexico</collectingCountry>
(
<bibRefCitation id="EFB7D5A3587FFFB98636188AFAC6FD7B" author="Gasca, R. &amp; Suarez-Morales, E. &amp; Franco-Gordo, L." box="[1084,1291,691,717]" pageId="20" pageNumber="21" pagination="927 - 940" refId="ref40207" refString="Gasca, R., Suarez-Morales, E. &amp; Franco-Gordo, L. (2010) New records of hyperiids (Amphipoda, Hyperiidea) from surface waters of the Central Mexican Pacific. Crustaceana, 83 (8), 927 - 940. https: // doi. org / 10.1163 / 001121610 X 504298" type="journal article" year="2010">
Gasca
<emphasis id="B9527440587FFFB98681188AFB73FD7B" box="[1163,1214,691,717]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
. 2010
</bibRefCitation>
). It seems to prefer shallow waters with records from hauls of 100–0 m and with 50–600 mw.
</paragraph>
</subSubSection>
</treatment>
</document>
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