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<document ID-CLB-Dataset="72036" ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-GBIF-Dataset="eafc7326-93e2-4d55-b708-2da4b77e2e5e" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan &amp; Machard, Patrice" docDate="2022" docId="9CCD5BDE97C15F7E9439578AB6C80B85" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) jordanensis Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard 2022, sp. nov." docType="treatment" docUuid="A95FB1A6-43F7-4FFC-A202-05073CC72F90" docUuidSource="ZooBank" docVersion="6" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1732679132351" updateUser="ExternalLinkService">
<mods:mods id="02D64A18104EEFBC8BBAF62F02187EB3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="38D2FF6A0E70A0DA81C0C1C6A58A5D26">
<mods:title id="E0E3D69D5D9ACF986DBBF0C9ACA5F652">Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
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<mods:role id="56BF1D9E7165AA19E455DD5D2CE997E7">
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<mods:namePart id="3739B75129DE102C36484AAE3477E6E4">Gueorguiev, Borislav</mods:namePart>
<mods:nameIdentifier id="9069C89842C3FB4B742FE979F43BABDE" type="ORCID">https://orcid.org/0000-0002-8532-0657</mods:nameIdentifier>
<mods:affiliation id="A9382778CA9E33F30AEC546590673F3B">National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
<mods:nameIdentifier id="95982DF81F45FF88799B87EBE1DC24D6" type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
</mods:name>
<mods:name id="7EA4E2F5AFD8F5ECF5C9844E398A94D4" type="personal">
<mods:role id="61E3A74118A3FC39C36601A7F11E63BC">
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<mods:namePart id="B765BE74CCF56C07AC322628C75EAE6E">Wrase, David W.</mods:namePart>
<mods:nameIdentifier id="0BD7839873C21595C37DBED101C4902F" type="ORCID">https://orcid.org/0000-0003-1779-2684</mods:nameIdentifier>
<mods:affiliation id="E254966583583838B3966A7A2130091C">Oderstr. 2, 15306, Gusow-Platkow, Germany</mods:affiliation>
</mods:name>
<mods:name id="60AAD8818097275A1021388431C63D2C" type="personal">
<mods:role id="764F2DB7DAF243EAB073B0C905BC77C2">
<mods:roleTerm id="B8DC88858986AA400A8D53F1C8CED10C">Author</mods:roleTerm>
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<mods:namePart id="3ABFA568913990818FA847ABD096C6F9">Assmann, Thorsten</mods:namePart>
<mods:nameIdentifier id="6B25714E5C7D5437CEE6406350F5B2A0" type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
<mods:affiliation id="334D73D7945E8C315C109A3596FF7D3E">Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
</mods:name>
<mods:name id="12CD3A6E9A0651DF99633B4A90017A30" type="personal">
<mods:role id="F16CAD52D6F45BB08CDF49D3ADA5D58A">
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<mods:namePart id="9BF8CB2699D13005D748AADD53D2E45E">Muilwijk, Jan</mods:namePart>
<mods:affiliation id="EC8DD0A8556E32C3660380B12B4C94B0">Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="D6868755D2720A1B87684B820B1F4493">Machard, Patrice</mods:namePart>
<mods:affiliation id="89F055BAB28D6FAE6A57083CF303742A">29 Chemin de Champigny, Molineuf, F- 41190 Valencisse, France</mods:affiliation>
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<mods:title id="5384D26890468F7BD46E7E34F56870D1">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="18AC58831E6018DCB47E02A8A11C8B56">2022</mods:date>
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<mods:number id="C3CBB83C59A324AC078D5EB72AF63021">2022-06-13</mods:number>
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<mods:number id="148D394011E5F349203949D73A20DD6F">69</mods:number>
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<treatment id="9CCD5BDE97C15F7E9439578AB6C80B85" ID-GBIF-Taxon="195868751" LSID="urn:lsid:zoobank.org:act:A95FB1A6-43F7-4FFC-A202-05073CC72F90" httpUri="http://treatment.plazi.org/id/9CCD5BDE97C15F7E9439578AB6C80B85" lastPageNumber="71" pageId="0" pageNumber="71" scope_family="Carabidae" scope_order="Coleoptera" scope_subGenus="Platyderus" scope_tribe="Sphodrini">
<subSubSection id="B8D08981679E058C1756219DB2049AE1" pageId="0" pageNumber="71" type="nomenclature">
<paragraph id="15B66FF0EE29CFCE45BDA51453ACF6F4" pageId="0" pageNumber="71">
18.
<taxonomicName id="5536E0A4C18A6E8D040A0A8CE85B8B92" LSID="http://zoobank.org/A95FB1A6-43F7-4FFC-A202-05073CC72F90" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) jordanensis</taxonomicName>
<taxonomicNameLabel id="CD4A51EADB9B77E79A082E8D2619D7B6" pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="75AC2E195E2750BC6C9B5FD4E737EE82" pageId="0" pageNumber="71" type="description">
<paragraph id="1BDD4E6DE059C0A79A029613B6C3B4C0" pageId="0" pageNumber="71">
<figureCitation id="D740A7218F3203A9EADDEF68DDECFDD8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">Figs 4C</figureCitation>
<figureCitation id="554FF20CDEC9A0F1D1C400B57D4937FF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6D</figureCitation>
<figureCitation id="E58512A772DE56D34E3EFC62F02039E8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7K</figureCitation>
<figureCitation id="A4B079B1C3BCE029E81B4B712244FEDA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9F</figureCitation>
<figureCitation id="FD41A09C2A1BA88F2982DD1151924F94" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11E</figureCitation>
<figureCitation id="8B605D179201F893F040625504BA659B" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12L</figureCitation>
<figureCitation id="E786FAC1E5108AB5E793837E58DE2D8A" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15E</figureCitation>
<figureCitation id="E5CFC5D6C5FE803022487299D2874D8D" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A58595AAE648CADC2E462787418BD75C" pageId="0" pageNumber="71" type="type locality">
<paragraph id="01A6122F966CC79843E241C5D2D75772" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="904BE4063025F30E649104861E38C53F" pageId="0" pageNumber="71">
Jordan,
<normalizedToken id="AC1A5F3369F9C60C75D6555E815E5F6E" originalValue="Ma’an">Ma'an</normalizedToken>
Governorate,
<normalizedToken id="3D37314762633E74978CED9AA0DA27B8" originalValue="Al-Betrā’">Al-Betra'</normalizedToken>
District, Little Petra, archaeological area, 30.3667, 35.4333.
</paragraph>
</subSubSection>
<subSubSection id="643CBAACA18958D455D521E1D9AAB299" pageId="0" pageNumber="71" type="type material">
<paragraph id="312118DEAF029B1766881243DEB0FAB3" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="480F23427972376FD5C9855A59C51B52" pageId="0" pageNumber="71">
<emphasis id="1F6E41DF750DFB85843E121F4EF6550E" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂, 'JORDAN (
<normalizedToken id="A475E49587029C00D853398880F2F0B4" originalValue="Ma’an">Ma'an</normalizedToken>
) / Little Petra, ca. 1000 m / archaeological area /
<geoCoordinate id="5ED697446003176516DB25DA26EEF967" degrees="30" direction="north" minutes="22" orientation="latitude" precision="925" value="30.366667">30°22'N</geoCoordinate>
/
<geoCoordinate id="2FC3BD2D078CE0DE65E06EA16FFF1F8F" degrees="35" direction="east" minutes="26" orientation="longitude" precision="925" value="35.433334">35°26'E</geoCoordinate>
/ (narrow shaft beside / stony staircase) / 25.III.2016 Wrase &amp; Laser [12B] [w, p]' (cWR). Paratypes: 1♀, 'Jordanien 1024 m / Prov
<normalizedToken id="69ACD8AE49BD0E95A1D8F3201212B2F8" originalValue="Ma’an">Ma'an</normalizedToken>
/ W. Ziegler 9.3.2015 [w, p] // N Wadi Musa / 30°24'.15&quot;N, 35°26'.53&quot;E [w, p]' (cZIEG); 1♂, 'Jordania / Prov.
<normalizedToken id="518264D0A9252BE46B879EB4F4D5C234" originalValue="Ma’an">Ma'an</normalizedToken>
/ n Wadi Musa 1023m / W.Ziegler 25.3.2016 [w, p] // Jabal al Bayda (Project / 30°24'291&quot;N, 35°26'743&quot;E [w, p]' (NMNHS); 1♂ 'S-Jordan: Wadi Rum / Qatar Spring /
<geoCoordinate id="82C74598DA026D17F6767C98F8D96AFE" degrees="29.51" direction="north" orientation="latitude" precision="555" value="29.51">29.51°N</geoCoordinate>
/
<geoCoordinate id="995E56C72EEDCAB947B7577311940DCA" degrees="35.41" direction="east" orientation="longitude" precision="555" value="35.41">35.41°E</geoCoordinate>
/ 1000-1100 m asl [w, p] // dry wadi with
<taxonomicName id="1990C5A0A484344FEEF754BF46B33342" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" lsidName="Ficus" order="Rosales" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="5DE71A666BCF7076DB3E2CC939149CDC" italics="true" pageId="0" pageNumber="71">Ficus</emphasis>
</taxonomicName>
trees, 21.III.2017 / leg. Th. Assmann [w, p]&quot; (cAL); 1♂,
<normalizedToken id="C0127CEDE6215AC464F5471AF068A6A2" originalValue="ʻS-Jordan">ʻS-Jordan</normalizedToken>
: Little Petra, ca. 1000m / wadis with Mediterranean trees in vicinity to the archaeological site /
<geoCoordinate id="C2D45A52336D17D01BE46FFD6BED44CF" degrees="30.34" direction="north" orientation="latitude" precision="555" value="30.34">30.34°N</geoCoordinate>
,
<geoCoordinate id="996F10970C7B88F96FD82E240FD5CCAB" degrees="035.46" direction="east" orientation="longitude" precision="555" value="35.46">035.46°E</geoCoordinate>
/ 25.III.2016 / leg. Th. Assmann [w, p]' (cAL).
</paragraph>
<paragraph id="B19E00D01EE6F53C54C3E9CC9CFAA167" pageId="0" pageNumber="71">TME: 5 specimens. TGE: 4♂♂, 1♀.</paragraph>
</subSubSection>
<subSubSection id="19F898B408994978C067A86A66802474" pageId="0" pageNumber="71" type="etymology">
<paragraph id="A1EA26A91B7A584676CC9C3DF4202319" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="25170663BBD0F7FFD330341498E12C61" pageId="0" pageNumber="71">The specific epithet is a Latinized adjective, based on the name of the country in which this species was found.</paragraph>
</subSubSection>
<subSubSection id="EA391AF010259535EB9CA92FD42C1730" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="0C1BE937956BBA3B96A30813520E70B6" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="A798CE5A067304A4EEFE3392AD38B224" pageId="0" pageNumber="71">
Differs from all other species of the &quot;
<taxonomicName id="2E89BE732E04E98CBE998B9BDEF37D3A" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="BABB02B9BE7FF5CA8AF3ABD88D4838A9" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group by the following set of characters: (1) meso- and metatarsomeres dorsally convex and smooth (Fig.
<figureCitation id="991DDED325AFCE75B11638214EA0EE87" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6D</figureCitation>
); (2) posterior angles of pronotum incompletely rounded, with tips protruding laterally (Fig.
<figureCitation id="37B9A0F1AA0A915B1B0BD6CFA1C49FBC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4C</figureCitation>
); and (3) ventral margin of apex of median lobe (lateral view) evidently convex thus apex appears bent up (Fig.
<figureCitation id="552D7F7BF2C325EBFDEFDA50B6E6F6CF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9F</figureCitation>
).
</paragraph>
<paragraph id="489FAB09CA961FC983491EEC955FFAB8" pageId="0" pageNumber="71">
<taxonomicName id="0E3DC674BA05A678FBE11C4222BB3D12" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="67F57B8A3A5189B89C55BB8C0775EEB8" italics="true" pageId="0" pageNumber="71">Platyderus jordanensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="6F7F8BDF7D316733210BBA953065C162" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="91410F42895E5A2B35D778AC3F912FB4" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
share a trait that can be a mark for their close relationships, the apical gonocoxite without nematiform setae.
</paragraph>
</subSubSection>
<subSubSection id="3D19800C3BDCF03A2AB7A9B33AD9D048" pageId="0" pageNumber="71" type="description">
<paragraph id="AC5E6E7168E1B9AE71767BB1DE8BE0D8" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="C2550C5F1249F65300F7061735590E97" pageId="0" pageNumber="71">
<emphasis id="8F88887C869182D9CC3D76F3F8136980" bold="true" pageId="0" pageNumber="71">
<emphasis id="A438108C24AD4F64F004F27C43DF93C7" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of large size for
<taxonomicName id="99B144BAF2E8D9B2A3BF2A4C4EC1FE92" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="30CF337387E1F6E84761716B426777FA" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 8.70-9.90 mm; BW: 3.00-3.45 mm), with rather elongate and moderately convex body (Fig.
<figureCitation id="D3B5D2304BD998C1C81983E561D292AE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4C</figureCitation>
).
<emphasis id="44DBAEA864FDC38106F971123BCFEE87" bold="true" pageId="0" pageNumber="71">
<emphasis id="6EE1A2EF4FAB99671F24309F616CD1EC" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation id="BC39A972EC8B6321A2161863C03D363C" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
.
<emphasis id="D5BB185DE6DB3E827CFA501C4B63E446" bold="true" pageId="0" pageNumber="71">
<emphasis id="D297A9C5028A717B09EEC0051B60B359" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body and appendages uniformly light reddish-brown, palpomeres and legs somewhat lighter than rest of body. Integument slightly to moderately shiny, head and pronotum shinier than elytra.
<emphasis id="6073F1E542EF6751C6702FD7E8F74FC2" bold="true" pageId="0" pageNumber="71">
<emphasis id="F7F7FC4779D10D0C8CCDE822658E4BE3" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Surface of pronotum with evident microreticulation, sculpticells regular isodiametric. Elytra (intervals, scutellum, basal margin and lateral gutter) with distinct isodiametric sculpticells. Ventral surface largely with more or less impressed isodiametric or slight transverse sculpticells. Head impunctate, nearly smooth, with a few, fine and shallow wrinkles in and laterally of frontal furrows. Pronotum surface mostly smooth, only basal area between foveae longitudinally wrinkled and adjacent lateral areas with rare and superficial punctures that do not reach anterior half. Elytral intervals without apparent punctation. Abdominal ventrite 1 finely wrinkled medially, smooth laterally, other ventrites neither wrinkled nor punctate.
<emphasis id="BF496CA73DE520734FACCFF616B1C146" bold="true" pageId="0" pageNumber="71">
<emphasis id="B1216CDB5EF66AFB60127142A5A7514A" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
One-third or more as wide as pronotum wide (PW/HW= 1.35-1.42). Eyes long, moderately convex. Labrum subrectangular, shorter than clypeus, with anterior margin concave medially. Frontoclypeal suture distinct in middle, indistinct at sides. Frontal furrows small, punctiform. Paraorbital sulci straight, backward surpassing posterior margin of eye, hardly reaching level of posterior supraorbital pore.
<emphasis id="FC80E526AD50D089377B8668073EE74C" bold="true" pageId="0" pageNumber="71">
<emphasis id="563FF8AE512B8FF198D7B3EA7B403871" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum about a tenth wider than long (PW/PL= 1.08-1.13), with widest point at medial third. Anterior transverse impression absent; posterior transverse impression barely distinct. Sides sinuate, convex medially and anteriorly, slightly concave posteriorly; anterior bead present laterally, lacking or present in medial 1/10; basal bead present laterally, reduced to absent in medial third or medial half. Metepisternum somewhat longer than wide, MA/MI= 0.88-0.91.
<emphasis id="22E683FDFE2F117E9C90D93B26329F74" bold="true" pageId="0" pageNumber="71">
<emphasis id="F49567D1064B260DE4F2F3FBF34A9D14" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Elongate, about one and a half times or little more as long as elytra width (EL/EW= 1.56-1.60), two-and-a-half-times as long as pronotum (EL/PL= 2.41-2.60), and a bit less than a half as wide as pronotum (EW/PW= 1.39-1.46), with widest point at medial third. Parascutellar striole linear, as impressed as other striae, not joining stria 1; striae 1-8 and striole slightly to indistinctly punctate; base of stria 1 ending in parascutellar pore, striae 2-5 reaching or barely not reaching basal bead, 6 and 7 ending little before basal bead. Interval 3 with three discal setiferous punctures. Umbilicate setiferous series with 16 punctures on each elytron in three specimens examined, including holotype.
<emphasis id="D5EDBE95AE75A76BC985E9447FCBF806" bold="true" pageId="0" pageNumber="71">
<emphasis id="403DB74F2F190105C146A8CE0C283195" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one or two setae in basal third and one in medial third. Mesofemora with 6 (holotype) or 5 (both paratypes) setiferous punctures on anterior side ventrally. Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis id="685FB4E07941462CB4BD05FCD29FC649" bold="true" pageId="0" pageNumber="71">
<emphasis id="C30B7CD030BD21632A6C5325AE377C32" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX subtriangular, with proximal margin symmetrical and pointed at apex (Fig.
<figureCitation id="3E9DB51B4AD7168E711143AFB041D2CD" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7K</figureCitation>
). Median lobe of aedeagus in lateral view slender, with narrow basal bulb, shaft broad with proximal part slightly constricted, and short, straight apex (Fig.
<figureCitation id="B574ACB2686647EA04ADD9CC1DD28291" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9F</figureCitation>
); median lobe in ventral view straight, 3.7 times longer than wide; apical lamella (dorsal view) rather short, nearly symmetrical, rounded at tip, with sides straight to slightly concave. Internal sac in lateral view (Fig.
<figureCitation id="897D6E91513A2C276C582CE551DBDC8F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9F</figureCitation>
) with ventral sclerite slightly broadened and rounded distally; same in ventral view (Fig.
<figureCitation id="2E916BB7EE326A1CB84FA60E2BC5C4C3" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11E</figureCitation>
), with dorsal sclerite large, having small left-sided and large right-sided protuberance, and ventral sclerite straight and rather narrow. Right paramere as on Fig.
<figureCitation id="A6AF9C5FBE8D5CCF98BEE81105BFF403" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12L</figureCitation>
.
<emphasis id="9838D52B0C3698E9BF3BF8E3A930289B" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation id="9423DE90DECE2A7F20F131EAE3E7B66F" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15E</figureCitation>
). Apical gonocoxite with widely rounded apex and one dorsolateral ensiform seta. Spermathecal canal connected in basal third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection id="20B84D72FD16B24E6F2391C66F54F6B2" pageId="0" pageNumber="71" type="comparisons">
<paragraph id="BF2CC1D2D310383B19D40122303E4840" pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph id="4F2687583811CC63E2DC3BC706AB700C" pageId="0" pageNumber="71">
In additions to characters mentioned in
<normalizedToken id="E0B13E9DDDE8372BF096C0C0354E44F7" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
,
<taxonomicName id="8AFB5084DE21D67229A5B7F04D5FC0AA" class="Insecta" kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="FFA496FDF87CC1C46AC9BC69C5508C42" italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov. differs from geographically close
<taxonomicName id="DDA64799775BC6EABC57B14B5E0067A8" class="Insecta" kingdom="Animalia" lsidName="P. brunneus subsp. ferranei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferranei">
<emphasis id="14DC3CACD978E4D14F0C8F46E6A1A796" italics="true" pageId="0" pageNumber="71">P. brunneus ferranei</emphasis>
</taxonomicName>
by pronotum less wide (PW/PL: 1.08-1.13, vs. PW/PL: 1.14-1.21), with anterior margin narrower than posterior margin (PA/PB: 0.78-0.87, vs. PA/PB: 0.88-0.92).
</paragraph>
<paragraph id="1E099A3323331FAF86CE28FE26814F74" pageId="0" pageNumber="71">
For differences between
<taxonomicName id="DD858AA964EEDF06BB96AD9FB1842A03" class="Insecta" kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="7ADC2EF0239B8598EA83F6F43A2E9985" italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
, from one side, and
<taxonomicName id="E14BAA8F709ED9187E99838CA21037AD" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="A66F73656DD8F0E924E02049472DA457" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
,
<taxonomicName id="541801AA0B039C6F334F400EDE38ECA7" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="D97553E51ED7968AB8E6889545D5900B" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
and
<taxonomicName id="A1D1004FAD66E60AA305822A96258D22" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="2B5825E75B3C16AB50D78CC88D658642" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
, from other side, see sections
<normalizedToken id="B25D1371AEF60DDFC70743C0E71274A2" originalValue="“Comparisons”">&quot;Comparisons&quot;</normalizedToken>
(under the first two species) and
<normalizedToken id="DE1C19CBE62FD77774CBD23BEF8DB2AD" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
(under
<taxonomicName id="ABE20D5AD94634996C3DDD5E2E67FC55" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="AE24CA45E6517435B963D56741CEC9A6" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="A800C22679152010F8BF8894D160F344" pageId="0" pageNumber="71" type="habitat">
<paragraph id="F7FD875D869B3A03230F7BF808DD841C" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="0E2520D5ABD0644499E20F265F4FFA7C" pageId="0" pageNumber="71">
In Little Petra
<taxonomicName id="11A9A4F96641CD61FABFED298F7288F4" class="Insecta" kingdom="Animalia" lsidName="P. jordanansis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanansis">
<emphasis id="F497385A380149D1F95B7E587709D4BC" italics="true" pageId="0" pageNumber="71">P. jordanansis</emphasis>
</taxonomicName>
sp. nov. lives in wadis that have a canyon-like profile and some Mediterranean tree species. There it lives together with
<taxonomicName id="D9EE775D0B548ECE51EEFEC1EC0A6CBA" class="Insecta" family="Carabidae" genus="Laemostenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laemostenus quadricollisquadricollis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="quadricollisquadricollis">
<emphasis id="E5005306A3063C0BF6EBD81A42DF9E28" italics="true" pageId="0" pageNumber="71">Laemostenus quadricollisquadricollis</emphasis>
</taxonomicName>
L. Redtenbacher, 1843,
<taxonomicName id="6E24631D8E9EBCC28DAC72D2C88CA526" authorityName="Piochard de la Brulerie" authorityYear="1876" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" lsidName="Trechus crucifer" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="crucifer">
<emphasis id="7BCD112ED0A593C41FC314E87E405A5E" italics="true" pageId="0" pageNumber="71">Trechus crucifer</emphasis>
</taxonomicName>
Piochard de la
<normalizedToken id="81517CA01572C7D9284FAA998F969C76" originalValue="Brûlerie">Brulerie</normalizedToken>
, 1876,
<taxonomicName id="07A8501B6A4824AAA546AEBA695FFEB7" authorityName="Motschulsky" authorityYear="1850" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" lsidName="Calathus cinctus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="cinctus">
<emphasis id="CEBF5F39B8C2CAED667515964787B494" italics="true" pageId="0" pageNumber="71">Calathus cinctus</emphasis>
</taxonomicName>
Motschulsky, 1850,
<taxonomicName id="3E51DE434460013CF8102ED2849C063E" authorityName="Menetries" authorityYear="1832" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" lsidName="Cymindis andreae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="andreae">
<emphasis id="F3FE9B0706EA08D68C9806997D57125E" italics="true" pageId="0" pageNumber="71">Cymindis andreae</emphasis>
</taxonomicName>
<normalizedToken id="09328264157CA6C882140C2C6375BB45" originalValue="Ménétriés">Menetries</normalizedToken>
, 1832, and
<taxonomicName id="5FF0408293E03BB7627FDE82EDB2DC3F" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus impressus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="0C530A9D901DC52F56AB4FD4870E24D3" italics="true" pageId="0" pageNumber="71">Carabus impressus</emphasis>
</taxonomicName>
Klug, 1832. In Wadi Rum the single specimen was found in a dry wadi with
<taxonomicName id="0CC878B092BC7622E842E1505E156152" class="Magnoliopsida" family="Fabaceae" genus="Retama" kingdom="Plantae" lsidName="Retama" order="Fabales" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="F26293DD759B1409B4E5D56E9A29B2B1" italics="true" pageId="0" pageNumber="71">Retama</emphasis>
</taxonomicName>
shrubs and single
<taxonomicName id="0EE6353674A91ADB611609A3FCAE1B7A" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" lsidName="Ficus" order="Rosales" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="8A6B181D1F97B5EBCD4E705F971969C8" italics="true" pageId="0" pageNumber="71">Ficus</emphasis>
</taxonomicName>
trees in the shade of the walls of the head of the canyon-like valley. It occurs together with ground beetles that are mainly typical for desert habitats (
<taxonomicName id="B1AAE5D30699CC0976A02B5403830F4A" authorityName="Schatzmayr" authorityYear="1936" class="Insecta" family="Carabidae" genus="Laemostenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laemostenus aegyptiacus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="aegyptiacus">
<emphasis id="9D4AB572F82F81139928469D0D1CDCA7" italics="true" pageId="0" pageNumber="71">Laemostenus aegyptiacus</emphasis>
</taxonomicName>
Schatzmayr, 1936,
<taxonomicName id="C079019AEF83F02DC06A366E19329B92" authorityName="Peyerimhoff" authorityYear="1907" class="Insecta" family="Carabidae" genus="Singilis" kingdom="Animalia" lsidName="Singilis filicornis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis id="36F7228DE3D31474FBB9B9BA5B72B8FC" italics="true" pageId="0" pageNumber="71">Singilis filicornis</emphasis>
</taxonomicName>
Peyerimhoff, 1907,
<taxonomicName id="9B17A5152F6BACDF69673B80FC913CF6" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" lsidName="Cymindis hierichontica" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="hierichontica">
<emphasis id="D4A1B853AC0220DF2BDE76FD32EAA0CE" italics="true" pageId="0" pageNumber="71">Cymindis hierichontica</emphasis>
</taxonomicName>
Reiche &amp; Saulcy, 1855,
<taxonomicName id="F63FFEF4C06C32F990E2E3AE8DE97EFC" baseAuthorityName="Klug" baseAuthorityYear="1832" class="Insecta" family="Carabidae" genus="Merizomena" kingdom="Animalia" lsidName="Merizomena castanea" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="6CC4670815229D0B583B7AB14A246CF3" italics="true" pageId="0" pageNumber="71">Merizomena castanea</emphasis>
</taxonomicName>
Klug, 1832, and
<taxonomicName id="1F19F46C0B2F256DC451FDFD018ED038" authorityName="Bedel" authorityYear="1907" class="Insecta" family="Carabidae" genus="Amara" kingdom="Animalia" lsidName="Amara maindroni" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis id="90D972FEBB8D02FB4F3E21C856089680" italics="true" pageId="0" pageNumber="71">Amara maindroni</emphasis>
</taxonomicName>
Bedel, 1907). A photograph of the habitat of the latter is given by
<bibRefCitation id="3D0B02C88ADE9681507B292C6BAF236D" DOI="https://doi.org/10.4081/fe.2017.226" author="Casale, A" journalOrPublisher="Fragmenta Entomologica" pageId="0" pageNumber="71" pagination="13 - 24" refId="B14" refString="Casale, A, Assmann, T, 2017. The Sphodrina of the southern Levant (Coleoptera: Carabidae, Sphodrini). Fragmenta Entomologica 49 (1): 13 - 24, DOI: https://doi.org/10.4081/fe.2017.226" title="The Sphodrina of the southern Levant (Coleoptera: Carabidae, Sphodrini)." url="https://doi.org/10.4081/fe.2017.226" volume="49" year="2017">Casale and Assmann (2017</bibRefCitation>
: 22, fig. 35).
</paragraph>
<caption id="B7C2D1EF7B8A4877358839DC3BDD8F62" doi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71" start="Figure 16" startId="F16">
<paragraph id="483983A43D776E02E1EE56E7D509A132" pageId="0" pageNumber="71">
<emphasis id="A82F0EC46609CDFBB8F678D69495C335" bold="true" pageId="0" pageNumber="71">Figure 16.</emphasis>
Distribution of species from subgenus
<taxonomicName id="0CF5AE6AA23714C19306BE0F9F4E8C35" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus Eremoderus</taxonomicName>
in the western Aegean Sea Region and Turkey:
<taxonomicName id="55598F8B258B4CB300CD1F4260201A78" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis id="0ABF9738A31581663F502B848848EE96" italics="true" pageId="0" pageNumber="71">Platyderus chatzakiae</emphasis>
</taxonomicName>
sp. nov. (red circle),
<taxonomicName id="A2DB52AA7BEC20A2A6F14F82114F409E" class="Insecta" kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="DE821850421226482554E6315F7BA44B" italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
</taxonomicName>
(blue circles),
<taxonomicName id="1DBEC94B7AD070B35B2058CD5FDE6A7A" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="6661663CC6C806C50E94CC7E5847CACD" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov. (brown circles), and
<taxonomicName id="52987D2EFE606E6315F0A0B2D597C6E8" class="Insecta" kingdom="Animalia" lsidName="P. vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci">
<emphasis id="BAF8B982B86568A17EFFFFEFB018E2E2" italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
</taxonomicName>
sp. nov. (yellow circle).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="A30A76F02F89038680F6B502C8535FFC" pageId="0" pageNumber="71" type="distribution">
<paragraph id="E011C47E6EBED90C8239CDA528ED8A75" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="0BC5D2F713C7E1659C15AAFB4F4018BE" pageId="0" pageNumber="71">
According to the data available, the species is the representative of the group &quot;
<taxonomicName id="3100D88B314CC03E6067393CE804D99D" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="52087D64249F0557D6D89A3BBC8D2E21" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; in the southern part of Jordan, stretching from the vicinity of Little Petra to areas close to the Saudi Arabian border (Fig.
<figureCitation id="A560F786574B40FFFAE66A411A8695B8" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
). As Wadi Rum is in the most southern part of Jordan, the species may also occur in Northwest Saudi Arabia.
</paragraph>
</subSubSection>
</treatment>
</document>
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