272 lines
26 KiB
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272 lines
26 KiB
XML
<document ID-CLB-Dataset="7324" ID-DOI="http://dx.doi.org/10.3897/subtbiol.38.64762" ID-GBIF-Dataset="46b559fa-ada2-4f87-b041-ca1836d2f9bf" ID-Pensoft-Pub="1314-2615-38-65" ID-Pensoft-UUID="85A862E2FE78530AAB249C52CD1FE9B8" ID-ZooBank="6A604C2355244D068ED1D71344627018" ModsDocID="1314-2615-38-65" checkinTime="1618544333050" checkinUser="pensoft" docAuthor="Grego 1, Jozef & Pesic 2, Vladimir" docDate="2021" docId="D5A01BF94323595295F04F08B12ACFB7" docLanguage="en" docName="SubterBiol 38: 65-76" docOrigin="Subterranean Biology 38" docPubDate="2021-04-15" docSource="http://dx.doi.org/10.3897/subtbiol.38.64762" docTitle="Travunijana djokovici Grego 1 & Pešić 2 2021, sp. nov." docType="treatment" docUuid="93FAB9F2-2C62-46B8-92D0-8494695354E5" docUuidSource="ZooBank" docVersion="5" id="85A862E2FE78530AAB249C52CD1FE9B8" lastPageNumber="65" masterDocId="85A862E2FE78530AAB249C52CD1FE9B8" masterDocTitle="First record of stygobiotic gastropod genus Travunijana Grego & Gloeer, 2019 (Mollusca, Hydrobiidae) from Montenegro" masterLastPageNumber="76" masterPageNumber="65" pageNumber="65" updateTime="1732870802581" updateUser="ExternalLinkService">
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<mods:titleInfo id="A91529077B33977FDAFBF11A14F2C4CC">
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<mods:title id="04893DCBA381142E54F3FBBFA130FCF7">First record of stygobiotic gastropod genus Travunijana Grego & Gloeer, 2019 (Mollusca, Hydrobiidae) from Montenegro</mods:title>
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<mods:name id="C6B6A25DBA47CAF15089089A7D516AB9" type="personal">
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<mods:role id="EA6C0B8C51EEFFA92AA78B0BFCCB77A5">
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<mods:roleTerm id="8E303C1978B5C612BD058335C02169CB">Author</mods:roleTerm>
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<mods:namePart id="646BC0A299CC7AA26A17A821BAED0DD9">Grego 1, Jozef</mods:namePart>
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<mods:affiliation id="0B8CBB1DD38B2E85EF0141BBEAE792D9">Horna Micina, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
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<mods:roleTerm id="E272242806BB87F8C46CC576428F05C3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3AE5B076C8741B462A3CA1F7D86925D3">Pesic 2, Vladimir</mods:namePart>
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<mods:affiliation id="CC60E2D95A044CD81CC2AFDA5D8FE7C8">Horna Micina, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
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<mods:typeOfResource id="88BFEF8B137653BF3456976BD703B309">text</mods:typeOfResource>
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<mods:titleInfo id="B2A2DB1E8F3171FF267A611B393C8DE4">
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<mods:title id="CD27435A2CF7014DE9E2EBA50CB0803C">Subterranean Biology</mods:title>
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<mods:part id="BDB37A2A20DAE6F4DF81AE78932EF20F">
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<mods:date id="96A372A88DFDFC17574C9C87D3AC3FE5">2021</mods:date>
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<mods:number id="3AB55D4A3872F52F9E840C8073775362">2021-04-15</mods:number>
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<mods:detail id="F2D18549C46FE71713344990ECBE27C9" type="volume">
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<mods:number id="C4194B8A53356FC007074A51BA5CFC56">38</mods:number>
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<mods:start id="D7CD9BC4A0EADA66F57A79660CD264E3">65</mods:start>
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<mods:end id="82355442A6D299F929040729E5AA3A0A">76</mods:end>
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<mods:url id="6DCDDDB74ED9E31E1457C0979708DA16">http://dx.doi.org/10.3897/subtbiol.38.64762</mods:url>
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<mods:classification id="4902C7DDCDDC2C0BF37D9300071A5438">journal article</mods:classification>
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<mods:identifier id="1DEBA0B32D86D95B71AD03B5DC9D1475" type="DOI">http://dx.doi.org/10.3897/subtbiol.38.64762</mods:identifier>
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<treatment id="D5A01BF94323595295F04F08B12ACFB7" ID-GBIF-Taxon="181019285" LSID="urn:lsid:zoobank.org:act:93FAB9F2-2C62-46B8-92D0-8494695354E5" httpUri="http://treatment.plazi.org/id/D5A01BF94323595295F04F08B12ACFB7" lastPageNumber="65" pageId="0" pageNumber="65" scope_family="Hydrobiidae" scope_phylum="Mollusca">
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<subSubSection id="741E0377B6477288E9DEE677905E5A58" pageId="0" pageNumber="65" type="nomenclature">
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<paragraph id="079DB56443773020459294265C8B9144" pageId="0" pageNumber="65">
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<taxonomicName id="0C04A4CED8C4939F87093A6CFACD70DD" ID-CoL="8762N" LSID="http://zoobank.org/93FAB9F2-2C62-46B8-92D0-8494695354E5" authority="Grego 1 & Pešić 2, 2021" authorityName="Grego 1 & Pešić 2" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Travunijana djokovici" order="Littorinimorpha" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="djokovici" status="sp. nov.">Travunijana djokovici</taxonomicName>
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<taxonomicNameLabel id="A0C847C276A66EECDB5302988CC0AE30" pageId="0" pageNumber="65">sp. nov.</taxonomicNameLabel>
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<figureCitation id="F08BBC35B472ED346405A61F11F03BCE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Travunijana djokovici sp. nov. A holotype NHMW 1132628 B paratype, coll. JG F 12393 (photo P. Gloeer) C T. vruljakensis Grego & Gloeer, 2019, holotype, HNHM Moll 100 174 D T. ovalis (Kuscer, 1933) E, F T. gloeri Grego, 2020 E holotype HNHM-Moll 104 418 F paratype JG F 1340." figureDoi="10.3897/subtbiol.38.64762.figure3" httpUri="https://binary.pensoft.net/fig/531343" pageId="0" pageNumber="65">Figs 3A, B</figureCitation>
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<figureCitation id="D9DED22E00D3D7881841D007D08935B8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. M protoconch SEM images of Travunijana A, B Travunijana djokovici sp. nov., paratype, SBMNH 632721) C T. vruljakensis Grego and Gloeer, paratype SBMNH 625961 D T. gloeri Grego, 2020, paratype SBMNH 632722. Scale bars: 1 mm. (SEM SBMNH Vanessa Delnavaz)." figureDoi="10.3897/subtbiol.38.64762.figure4" httpUri="https://binary.pensoft.net/fig/531344" pageId="0" pageNumber="65">, 4A, B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D90F5E9286B0F6223970B0732321C879" pageId="0" pageNumber="65" type="type locality">
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<paragraph id="DD793FAF6E3CC49238C056575A1ABB4D" pageId="0" pageNumber="65">Type locality.</paragraph>
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<paragraph id="1BF9FD9C1E817469D201BB3CA625AEBC" pageId="0" pageNumber="65">
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Montenegro • Danilovgrad district,
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<normalizedToken id="C91C9773D0BCA6F4E670CE977DFFC7DD" originalValue="Bandići">Bandici</normalizedToken>
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Village,
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<normalizedToken id="04CAEEF8FE5221E8FEDA6F79DB95BA5B" originalValue="Vriješko">Vrijesko</normalizedToken>
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Vrelo spring under road to Modro Oko Lake, left tributary of Matica River,
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<geoCoordinate id="2033C3597767C70B653D261B444531A3" degrees="42.481206" direction="north" orientation="latitude" precision="1" value="42.481205">42.481206°N</geoCoordinate>
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,
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<geoCoordinate id="25FEB4E22FBFECE3AF587590C30A6169" degrees="19.145484" direction="east" orientation="longitude" precision="1" value="19.145485">19.145484°E</geoCoordinate>
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(Fig.
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<figureCitation id="6744CFEC76AD7F623360AA386F60C7B4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the Genus Travunijana Grego & Gloeer, 2019 (Green dots 1 - 17) and type locality of Travunijana djokovici sp. nov. (Red dot 18). Grey: Permeable karstified carbonates; White: impermeable unkarstified bedrocks; Yellow: Permeable alluvial deposits; Orange: sand deposits with limited permeability for subterranean gastropod dispersal. Light blue ring: karst springs; Blue ring with green centre and black X: estavelles; Dark green diamond: submarine or brackish karst spring." figureDoi="10.3897/subtbiol.38.64762.figure1" httpUri="https://binary.pensoft.net/fig/531341" pageId="0" pageNumber="65">1</figureCitation>
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and
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<figureCitation id="45B593F81B52AB618731FFD25EB3D132" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photo of the type locality of Travunijana djokovici sp. nov. (photo M. Olsavsky)." figureDoi="10.3897/subtbiol.38.64762.figure2" httpUri="https://binary.pensoft.net/fig/531342" pageId="0" pageNumber="65">2</figureCitation>
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).
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</paragraph>
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<caption id="E6E5749F6F5FE06D2893DCDB7E03A4ED" doi="10.3897/subtbiol.38.64762.figure2" httpUri="https://binary.pensoft.net/fig/531342" pageId="0" pageNumber="65" start="Figure 2" startId="F2">
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<paragraph id="C2EC75742DD1DCF08CC52E6932C20BFD" pageId="0" pageNumber="65">
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<emphasis id="EF628245157BA6CAE80F255F40EEE8CB" bold="true" pageId="0" pageNumber="65">Figure 2.</emphasis>
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Photo of the type locality of
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<taxonomicName id="98F2866AFAC504E69A05B2E4121E496F" authorityName="Grego 1 & Pešić 2" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" kingdom="Animalia" lsidName="Travunijana djokovici" order="Littorinimorpha" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="djokovici">
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<emphasis id="78F00CEBF7D5383AAADDE46B143018F8" italics="true" pageId="0" pageNumber="65">Travunijana djokovici</emphasis>
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</taxonomicName>
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sp. nov. (photo M.
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<normalizedToken id="B550FBDC1219C8DBE160F4309D78563A" originalValue="Olšavský">Olsavsky</normalizedToken>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="117221191DAFE6D3BA5E0CD3EC0ED324" pageId="0" pageNumber="65" type="type material">
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<paragraph id="FE2DE360AAB9405A6F1C5A168E294A9A" pageId="0" pageNumber="65">Type material.</paragraph>
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<paragraph id="A4334D438777013D2B0EF662E1C09177" pageId="0" pageNumber="65">
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<emphasis id="F470A9E5201E4FFE2C8FB5EA463F90AA" bold="true" pageId="0" pageNumber="65">
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<emphasis id="E0B9B709DF0153075F308869F8F61B90" italics="true" pageId="0" pageNumber="65">Holotype</emphasis>
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:
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</emphasis>
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Type locality: J. Grego, G. Jakab. M.
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<normalizedToken id="FC47B3DE805C1B25A51F87CD0D7C6202" originalValue="Olšavský">Olsavsky</normalizedToken>
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, M.
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<normalizedToken id="F9D65A615A4831256A1ECC1E6CF198D9" originalValue="Kováčiková">Kovacikova</normalizedToken>
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leg. 20.04.2019, NHMW1132628.
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</paragraph>
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<paragraph id="A5F40E661C7031D81FD0C833A1F977E2" pageId="0" pageNumber="65">
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<emphasis id="26F5F2373F1E2518779841FB92669375" bold="true" pageId="0" pageNumber="65">
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<emphasis id="BDF6EA4AAAD5A6466FDFBC1A3AD5FB40" italics="true" pageId="0" pageNumber="65">Paratypes</emphasis>
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:
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</emphasis>
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same data: NHMW113629/ 2 dry specimens, HNHM105300/ 2 dry specimens and 37 dry specimens T 1293 in coll. Grego.
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</paragraph>
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</subSubSection>
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<subSubSection id="04D74BF45F515F1C9F35058AA911EA56" pageId="0" pageNumber="65" type="description">
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<paragraph id="3678DCB4B585AFF41618BDCFEC6232AA" pageId="0" pageNumber="65">Measurements.</paragraph>
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<paragraph id="F5D73C7F7DB083A6601B188A180254E8" pageId="0" pageNumber="65">
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<emphasis id="79DAA9E1DD04AEC06471D8773B86D7A6" bold="true" italics="true" pageId="0" pageNumber="65">Holotype</emphasis>
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H 3.23 mm; W 1.77 mm; WB 1.46 mm; HA 1.44; WA 1.14.
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</paragraph>
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<paragraph id="61607FA7D0E237D0D2D9767A5381A684" pageId="0" pageNumber="65">
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<emphasis id="6F971FEAD53CD609BF932F8F9F812671" bold="true" italics="true" pageId="0" pageNumber="65">Paratype</emphasis>
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H 3.33 mm; W 1.86 mm; WB 1.46 mm; HA 1.49; WA 1.26.
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</paragraph>
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</subSubSection>
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<subSubSection id="B22F0478E02820DE6E635E1C49DFF911" pageId="0" pageNumber="65" type="materials_examined">
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<paragraph id="F57292B84E2F6CAD37AA5C5BD9547755" pageId="0" pageNumber="65">Other material.</paragraph>
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<paragraph id="9261C9B0F48EC06B5889584B357EEA19" pageId="0" pageNumber="65">Same locality data: 12 fragmented shells in coll. JG T 1294.</paragraph>
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</subSubSection>
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<subSubSection id="59D06F07A9CF22CC0C273C58276C9CC2" pageId="0" pageNumber="65" type="differential diagnosis">
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<paragraph id="94F6F68AAF7A49CFAADEF2885B39D4B3" pageId="0" pageNumber="65">Differential diagnosis.</paragraph>
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<paragraph id="6930AE76C244246ED99690C92148CC86" pageId="0" pageNumber="65">
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The faintly pitted protoconch clearly distinguishes the new species from the members of the genus
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<taxonomicName id="89D0B1B4307516893E5103F295220451" class="Gastropoda" family="Hydrobiidae" genus="Plagigeyeria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Plagigeyeria" order="Littorinimorpha" pageId="0" pageNumber="65" phylum="Mollusca" rank="genus">
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<emphasis id="78ADD8A1B3EB25C6FE12C19EAC0E99BD" italics="true" pageId="0" pageNumber="65">Plagigeyeria</emphasis>
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</taxonomicName>
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and indicates its position within the genus
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<taxonomicName id="EF243A6862BD28AB65CF2EE3D1974A73" authorityName="Grego & Gloer" authorityYear="2019" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" kingdom="Animalia" lsidName="Travunijana" order="Littorinimorpha" pageId="0" pageNumber="65" phylum="Mollusca" rank="genus">
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<emphasis id="5E7C0082C89A8126DCDEC9E526CD3927" italics="true" pageId="0" pageNumber="65">Travunijana</emphasis>
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</taxonomicName>
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. Moreover, the weakly sinuated columellar margin also supports its generic assignment.
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</paragraph>
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<paragraph id="4874626967706D6C58D556BA63CA7599" pageId="0" pageNumber="65">
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Compared to the most closely related
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<taxonomicName id="A3822DBFD647591022F4A3C678A02CFB" authorityName="Grego" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" kingdom="Animalia" lsidName="Travunijana gloeri" order="Littorinimorpha" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="gloeri">
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<emphasis id="9F386887F5C097A2D9BB0533A4E8011E" italics="true" pageId="0" pageNumber="65">Travunijana gloeri</emphasis>
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</taxonomicName>
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Grego, 2020,
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<taxonomicName id="6C76E9ADB9588F184A47051FD0A10B25" lsidName="T. vruljakensis" pageId="0" pageNumber="65" rank="species" species="vruljakensis">
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<emphasis id="12ED9A34EACA22F04A208F37AEC43B4E" italics="true" pageId="0" pageNumber="65">T. vruljakensis</emphasis>
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</taxonomicName>
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Grego &
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<normalizedToken id="25A3A55FD3FFC331D056529755979D30" originalValue="Glöer">Gloeer</normalizedToken>
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, 2019 and
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<taxonomicName id="2A8669BE9CF9F2DF3F212ECD8FDBC23C" lsidName="T. ovalis" pageId="0" pageNumber="65" rank="species" species="ovalis">
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<emphasis id="2D68C43A09C6C4EB007BE00CF94450C4" italics="true" pageId="0" pageNumber="65">T. ovalis</emphasis>
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</taxonomicName>
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(
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<normalizedToken id="2A44CD887C6E60DFC251E4D5301B61B5" originalValue="Kuščer">Kuscer</normalizedToken>
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, 1937), all from
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<normalizedToken id="2EF004C0BA8BA7611B5D519F9CF7B9F9" originalValue="Trebišnjica">Trebisnjica</normalizedToken>
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Basin (Fig.
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<figureCitation id="565C3C3F4B2953C72779CCFF559758DE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Travunijana djokovici sp. nov. A holotype NHMW 1132628 B paratype, coll. JG F 12393 (photo P. Gloeer) C T. vruljakensis Grego & Gloeer, 2019, holotype, HNHM Moll 100 174 D T. ovalis (Kuscer, 1933) E, F T. gloeri Grego, 2020 E holotype HNHM-Moll 104 418 F paratype JG F 1340." figureDoi="10.3897/subtbiol.38.64762.figure3" httpUri="https://binary.pensoft.net/fig/531343" pageId="0" pageNumber="65">3C-F</figureCitation>
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), the new species differs by its shell morphology, which has more elongate-pyramidal shape, more distinct regular ribs on the shell surface and an elongate lens-like aperture, the latter with more distinct siphonal and anal grove.
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</paragraph>
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<caption id="E8197B31AF12CF9CABD5539A9A1234F9" doi="10.3897/subtbiol.38.64762.figure3" httpUri="https://binary.pensoft.net/fig/531343" pageId="0" pageNumber="65" start="Figure 3" startId="F3">
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<paragraph id="C1EFDF3C4A25B13E2974FE906E223A6F" pageId="0" pageNumber="65">
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<emphasis id="31EF6B45A2FD868ED96030490F5C3C61" bold="true" pageId="0" pageNumber="65">Figure 3.</emphasis>
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<emphasis id="CA07AFDEB449B4974AC78F2ADCC7D275" bold="true" pageId="0" pageNumber="65">A, B</emphasis>
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<taxonomicName id="5436F9B384F501C9C477D092F5E3441C" authorityName="Grego 1 & Pešić 2" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" kingdom="Animalia" lsidName="Travunijana djokovici" order="Littorinimorpha" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="djokovici">
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<emphasis id="AD8A476DD475DDA094C6D3E270B55B3A" italics="true" pageId="0" pageNumber="65">Travunijana djokovici</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="AAA9A6C7EDD700BFB83D04D3870D7CD6" bold="true" pageId="0" pageNumber="65">A</emphasis>
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holotype NHMW 1132628
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<emphasis id="C793E20F467D48F94A990AA5BE437577" bold="true" pageId="0" pageNumber="65">B</emphasis>
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paratype, coll. JG F12393 (photo P.
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<normalizedToken id="55A13B8E27FA94FBEF9B747EB4CBC774" originalValue="Glöer">Gloeer</normalizedToken>
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)
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<emphasis id="6423441DEBE612E18D156F8F8533B302" bold="true" pageId="0" pageNumber="65">C</emphasis>
|
|
<taxonomicName id="FDFF6930FB3DD47AE796C7E0F8B1D69C" lsidName="T. vruljakensis" pageId="0" pageNumber="65" rank="species" species="vruljakensis">
|
|
<emphasis id="3BFAC0F08101C5030D8D652113A5FCEE" italics="true" pageId="0" pageNumber="65">T. vruljakensis</emphasis>
|
|
</taxonomicName>
|
|
Grego &
|
|
<normalizedToken id="1612F311AD480EF202651742D57322D1" originalValue="Glöer">Gloeer</normalizedToken>
|
|
, 2019, holotype, HNHM Moll 100 174
|
|
<emphasis id="5D2070202790EE057630BF8892D8496C" bold="true" pageId="0" pageNumber="65">D</emphasis>
|
|
<taxonomicName id="6956015160BCFB4E081847869E73F6A6" lsidName="T. ovalis" pageId="0" pageNumber="65" rank="species" species="ovalis">
|
|
<emphasis id="D17F67336A4EAC94AD3112CE24076A71" italics="true" pageId="0" pageNumber="65">T. ovalis</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<normalizedToken id="081E1698D8274E6245D31E0E4974F4AC" originalValue="Kuščer">Kuscer</normalizedToken>
|
|
, 1933)
|
|
<emphasis id="2B3D1040C81C0C890D0BEE870E26ED32" bold="true" pageId="0" pageNumber="65">E, F</emphasis>
|
|
<taxonomicName id="E8A0828A69A616AA369E764B6D0650B4" lsidName="T. gloeri" pageId="0" pageNumber="65" rank="species" species="gloeri">
|
|
<emphasis id="1696E70908BF9F4D016618D6CCC937E3" italics="true" pageId="0" pageNumber="65">T. gloeri</emphasis>
|
|
</taxonomicName>
|
|
Grego, 2020
|
|
<emphasis id="640DC83491E5A11777984379F5DE327D" bold="true" pageId="0" pageNumber="65">E</emphasis>
|
|
holotype HNHM-Moll 104 418
|
|
<emphasis id="B3C888118D754A91AFA798B69E93FD26" bold="true" pageId="0" pageNumber="65">F</emphasis>
|
|
paratype JG F1340.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="E7EA88AD1246375FF9BD1B3FB6A4C589" pageId="0" pageNumber="65" type="description">
|
|
<paragraph id="CEB913EBF06E1E23BCA405C38A786F38" pageId="0" pageNumber="65">Description.</paragraph>
|
|
<paragraph id="FE75D035B6460E2972E5776E56EEFD25" pageId="0" pageNumber="65">
|
|
<emphasis id="11B713C85A0D0C3F4A3B2F6E3A1F3149" bold="true" pageId="0" pageNumber="65">
|
|
<emphasis id="5D74E92D166E01088B2D7FF9E72BE8A5" italics="true" pageId="0" pageNumber="65">Shell</emphasis>
|
|
:
|
|
</emphasis>
|
|
elongate-conical, milky whitish colour, periostracum yellowish, consisted of five moderately convex whorls with a semi-deep suture. The surface covered by dense, regular weak axial ribs. Aperture elongate, lens-like, its axis declined from the columella; the labral peristome straight in its lateral profile and slightly outward curved and callous. The columella peristome weakly sinuated at its middle part. The siphon and anal groves clearly indicated at the tips of elongated aperture. The umbilicus is tiny, almost closed and hidden behind the reflected columellar margin.
|
|
</paragraph>
|
|
<paragraph id="7D3DC1D6CC617F560D930B6FE4AF58C2" pageId="0" pageNumber="65">
|
|
<emphasis id="44AFB9135908704C150ED768BDB60F25" bold="true" italics="true" pageId="0" pageNumber="65">Protoconch</emphasis>
|
|
very faintly and densely pitted, at the penultimate whorls smoothly transferring through weak malleated structures, faint regular growth lines into a regular weak ribbing covering the rest of the teleoconch.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="28F177724B570F96CF3C3598BC57AF38" pageId="0" pageNumber="65" type="etymology">
|
|
<paragraph id="984E18D62E562E12A02710E9645CC197" pageId="0" pageNumber="65">Etymology.</paragraph>
|
|
<paragraph id="5CE856F8855CC6A19EA981DF448C8006" pageId="0" pageNumber="65">The new species is named after Novak Djokovic a famous Serbian tennis player to acknowledge his inspiring enthusiasm and energy.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B9254975BC271A5DAFCE97B355980112" pageId="0" pageNumber="65" type="distribution">
|
|
<paragraph id="F8F0CD92411B994691AFE972139D38DD" pageId="0" pageNumber="65">Distribution.</paragraph>
|
|
<paragraph id="D4E90324F7D98A29609E678E50581299" pageId="0" pageNumber="65">Montenegro; only known from the type locality.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="82BB68A5CE66CBA73D5CAF307F69643C" pageId="0" pageNumber="65" type="ecology">
|
|
<paragraph id="2DD6CFEE89ED05611F8AF66DAEAC3A19" pageId="0" pageNumber="65">Ecology.</paragraph>
|
|
<paragraph id="15DF5A59CC27B79B45E0B1B1CD03434D" pageId="0" pageNumber="65">
|
|
The type locality is a medium sized karst spring with stable water outflow, situated at the border of a limestone massif and alluvium of the Matica River. The status of empty shells washed out of the spring-head suggests its stygobiont habitat inside the karst conduit. The new species was syntopic with
|
|
<taxonomicName id="1A790839870856DAA3E203F81A199B21" authorityName="Bole" authorityYear="1961" class="Gastropoda" family="Hydrobiidae" genus="Plagigeyeria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Plagigeyeria montenegrina" order="Littorinimorpha" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="montenegrina">Plagigeyeria cf. montenegrina</taxonomicName>
|
|
Bole, 1961,
|
|
<taxonomicName id="A8B87A03144B8B9F898B04D8A4866461" authorityName="Radoman" authorityYear="1973" class="Gastropoda" family="Hydrobiidae" genus="Bracenica" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Bracenica spiridoni" order="Littorinimorpha" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="spiridoni">Bracenica cf. spiridoni</taxonomicName>
|
|
Radoman, 1973,
|
|
<taxonomicName id="7FEB2CFC879B61492C14C4525F4E292F" authorityName="Gloer & Pesic" authorityYear="2014" class="Gastropoda" family="Hydrobiidae" genus="Zeteana" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zeteana" order="Littorinimorpha" pageId="0" pageNumber="65" phylum="Mollusca" rank="genus">
|
|
<emphasis id="D696C723D10151F486E5AED55BC501D4" italics="true" pageId="0" pageNumber="65">Zeteana</emphasis>
|
|
</taxonomicName>
|
|
sp. and
|
|
<taxonomicName id="7DC63DD31F4061FBF5F69728CDA13B00" baseAuthorityName="Pešić & Glöer" baseAuthorityYear="2012" class="Gastropoda" family="Hydrobiidae" genus="Montenegrospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Montenegrospeum bogici" order="Littorinimorpha" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="bogici">Montenegrospeum cf. bogici</taxonomicName>
|
|
(
|
|
<normalizedToken id="A2897D03C0662FDD145F27D3779CAC2D" originalValue="Pešić">Pesic</normalizedToken>
|
|
&
|
|
<normalizedToken id="6D30BF6BF48F50C933E46E3470D2A528" originalValue="Glöer">Gloeer</normalizedToken>
|
|
, 2012) at its type locality thanatocoenosis.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C423B7AFA97E924802AFF3D72E025B28" pageId="0" pageNumber="65" type="conservation status">
|
|
<paragraph id="89272DB77A4C6113AB4BE893F586881B" pageId="0" pageNumber="65">Conservation status.</paragraph>
|
|
<paragraph id="4433FA7101297FB60AC91025923BF54E" pageId="0" pageNumber="65">Number of known locations (1) fewer than 5 and AOO (area of occupancy) smaller than 20 km2. There is no reason to suppose that AOO, EOO (extent of occurrence), number of locations, number of subpopulations or the number or mature individuals are declining or extremely fluctuating. However, due to small AOO it is assessed as Vulnerable (VU) D2 species.</paragraph>
|
|
<caption id="7FB06F16C56ADCB48A14D2F1500E5960" doi="10.3897/subtbiol.38.64762.figure4" httpUri="https://binary.pensoft.net/fig/531344" pageId="0" pageNumber="65" start="Figure 4" startId="F4">
|
|
<paragraph id="9F006F945B92DC6AF26D664D3B0BE2FF" pageId="0" pageNumber="65">
|
|
<emphasis id="43A39DED48BD59008A6D0004A2FC74FE" bold="true" pageId="0" pageNumber="65">Figure 4.</emphasis>
|
|
M protoconch SEM images of
|
|
<taxonomicName id="76EB25E02827F5A2CFEC6A92A1B0FFC8" authorityName="Grego & Gloer" authorityYear="2019" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" kingdom="Animalia" lsidName="Travunijana" order="Littorinimorpha" pageId="0" pageNumber="65" phylum="Mollusca" rank="genus">
|
|
<emphasis id="DFAF42727BC32B40236214A83066B63F" italics="true" pageId="0" pageNumber="65">Travunijana</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="B60F3E6E8EBF775D2671FAA9F9D2CA09" bold="true" pageId="0" pageNumber="65">A, B</emphasis>
|
|
<taxonomicName id="060F48B69CADEE8DDB277EB9D5AEEE5F" authorityName="Grego 1 & Pešić 2" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" kingdom="Animalia" lsidName="Travunijana djokovici" order="Littorinimorpha" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="djokovici">
|
|
<emphasis id="E700E065058F42BD0E5E15EE58B4553B" italics="true" pageId="0" pageNumber="65">Travunijana djokovici</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., paratype, SBMNH 632721)
|
|
<emphasis id="F5EC910567744D169B452F12BAE26DF7" bold="true" pageId="0" pageNumber="65">C</emphasis>
|
|
<taxonomicName id="92A74D3F5966ECEAD8C73124E569E63B" lsidName="T. vruljakensis" pageId="0" pageNumber="65" rank="species" species="vruljakensis">
|
|
<emphasis id="E72486C2EC3AEE2898A609662E8C6917" italics="true" pageId="0" pageNumber="65">T. vruljakensis</emphasis>
|
|
</taxonomicName>
|
|
Grego and
|
|
<normalizedToken id="281E87D353677CDC978B3957A276DAA9" originalValue="Glöer">Gloeer</normalizedToken>
|
|
, paratype SBMNH 625961
|
|
<emphasis id="7E348FBC72C16FEF798EC817B646DE20" bold="true" pageId="0" pageNumber="65">D</emphasis>
|
|
<taxonomicName id="D1D2A7199B2CBC7FABF24A81B285A2FE" lsidName="T. gloeri" pageId="0" pageNumber="65" rank="species" species="gloeri">
|
|
<emphasis id="99255E27061D066B89C748D3A6698D8B" italics="true" pageId="0" pageNumber="65">T. gloeri</emphasis>
|
|
</taxonomicName>
|
|
Grego, 2020, paratype SBMNH 632722. Scale bars: 1 mm. (SEM SBMNH Vanessa Delnavaz).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |