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<document id="AB6191594F109A77C593D9E71E1F5A90" ID-DOI="10.11646/zootaxa.2849.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5288200" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629720910444" checkinUser="felipe" docAuthor="Ferrington Jr, Leonard C. &amp; Saether, Ole A." docDate="2011" docId="EE7687BAFF99FF98BDAABC3817688B6D" docLanguage="en" docName="zt02849p314.pdf" docOrigin="Zootaxa 2849 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2849.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydrosmittia togadistalis Ferrington Jr &amp; Saether 2011, comb. n." docType="treatment" docVersion="4" lastPageNumber="111" masterDocId="124FFFC2FFF7FFF6BD3DBD6214788F3C" masterDocTitle="A revision of the genera Pseudosmittia Edwards, 1932, Allocladius Kieffer, 1913, and Hydrosmittia gen. n. (Diptera: Chironomidae, Orthocladiinae) 2849" masterLastPageNumber="314" masterPageNumber="1" pageNumber="111" updateTime="1699101708276" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A1CED2296315B639F17CC5FCACA2FBCC">A revision of the genera Pseudosmittia Edwards, 1932, Allocladius Kieffer, 1913, and Hydrosmittia gen. n. (Diptera: Chironomidae, Orthocladiinae) 2849</mods:title>
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<mods:namePart id="1761BE8F2FB727270FCCFBC01DC29C7E">Ferrington Jr, Leonard C.</mods:namePart>
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<mods:namePart id="F88585F2008AEEFF94C1D3C1B250DAFF">Saether, Ole A.</mods:namePart>
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<mods:title id="55498D0AB224D9941E2728E3CFABE5CD">Zootaxa</mods:title>
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<mods:date id="2E7922A89111437FE3CCA4FE6052EF36">2011</mods:date>
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<mods:number id="250FE15FDEE3994E7C0BC08583F9674F">2011-04-29</mods:number>
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<mods:number id="17BC88B7009C155C7E151456AD1315A1">2849</mods:number>
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<mods:classification id="EEB05A9DDD08FF9652FDB1975573A17F">journal article</mods:classification>
<mods:identifier id="B591DEB78117716CE5C39B8C451FA4B8" type="DOI">10.11646/zootaxa.2849.1.1</mods:identifier>
<mods:identifier id="583083C4F9891BA107FEF05D76E945E9" type="ISSN">1175­5334</mods:identifier>
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<treatment id="EE7687BAFF99FF98BDAABC3817688B6D" LSID="urn:lsid:plazi:treatment:EE7687BAFF99FF98BDAABC3817688B6D" httpUri="http://treatment.plazi.org/id/EE7687BAFF99FF98BDAABC3817688B6D" lastPageNumber="111" pageId="110" pageNumber="111">
<subSubSection id="2EC56527FF99FF98BDAABC3817D78E48" box="[151,943,346,372]" pageId="110" pageNumber="111" type="nomenclature">
<paragraph id="666036ACFF99FF98BDAABC3817D78E48" blockId="110.[151,943,346,411]" box="[151,943,346,372]" pageId="110" pageNumber="111">
<heading id="3D2881C0FF99FF98BDAABC3817D78E48" bold="true" box="[151,943,346,372]" fontSize="11" level="1" pageId="110" pageNumber="111" reason="1">
<emphasis id="54ABEABEFF99FF98BDAABC3817D78E48" bold="true" box="[151,943,346,372]" pageId="110" pageNumber="111">
<taxonomicName id="A1DF4D2FFF99FF98BDAABC3817458E48" ID-CoL="9HTRX" authority="(Sasa, Watanabe et Arakawa)" authorityName="Ferrington Jr &amp; Saether" authorityYear="2011" baseAuthorityName="Sasa, Watanabe et Arakawa" baseAuthorityYear="2011" box="[151,829,346,372]" class="Insecta" family="Chironomidae" genus="Hydrosmittia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="togadistalis" status="comb. nov.">
<emphasis id="54ABEABEFF99FF98BDAABC3815BC8E48" bold="true" box="[151,452,346,372]" italics="true" pageId="110" pageNumber="111">Hydrosmittia togadistalis</emphasis>
(Sasa, Watanabe
<emphasis id="54ABEABEFF99FF98BF9EBC3816C08E48" bold="true" box="[675,696,346,372]" italics="true" pageId="110" pageNumber="111">et</emphasis>
Arakawa)
</taxonomicName>
<taxonomicNameLabel id="4F9857C5FF99FF98BE79BC3817D78E48" box="[836,943,346,372]" pageId="110" pageNumber="111" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="2EC56527FF99FF98BDAABCE115778EA7" box="[151,271,387,411]" pageId="110" pageNumber="111" type="description">
<paragraph id="666036ACFF99FF98BDAABCE115778EA7" blockId="110.[151,943,346,411]" box="[151,271,387,411]" pageId="110" pageNumber="111">
(
<figureCitation id="FEE42A29FF99FF98BDA2BCE1157E8EA7" box="[159,262,387,411]" captionStart="FIGURE 36" captionStartId="109.[151,250,1955,1978]" captionTargetBox="[379,1211,457,1869]" captionTargetId="figure-112@109.[364,1223,432,1931]" captionTargetPageId="109" captionText="FIGURE 36. Hydrosmittia spp., male hypopygium, left dorsal aspect, right ventral aspect. A—H. kisotriangulata (Sasa et Kondo) comb. n.; B—Hydrosmittia togadistalis (Sasa, Watanabe et Arakawa) comb. n." figureDoi="http://doi.org/10.5281/zenodo.5288369" httpUri="https://zenodo.org/record/5288369/files/figure.png" pageId="110" pageNumber="111">Fig. 36 B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="2EC56527FF99FF98BDAABCA910D28EDD" box="[151,1194,459,481]" pageId="110" pageNumber="111" type="reference_group">
<paragraph id="666036ACFF99FF98BDAABCA910D28EDD" blockId="110.[151,1194,459,481]" box="[151,1194,459,481]" pageId="110" pageNumber="111">
<treatmentCitationGroup id="46CF1182FF99FF98BDAABCA910D28EDD" box="[151,1194,459,481]" pageId="110" pageNumber="111">
<treatmentCitation id="E77E10BDFF99FF98BDAABCA917608EDD" author="Sasa, M. &amp; Watanabe, M. &amp; Arakawa, R." box="[151,792,459,481]" page="231" pageId="110" pageNumber="111" year="1992">
<taxonomicName id="A1DF4D2FFF99FF98BDAABCA917608EDD" ID-CoL="4PBNB" authority="Sasa, Watanabe et Arakawa 1992: 231" authorityName="Sasa, Watanabe et Arakawa" authorityPageNumber="231" authorityYear="1992" box="[151,792,459,481]" class="Insecta" family="Chironomidae" genus="Pseudosmittia" kingdom="Animalia" order="Diptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="togadistalis">
<emphasis id="54ABEABEFF99FF98BDAABCA915E18EDD" box="[151,409,459,481]" italics="true" pageId="110" pageNumber="111">Pseudosmittia togadistalis</emphasis>
<bibRefCitation id="024E4B5DFF99FF98BCA3BCA917608EDD" author="Sasa, M. &amp; Watanabe, M. &amp; Arakawa, R." box="[414,792,459,481]" pageId="110" pageNumber="111" pagination="231 - 246" refId="ref244046" refString="Sasa, M., Watanabe, M. &amp; Arakawa, R. (1992) Additional records of Chironomidae from Togamura, 1992. Research Report from Toyama Prefectural Environmental Pollution Research Centre, 1992, 231 - 246." type="journal article" year="1992">
Sasa, Watanabe
<emphasis id="54ABEABEFF99FF98BF00BCAE16378EDD" box="[573,591,460,481]" italics="true" pageId="110" pageNumber="111">et</emphasis>
Arakawa 1992: 231
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="E77E10BDFF99FF98BE1EBCAE178A8EDD" author="Yamamoto, M." box="[803,1010,459,481]" page="89" pageId="110" pageNumber="111" year="2004">
<bibRefCitation id="024E4B5DFF99FF98BE1EBCAE178A8EDD" author="Yamamoto, M." box="[803,1010,459,481]" pageId="110" pageNumber="111" pagination="1 - 121" refId="ref244823" refString="Yamamoto, M. (2004) A catalog of Japanese Orthocladiinae (Diptera: Chironomidae). Makunagi, Acta Dipterologica, 21, 1 - 121." type="journal article" year="2004">Yamamoto 2004: 89</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="E77E10BDFF99FF98BEC0BCA910DE8EDD" author="Saether, O. A." box="[1021,1190,459,481]" httpUri="http://treatment.plazi.org/id/F46687CDFFF8FFE1147EDBB87A853D92" page="41" pageId="110" pageNumber="111" year="2006">
<bibRefCitation id="024E4B5DFF99FF98BEC0BCA910DE8EDD" author="Saether, O. A." box="[1021,1190,459,481]" pageId="110" pageNumber="111" pagination="21 - 51" refId="ref242508" refString="Saether, O. A. (2006) Japanese Pseudosmittia Edwards (Diptera: Chironomidae). Zootaxa, 1198: 21 - 51." type="journal article" year="2006">Saether 2006: 41</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="2EC56527FF99FF98BDAABF7517688B6D" pageId="110" pageNumber="111" type="materials_examined">
<paragraph id="666036ACFF99FF98BDAABF75115F8D6A" blockId="110.[151,1436,535,676]" pageId="110" pageNumber="111">
<emphasis id="54ABEABEFF99FF98BDAABF75151E8D13" box="[151,358,535,559]" italics="true" pageId="110" pageNumber="111">Material examined</emphasis>
:
<materialsCitation id="D6B73CF1FF99FF98BC49BF75115B8D6A" collectingDate="1992-12-08" collectorName="M. Watanabe &amp; R. Arakawa &amp; No. A &amp; Nos. A" country="Japan" county="Honshu" location="Toga-Mura" municipality="Pref" pageId="110" pageNumber="111" specimenCount="3" stateProvince="Toyama" typeStatus="holotype">
<collectingCountry id="1EC8763CFF99FF98BC49BF7515B68D13" box="[372,462,535,559]" name="Japan" pageId="110" pageNumber="111">JAPAN</collectingCountry>
:
<collectingCounty id="8F014E20FF99FF98BCE5BF7516488D13" box="[472,560,535,559]" pageId="110" pageNumber="111">Honshu</collectingCounty>
,
<collectingRegion id="A41BF84EFF99FF98BF06BF7A16ED8D0C" box="[571,661,536,560]" country="Japan" name="Toyama" pageId="110" pageNumber="111">Toyama</collectingRegion>
<collectingMunicipality id="8604ACD6FF99FF98BFA6BF7516B48D13" box="[667,716,535,559]" pageId="110" pageNumber="111">Pref</collectingMunicipality>
..
<location id="63006077FF99FF98BFE0BF7A17248D0C" LSID="urn:lsid:plazi:treatment:EE7687BAFF99FF98BDAABC3817688B6D:63006077FF99FF98BFE0BF7A17248D0C" box="[733,860,536,560]" country="Japan" county="Honshu" municipality="Pref" name="Toga-Mura" pageId="110" pageNumber="111" stateProvince="Toyama">Toga-Mura</location>
,
<location id="63006077FF99FF98BE5BBF7517A18D13" LSID="urn:lsid:plazi:treatment:EE7687BAFF99FF98BDAABC3817688B6D:63006077FF99FF98BE5BBF7517A18D13" box="[870,985,535,559]" country="Japan" county="Honshu" municipality="Pref" name="Yachidani" pageId="110" pageNumber="111" stateProvince="Toyama">Yachidani</location>
,
<typeStatus id="B964880EFF99FF98BED9BF75103C8D13" box="[996,1092,535,559]" pageId="110" pageNumber="111" type="holotype">holotype</typeStatus>
male,
<date id="1261106CFF99FF98B9B0BF7510868D13" box="[1165,1278,535,559]" pageId="110" pageNumber="111" value="1992-12-08">
<collectingDate id="0225E984FF99FF98B9B0BF7510868D13" box="[1165,1278,535,559]" pageId="110" pageNumber="111" value="1992-12-08">8 xii 1992</collectingDate>
</date>
,
<collectorName id="CB2A537AFF99FF98B834BF7A11E48D13" box="[1289,1436,535,559]" pageId="110" pageNumber="111">M. Watanabe</collectorName>
&amp;
<collectorName id="CB2A537AFF99FF98BD8EBF5D154E8D6A" box="[179,310,574,598]" pageId="110" pageNumber="111">R. Arakawa</collectorName>
(NMST
<collectorName id="CB2A537AFF99FF98BCA5BF5D15A38D6A" box="[408,475,574,598]" pageId="110" pageNumber="111">No. A</collectorName>
191: 95).
<typeStatus id="B964880EFF99FF98BF77BF5D16CC8D6B" box="[586,692,575,599]" pageId="110" pageNumber="111" type="paratype">Paratypes</typeStatus>
males, females, as
<typeStatus id="B964880EFF99FF98BEBBBF5C179D8D6A" box="[902,997,574,598]" pageId="110" pageNumber="111" type="holotype">holotype</typeStatus>
(NMST
<collectorName id="CB2A537AFF99FF98B975BF5D10ED8D6A" box="[1096,1173,574,598]" pageId="110" pageNumber="111">Nos. A</collectorName>
191: 76, 96)
</materialsCitation>
.
</paragraph>
<paragraph id="666036ACFF99FF98BDF2BF04112F8D98" blockId="110.[151,1436,535,676]" pageId="110" pageNumber="111">
<emphasis id="54ABEABEFF99FF98BDF2BF0415BC8D41" box="[207,452,613,637]" italics="true" pageId="110" pageNumber="111">Diagnostic characters</emphasis>
: The well sclerotized hook-like apical projection of the phallapodeme ending in 2–3 distinct, sharp teeth (
<figureCitation id="FEE42A29FF99FF98BCBCBFEE15938D98" box="[385,491,652,676]" captionStart="FIGURE 34" captionStartId="101.[151,250,1628,1651]" captionTargetBox="[265,1323,274,1605]" captionTargetId="figure-132@101.[239,1373,222,1658]" captionTargetPageId="101" captionText="FIGURE 34. Allocladius sp., immmatures. A–B, A. sp. Chile”. A—male pupa, dorsal view; B—male pupa, ventral view. C–F, A. sp. “Falkland Islands”, larva. C—labrum and epipharynx; D—antenna; E—mandible; F—mentum. (From Andersen et al. 2010.)" figureDoi="http://doi.org/10.5281/zenodo.5288357" httpUri="https://zenodo.org/record/5288357/files/figure.png" pageId="110" pageNumber="111">Fig. 34 B</figureCitation>
) will separate the species from all other species of the
<taxonomicName id="A1DF4D2FFF99FF98B96FBFEE10898D98" authorityName="Edwards. This" authorityYear="1932" box="[1106,1265,652,676]" class="Insecta" family="Chironomidae" genus="Pseudosmittia" kingdom="Animalia" order="Diptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="54ABEABEFF99FF98B96FBFEE10898D98" box="[1106,1265,652,676]" italics="true" pageId="110" pageNumber="111">Pseudosmittia</emphasis>
</taxonomicName>
complex
</paragraph>
<paragraph id="666036ACFF99FF98BDAABFB810B18CB2" blockId="110.[151,1436,730,910]" pageId="110" pageNumber="111">
Although no lateral spinules of the virga were observed they could well be present, but weak as in the rest of the
<taxonomicName id="A1DF4D2FFF99FF98BDAABE6314828C25" authorityName="Ferrington Jr &amp; Saether" authorityYear="2011" box="[151,250,769,793]" class="Insecta" family="Chironomidae" genus="Hydrosmittia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="annulata">
<emphasis id="54ABEABEFF99FF98BDAABE6314828C25" box="[151,250,769,793]" italics="true" pageId="110" pageNumber="111">annulata</emphasis>
</taxonomicName>
group. However, the placement within
<taxonomicName id="A1DF4D2FFF99FF98BF83BE63172A8C25" authorityName="Ferrington Jr &amp; Saether" authorityYear="2011" box="[702,850,769,793]" class="Insecta" family="Chironomidae" genus="Hydrosmittia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="54ABEABEFF99FF98BF83BE63172A8C25" box="[702,850,769,793]" italics="true" pageId="110" pageNumber="111">Hydrosmittia</emphasis>
</taxonomicName>
is tentative as the species falls outside the genus in some parsimony tests and the immatures are unknown. We regard the absence of oral projections on the transverse sternapodeme as a synapomorphy with the rest of the
<taxonomicName id="A1DF4D2FFF99FF98BFC3BE2D171A8C5B" authorityName="Ferrington Jr &amp; Saether" authorityYear="2011" box="[766,866,847,871]" class="Insecta" family="Chironomidae" genus="Hydrosmittia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="annulata">
<emphasis id="54ABEABEFF99FF98BFC3BE2D171A8C5B" box="[766,866,847,871]" italics="true" pageId="110" pageNumber="111">annulata</emphasis>
</taxonomicName>
group and the presence of two apical teeth on the phallapodeme, although unsclerotized, present also in
<taxonomicName id="A1DF4D2FFF99FF98BFCABE1517E48CB2" authorityName="Ferrington Jr &amp; Saether" authorityYear="2011" box="[759,924,886,910]" class="Insecta" family="Chironomidae" genus="Hydrosmittia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="tenuistylata">
<emphasis id="54ABEABEFF99FF98BFCABE1517E48CB2" box="[759,924,886,910]" italics="true" pageId="110" pageNumber="111">H. tenuistylata</emphasis>
</taxonomicName>
as a likely synapomorphy.
</paragraph>
<paragraph id="666036ACFF99FF98BDAABEA616DF8CE0" blockId="110.[151,679,964,988]" box="[151,679,964,988]" pageId="110" pageNumber="111">
The species is redescribed by
<bibRefCitation id="024E4B5DFF99FF98BCE1BEA616DB8CE0" author="Saether, O. A." box="[476,675,964,988]" pageId="110" pageNumber="111" pagination="21 - 51" refId="ref242508" refString="Saether, O. A. (2006) Japanese Pseudosmittia Edwards (Diptera: Chironomidae). Zootaxa, 1198: 21 - 51." type="journal article" year="2006">Saether (2006: 41)</bibRefCitation>
.
</paragraph>
<paragraph id="666036ACFF99FF98BDAAB97015648B16" blockId="110.[151,784,1042,1106]" box="[151,284,1042,1066]" pageId="110" pageNumber="111">
<emphasis id="54ABEABEFF99FF98BDAAB97015648B16" box="[151,284,1042,1066]" italics="true" pageId="110" pageNumber="111">Distribution</emphasis>
</paragraph>
<paragraph id="666036ACFF99FF98BDAAB95B17688B6D" blockId="110.[151,784,1042,1106]" box="[151,784,1081,1106]" pageId="110" pageNumber="111">
<materialsCitation id="D6B73CF1FF99FF98BDAAB95B17748B6D" box="[151,780,1081,1106]" country="Japan" location="Japan" pageId="110" pageNumber="111" specimenCount="1">
The species is known only from the
<typeStatus id="B964880EFF99FF98BF1CB95816288B6E" box="[545,592,1082,1106]" pageId="110" pageNumber="111">type</typeStatus>
locality in
<collectingCountry id="1EC8763CFF99FF98BFF7B95817748B6D" box="[714,780,1082,1105]" name="Japan" pageId="110" pageNumber="111">Japan</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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