573 lines
71 KiB
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573 lines
71 KiB
XML
<document id="C1DBCFF30EFE2CAFD2DCCEC42CE2E166" ID-DOI="10.11646/zootaxa.4747.2.3" ID-GBIF-Dataset="a1407eed-b24d-4132-b411-ee8da970d2d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="3694804" ID-ZooBank="6D32355F-DBFD-44C6-AC13-086F13A35066" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583220544419" checkinUser="plazi" docAuthor="Denzer, Wolfgang, Campbell, Patrick D., Manthey, Ulrich, Glässer-Trobisch, Andrea & Koch, André" docDate="2020" docId="FF530825FF907B48FF7A7F70750BFE4A" docLanguage="en" docName="zootaxa.4747.2.3.pdf" docOrigin="Zootaxa 4747 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrosaurus celebensis" docType="treatment" docVersion="7" lastPageNumber="292" masterDocId="036A705DFF9F7B5AFFED7B31774FFFBD" masterDocTitle="Dragons in neglect: Taxonomic revision of the Sulawesi sailfin lizards of the genus Hydrosaurus Kaup, 1828 (Squamata, Agamidae)" masterLastPageNumber="301" masterPageNumber="275" pageNumber="290" updateTime="1698724071920" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1F3E9CD69E3678B0D0108BD303AB9A2A">Dragons in neglect: Taxonomic revision of the Sulawesi sailfin lizards of the genus Hydrosaurus Kaup, 1828 (Squamata, Agamidae)</mods:title>
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<mods:namePart id="FEE4B6F000D8694EB2FF973C8723E206">Denzer, Wolfgang</mods:namePart>
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<mods:namePart id="3B1C77538CB1EADD97028C5893B121CD">Campbell, Patrick D.</mods:namePart>
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<mods:namePart id="E30C0E1DCCBC0C6A40556AB2C95FEB59">Manthey, Ulrich</mods:namePart>
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<mods:namePart id="D5D17465B8718A06AAF446D28D069E2B">Koch, André</mods:namePart>
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<treatment id="FF530825FF907B48FF7A7F70750BFE4A" ID-DOI="http://doi.org/10.5281/zenodo.3703626" ID-GBIF-Taxon="162381186" ID-Zenodo-Dep="3703626" LSID="urn:lsid:plazi:treatment:FF530825FF907B48FF7A7F70750BFE4A" httpUri="http://treatment.plazi.org/id/FF530825FF907B48FF7A7F70750BFE4A" lastPageId="18" lastPageNumber="292" pageId="15" pageNumber="290">
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<subSubSection id="3FE0EAB8FF907B55FF7A7F707665FB23" pageId="15" pageNumber="290" type="nomenclature">
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<paragraph id="7745B933FF907B55FF7A7F707526FBE6" blockId="15.[151,617,1089,1116]" box="[151,617,1089,1116]" pageId="15" pageNumber="290">
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<heading id="2C0D0E5FFF907B55FF7A7F707526FBE6" bold="true" box="[151,617,1089,1116]" fontSize="11" level="1" pageId="15" pageNumber="290" reason="1">
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<taxonomicName id="B0FAC2B0FF907B55FF7A7F707526FBE6" authority="(Peters, 1872)" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[151,617,1089,1116]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="celebensis">
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<emphasis id="458E6521FF907B55FF7A7F707526FBE6" bold="true" box="[151,617,1089,1116]" pageId="15" pageNumber="290">
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<emphasis id="458E6521FF907B55FF7A7F7076F8FBE1" bold="true" box="[151,439,1089,1116]" italics="true" pageId="15" pageNumber="290">Hydrosaurus celebensis</emphasis>
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(
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<bibRefCitation id="136BC4C2FF907B55FE2B7F70752EFBE6" author="Peters, W." box="[454,609,1089,1115]" pageId="15" pageNumber="290" pagination="581 - 585" refId="ref19232" refString="Peters, W. (1872) Uber einige von Hrn Dr. A. B. Meyer bei Gorontalo und auf den Togian-Inseln gesammelte Amphibien. Monatsberichte der koniglichen Akademie der Wissenschaften zu Berlin, 1872, 581 - 585." type="journal article" year="1872">Peters, 1872</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="7745B933FF907B55FF7A7FB47665FB23" blockId="15.[151,1311,1157,1396]" box="[151,298,1157,1182]" pageId="15" pageNumber="290">
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<heading id="2C0D0E5FFF907B55FF7A7FB47665FB23" bold="true" box="[151,298,1157,1182]" fontSize="10" level="2" pageId="15" pageNumber="290" reason="2">
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<emphasis id="458E6521FF907B55FF7A7FB47665FB23" bold="true" box="[151,298,1157,1182]" pageId="15" pageNumber="290">Chresonymy:</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="3FE0EAB8FF907B55FF7A7F95753AFAC9" pageId="15" pageNumber="290" type="reference_group">
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<paragraph id="7745B933FF907B55FF7A7F957505FB01" blockId="15.[151,1311,1157,1396]" box="[151,586,1187,1212]" pageId="15" pageNumber="290">
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<treatmentCitationGroup id="57EA9E1DFF907B55FF7A7F957505FB01" box="[151,586,1187,1212]" pageId="15" pageNumber="290">
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<treatmentCitation id="F65B9F22FF907B55FF7A7F957505FB01" author="Gunther, A." box="[151,586,1187,1212]" page="168" pageId="15" pageNumber="290" year="1873">
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<taxonomicName id="B0FAC2B0FF907B55FF7A7F957505FB01" authority="(Gunther 1873: 168)" baseAuthorityName="Gunther" baseAuthorityPageNumber="168" baseAuthorityYear="1873" box="[151,586,1187,1212]" class="Aves" family="Phasianidae" genus="Lophura" kingdom="Animalia" order="Galliformes" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="amboinensis">
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<emphasis id="458E6521FF907B55FF7A7F95763BFB01" box="[151,372,1188,1212]" italics="true" pageId="15" pageNumber="290">Lophura amboinensis</emphasis>
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(
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<bibRefCitation id="136BC4C2FF907B55FE6C7F92750CFB01" author="Gunther, A." box="[385,579,1187,1212]" pageId="15" pageNumber="290" pagination="165 - 172" refId="ref17972" refString="Gunther, A. (1873) Notes on some reptiles and batrachians obtained by Dr. Bernhard Meyer in Celebes and the Philippine Islands. Proceedings of the Zoological Society London, 1873, 165 - 172." type="journal article" year="1873">Günther 1873: 168</bibRefCitation>
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)
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</taxonomicName>
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</treatmentCitation>
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</paragraph>
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<paragraph id="7745B933FF907B55FF7A7FF27250FB66" blockId="15.[151,1311,1157,1396]" box="[151,1311,1218,1243]" pageId="15" pageNumber="290">
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<treatmentCitationGroup id="57EA9E1DFF907B55FF7A7FF27250FB66" box="[151,1311,1218,1243]" pageId="15" pageNumber="290">
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<taxonomicName id="B0FAC2B0FF907B55FF7A7FF2763BFB66" baseAuthorityName="Gunther" baseAuthorityYear="1873" box="[151,372,1219,1243]" class="Aves" family="Phasianidae" genus="Lophura" kingdom="Animalia" order="Galliformes" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="amboinensis">
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<emphasis id="458E6521FF907B55FF7A7FF2763BFB66" box="[151,372,1219,1243]" italics="true" pageId="15" pageNumber="290">Lophura amboinensis</emphasis>
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</taxonomicName>
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(part) (
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<treatmentCitation id="F65B9F22FF907B55FE527FF375D5FB66" author="Boulenger, G. A." box="[447,666,1218,1243]" page="402" pageId="15" pageNumber="290" year="1885">
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<bibRefCitation id="136BC4C2FF907B55FE527FF375D5FB66" author="Boulenger, G. A." box="[447,666,1218,1243]" pageId="15" pageNumber="290" refId="ref16701" refString="Boulenger, G. A. (1885) Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. Trustees of the British Museum, London, 450 pp." type="book" year="1885">Boulenger 1885: 402</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="F65B9F22FF907B55FD497FF27473FB66" author="Meyer, A. B." box="[676,828,1218,1243]" page="6" pageId="15" pageNumber="290" year="1887">
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<bibRefCitation id="136BC4C2FF907B55FD497FF27473FB66" author="Meyer, A. B." box="[676,828,1218,1243]" pageId="15" pageNumber="290" pagination="1 - 16" refId="ref19040" refString="Meyer, A. B. (1887) Verzeichnis der von mir in den Jahren 1870 - 1873 im ostindischen Archipel gesammelten Reptilien und Batrachier. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch-Ethnographischen Museums zu Dresden, 2, 1 - 16." type="journal article" year="1887">Meyer 1887: 6</bibRefCitation>
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</treatmentCitation>
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; Casto de Elera 1895: 417;
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<treatmentCitation id="F65B9F22FF907B55FBB97FF37257FB66" author="de Rooij, N." box="[1108,1304,1218,1243]" page="128" pageId="15" pageNumber="290" year="1915">
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<bibRefCitation id="136BC4C2FF907B55FBB97FF37257FB66" author="de Rooij, N." box="[1108,1304,1218,1243]" pageId="15" pageNumber="290" refId="ref17185" refString="de Rooij, N. (1915) The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. E. J. Brill, Leiden, xiv + 384 pp. https: // doi. org / 10.5962 / bhl. title. 5069" type="book" year="1915">de Rooij 1915: 128</bibRefCitation>
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)
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</paragraph>
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<paragraph id="7745B933FF907B55FF7A7FD0744FFB47" blockId="15.[151,1311,1157,1396]" box="[151,768,1249,1274]" pageId="15" pageNumber="290">
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<treatmentCitationGroup id="57EA9E1DFF907B55FF7A7FD0744FFB47" box="[151,768,1249,1274]" pageId="15" pageNumber="290">
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<taxonomicName id="B0FAC2B0FF907B55FF7A7FD07547FB44" authority="Weber 1890" box="[151,520,1249,1273]" class="Aves" family="Phasianidae" genus="Lophura" kingdom="Animalia" order="Galliformes" pageId="15" pageNumber="290" phylum="Chordata" rank="variety" species="amboinensis" variety="celebensis">
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<emphasis id="458E6521FF907B55FF7A7FD0763BFB44" box="[151,372,1249,1273]" italics="true" pageId="15" pageNumber="290">Lophura amboinensis</emphasis>
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var
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<emphasis id="458E6521FF907B55FE4D7FD07547FB44" box="[416,520,1249,1273]" italics="true" pageId="15" pageNumber="290">celebensis</emphasis>
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</taxonomicName>
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(
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<treatmentCitation id="F65B9F22FF907B55FDF87FD07588FB47" author="Weber, M." box="[533,711,1249,1274]" page="160" pageId="15" pageNumber="290" year="1890">
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<bibRefCitation id="136BC4C2FF907B55FDF87FD07588FB47" author="Weber, M." box="[533,711,1249,1274]" pageId="15" pageNumber="290" pagination="159 - 177" refId="ref20299" refString="Weber, M. (1890) Reptilia from the Malay Archipelago. I. Sauria, Crocodilidae, Chelonia. In: Weber, M. (Ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. E. J. Brill, Leiden, pp. 159 - 177. https: // doi. org / 10.5962 / bhl. title. 52289" type="book chapter" year="1890">Weber 1890: 160</bibRefCitation>
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, 167)
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<paragraph id="7745B933FF907B55FF7A7E3175FDFAA5" blockId="15.[151,1311,1157,1396]" box="[151,690,1279,1304]" pageId="15" pageNumber="290">
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<treatmentCitationGroup id="57EA9E1DFF907B55FF7A7E3175FDFAA5" box="[151,690,1279,1304]" pageId="15" pageNumber="290">
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<taxonomicName id="B0FAC2B0FF907B55FF7A7E3176EFFAA5" baseAuthorityName="Schlosser" baseAuthorityYear="1768" box="[151,416,1280,1304]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="amboinensis">
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<emphasis id="458E6521FF907B55FF7A7E3176EFFAA5" box="[151,416,1280,1304]" italics="true" pageId="15" pageNumber="290">Hydrosaurus amboinensis</emphasis>
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</taxonomicName>
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(part) (
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<treatmentCitation id="F65B9F22FF907B55FE067FCE75E3FAA5" author="Wermuth, H." box="[491,684,1279,1304]" page="64" pageId="15" pageNumber="290" year="1967">
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<bibRefCitation id="136BC4C2FF907B55FE067FCE75E3FAA5" author="Wermuth, H." box="[491,684,1279,1304]" pageId="15" pageNumber="290" pagination="1 - 127" refId="ref20367" refString="Wermuth, H. (1967) Liste der rezenten Amphibien und Reptilien. Agamidae. Das Tierreich, 86, i-xiv + 1 - 127." type="journal article" year="1967">Wermuth 1967: 64</bibRefCitation>
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)
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</paragraph>
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<paragraph id="7745B933FF907B55FF7A7E2E7535FA8A" blockId="15.[151,1311,1157,1396]" box="[151,634,1310,1335]" pageId="15" pageNumber="290">
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<treatmentCitationGroup id="57EA9E1DFF907B55FF7A7E2E7535FA8A" box="[151,634,1310,1335]" pageId="15" pageNumber="290">
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<treatmentCitation id="F65B9F22FF907B55FF7A7E2E7535FA8A" author="Koch, A." box="[151,634,1310,1335]" page="149" pageId="15" pageNumber="290" year="2012">
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<taxonomicName id="B0FAC2B0FF907B55FF7A7E2E7535FA8A" authority="(Koch 2012: 149)" baseAuthorityName="Koch" baseAuthorityPageNumber="149" baseAuthorityYear="2012" box="[151,634,1310,1335]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="amboinensis">
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<emphasis id="458E6521FF907B55FF7A7E2E7655FA8A" box="[151,282,1311,1335]" italics="true" pageId="15" pageNumber="290">Hydrosaurus</emphasis>
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cf.
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<emphasis id="458E6521FF907B55FEAC7E2E768FFA8A" box="[321,448,1311,1335]" italics="true" pageId="15" pageNumber="290">amboinensis</emphasis>
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(
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<bibRefCitation id="136BC4C2FF907B55FE207E2F753CFA8A" author="Koch, A." box="[461,627,1310,1335]" pageId="15" pageNumber="290" refId="ref18570" refString="Koch, A. (2012) Discovery, Diversity, and Distribution of the Amphibians and Reptiles of Sulawesi and its Offshore Islands. Edition Chimaira, Frankfurt a. M., 374 pp." type="book" year="2012">Koch 2012: 149</bibRefCitation>
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)
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<paragraph id="7745B933FF907B55FF7A7E0C75D3FAE8" blockId="15.[151,1311,1157,1396]" box="[151,668,1341,1366]" pageId="15" pageNumber="290">
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<treatmentCitationGroup id="57EA9E1DFF907B55FF7A7E0C75D3FAE8" box="[151,668,1341,1366]" pageId="15" pageNumber="290">
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<taxonomicName id="B0FAC2B0FF907B55FF7A7E0C7655FAE8" authorityName="Kaup" authorityYear="1828" box="[151,282,1341,1365]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="genus">
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<emphasis id="458E6521FF907B55FF7A7E0C7655FAE8" box="[151,282,1341,1365]" italics="true" pageId="15" pageNumber="290">Hydrosaurus</emphasis>
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</taxonomicName>
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sp. [
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|
<collectingRegion id="B53E77D1FF907B55FEA77E0C76E5FAEB" box="[330,426,1341,1366]" country="Indonesia" name="Sulawesi Utara" pageId="15" pageNumber="290">Sulawesi</collectingRegion>
|
|
] (
|
|
<treatmentCitation id="F65B9F22FF907B55FE537E0C75DAFAE8" author="Siler, C. D. & Lira-Noriega, A. & Brown, R. M." box="[446,661,1341,1366]" page="102" pageId="15" pageNumber="290" year="2014">
|
|
<bibRefCitation id="136BC4C2FF907B55FE537E0C75DAFAE8" author="Siler, C. D. & Lira-Noriega, A. & Brown, R. M." box="[446,661,1341,1366]" pageId="15" pageNumber="290" pagination="100 - 108" refId="ref20070" refString="Siler, C. D., Lira-Noriega, A. & Brown, R. M. (2014) Conservation genetics of Australasian sailfin lizards: Flagship species threatened by coastal development and insufficient protected area coverage. Biological Conservation, 169, 100 - 108 https: // doi. org / 10.1016 / j. biocon. 2013.10.014" type="journal article" year="2014">
|
|
Siler
|
|
<emphasis id="458E6521FF907B55FE187E0F756DFAE8" box="[501,546,1341,1365]" italics="true" pageId="15" pageNumber="290">et al</emphasis>
|
|
. 2014: 102
|
|
</bibRefCitation>
|
|
</treatmentCitation>
|
|
)
|
|
</treatmentCitationGroup>
|
|
</paragraph>
|
|
<paragraph id="7745B933FF907B55FF7A7E6D753AFAC9" blockId="15.[151,1311,1157,1396]" box="[151,629,1371,1396]" pageId="15" pageNumber="290">
|
|
<treatmentCitationGroup id="57EA9E1DFF907B55FF7A7E6D753AFAC9" box="[151,629,1371,1396]" pageId="15" pageNumber="290">
|
|
<treatmentCitation id="F65B9F22FF907B55FF7A7E6D753AFAC9" author="Siler, C. D. & Lira-Noriega, A. & Brown, R. M." box="[151,629,1371,1396]" page="104" pageId="15" pageNumber="290" year="2014">
|
|
<taxonomicName id="B0FAC2B0FF907B55FF7A7E6D753AFAC9" authority="(Siler et al. 2014: 104)" baseAuthorityName="Siler" baseAuthorityPageNumber="104" baseAuthorityYear="2014" box="[151,629,1371,1396]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF907B55FF7A7E6D76C6FAC9" box="[151,393,1372,1396]" italics="true" pageId="15" pageNumber="290">Hydrosaurus celebensis</emphasis>
|
|
(
|
|
<bibRefCitation id="136BC4C2FF907B55FE7B7E6A7521FAC9" author="Siler, C. D. & Lira-Noriega, A. & Brown, R. M." box="[406,622,1371,1396]" pageId="15" pageNumber="290" pagination="100 - 108" refId="ref20070" refString="Siler, C. D., Lira-Noriega, A. & Brown, R. M. (2014) Conservation genetics of Australasian sailfin lizards: Flagship species threatened by coastal development and insufficient protected area coverage. Biological Conservation, 169, 100 - 108 https: // doi. org / 10.1016 / j. biocon. 2013.10.014" type="journal article" year="2014">
|
|
Siler
|
|
<emphasis id="458E6521FF907B55FE237E6C76B4FAC9" box="[462,507,1372,1396]" italics="true" pageId="15" pageNumber="290">et al</emphasis>
|
|
. 2014: 104
|
|
</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
</treatmentCitation>
|
|
</treatmentCitationGroup>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="3FE0EAB8FF907B55FF7A7E9375F8F9B9" pageId="15" pageNumber="290" type="materials_examined">
|
|
<paragraph id="7745B933FF907B55FF7A7E937494FA01" blockId="15.[151,1437,1442,2045]" box="[151,987,1442,1469]" pageId="15" pageNumber="290">
|
|
<emphasis id="458E6521FF907B55FF7A7E937794FA01" bold="true" box="[151,219,1442,1468]" pageId="15" pageNumber="290">Type:</emphasis>
|
|
<materialsCitation id="C792B36EFF907B55FF0F7E937494FA01" ID-GBIF-Occurrence="2573218758" box="[226,987,1442,1469]" collectionCode="ZMB" collectorName="A. B. Meyer" pageId="15" pageNumber="290" specimenCode="ZMB 7393" specimenCount="1" typeStatus="holotype">
|
|
<typeStatus id="A8410791FF907B55FF0F7E937605FA01" box="[226,330,1442,1468]" pageId="15" pageNumber="290" type="holotype">Holotype</typeStatus>
|
|
<specimenCode id="275C1148FF907B55FEBC7E92769FFA01" box="[337,464,1442,1469]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="15" pageNumber="290" type="Museum">ZMB 7393</specimenCode>
|
|
, adult male, collected by
|
|
<collectorName id="DA0FDCE5FF907B55FD017E937437FA00" box="[748,888,1442,1469]" pageId="15" pageNumber="290">A. B. Meyer</collectorName>
|
|
in 1870.
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<paragraph id="7745B933FF907B55FF2A7EF775F8F9B9" blockId="15.[151,1437,1442,2045]" pageId="15" pageNumber="290">
|
|
<materialsCitation id="C792B36EFF907B55FF2A7EF775FDF9B9" ID-GBIF-Occurrence="2573218759" country="Indonesia" location="Poso river" municipality="Bay of Tomini" pageId="15" pageNumber="290" specimenCount="1" stateProvince="Sulawesi Tengah" typeStatus="holotype">
|
|
<emphasis id="458E6521FF907B55FF2A7EF776CEFA5D" bold="true" box="[199,385,1477,1504]" pageId="15" pageNumber="290">
|
|
<typeStatus id="A8410791FF907B55FF2A7EF7764DFA5D" box="[199,258,1478,1504]" pageId="15" pageNumber="290">Type</typeStatus>
|
|
locality: “
|
|
</emphasis>
|
|
Ufer des Flusses von Posso in der Bai von Tomini” (= banks of the
|
|
<location id="7225EFE8FF907B55FB677EF673B1FA5D" LSID="urn:lsid:plazi:treatment:FF530825FF907B48FF7A7F70750BFE4A:7225EFE8FF907B55FB677EF673B1FA5D" box="[1162,1278,1478,1505]" country="Indonesia" municipality="Bay of Tomini" name="Poso river" pageId="15" pageNumber="290" stateProvince="Sulawesi Tengah">Poso river</location>
|
|
in the
|
|
<collectingMunicipality id="97212349FF907B55FAA27EF677A3F9B8" pageId="15" pageNumber="290">Bay of Tomini</collectingMunicipality>
|
|
),
|
|
<collectingRegion id="B53E77D1FF907B55FF127EDB768FF9B9" box="[255,448,1514,1540]" country="Indonesia" name="Sulawesi Tengah" pageId="15" pageNumber="290">Central Sulawesi</collectingRegion>
|
|
,
|
|
<collectingCountry id="0FEDF9A3FF907B55FE217EDB7576F9B9" box="[460,569,1514,1540]" name="Indonesia" pageId="15" pageNumber="290">Indonesia</collectingCountry>
|
|
(
|
|
<figureCitation id="EFC1A5B6FF907B55FDAA7EDB75E6F9B9" box="[583,681,1514,1540]" captionStart="FIGURE 8" captionStartId="18.[151,250,1846,1871]" captionTargetBox="[151,1436,524,1826]" captionTargetId="figure@18.[151,1436,524,1829]" captionTargetPageId="18" captionText="FIGURE 8. Map of Sulawesi showing the distribution of H. microlophus (red) and H. celebensis (grey). Type localities are indicated by stars.. 0 Makassar; 1 Bantimurung; 2 Pattunuang; 3 Bengo-Bengo & Rompegading; 4 Camba; 5 Ujung Lamuru; 6 Belopa; 7 Palopo-Pompengan & Palopo-Siguntu; 8 Toraja; 9 Malele; 10 Poso River, Bay of Tomini; 11 Desa Jelaja; 12 Wasponda; 13 Desa Limbatu; 14 Morowali Nature Reserve; W1 Tempe („at the Minralang river near Tempe“ after Weber1890: 167); W2 Pampanua („at Pampanua on the river Tjinrana“ after Weber 1890: 167)" figureDoi="http://doi.org/10.5281/zenodo.3694822" httpUri="https://zenodo.org/record/3694822/files/figure.png" pageId="15" pageNumber="290">Figure 8</figureCitation>
|
|
)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="3FE0EAB8FF907B55FF2A7D3C7527F995" box="[199,616,1549,1576]" pageId="15" pageNumber="290" type="description">
|
|
<paragraph id="7745B933FF907B55FF2A7D3C7527F995" blockId="15.[151,1437,1442,2045]" box="[199,616,1549,1576]" pageId="15" pageNumber="290">
|
|
<emphasis id="458E6521FF907B55FF2A7D3C76A8F995" bold="true" box="[199,487,1549,1576]" pageId="15" pageNumber="290">
|
|
Description of
|
|
<typeStatus id="A8410791FF907B55FE9A7D3F76ADF995" box="[375,482,1550,1576]" pageId="15" pageNumber="290" type="holotype">holotype</typeStatus>
|
|
:
|
|
</emphasis>
|
|
See above.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="3FE0EAB8FF907B4AFF2A7D0075A4FCE2" lastPageId="16" lastPageNumber="291" pageId="15" pageNumber="290" type="diagnosis">
|
|
<paragraph id="7745B933FF907B55FF2A7D0075F1F8D1" blockId="15.[151,1437,1442,2045]" pageId="15" pageNumber="290">
|
|
<emphasis id="458E6521FF907B55FF2A7D00760BF9F1" bold="true" box="[199,324,1585,1612]" pageId="15" pageNumber="290">Diagnosis:</emphasis>
|
|
Large species, exceeding
|
|
<quantity id="B00214D6FF907B55FD807D037596F9F0" box="[621,729,1586,1613]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="290" unit="mm" value="1000.0">1000 mm</quantity>
|
|
in total length, possibly
|
|
<quantity id="B00214D6FF907B55FC1F7D037311F9F0" box="[1010,1118,1586,1613]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="290" unit="mm" value="1200.0">1200 mm</quantity>
|
|
(assuming TL/SVL = 2.4, a complete tail of
|
|
<typeStatus id="A8410791FF907B55FEBE7D6776FAF9CD" box="[339,437,1622,1648]" pageId="15" pageNumber="290" type="holotype">holotype</typeStatus>
|
|
amounts to
|
|
<quantity id="B00214D6FF907B55FDA97D6775E8F9CC" box="[580,679,1622,1649]" metricMagnitude="-1" metricUnit="m" metricValue="8.64" pageId="15" pageNumber="290" unit="mm" value="864.0">864 mm</quantity>
|
|
); largest specimen examined,
|
|
<typeStatus id="A8410791FF907B55FC127D67732EF9CD" box="[1023,1121,1622,1648]" pageId="15" pageNumber="290" type="holotype">holotype</typeStatus>
|
|
SVL
|
|
<quantity id="B00214D6FF907B55FB4B7D677248F9CC" box="[1190,1287,1622,1649]" metricMagnitude="-1" metricUnit="m" metricValue="3.6" pageId="15" pageNumber="290" unit="mm" value="360.0">360 mm</quantity>
|
|
, TL
|
|
<quantity id="B00214D6FF907B55FAD37D6472D2F9CC" box="[1342,1437,1621,1649]" metricMagnitude="-1" metricUnit="m" metricValue="5.8" pageId="15" pageNumber="290" unit="mm" value="580.0">580 mm</quantity>
|
|
(TL/SVL = 1.61), part of the tail missing; complete specimen, MZB Lac271, SVL
|
|
<quantity id="B00214D6FF907B55FBC17D4B73C4F928" box="[1068,1163,1658,1685]" metricMagnitude="-1" metricUnit="m" metricValue="2.33" pageId="15" pageNumber="290" unit="mm" value="233.0">233 mm</quantity>
|
|
, TL
|
|
<quantity id="B00214D6FF907B55FB507D487256F928" box="[1213,1305,1657,1685]" metricMagnitude="-1" metricUnit="m" metricValue="5.63" pageId="15" pageNumber="290" unit="mm" value="563.0">563 mm</quantity>
|
|
(TL/SVL = 2.42). Head, neck, gular region and shoulder completely black; a row of enlarged flat, sometimes conical scales on either side of the neck; nuchal and dorsal crests continuous; a group of dirty white enlarged flat scales on the anterior part of the dorsum; few (<10) additional enlarged scales approximately at midbody and before the hindlimbs. Dorsal colouration typically yellowish, sometimes dark orange, interspersed with black spots; ventrally beige, limbs black with a few yellow spots; scales under fourth and fifth toes broad with several keels from near the base of the toe; tail black, sail black or dark violet with black stripes.
|
|
</paragraph>
|
|
<paragraph id="7745B933FF907B4AFF2A7C447785FEA2" blockId="15.[151,1437,1442,2045]" lastBlockId="16.[151,1437,153,2015]" lastPageId="16" lastPageNumber="291" pageId="15" pageNumber="290">
|
|
<emphasis id="458E6521FF907B55FF2A7C447690F82D" bold="true" box="[199,479,1909,1936]" pageId="15" pageNumber="290">Intraspecific Variation:</emphasis>
|
|
The number of enlarged scales along the neck varies from three to five. Equally, the number of enlarged scales on the dorsum varies in the anterior group from two to seven. At midbody and near the base of the tail there are typically only one or two enlarged scales. One male from Pattunuang had the midbody group split into two groups of two scales giving the impression of a crossband. While the ventral scales in
|
|
<emphasis id="458E6521FF907B55FABA7CD27224F841" box="[1367,1387,2019,2044]" italics="true" pageId="15" pageNumber="290">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF907B4AFA977CD277BEFF0E" italics="true" lastPageId="16" lastPageNumber="291" pageId="15" pageNumber="290">ce- lebensis</emphasis>
|
|
are typically smaller than the dorsal scales, a photograph (courtesy of J. McGuire) of the specimen (tissue sample JAM 6859) from Desa Jelaja, Southwest
|
|
<collectingRegion id="B53E77D1FF8F7B4AFD537B8C7469FF6A" box="[702,806,189,215]" country="Indonesia" name="Sulawesi Utara" pageId="16" pageNumber="291">Sulawesi</collectingRegion>
|
|
, used in the phylogenetic analysis conducted by
|
|
<bibRefCitation id="136BC4C2FF8F7B4AFAA17B8C7648FF46" author="Siler, C. D. & Lira-Noriega, A. & Brown, R. M." pageId="16" pageNumber="291" pagination="100 - 108" refId="ref20070" refString="Siler, C. D., Lira-Noriega, A. & Brown, R. M. (2014) Conservation genetics of Australasian sailfin lizards: Flagship species threatened by coastal development and insufficient protected area coverage. Biological Conservation, 169, 100 - 108 https: // doi. org / 10.1016 / j. biocon. 2013.10.014" type="journal article" year="2014">
|
|
Siler
|
|
<emphasis id="458E6521FF8F7B4AFA657B8F77E2FF46" italics="true" pageId="16" pageNumber="291">et al</emphasis>
|
|
. (2014)
|
|
</bibRefCitation>
|
|
shows that the size of the ventral scales appear to be larger than or equal to the size of the dorsal scale size.
|
|
</paragraph>
|
|
<paragraph id="7745B933FF8F7B4AFF2A7A19721FFE4A" blockId="16.[151,1437,153,2015]" pageId="16" pageNumber="291">
|
|
<emphasis id="458E6521FF8F7B4AFF2A7A1976F7FEFE" bold="true" box="[199,440,296,323]" pageId="16" pageNumber="291">Sexual dimorphism:</emphasis>
|
|
Adult males of
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFD857A1B75B3FEFE" box="[616,764,297,323]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFD857A1B75B3FEFE" box="[616,764,297,323]" italics="true" pageId="16" pageNumber="291">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
have a considerably higher dorsal sail and a more developed nasal crest than in females. In general, the colouration and pattern is more contrasted in males while females appear to be duller in colour. Typically males of
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFD9C7A437447FE36" box="[625,776,369,395]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFD9C7A437447FE36" box="[625,776,369,395]" italics="true" pageId="16" pageNumber="291">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
are black and yellow (or light orange) spotted whereas in females the spotted pattern is less pronounced consisting of an olive-brown, greyish ground colour with black spots in the Pattunuang population and rather greenish with light spots in the Palopo population. The colour of the head and anterior part of the neck is black in males while it is dark brown or grey with white speckles in females.
|
|
</paragraph>
|
|
<paragraph id="7745B933FF8F7B4AFF2A793175A4FCE2" blockId="16.[151,1437,153,2015]" pageId="16" pageNumber="291">
|
|
<emphasis id="458E6521FF8F7B4AFF2A79317620FDA6" bold="true" box="[199,367,512,539]" pageId="16" pageNumber="291">Comparisons:</emphasis>
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFE9B793075CFFDA6" box="[374,640,513,539]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFE9B793075CFFDA6" box="[374,640,513,539]" italics="true" pageId="16" pageNumber="291">Hydrosaurus celebensis</emphasis>
|
|
</taxonomicName>
|
|
differs from
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFCFE7933748FFDA6" baseAuthorityName="Koch" baseAuthorityYear="2012" box="[787,960,513,539]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="amboinensis">
|
|
<emphasis id="458E6521FF8F7B4AFCFE7933748FFDA6" box="[787,960,513,539]" italics="true" pageId="16" pageNumber="291">H. amboinensis</emphasis>
|
|
</taxonomicName>
|
|
by the possession of the following characters: a group of strongly enlarged, flat scales on the anterior part of the dorsum. The colour on the dorsum is yellowish (vs. yellowish green in male
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFDEB797B75FAFDDE" baseAuthorityName="Koch" baseAuthorityYear="2012" box="[518,693,585,611]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="amboinensis">
|
|
<emphasis id="458E6521FF8F7B4AFDEB797B75FAFDDE" box="[518,693,585,611]" italics="true" pageId="16" pageNumber="291">H. amboinensis</emphasis>
|
|
</taxonomicName>
|
|
and green in females). Head, limbs and tail of male
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFAEB797B72D3FDDE" box="[1286,1436,585,611]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFAEB797B72D3FDDE" box="[1286,1436,585,611]" italics="true" pageId="16" pageNumber="291">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
are completely or nearly completely black whereas both sexes in
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFC6E795F737CFD3A" baseAuthorityName="Koch" baseAuthorityYear="2012" box="[899,1075,621,647]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="amboinensis">
|
|
<emphasis id="458E6521FF8F7B4AFC6E795F737CFD3A" box="[899,1075,621,647]" italics="true" pageId="16" pageNumber="291">H. amboinensis</emphasis>
|
|
</taxonomicName>
|
|
have an entirely black marbled body. Additionally, the gular scales of
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFDA579A375B8FD16" baseAuthorityName="Koch" baseAuthorityYear="2012" box="[584,759,657,683]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="amboinensis">
|
|
<emphasis id="458E6521FF8F7B4AFDA579A375B8FD16" box="[584,759,657,683]" italics="true" pageId="16" pageNumber="291">H. amboinensis</emphasis>
|
|
</taxonomicName>
|
|
are round and irregular in size, while those of
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFAEB79A372D3FD16" box="[1286,1436,657,683]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFAEB79A37255FD16" box="[1286,1306,658,683]" italics="true" pageId="16" pageNumber="291">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8F7B4AFAC479A072D3FD16" box="[1321,1436,657,683]" italics="true" pageId="16" pageNumber="291">celebensis</emphasis>
|
|
</taxonomicName>
|
|
are minute and decreasing in size from the mental towards the onset of the gular pouch.
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFBB0798773BEFD72" box="[1117,1265,693,719]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFBB0798773BEFD72" box="[1117,1265,693,719]" italics="true" pageId="16" pageNumber="291">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
differs from
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFA6C798777ACFD4E" authorityName="Barbour" authorityYear="1911" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="weberi">
|
|
<emphasis id="458E6521FF8F7B4AFA6C798777ACFD4E" italics="true" pageId="16" pageNumber="291">H. weberi</emphasis>
|
|
</taxonomicName>
|
|
in its general colour and pattern on the dorsum which is yellowish with black dots (vs. grey green in
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFAAD79E872C3FD4E" authorityName="Barbour" authorityYear="1911" box="[1344,1420,729,755]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="weberi">
|
|
<emphasis id="458E6521FF8F7B4AFAAD79E872C3FD4E" box="[1344,1420,729,755]" italics="true" pageId="16" pageNumber="291">weberi</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFF7A79CF7661FCAA" box="[151,302,765,791]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFF7A79CF77E4FCAA" box="[151,171,766,791]" italics="true" pageId="16" pageNumber="291">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8F7B4AFF5679CC7661FCAA" box="[187,302,765,791]" italics="true" pageId="16" pageNumber="291">celebensis</emphasis>
|
|
</taxonomicName>
|
|
differs from
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFE2B79CF7579FCAA" authorityName="Barbour" authorityYear="1911" box="[454,566,765,791]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="weberi">
|
|
<emphasis id="458E6521FF8F7B4AFE2B79CF7695FCAA" box="[454,474,766,791]" italics="true" pageId="16" pageNumber="291">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8F7B4AFE0779CC7579FCAA" box="[490,566,765,791]" italics="true" pageId="16" pageNumber="291">weberi</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFD9D79CF7450FCAA" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[624,799,765,791]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8F7B4AFD9D79CF75CBFCAA" box="[624,644,766,791]" italics="true" pageId="16" pageNumber="291">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8F7B4AFD7979CC7450FCAA" box="[660,799,765,791]" italics="true" pageId="16" pageNumber="291">microlophus</emphasis>
|
|
</taxonomicName>
|
|
by less pronounced enlarged (i.e. armour-like) scales at the front of the forelimbs. In
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFE337813753CFC86" box="[478,627,801,827]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFE33781376BDFC86" box="[478,498,802,827]" italics="true" pageId="16" pageNumber="291">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8F7B4AFDED7810753CFC86" box="[512,627,801,827]" italics="true" pageId="16" pageNumber="291">celebensis</emphasis>
|
|
</taxonomicName>
|
|
the nuchal and dorsal crests are continuous, while they are separated by considerably smaller scales on the neck in
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFD98787775ABFCE2" authorityName="Barbour" authorityYear="1911" box="[629,740,837,863]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="weberi">
|
|
<emphasis id="458E6521FF8F7B4AFD98787775C6FCE2" box="[629,649,838,863]" italics="true" pageId="16" pageNumber="291">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8F7B4AFD75787475ABFCE2" box="[664,740,837,863]" italics="true" pageId="16" pageNumber="291">weberi</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="3FE0EAB8FF8F7B4AFF2A785975E5FB1E" pageId="16" pageNumber="291" type="biology_ecology">
|
|
<paragraph id="7745B933FF8F7B4AFF2A785975E5FB1E" blockId="16.[151,1437,153,2015]" pageId="16" pageNumber="291">
|
|
<emphasis id="458E6521FF8F7B4AFF2A78597664FC3E" bold="true" box="[199,299,872,899]" pageId="16" pageNumber="291">Biology:</emphasis>
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFEDD785B76F3FC3E" box="[304,444,873,899]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFEDD785B76F3FC3E" box="[304,444,873,899]" italics="true" pageId="16" pageNumber="291">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
was observed in a variety of habitats. In particular, the habitat of
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFBBA785B73ADFC3E" box="[1111,1250,873,899]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFBBA785B7324FC3E" box="[1111,1131,874,899]" italics="true" pageId="16" pageNumber="291">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8F7B4AFB98785873ADFC3E" box="[1141,1250,873,899]" italics="true" pageId="16" pageNumber="291">celebensis</emphasis>
|
|
</taxonomicName>
|
|
in the Pattunuang canyon differs from that in Bantimurung and near Palopo. The Pattunuang river is very stony and during the rainy season flows at a rapid pace. Its riverbank vegetation mainly consists of trees and only a few bushes. In Bantimurung specimens of
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFF5978E7760CFC52" box="[180,323,981,1007]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFF5978E7760CFC52" box="[180,323,981,1007]" italics="true" pageId="16" pageNumber="291">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
were spotted in certain places where
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFD3E78E77437FC52" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[723,888,981,1007]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8F7B4AFD3E78E77437FC52" box="[723,888,981,1007]" italics="true" pageId="16" pageNumber="291">H. microlophus</emphasis>
|
|
</taxonomicName>
|
|
(see above) was not found.
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFB4878E7727CFC52" box="[1189,1331,981,1007]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFB4878E7727CFC52" box="[1189,1331,981,1007]" italics="true" pageId="16" pageNumber="291">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
inhabited more inaccessible areas with dense vegetation on the banks and was often seen on branches above the water. Near Palopo
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFF7A7F2F766CFB8A" box="[151,291,1053,1079]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFF7A7F2F766CFB8A" box="[151,291,1053,1079]" italics="true" pageId="16" pageNumber="291">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
inhabited an area near a slow flowing river without stones or rocks. Most of the individuals were found sitting in dense, low vegetation. Some of the subadults were spotted on thin branches above the water. In some places they were also spotted high up in trees where adults could be seen resting on branches. An adult female carrying eggs could also be observed in Pattanuang towards the end of March.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="3FE0EAB8FF8F7B48FF2A7F9D750BFE4A" lastPageId="18" lastPageNumber="293" pageId="16" pageNumber="291" type="distribution">
|
|
<paragraph id="7745B933FF8F7B4AFF2A7F9D7560FAEA" blockId="16.[151,1437,153,2015]" pageId="16" pageNumber="291">
|
|
<emphasis id="458E6521FF8F7B4AFF2A7F9D762CFB7A" bold="true" box="[199,355,1196,1223]" pageId="16" pageNumber="291">Distribution:</emphasis>
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFE847F9F76B9FB7A" box="[361,502,1197,1223]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFE847F9F76B9FB7A" box="[361,502,1197,1223]" italics="true" pageId="16" pageNumber="291">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
is endemic to
|
|
<collectingRegion id="B53E77D1FF8F7B4AFD607F9C75BFFB7A" box="[653,752,1197,1223]" country="Indonesia" name="Sulawesi Utara" pageId="16" pageNumber="291">Sulawesi</collectingRegion>
|
|
. It appears to inhabit the northern part of
|
|
<collectingRegion id="B53E77D1FF8F7B4AFB737F9C720BFB7A" box="[1182,1348,1197,1223]" country="Indonesia" name="Sulawesi Selatan" pageId="16" pageNumber="291">South Sulawesi</collectingRegion>
|
|
,
|
|
<collectingRegion id="B53E77D1FF8F7B4AFAA37F9C77B8FB56" country="Indonesia" name="Sulawesi Tengah" pageId="16" pageNumber="291">Central Sulawesi</collectingRegion>
|
|
and South East
|
|
<collectingRegion id="B53E77D1FF8F7B4AFE4E7FE0754CFB56" box="[419,515,1233,1259]" country="Indonesia" name="Sulawesi Utara" pageId="16" pageNumber="291">Sulawesi</collectingRegion>
|
|
(
|
|
<figureCitation id="EFC1A5B6FF8F7B4AFDFC7FE07522FB56" box="[529,621,1233,1259]" captionStart="FIGURE 8" captionStartId="18.[151,250,1846,1871]" captionTargetBox="[151,1436,524,1826]" captionTargetId="figure@18.[151,1436,524,1829]" captionTargetPageId="18" captionText="FIGURE 8. Map of Sulawesi showing the distribution of H. microlophus (red) and H. celebensis (grey). Type localities are indicated by stars.. 0 Makassar; 1 Bantimurung; 2 Pattunuang; 3 Bengo-Bengo & Rompegading; 4 Camba; 5 Ujung Lamuru; 6 Belopa; 7 Palopo-Pompengan & Palopo-Siguntu; 8 Toraja; 9 Malele; 10 Poso River, Bay of Tomini; 11 Desa Jelaja; 12 Wasponda; 13 Desa Limbatu; 14 Morowali Nature Reserve; W1 Tempe („at the Minralang river near Tempe“ after Weber1890: 167); W2 Pampanua („at Pampanua on the river Tjinrana“ after Weber 1890: 167)" figureDoi="http://doi.org/10.5281/zenodo.3694822" httpUri="https://zenodo.org/record/3694822/files/figure.png" pageId="16" pageNumber="291">Figure 8</figureCitation>
|
|
). The populations in Pattunuang and Bantimurung (
|
|
<collectingRegion id="B53E77D1FF8F7B4AFB7E7FE07275FB51" box="[1171,1338,1233,1260]" country="Indonesia" name="Sulawesi Selatan" pageId="16" pageNumber="291">South Sulawesi</collectingRegion>
|
|
) are possibly a result of recent introductions according to a local source (A. Siady Hamzah, Universitas Hasanuddin, Makassar, UNHAS, pers. comm.) and information on the website of the Bantimurung-Bulusaraung National Park by Taufik
|
|
<bibRefCitation id="136BC4C2FF8F7B4AFAB77E2877AEFAEA" author="Ismail T" pageId="16" pageNumber="291" refId="ref18342" refString="Ismail T (2013) Available from: http: // www. tn-babul. org / index. php? option = com _ content & view = article & id = 511: hydrosaurusseperti-dinosaurus-yah & catid = 49: artikel & Itemid = 1 (last accessed 12 January 2019)" type="url" year="2013">Ismail (2013)</bibRefCitation>
|
|
. See also the Discussion below.
|
|
</paragraph>
|
|
<paragraph id="7745B933FF8F7B4AFF2A7E517450F9CA" blockId="16.[151,1437,153,2015]" pageId="16" pageNumber="291">
|
|
<emphasis id="458E6521FF8F7B4AFF2A7E517361FAC6" bold="true" box="[199,1070,1376,1403]" pageId="16" pageNumber="291">
|
|
Comparison between live specimens of
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFD707E50741DFAC6" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[669,850,1376,1403]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8F7B4AFD707E50741DFAC6" bold="true" box="[669,850,1376,1403]" italics="true" pageId="16" pageNumber="291">H. microlophus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFC637E507367FAC6" box="[910,1064,1376,1403]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFC637E507367FAC6" bold="true" box="[910,1064,1376,1403]" italics="true" pageId="16" pageNumber="291">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
Based on available photographs of live specimens from
|
|
<collectingRegion id="B53E77D1FF8F7B4AFE497EB47546FA22" box="[420,521,1413,1439]" country="Indonesia" name="Sulawesi Utara" pageId="16" pageNumber="291">Sulawesi</collectingRegion>
|
|
both species can be further distinguished by pholidosis, colouration and pattern. While some of the pictured specimens (mostly males) can be directly assigned to
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFBD87E9B723EFA7E" authority="(Peters, 1872)" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[1077,1393,1449,1475]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFBD87E9B7384FA7E" box="[1077,1227,1449,1475]" italics="true" pageId="16" pageNumber="291">H. celebensis</emphasis>
|
|
(
|
|
<bibRefCitation id="136BC4C2FF8F7B4AFB377E987227FA7E" author="Peters, W." box="[1242,1384,1449,1475]" pageId="16" pageNumber="291" pagination="581 - 585" refId="ref19232" refString="Peters, W. (1872) Uber einige von Hrn Dr. A. B. Meyer bei Gorontalo und auf den Togian-Inseln gesammelte Amphibien. Monatsberichte der koniglichen Akademie der Wissenschaften zu Berlin, 1872, 581 - 585." type="journal article" year="1872">Peters, 1872</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
because they have a near identical appearance to that of the
|
|
<typeStatus id="A8410791FF8F7B4AFCF27EFC7400FA5A" box="[799,847,1485,1511]" pageId="16" pageNumber="291">type</typeStatus>
|
|
specimen, others were quite difficult to identify and could only be assigned through inference to
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFD657EC3747AF9B6" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[648,821,1521,1547]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8F7B4AFD657EC3747AF9B6" box="[648,821,1521,1547]" italics="true" pageId="16" pageNumber="291">H. microlophus</emphasis>
|
|
</taxonomicName>
|
|
. In particular, juveniles within the presumed
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFADD7EC377ADF992" baseAuthorityName="Bleeker" baseAuthorityYear="1860" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8F7B4AFADD7EC3720BF9B6" box="[1328,1348,1522,1547]" italics="true" pageId="16" pageNumber="291">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8F7B4AFABE7EC377ADF992" italics="true" pageId="16" pageNumber="291">microlophus</emphasis>
|
|
</taxonomicName>
|
|
populations possess the same arrangements of groups of enlarged dorsal scales as the
|
|
<typeStatus id="A8410791FF8F7B4AFB2A7D2473B8F992" box="[1223,1271,1557,1583]" pageId="16" pageNumber="291">type</typeStatus>
|
|
specimen. All adults within the same populations also show these characteristic groups of enlarged lateral scales and we therefore consider these specimens as belonging to
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFD867D6F7456F9CA" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[619,793,1629,1655]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8F7B4AFD867D6F7530F9CA" box="[619,639,1630,1655]" italics="true" pageId="16" pageNumber="291">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8F7B4AFD637D6C7456F9CA" box="[654,793,1629,1655]" italics="true" pageId="16" pageNumber="291">microlophus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="7745B933FF8F7B4BFF2A7DB07635FE12" blockId="16.[151,1437,153,2015]" lastBlockId="17.[151,1437,153,431]" lastPageId="17" lastPageNumber="292" pageId="16" pageNumber="291">
|
|
Specimens in various developmental stages of
|
|
<emphasis id="458E6521FF8F7B4AFD2A7DB3743AF926" box="[711,885,1665,1691]" italics="true" pageId="16" pageNumber="291">
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFD2A7DB3743EF926" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[711,881,1665,1691]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="microlophus">H. microlophus</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
were observed near Bantimurung, Bengo-Bengo / Camba and Rompegading on the southwestern peninsula of
|
|
<collectingRegion id="B53E77D1FF8F7B4AFCE17D947423F902" box="[780,876,1701,1727]" country="Indonesia" name="Sulawesi Utara" pageId="16" pageNumber="291">Sulawesi</collectingRegion>
|
|
(
|
|
<figureCitation id="EFC1A5B6FF8F7B4AFC957D94749CF902" box="[888,979,1701,1727]" captionStart="FIGURE 8" captionStartId="18.[151,250,1846,1871]" captionTargetBox="[151,1436,524,1826]" captionTargetId="figure@18.[151,1436,524,1829]" captionTargetPageId="18" captionText="FIGURE 8. Map of Sulawesi showing the distribution of H. microlophus (red) and H. celebensis (grey). Type localities are indicated by stars.. 0 Makassar; 1 Bantimurung; 2 Pattunuang; 3 Bengo-Bengo & Rompegading; 4 Camba; 5 Ujung Lamuru; 6 Belopa; 7 Palopo-Pompengan & Palopo-Siguntu; 8 Toraja; 9 Malele; 10 Poso River, Bay of Tomini; 11 Desa Jelaja; 12 Wasponda; 13 Desa Limbatu; 14 Morowali Nature Reserve; W1 Tempe („at the Minralang river near Tempe“ after Weber1890: 167); W2 Pampanua („at Pampanua on the river Tjinrana“ after Weber 1890: 167)" figureDoi="http://doi.org/10.5281/zenodo.3694822" httpUri="https://zenodo.org/record/3694822/files/figure.png" pageId="16" pageNumber="291">Figure 8</figureCitation>
|
|
).
|
|
<figureCitation id="EFC1A5B6FF8F7B4AFC0B7D947372F902" box="[998,1085,1701,1727]" captionStart="FIGURE 7" captionStartId="17.[151,246,1791,1816]" captionTargetBox="[151,1435,445,1772]" captionTargetId="figure@17.[151,1435,445,1772]" captionTargetPageId="17" captionText="FIGURE 7. Juvenile specimen of H. microlophus from Rompegading displaying the characters described by Bleeker (1860). Photo by Andrea Glässer-Trobisch & Dietmar Trobisch." figureDoi="http://doi.org/10.5281/zenodo.3694818" httpUri="https://zenodo.org/record/3694818/files/figure.png" pageId="16" pageNumber="291">Figure 7</figureCitation>
|
|
depicts a juvenile specimen from Rompegading displaying the characters described by Bleeker (1860). In particular the arrangement, size and shape of the enlarged dorsal scales fit well with the type specimen. Since the adult specimens of the southwestern peninsula differ vastly from both the
|
|
<typeStatus id="A8410791FF8F7B4AFEC37C2076C5F896" box="[302,394,1809,1835]" pageId="16" pageNumber="291" type="holotype">holotype</typeStatus>
|
|
and live specimens of
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFD907C237445F896" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[637,778,1809,1835]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFD907C237445F896" box="[637,778,1809,1835]" italics="true" pageId="16" pageNumber="291">H. celebensis</emphasis>
|
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</taxonomicName>
|
|
, hence we conclude that they represent
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFB5D7C23721BF896" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[1200,1364,1809,1835]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8F7B4AFB5D7C23721BF896" box="[1200,1364,1809,1835]" italics="true" pageId="16" pageNumber="291">H. microlophus</emphasis>
|
|
</taxonomicName>
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|
. Adult males (and most females) from these populations have a much more pronounced scalation on the dorsum. The enlarged scales on the dorsum of the adults are strongly keeled and have a tetrahedral appearance (vs. near flat and only slightly keeled in
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFF127C4F76C3F82A" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[255,396,1917,1943]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8F7B4AFF127C4F76C3F82A" box="[255,396,1917,1943]" italics="true" pageId="16" pageNumber="291">H. celebensis</emphasis>
|
|
</taxonomicName>
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|
). In males these scales are black in colour and whitish to yellow in females. An additional feature seen in adult
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFECD7C93768CF806" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[288,451,1953,1979]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8F7B4AFECD7C93768CF806" box="[288,451,1953,1979]" italics="true" pageId="16" pageNumber="291">H. microlophus</emphasis>
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|
</taxonomicName>
|
|
are the strongly enlarged scales on the anterior part of the forelimbs. This character very much resembles the description by
|
|
<bibRefCitation id="136BC4C2FF8F7B4AFE267CF47521F862" author="Barbour, T." box="[459,622,1989,2015]" pageId="16" pageNumber="291" pagination="15 - 22" refId="ref16465" refString="Barbour, T. (1911) New lizards and a new toad from the Dutch East Indies, with notes on other species. Proceedings of the Biological Society of Washington, 24, 15 - 22." type="journal article" year="1911">Barbour (1911)</bibRefCitation>
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|
for
|
|
<taxonomicName id="B0FAC2B0FF8F7B4AFD747CF7744CF862" authorityName="Barbour" authorityYear="1911" box="[665,771,1989,2015]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="291" phylum="Chordata" rank="species" species="weberi">
|
|
<emphasis id="458E6521FF8F7B4AFD747CF775E2F862" box="[665,685,1990,2015]" italics="true" pageId="16" pageNumber="291">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8F7B4AFD577CF4744CF862" box="[698,771,1989,2015]" italics="true" pageId="16" pageNumber="291">weberi</emphasis>
|
|
</taxonomicName>
|
|
who compared it to a piece of “armour”. The ground colour of adult
|
|
<taxonomicName id="B0FAC2B0FF8E7B4BFF3C7BAB763BFF0E" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[209,372,153,179]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="292" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8E7B4BFF3C7BAB77AAFF0E" box="[209,229,154,179]" italics="true" pageId="17" pageNumber="292">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8E7B4BFF1D7BA8763BFF0E" box="[240,372,153,179]" italics="true" pageId="17" pageNumber="292">microlophus</emphasis>
|
|
</taxonomicName>
|
|
males is generally dirty yellow, with the upper parts of the head, extremities and enlarged dorsal scales blackish. The gular region is yellowish. Adult females show a very different colouration (
|
|
<figureCitation id="EFC1A5B6FF8E7B4BFBB77B8C7387FF6A" box="[1114,1224,189,215]" captionStart="FIGURE 3" captionStartId="3.[151,250,1813,1838]" captionTargetBox="[151,1435,291,1762]" captionTargetId="figure@3.[151,1435,291,1763]" captionTargetPageId="3" captionText="FIGURE 3. Comparison of Hydrosaurus amboinensis with the two phenotypes observed on Sulawesi.A: Hydrosaurus amboinensis, male, Air besar (Passo), Ambon, B: Hydrosaurus amboinensis, female, Air besar (Passo), Ambon, C: Hydrosaurus celebensis, male, Pattunuang, Sulawesi, D: Hydrosaurus celebensis, female, Pattunuang, Sulawesi, E: Hydrosaurus microlophus, male, Rompegading, Sulawesi, F: Hydrosaurus microlophus, female, Ujung Lamuru, Sulawesi. Note the pronounced differences in colour pattern as well as number and arrangement of enlarged scales dorsolaterally. Photos by Andrea Glässer-Trobisch & Dietmar Trobisch." figureDoi="http://doi.org/10.5281/zenodo.3694810" httpUri="https://zenodo.org/record/3694810/files/figure.png" pageId="17" pageNumber="292">Figure 3F</figureCitation>
|
|
); the general dorsal ground colour is grey to dark grey or nearly black, while the enlarged scales are light. We did not observe or find a preserved adult male specimen with such a colouration nor did we find a female specimen with dark enlarged scales as described above. The colouration of some subadult females is very similar to that observed in juvenile specimens of both sexes (i.e., grey with cream coloured enlarged scales). The change in colour and development of the shape of the scales from one that is less to one that is more pronounced appears to be in line with ontogenetic development since only adult specimens show sexual dichromatism.
|
|
</paragraph>
|
|
<caption id="2385E9BBFF8E7B4BFF7A7DCE7580F881" ID-DOI="http://doi.org/10.5281/zenodo.3694818" ID-Zenodo-Dep="3694818" httpUri="https://zenodo.org/record/3694818/files/figure.png" pageId="17" pageNumber="292" startId="17.[151,246,1791,1816]" targetBox="[151,1435,445,1772]" targetPageId="17">
|
|
<paragraph id="7745B933FF8E7B4BFF7A7DCE7580F881" blockId="17.[151,1437,1791,1852]" pageId="17" pageNumber="292">
|
|
<emphasis id="458E6521FF8E7B4BFF7A7DCE765FF8A5" bold="true" box="[151,272,1791,1816]" pageId="17" pageNumber="292">FIGURE 7.</emphasis>
|
|
Juvenile specimen of
|
|
<taxonomicName id="B0FAC2B0FF8E7B4BFE027C3075C5F8A5" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[495,650,1792,1816]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="292" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8E7B4BFE027C3075C5F8A5" box="[495,650,1792,1816]" italics="true" pageId="17" pageNumber="292">H. microlophus</emphasis>
|
|
</taxonomicName>
|
|
from Rompegading displaying the characters described by Bleeker (1860). Photo by Andrea Glässer-Trobisch & Dietmar Trobisch.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="7745B933FF8E7B48FF2A7C5B750BFE4A" blockId="17.[151,1437,1898,2033]" lastBlockId="18.[151,1437,153,503]" lastPageId="18" lastPageNumber="293" pageId="17" pageNumber="292">
|
|
A second distinct phenotype (
|
|
<figureCitation id="EFC1A5B6FF8E7B4BFE127C5B75FEF838" box="[511,689,1898,1925]" captionStart="FIGURE 3" captionStartId="3.[151,250,1813,1838]" captionTargetBox="[151,1435,291,1762]" captionTargetId="figure@3.[151,1435,291,1763]" captionTargetPageId="3" captionText="FIGURE 3. Comparison of Hydrosaurus amboinensis with the two phenotypes observed on Sulawesi.A: Hydrosaurus amboinensis, male, Air besar (Passo), Ambon, B: Hydrosaurus amboinensis, female, Air besar (Passo), Ambon, C: Hydrosaurus celebensis, male, Pattunuang, Sulawesi, D: Hydrosaurus celebensis, female, Pattunuang, Sulawesi, E: Hydrosaurus microlophus, male, Rompegading, Sulawesi, F: Hydrosaurus microlophus, female, Ujung Lamuru, Sulawesi. Note the pronounced differences in colour pattern as well as number and arrangement of enlarged scales dorsolaterally. Photos by Andrea Glässer-Trobisch & Dietmar Trobisch." figureDoi="http://doi.org/10.5281/zenodo.3694810" httpUri="https://zenodo.org/record/3694810/files/figure.png" pageId="17" pageNumber="292">Figures 3 C & D</figureCitation>
|
|
) was photographed in the south of
|
|
<collectingRegion id="B53E77D1FF8E7B4BFBC87C5B7386F839" box="[1061,1225,1898,1924]" country="Indonesia" name="Sulawesi Selatan" pageId="17" pageNumber="292">South Sulawesi</collectingRegion>
|
|
(in the valley of the Pattunuang river and in Bantimurung) in the vicinity of a
|
|
<taxonomicName id="B0FAC2B0FF8E7B4BFD157CBE74D2F815" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[760,925,1934,1961]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="292" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8E7B4BFD157CBE7443F815" box="[760,780,1935,1960]" italics="true" pageId="17" pageNumber="292">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8E7B4BFCF47CBF74D2F815" box="[793,925,1934,1960]" italics="true" pageId="17" pageNumber="292">microlophus</emphasis>
|
|
</taxonomicName>
|
|
population and in the north of the Southwestern peninsula near Palopo (
|
|
<figureCitation id="EFC1A5B6FF8E7B4BFE787C8376BCF871" box="[405,499,1970,1996]" captionStart="FIGURE 8" captionStartId="18.[151,250,1846,1871]" captionTargetBox="[151,1436,524,1826]" captionTargetId="figure@18.[151,1436,524,1829]" captionTargetPageId="18" captionText="FIGURE 8. Map of Sulawesi showing the distribution of H. microlophus (red) and H. celebensis (grey). Type localities are indicated by stars.. 0 Makassar; 1 Bantimurung; 2 Pattunuang; 3 Bengo-Bengo & Rompegading; 4 Camba; 5 Ujung Lamuru; 6 Belopa; 7 Palopo-Pompengan & Palopo-Siguntu; 8 Toraja; 9 Malele; 10 Poso River, Bay of Tomini; 11 Desa Jelaja; 12 Wasponda; 13 Desa Limbatu; 14 Morowali Nature Reserve; W1 Tempe („at the Minralang river near Tempe“ after Weber1890: 167); W2 Pampanua („at Pampanua on the river Tjinrana“ after Weber 1890: 167)" figureDoi="http://doi.org/10.5281/zenodo.3694822" httpUri="https://zenodo.org/record/3694822/files/figure.png" pageId="17" pageNumber="292">Figure 8</figureCitation>
|
|
). Males of these populations agree completely with the
|
|
<typeStatus id="A8410791FF8E7B4BFBBB7C8373FDF871" box="[1110,1202,1970,1996]" pageId="17" pageNumber="292" type="holotype">holotype</typeStatus>
|
|
of
|
|
<taxonomicName id="B0FAC2B0FF8E7B4BFB377C827227F871" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[1242,1384,1970,1996]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="292" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8E7B4BFB377C827227F871" box="[1242,1384,1970,1996]" italics="true" pageId="17" pageNumber="292">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
(see
|
|
<figureCitation id="EFC1A5B6FF8E7B4BFF7A7CE7764BF84D" box="[151,260,2006,2032]" captionStart="FIGURE 4" captionStartId="10.[151,250,1885,1910]" captionTargetBox="[151,1436,324,1838]" captionTargetId="figure@10.[151,1436,323,1838]" captionTargetPageId="10" captionText="FIGURE 4. A: The juvenile holotype (NHMUK 1863.12.4.35) of Istiurus (= Hydrosaurus) microlophus Bleeker, 1860 from Makassar, Southwest Sulawesi. Note the groups of enlarged scales on the dorsum. Photo by Patrick D. Campbell. B: The adult male holotype (ZMB 7393) of Lophura (= Hydrosaurus) celebensis Peters, 1872 from the Poso River, Central Sulawesi. Note the very few light enlarged scales on the dorsum. Photo by André Koch." figureDoi="http://doi.org/10.5281/zenodo.3694812" httpUri="https://zenodo.org/record/3694812/files/figure.png" pageId="17" pageNumber="292">Figure 4B</figureCitation>
|
|
). They only have a few enlarged scales interspersed on the dorsum and their dorsal colouration is a pattern which is black and yellow spotted with white enlarged scales dorsolaterally. This particular colouration was observed during the dry season. During the wet season several males, but not all, displayed an even richer colouration, showing contrasts with the yellow coloured scales switching to a bright orange yellow colour. Head, gular region and extremities were completely black. Typically, females were slightly duller in colouration and less rich in contrast. Unfortunately, we were not able to find a juvenile within these populations that we could refer unambiguously to
|
|
<taxonomicName id="B0FAC2B0FF8D7B48FC2E7A1B7301FEFE" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[963,1102,297,323]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="293" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8D7B48FC2E7A1B7301FEFE" box="[963,1102,297,323]" italics="true" pageId="18" pageNumber="293">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
. In both Pattunuang and Palopo where populations of
|
|
<taxonomicName id="B0FAC2B0FF8D7B48FE6C7A7F755FFEDA" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[385,528,333,359]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="293" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8D7B48FE6C7A7F76DAFEDA" box="[385,405,334,359]" italics="true" pageId="18" pageNumber="293">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8D7B48FE4E7A7C755FFEDA" box="[419,528,333,359]" italics="true" pageId="18" pageNumber="293">celebensis</emphasis>
|
|
</taxonomicName>
|
|
were found, almost all juveniles had more or less slightly marked crossbands of enlarged scales. In particular, we found that the juveniles of Pattunuang agreed with the general characteristics described for the type of
|
|
<taxonomicName id="B0FAC2B0FF8D7B48FEE77AA776E3FE12" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[266,428,405,431]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="293" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8D7B48FEE77AA77651FE12" box="[266,286,406,431]" italics="true" pageId="18" pageNumber="293">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF8D7B48FEC47AA476E3FE12" box="[297,428,405,431]" italics="true" pageId="18" pageNumber="293">microlophus</emphasis>
|
|
</taxonomicName>
|
|
. Both in Pattunuang and Palopo adult
|
|
<taxonomicName id="B0FAC2B0FF8D7B48FCD47AA474F2FE12" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[825,957,405,431]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="293" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8D7B48FCD47AA474F2FE12" box="[825,957,405,431]" italics="true" pageId="18" pageNumber="293">microlophus</emphasis>
|
|
</taxonomicName>
|
|
have not been observed but in Pattunuang it is probable that they are in contact with
|
|
<taxonomicName id="B0FAC2B0FF8D7B48FDF07A8875C5FE6E" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[541,650,441,467]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="293" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF8D7B48FDF07A8875C5FE6E" box="[541,650,441,467]" italics="true" pageId="18" pageNumber="293">celebensis</emphasis>
|
|
</taxonomicName>
|
|
because the next localities of
|
|
<taxonomicName id="B0FAC2B0FF8D7B48FC537A88730DFE6E" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[958,1090,441,467]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="293" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF8D7B48FC537A88730DFE6E" box="[958,1090,441,467]" italics="true" pageId="18" pageNumber="293">microlophus</emphasis>
|
|
</taxonomicName>
|
|
are only ca.
|
|
<quantity id="B00214D6FF8D7B48FB297A8873B7FE6E" box="[1220,1272,441,467]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="293" unit="km" value="7.0">7 km</quantity>
|
|
(Bantimurung) resp. about
|
|
<quantity id="B00214D6FF8D7B48FEFC7AEC761BFE4A" box="[273,340,477,503]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="293" unit="km" value="10.0">10 km</quantity>
|
|
(Bengo-Bengo) away.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |