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<document id="03B59001D46273967E62E9CA2FADB4DB" ID-CLB-Dataset="59956" ID-DOI="10.11646/zootaxa.5323.4.6" ID-GBIF-Dataset="c4c28226-bd22-482b-99d4-6e595b21a239" ID-ISSN="1175-5326" ID-Zenodo-Dep="8212468" ID-ZooBank="0AD185E5-E2B1-44EE-8898-B3253904027E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691079772263" checkinUser="plazi" docAuthor="Sampaio, Vinícius Gomes &amp; Campos, Luiz Alexandre" docDate="2023" docId="8C3A87D9C501833DFF3426CD0020FCC2" docLanguage="en" docName="zootaxa.5323.4.6.pdf" docOrigin="Zootaxa 5323 (4)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paropsocolis divergens Sampaio &amp; Campos 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="565" masterDocId="7003FFA1C5088331FFA3265A0279FFD7" masterDocTitle="Paropsocolis, a new pentatomid genus with two new species from Brazil (Heteroptera, Pentatomidae, Ochlerini)" masterLastPageNumber="566" masterPageNumber="553" pageNumber="562" updateTime="1691080428098" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1EF3123ADAF5D70482F38B33C8CB2DD3">Paropsocolis, a new pentatomid genus with two new species from Brazil (Heteroptera, Pentatomidae, Ochlerini)</mods:title>
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<mods:namePart id="403FD6E91C44EA395013E14B70D449C1">Sampaio, Vinícius Gomes</mods:namePart>
<mods:affiliation id="F2C449476F622E2136832E8BFCF65DFE">Departamento de Zoologia, Instituto de Biociências, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, 91501 - 970, Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:namePart id="86BE4701C6A3015DF451213C8BDCB0EE">Campos, Luiz Alexandre</mods:namePart>
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<mods:affiliation id="EDC4A1B622441B94CB23D6C0DD7ADFEC">Departamento de Zoologia, Instituto de Biociências, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, 91501 - 970, Porto Alegre, RS, Brazil. &amp; luiz. campos @ ufrgs. br; https: // orcid. org / 0000 - 0001 - 5414 - 8746</mods:affiliation>
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<subSubSection id="4C896544C5018338FF3426CD0165FF66" box="[151,796,151,178]" pageId="9" pageNumber="562" type="nomenclature">
<paragraph id="042C36CFC5018338FF3426CD0165FF66" blockId="9.[151,796,151,214]" box="[151,796,151,178]" pageId="9" pageNumber="562">
<heading id="5F6481A3C5018338FF3426CD0165FF66" bold="true" box="[151,796,151,178]" fontSize="11" level="1" pageId="9" pageNumber="562" reason="1">
<emphasis id="36E7EADDC5018338FF3426CD0165FF66" bold="true" box="[151,796,151,178]" pageId="9" pageNumber="562">
<taxonomicName id="C3934D4CC5018338FF3426CD00C8FF66" authority="Sampaio &amp; Campos" authorityName="Sampaio &amp; Campos" authorityYear="2023" box="[151,689,151,178]" class="Insecta" family="Pentatomidae" genus="Paropsocolis" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="divergens" status="sp. nov.">
<emphasis id="36E7EADDC5018338FF3426CD03D4FF65" bold="true" box="[151,429,151,178]" italics="true" pageId="9" pageNumber="562">Paropsocolis divergens</emphasis>
Sampaio &amp; Campos
</taxonomicName>
,
<taxonomicNameLabel id="2DD457A6C5018338FD1D26CD0165FF66" box="[702,796,151,177]" pageId="9" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4C896544C5018338FF3426E10333FF01" box="[151,330,187,214]" pageId="9" pageNumber="562" type="description">
<paragraph id="042C36CFC5018338FF3426E10333FF01" blockId="9.[151,796,151,214]" box="[151,330,187,214]" pageId="9" pageNumber="562">
(
<figureCitation id="9CA82A4AC5018338FF3C26E10294FF02" box="[159,237,187,213]" captionStart="FIGURE 1" captionStartId="2.[152,255,1829,1853]" captionTargetBox="[163,1427,182,1792]" captionTargetId="figure-20@2.[158,1430,179,1803]" captionTargetPageId="2" captionText="FIGURE 1. Hypothesis for the phylogenetic relationships of Paropsocolis Sampaio &amp; Campos, gen. nov. generated under implied weighting (K = 16.889) from a data matrix containing the 60 morphological characters from Roell &amp; Campos (2019). White circles = homoplasious transformations; black circles = non homoplasious transformations. Numbers to the left of nodes indicate relative Bremer support and GC frequencies, respectively." figureDoi="http://doi.org/10.5281/zenodo.8212470" httpUri="https://zenodo.org/record/8212470/files/figure.png" pageId="9" pageNumber="562">Figs. 1</figureCitation>
,
<figureCitation id="9CA82A4AC5018338FF5826E1033BFF01" box="[251,322,187,214]" captionStart-0="FIGURES 2528" captionStart-1="FIGURES 2931" captionStart-2="FIGURES 3237" captionStart-3="FIGURES 3842" captionStartId-0="9.[152,270,1981,2005]" captionStartId-1="10.[152,270,1646,1670]" captionStartId-2="11.[152,270,1047,1071]" captionStartId-3="11.[152,270,1875,1899]" captionTargetBox-0="[161,1425,353,1947]" captionTargetBox-1="[162,1425,285,1613]" captionTargetBox-2="[162,1425,285,1014]" captionTargetBox-3="[161,1426,1143,1842]" captionTargetId-0="figure-47@9.[151,1436,342,1957]" captionTargetId-1="figure-47@10.[151,1436,274,1624]" captionTargetId-2="figure-64@11.[151,1436,274,1025]" captionTargetId-3="figure-101@11.[151,1436,1133,1852]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="11" captionText-0="FIGURES 2528. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male holotype, 25, dorsal; 26, ventral; 27, lateral; 28, labels. Scale bar = 5 mm." captionText-1="FIGURES 2931. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male holotype, 29, head and pronotum; 30, labial intercalary segment (bracket); 31, external scent efferent system. Scale bars: 9 = 5 mm; 30, 31 = 1 mm." captionText-2="FIGURES 3237. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, pygophore, 32, dorsal; 33, dorsoposterior; 34, dorsolateral; 35, posterior; 36, lateral; 37, ventral. Scale bar = 0.5 mm." captionText-3="FIGURES 3842. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, 38, phallus (lateral); 39, dorsal process of endosoma; 4042, right paramere in dorsal, lateral and medial views, respectively. Abbreviations: dpe = dorsal process of endosoma; th = theca; vpe = ventral process of endosoma; vpt = ventral process of theca; vtt = ventral tubercle of theca. Scale bars = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8212492" figureDoi-1="http://doi.org/10.5281/zenodo.8212494" figureDoi-2="http://doi.org/10.5281/zenodo.8212498" figureDoi-3="http://doi.org/10.5281/zenodo.8212506" httpUri-0="https://zenodo.org/record/8212492/files/figure.png" httpUri-1="https://zenodo.org/record/8212494/files/figure.png" httpUri-2="https://zenodo.org/record/8212498/files/figure.png" httpUri-3="https://zenodo.org/record/8212506/files/figure.png" pageId="9" pageNumber="562">2542</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4C896544C5018338FF3426A50374FEEA" pageId="9" pageNumber="562" type="etymology">
<paragraph id="042C36CFC5018338FF3426A50374FEEA" blockId="9.[151,1436,255,317]" pageId="9" pageNumber="562">
<emphasis id="36E7EADDC5018338FF3426A50367FECE" bold="true" box="[151,286,255,281]" pageId="9" pageNumber="562">Etymology.</emphasis>
The epithet refers to the diverging apex of the paramere bifurcation. Latin:
<emphasis id="36E7EADDC5018338FBFD26A506BEFECE" box="[1118,1223,255,281]" italics="true" pageId="9" pageNumber="562">divergere</emphasis>
, turned to different directions.
</paragraph>
</subSubSection>
<caption id="50EC6647C5018338FF3B21E703E0F82F" ID-DOI="http://doi.org/10.5281/zenodo.8212492" ID-Zenodo-Dep="8212492" httpUri="https://zenodo.org/record/8212492/files/figure.png" pageId="9" pageNumber="562" startId="9.[152,270,1981,2005]" targetBox="[161,1425,353,1947]" targetPageId="9" targetType="figure">
<paragraph id="042C36CFC5018338FF3B21E703E0F82F" blockId="9.[151,1436,1981,2041]" pageId="9" pageNumber="562">
<emphasis id="36E7EADDC5018338FF3B21E70323F802" bold="true" box="[152,346,1981,2005]" pageId="9" pageNumber="562">FIGURES 2528.</emphasis>
<taxonomicName id="C3934D4CC5018338FEFC21E70159F802" authority="Sampaio &amp; Campos" authorityName="Sampaio &amp; Campos" authorityYear="2023" box="[351,800,1981,2005]" class="Insecta" family="Pentatomidae" genus="Paropsocolis" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="divergens" status="sp. nov.">
<emphasis id="36E7EADDC5018338FEFC21E70035F802" box="[351,588,1981,2005]" italics="true" pageId="9" pageNumber="562">Paropsocolis divergens</emphasis>
Sampaio &amp; Campos
</taxonomicName>
,
<emphasis id="36E7EADDC5018338FC8A21E70100F802" bold="true" box="[809,889,1981,2005]" pageId="9" pageNumber="562">
<taxonomicNameLabel id="2DD457A6C5018338FC8A21E70100F802" box="[809,889,1981,2005]" pageId="9" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype, 25, dorsal; 26, ventral; 27, lateral; 28, labels. Scale bar = 5 mm.
</paragraph>
</caption>
<subSubSection id="4C896544C502833BFF6426CD01C9FF2E" pageId="10" pageNumber="563" type="materials_examined">
<paragraph id="042C36CFC502833BFF6426CD005FFF02" blockId="10.[151,1437,150,249]" pageId="10" pageNumber="563">
<materialsCitation id="B4FB3C92C502833BFF6426CD005FFF02" collectingDate="1968-12" collectionCode="CEIOC" collectorName="M. Alvarenga. Deposited" country="Brazil" location="Jacareacanga" municipality="Male" pageId="10" pageNumber="563" specimenCount="1" stateProvince="Para" typeStatus="holotype">
<emphasis id="36E7EADDC502833BFF6426CD0342FF66" bold="true" box="[199,315,151,177]" pageId="10" pageNumber="563">
<typeStatus id="DB28886DC502833BFF6426CD034FFF66" box="[199,310,151,177]" pageId="10" pageNumber="563">Holotype</typeStatus>
.
</emphasis>
<collectingMunicipality id="E448ACB5C502833BFEEB26CD03FDFF66" box="[328,388,151,177]" pageId="10" pageNumber="563">Male</collectingMunicipality>
, here designated.
<collectingCountry id="7C84765FC502833BFDFE26CD00BCFF66" box="[605,709,151,177]" name="Brazil" pageId="10" pageNumber="563">BRAZIL</collectingCountry>
,
<collectingRegion id="C657F82DC502833BFD7526CD0175FF66" box="[726,780,151,177]" country="Brazil" name="Para" pageId="10" pageNumber="563">Pará</collectingRegion>
:
<location id="014C6014C502833BFCBF26CD01C9FF66" LSID="urn:lsid:plazi:treatment:8C3A87D9C501833DFF3426CD0020FCC2:014C6014C502833BFCBF26CD01C9FF66" box="[796,944,151,177]" country="Brazil" municipality="Male" name="Jacareacanga" pageId="10" pageNumber="563" stateProvince="Para">Jacareacanga</location>
(-6.22, -57.80),
<date id="702D100FC502833BFBD926CD069DFF66" box="[1146,1252,151,177]" pageId="10" pageNumber="563" value="1968-12">
<collectingDate id="6069E9E7C502833BFBD926CD069DFF66" box="[1146,1252,151,177]" pageId="10" pageNumber="563" value="1968-12">XII-1968</collectingDate>
</date>
,
<collectorName id="A9665319C502833BFB5626CD0370FF02" pageId="10" pageNumber="563">M. Alvarenga. Deposited</collectorName>
in:
<collectionCode id="6282AE0AC502833BFE9726E103F1FF02" box="[308,392,187,213]" pageId="10" pageNumber="563">CEIOC</collectionCode>
(
<figureCitation id="9CA82A4AC502833BFE3B26E10067FF02" box="[408,542,186,213]" captionStart-0="FIGURES 2528" captionStart-1="FIGURES 2931" captionStartId-0="9.[152,270,1981,2005]" captionStartId-1="10.[152,270,1646,1670]" captionTargetBox-0="[161,1425,353,1947]" captionTargetBox-1="[162,1425,285,1613]" captionTargetId-0="figure-47@9.[151,1436,342,1957]" captionTargetId-1="figure-47@10.[151,1436,274,1624]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 2528. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male holotype, 25, dorsal; 26, ventral; 27, lateral; 28, labels. Scale bar = 5 mm." captionText-1="FIGURES 2931. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male holotype, 29, head and pronotum; 30, labial intercalary segment (bracket); 31, external scent efferent system. Scale bars: 9 = 5 mm; 30, 31 = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8212492" figureDoi-1="http://doi.org/10.5281/zenodo.8212494" httpUri-0="https://zenodo.org/record/8212492/files/figure.png" httpUri-1="https://zenodo.org/record/8212494/files/figure.png" pageId="10" pageNumber="563">Figs. 2531</figureCitation>
)
</materialsCitation>
</paragraph>
<paragraph id="042C36CFC502833BFF64268501C9FF2E" blockId="10.[151,1437,150,249]" box="[199,944,223,249]" pageId="10" pageNumber="563">
<materialsCitation id="B4FB3C92C502833BFF64268501D5FF2E" box="[199,940,223,249]" collectingDate="1968-12" collectionCode="CEIOC" collectorName="M. Alvarenga. Deposited" country="Brazil" location="Jacareacanga" municipality="Male" pageId="10" pageNumber="563" specimenCount="1" stateProvince="Para" typeStatus="paratype">
<emphasis id="36E7EADDC502833BFF6426850342FF2E" bold="true" box="[199,315,223,249]" pageId="10" pageNumber="563">
<typeStatus id="DB28886DC502833BFF642685034FFF2E" box="[199,310,223,249]" pageId="10" pageNumber="563">Paratype</typeStatus>
.
</emphasis>
Male, same data as the holotype. Deposited in:
<collectionCode id="6282AE0AC502833BFCF7268501D5FF2E" box="[852,940,223,249]" pageId="10" pageNumber="563">CEIOC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="50EC6647C502833BFF3B203406D4F97D" ID-DOI="http://doi.org/10.5281/zenodo.8212494" ID-Zenodo-Dep="8212494" httpUri="https://zenodo.org/record/8212494/files/figure.png" pageId="10" pageNumber="563" startId="10.[152,270,1646,1670]" targetBox="[162,1425,285,1613]" targetPageId="10" targetType="figure">
<paragraph id="042C36CFC502833BFF3B203406D4F97D" blockId="10.[151,1436,1646,1707]" pageId="10" pageNumber="563">
<emphasis id="36E7EADDC502833BFF3B20340325F950" bold="true" box="[152,348,1646,1671]" pageId="10" pageNumber="563">FIGURES 2931.</emphasis>
<taxonomicName id="C3934D4CC502833BFEC720350156F950" authority="Sampaio &amp; Campos" authorityName="Sampaio &amp; Campos" authorityYear="2023" box="[356,815,1647,1671]" class="Insecta" family="Pentatomidae" genus="Paropsocolis" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="divergens" status="sp. nov.">
<emphasis id="36E7EADDC502833BFEC72035002DF950" box="[356,596,1647,1671]" italics="true" pageId="10" pageNumber="563">Paropsocolis divergens</emphasis>
Sampaio &amp; Campos
</taxonomicName>
,
<emphasis id="36E7EADDC502833BFC99203501F4F950" bold="true" box="[826,909,1647,1671]" pageId="10" pageNumber="563">
<taxonomicNameLabel id="2DD457A6C502833BFC99203501F4F950" box="[826,909,1647,1671]" pageId="10" pageNumber="563" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype, 29, head and pronotum; 30, labial intercalary segment (bracket); 31, external scent efferent system. Scale bars: 9 = 5 mm; 30, 31 = 1 mm.
</paragraph>
</caption>
<subSubSection id="4C896544C502833BFF6420A706E3F854" pageId="10" pageNumber="563" type="diagnosis">
<paragraph id="042C36CFC502833BFF6420A706E3F854" blockId="10.[151,1437,1789,2032]" pageId="10" pageNumber="563">
<emphasis id="36E7EADDC502833BFF6420A7033BF8C0" bold="true" box="[199,322,1789,1815]" pageId="10" pageNumber="563">Diagnosis.</emphasis>
Body convex ventrally (
<figureCitation id="9CA82A4AC502833BFDD220A700B2F8C0" box="[625,715,1789,1815]" captionStart="FIGURES 2528" captionStartId="9.[152,270,1981,2005]" captionTargetBox="[161,1425,353,1947]" captionTargetId="figure-47@9.[151,1436,342,1957]" captionTargetPageId="9" captionText="FIGURES 2528. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male holotype, 25, dorsal; 26, ventral; 27, lateral; 28, labels. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212492" httpUri="https://zenodo.org/record/8212492/files/figure.png" pageId="10" pageNumber="563">Fig. 27</figureCitation>
); labium not attaining the posterior margin of metasternum; metasternal carina narrow; peritremal disc attaining the lateral third of metapleura (
<figureCitation id="9CA82A4AC502833BFB9B217B06F2F8EC" box="[1080,1163,1825,1851]" captionStart="FIGURES 2931" captionStartId="10.[152,270,1646,1670]" captionTargetBox="[162,1425,285,1613]" captionTargetId="figure-47@10.[151,1436,274,1624]" captionTargetPageId="10" captionText="FIGURES 2931. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male holotype, 29, head and pronotum; 30, labial intercalary segment (bracket); 31, external scent efferent system. Scale bars: 9 = 5 mm; 30, 31 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8212494" httpUri="https://zenodo.org/record/8212494/files/figure.png" pageId="10" pageNumber="563">Fig. 31</figureCitation>
); pygophore with lateral margins sinuous (
<figureCitation id="9CA82A4AC502833BFEFF211F03C9F888" box="[348,432,1861,1887]" captionStart="FIGURES 3237" captionStartId="11.[152,270,1047,1071]" captionTargetBox="[162,1425,285,1014]" captionTargetId="figure-64@11.[151,1436,274,1025]" captionTargetPageId="11" captionText="FIGURES 3237. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, pygophore, 32, dorsal; 33, dorsoposterior; 34, dorsolateral; 35, posterior; 36, lateral; 37, ventral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212498" httpUri="https://zenodo.org/record/8212498/files/figure.png" pageId="10" pageNumber="563">Fig. 32</figureCitation>
); superior layer of ventral rim forming a v-shaped posterior projection (
<figureCitation id="9CA82A4AC502833BFB6E211F0766F888" box="[1229,1311,1861,1887]" captionStart="FIGURES 3237" captionStartId="11.[152,270,1047,1071]" captionTargetBox="[162,1425,285,1014]" captionTargetId="figure-64@11.[151,1436,274,1025]" captionTargetPageId="11" captionText="FIGURES 3237. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, pygophore, 32, dorsal; 33, dorsoposterior; 34, dorsolateral; 35, posterior; 36, lateral; 37, ventral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212498" httpUri="https://zenodo.org/record/8212498/files/figure.png" pageId="10" pageNumber="563">Fig. 37</figureCitation>
); paramere dish-shaped process large and deep; apical bifurcation of paramere diverging (
<figureCitation id="9CA82A4AC502833BFBA7213306F7F854" box="[1028,1166,1897,1923]" captionStart="FIGURES 3842" captionStartId="11.[152,270,1875,1899]" captionTargetBox="[161,1426,1143,1842]" captionTargetId="figure-101@11.[151,1436,1133,1852]" captionTargetPageId="11" captionText="FIGURES 3842. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, 38, phallus (lateral); 39, dorsal process of endosoma; 4042, right paramere in dorsal, lateral and medial views, respectively. Abbreviations: dpe = dorsal process of endosoma; th = theca; vpe = ventral process of endosoma; vpt = ventral process of theca; vtt = ventral tubercle of theca. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212506" httpUri="https://zenodo.org/record/8212506/files/figure.png" pageId="10" pageNumber="563">Figs. 4042</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4C896544C502833DFF6421D703DBFD26" lastPageId="12" lastPageNumber="565" pageId="10" pageNumber="563" type="description">
<paragraph id="042C36CFC502833AFF6421D706CAFF2E" blockId="10.[151,1437,1789,2032]" lastBlockId="11.[151,1437,150,249]" lastPageId="11" lastPageNumber="564" pageId="10" pageNumber="563">
<emphasis id="36E7EADDC502833BFF6421D70320F870" bold="true" box="[199,345,1933,1959]" pageId="10" pageNumber="563">Description.</emphasis>
Body ventrally convex (
<figureCitation id="9CA82A4AC502833BFDCB21D700C2F870" box="[616,699,1933,1959]" captionStart="FIGURES 2528" captionStartId="9.[152,270,1981,2005]" captionTargetBox="[161,1425,353,1947]" captionTargetId="figure-47@9.[151,1436,342,1957]" captionTargetPageId="9" captionText="FIGURES 2528. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male holotype, 25, dorsal; 26, ventral; 27, lateral; 28, labels. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212492" httpUri="https://zenodo.org/record/8212492/files/figure.png" pageId="10" pageNumber="563">Fig. 27</figureCitation>
). Punctures black, minute on head, evenly distributed on pronotum and scutellum. Head 1.6 times wider than long; labium not attaining the posterior margin of metasternum. Thorax. Pronotal anterior margin shallowly concave, anterolateral angles in round projection, humeral angles reddish (
<figureCitation id="9CA82A4AC502833AFACC218F02CEFF66" captionStart="FIGURES 2931" captionStartId="10.[152,270,1646,1670]" captionTargetBox="[162,1425,285,1613]" captionTargetId="figure-47@10.[151,1436,274,1624]" captionTargetPageId="10" captionText="FIGURES 2931. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male holotype, 29, head and pronotum; 30, labial intercalary segment (bracket); 31, external scent efferent system. Scale bars: 9 = 5 mm; 30, 31 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8212494" httpUri="https://zenodo.org/record/8212494/files/figure.png" lastPageId="11" lastPageNumber="564" pageId="10" pageNumber="563">Fig. 29</figureCitation>
). Scutellar apex acute (
<figureCitation id="9CA82A4AC503833AFE1F26CD0069FF66" box="[444,528,150,177]" captionStart="FIGURES 2528" captionStartId="9.[152,270,1981,2005]" captionTargetBox="[161,1425,353,1947]" captionTargetId="figure-47@9.[151,1436,342,1957]" captionTargetPageId="9" captionText="FIGURES 2528. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male holotype, 25, dorsal; 26, ventral; 27, lateral; 28, labels. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212492" httpUri="https://zenodo.org/record/8212492/files/figure.png" pageId="11" pageNumber="564">Fig. 25</figureCitation>
). Metasternal carina narrow. Peritremal disc sinuous, attaining the lateral third of metapleura (
<figureCitation id="9CA82A4AC503833AFE8126E1030CFF02" box="[290,373,187,213]" captionStart="FIGURES 2931" captionStartId="10.[152,270,1646,1670]" captionTargetBox="[162,1425,285,1613]" captionTargetId="figure-47@10.[151,1436,274,1624]" captionTargetPageId="10" captionText="FIGURES 2931. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male holotype, 29, head and pronotum; 30, labial intercalary segment (bracket); 31, external scent efferent system. Scale bars: 9 = 5 mm; 30, 31 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8212494" httpUri="https://zenodo.org/record/8212494/files/figure.png" pageId="11" pageNumber="564">Fig. 31</figureCitation>
). Abdomen. Connexival segments dark brown, paler on middle (
<figureCitation id="9CA82A4AC503833AFBE726E106E1FF02" box="[1092,1176,186,213]" captionStart="FIGURES 2528" captionStartId="9.[152,270,1981,2005]" captionTargetBox="[161,1425,353,1947]" captionTargetId="figure-47@9.[151,1436,342,1957]" captionTargetPageId="9" captionText="FIGURES 2528. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male holotype, 25, dorsal; 26, ventral; 27, lateral; 28, labels. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212492" httpUri="https://zenodo.org/record/8212492/files/figure.png" pageId="11" pageNumber="564">Fig. 25</figureCitation>
). Urosternites the same color of the dorsal surface; spiracles black, contrasting with the surrounding surface (
<figureCitation id="9CA82A4AC503833AFBF1268506DFFF2E" box="[1106,1190,223,249]" captionStart="FIGURES 2528" captionStartId="9.[152,270,1981,2005]" captionTargetBox="[161,1425,353,1947]" captionTargetId="figure-47@9.[151,1436,342,1957]" captionTargetPageId="9" captionText="FIGURES 2528. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male holotype, 25, dorsal; 26, ventral; 27, lateral; 28, labels. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212492" httpUri="https://zenodo.org/record/8212492/files/figure.png" pageId="11" pageNumber="564">Fig. 26</figureCitation>
).
</paragraph>
<caption id="50EC6647C503833AFF3B224D011BFB84" ID-DOI="http://doi.org/10.5281/zenodo.8212498" ID-Zenodo-Dep="8212498" httpUri="https://zenodo.org/record/8212498/files/figure.png" pageId="11" pageNumber="564" startId="11.[152,270,1047,1071]" targetBox="[162,1425,285,1014]" targetPageId="11" targetType="figure">
<paragraph id="042C36CFC503833AFF3B224D011BFB84" blockId="11.[151,1436,1047,1108]" pageId="11" pageNumber="564">
<emphasis id="36E7EADDC503833AFF3B224D0322FBF8" bold="true" box="[152,347,1047,1071]" pageId="11" pageNumber="564">FIGURES 3237.</emphasis>
<taxonomicName id="C3934D4CC503833AFEC22242015DFBE7" authority="Sampaio &amp; Campos" authorityName="Sampaio &amp; Campos" authorityYear="2023" box="[353,804,1048,1072]" class="Insecta" family="Pentatomidae" genus="Paropsocolis" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="564" phylum="Arthropoda" rank="species" species="divergens" status="sp. nov.">
<emphasis id="36E7EADDC503833AFEC222420037FBE7" box="[353,590,1048,1072]" italics="true" pageId="11" pageNumber="564">Paropsocolis divergens</emphasis>
Sampaio &amp; Campos
</taxonomicName>
,
<emphasis id="36E7EADDC503833AFC8D22420106FBE7" bold="true" box="[814,895,1048,1072]" pageId="11" pageNumber="564">
<taxonomicNameLabel id="2DD457A6C503833AFC8D22420106FBE7" box="[814,895,1048,1072]" pageId="11" pageNumber="564" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, pygophore, 32, dorsal; 33, dorsoposterior; 34, dorsolateral; 35, posterior; 36, lateral; 37, ventral. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="50EC6647C503833AFF3B21090348F800" ID-DOI="http://doi.org/10.5281/zenodo.8212506" ID-Zenodo-Dep="8212506" httpUri="https://zenodo.org/record/8212506/files/figure.png" pageId="11" pageNumber="564" startId="11.[152,270,1875,1899]" targetBox="[161,1426,1143,1842]" targetPageId="11" targetType="figure">
<paragraph id="042C36CFC503833AFF3B21090348F800" blockId="11.[151,1436,1875,2007]" pageId="11" pageNumber="564">
<emphasis id="36E7EADDC503833AFF3B21090324F8BC" bold="true" box="[152,349,1875,1899]" pageId="11" pageNumber="564">FIGURES 3842.</emphasis>
<taxonomicName id="C3934D4CC503833AFEC52109014DF8BC" authority="Sampaio &amp; Campos" authorityName="Sampaio &amp; Campos" authorityYear="2023" box="[358,820,1875,1899]" class="Insecta" family="Pentatomidae" genus="Paropsocolis" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="564" phylum="Arthropoda" rank="species" species="divergens" status="sp. nov.">
<emphasis id="36E7EADDC503833AFEC52109002FF8BC" box="[358,598,1875,1899]" italics="true" pageId="11" pageNumber="564">Paropsocolis divergens</emphasis>
Sampaio &amp; Campos
</taxonomicName>
,
<emphasis id="36E7EADDC503833AFCE2210901ECF8BC" bold="true" box="[833,917,1875,1899]" pageId="11" pageNumber="564">
<taxonomicNameLabel id="2DD457A6C503833AFCE2210901ECF8BC" box="[833,917,1875,1899]" pageId="11" pageNumber="564" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, 38, phallus (lateral); 39, dorsal process of endosoma; 4042, right paramere in dorsal, lateral and medial views, respectively. Abbreviations: dpe = dorsal process of endosoma; th = theca; vpe = ventral process of endosoma; vpt = ventral process of theca; vtt = ventral tubercle of theca. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="042C36CFC504833DFF6426CD01E6FEB2" blockId="12.[151,1437,150,789]" pageId="12" pageNumber="565">
<emphasis id="36E7EADDC504833DFF6426CD0372FF66" bold="true" box="[199,267,151,177]" pageId="12" pageNumber="565">Male.</emphasis>
Measurements (n=2): Total length 14.10 ± 0.14 (14.0014.20); width of abdomen 8.65 ± 0.46 (8.30 9.00); head length 1.85 ± 0.07 (1.801.90); width 3.00 ± 0.14 (2.903.10); pronotum length 3.55 ± 0.07 (3.503.60); width 7.85 ± 0.07 (7.807.90); scutellum length 5.75 ± 0.07 (5.705.80); width 4.85 ± 0.21 (4.705.00); length of antennomeres: I 0.85 ± 0.07 (0.800.90), II 0.95 ± 0.07 (0.901.00), III 1.35 ± 0.07 (1.301.40), IV 2.05 ± 0.07 (2.002.10); V (n=1) 2.60 ± 0.00 (2.60); length of labiomeres: I 1.05 ± 0.07 (1.001.10), Int 0.10 ± 0.00 (0.10), II 1.50 ± 0.00 (1.50), III 1.35 ± 0.07 (1.301.40), IV 1.00 ± 0.00 (1.00).
</paragraph>
<paragraph id="042C36CFC504833DFF6427350108FD1A" blockId="12.[151,1437,150,789]" pageId="12" pageNumber="565">
Genitalia (
<figureCitation id="9CA82A4AC504833DFE9E273503BEFE5E" box="[317,455,367,393]" captionStart-0="FIGURES 3237" captionStart-1="FIGURES 3842" captionStartId-0="11.[152,270,1047,1071]" captionStartId-1="11.[152,270,1875,1899]" captionTargetBox-0="[162,1425,285,1014]" captionTargetBox-1="[161,1426,1143,1842]" captionTargetId-0="figure-64@11.[151,1436,274,1025]" captionTargetId-1="figure-101@11.[151,1436,1133,1852]" captionTargetPageId-0="11" captionTargetPageId-1="11" captionText-0="FIGURES 3237. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, pygophore, 32, dorsal; 33, dorsoposterior; 34, dorsolateral; 35, posterior; 36, lateral; 37, ventral. Scale bar = 0.5 mm." captionText-1="FIGURES 3842. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, 38, phallus (lateral); 39, dorsal process of endosoma; 4042, right paramere in dorsal, lateral and medial views, respectively. Abbreviations: dpe = dorsal process of endosoma; th = theca; vpe = ventral process of endosoma; vpt = ventral process of theca; vtt = ventral tubercle of theca. Scale bars = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8212498" figureDoi-1="http://doi.org/10.5281/zenodo.8212506" httpUri-0="https://zenodo.org/record/8212498/files/figure.png" httpUri-1="https://zenodo.org/record/8212506/files/figure.png" pageId="12" pageNumber="565">Figs. 3242</figureCitation>
). Pygophore. Lateral margins sinuous (
<figureCitation id="9CA82A4AC504833DFCDD273501ABFE5E" box="[894,978,367,393]" captionStart="FIGURES 3237" captionStartId="11.[152,270,1047,1071]" captionTargetBox="[162,1425,285,1014]" captionTargetId="figure-64@11.[151,1436,274,1025]" captionTargetPageId="11" captionText="FIGURES 3237. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, pygophore, 32, dorsal; 33, dorsoposterior; 34, dorsolateral; 35, posterior; 36, lateral; 37, ventral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212498" httpUri="https://zenodo.org/record/8212498/files/figure.png" pageId="12" pageNumber="565">Fig. 32</figureCitation>
); dorsal wall length on midline twice the segment X length (
<figureCitation id="9CA82A4AC504833DFECB27C903C2FE7A" box="[360,443,403,429]" captionStart="FIGURES 3237" captionStartId="11.[152,270,1047,1071]" captionTargetBox="[162,1425,285,1014]" captionTargetId="figure-64@11.[151,1436,274,1025]" captionTargetPageId="11" captionText="FIGURES 3237. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, pygophore, 32, dorsal; 33, dorsoposterior; 34, dorsolateral; 35, posterior; 36, lateral; 37, ventral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212498" httpUri="https://zenodo.org/record/8212498/files/figure.png" pageId="12" pageNumber="565">Fig. 32</figureCitation>
); dorsal rim roundly concave over segment X (
<figureCitation id="9CA82A4AC504833DFC6327C90630FE7A" box="[960,1097,403,429]" captionStart="FIGURES 3237" captionStartId="11.[152,270,1047,1071]" captionTargetBox="[162,1425,285,1014]" captionTargetId="figure-64@11.[151,1436,274,1025]" captionTargetPageId="11" captionText="FIGURES 3237. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, pygophore, 32, dorsal; 33, dorsoposterior; 34, dorsolateral; 35, posterior; 36, lateral; 37, ventral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212498" httpUri="https://zenodo.org/record/8212498/files/figure.png" pageId="12" pageNumber="565">Figs. 3234</figureCitation>
); posterolateral angles broadly excavated (
<figureCitation id="9CA82A4AC504833DFEBA27ED03DCFE06" box="[281,421,439,465]" captionStart="FIGURES 3237" captionStartId="11.[152,270,1047,1071]" captionTargetBox="[162,1425,285,1014]" captionTargetId="figure-64@11.[151,1436,274,1025]" captionTargetPageId="11" captionText="FIGURES 3237. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, pygophore, 32, dorsal; 33, dorsoposterior; 34, dorsolateral; 35, posterior; 36, lateral; 37, ventral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212498" httpUri="https://zenodo.org/record/8212498/files/figure.png" pageId="12" pageNumber="565">Figs. 3234</figureCitation>
); ventral rim with sparse setae, middle third concave, superior layer forming a v-shaped posterior projection (
<figureCitation id="9CA82A4AC504833DFE26278103A2FE22" box="[389,475,475,501]" captionStart="FIGURES 3237" captionStartId="11.[152,270,1047,1071]" captionTargetBox="[162,1425,285,1014]" captionTargetId="figure-64@11.[151,1436,274,1025]" captionTargetPageId="11" captionText="FIGURES 3237. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, pygophore, 32, dorsal; 33, dorsoposterior; 34, dorsolateral; 35, posterior; 36, lateral; 37, ventral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212498" httpUri="https://zenodo.org/record/8212498/files/figure.png" pageId="12" pageNumber="565">Fig. 37</figureCitation>
). Segment X with lateral short pale setae (
<figureCitation id="9CA82A4AC504833DFC1C27810633FE22" box="[959,1098,475,501]" captionStart="FIGURES 3237" captionStartId="11.[152,270,1047,1071]" captionTargetBox="[162,1425,285,1014]" captionTargetId="figure-64@11.[151,1436,274,1025]" captionTargetPageId="11" captionText="FIGURES 3237. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, pygophore, 32, dorsal; 33, dorsoposterior; 34, dorsolateral; 35, posterior; 36, lateral; 37, ventral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212498" httpUri="https://zenodo.org/record/8212498/files/figure.png" pageId="12" pageNumber="565">Figs. 3234</figureCitation>
). Parameres laterally sinuous, dish-shaped process large and deep; apical bifurcation of paramere diverging, branches subequal in length, crossing each other (
<figureCitation id="9CA82A4AC504833DFEB8247903DFFDEA" box="[283,422,547,573]" captionStart="FIGURES 3842" captionStartId="11.[152,270,1875,1899]" captionTargetBox="[161,1426,1143,1842]" captionTargetId="figure-101@11.[151,1436,1133,1852]" captionTargetPageId="11" captionText="FIGURES 3842. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, 38, phallus (lateral); 39, dorsal process of endosoma; 4042, right paramere in dorsal, lateral and medial views, respectively. Abbreviations: dpe = dorsal process of endosoma; th = theca; vpe = ventral process of endosoma; vpt = ventral process of theca; vtt = ventral tubercle of theca. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212506" httpUri="https://zenodo.org/record/8212506/files/figure.png" pageId="12" pageNumber="565">Figs. 4042</figureCitation>
). Phallus strongly sclerotized; proximal thecal width almost ten times lesser the maximum thecal width in lateral view (
<figureCitation id="9CA82A4AC504833DFE79241D0054FDB6" box="[474,557,583,609]" captionStart="FIGURES 3842" captionStartId="11.[152,270,1875,1899]" captionTargetBox="[161,1426,1143,1842]" captionTargetId="figure-101@11.[151,1436,1133,1852]" captionTargetPageId="11" captionText="FIGURES 3842. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, 38, phallus (lateral); 39, dorsal process of endosoma; 4042, right paramere in dorsal, lateral and medial views, respectively. Abbreviations: dpe = dorsal process of endosoma; th = theca; vpe = ventral process of endosoma; vpt = ventral process of theca; vtt = ventral tubercle of theca. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212506" httpUri="https://zenodo.org/record/8212506/files/figure.png" pageId="12" pageNumber="565">Fig. 38</figureCitation>
, th); dorsal process of endosoma more than twice longer than the rigid part of the ventral process of endosoma (
<figureCitation id="9CA82A4AC504833DFDB524310010FD52" box="[534,617,619,645]" captionStart="FIGURES 3842" captionStartId="11.[152,270,1875,1899]" captionTargetBox="[161,1426,1143,1842]" captionTargetId="figure-101@11.[151,1436,1133,1852]" captionTargetPageId="11" captionText="FIGURES 3842. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, 38, phallus (lateral); 39, dorsal process of endosoma; 4042, right paramere in dorsal, lateral and medial views, respectively. Abbreviations: dpe = dorsal process of endosoma; th = theca; vpe = ventral process of endosoma; vpt = ventral process of theca; vtt = ventral tubercle of theca. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212506" httpUri="https://zenodo.org/record/8212506/files/figure.png" pageId="12" pageNumber="565">Fig. 38</figureCitation>
, dpe, vpe); apical lobes of the dorsal process of endosoma large, angled ventrally, and much longer than the lobes of the ventral process of theca (
<figureCitation id="9CA82A4AC504833DFC6D24D5062FFD7E" box="[974,1110,655,681]" captionStart="FIGURES 3842" captionStartId="11.[152,270,1875,1899]" captionTargetBox="[161,1426,1143,1842]" captionTargetId="figure-101@11.[151,1436,1133,1852]" captionTargetPageId="11" captionText="FIGURES 3842. Paropsocolis divergens Sampaio &amp; Campos, sp. nov., male, 38, phallus (lateral); 39, dorsal process of endosoma; 4042, right paramere in dorsal, lateral and medial views, respectively. Abbreviations: dpe = dorsal process of endosoma; th = theca; vpe = ventral process of endosoma; vpt = ventral process of theca; vtt = ventral tubercle of theca. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8212506" httpUri="https://zenodo.org/record/8212506/files/figure.png" pageId="12" pageNumber="565">Figs. 38, 39</figureCitation>
, dpe, vpt); ventral process of endosoma ruptured, making it impossible to determine its length.
</paragraph>
<paragraph id="042C36CFC504833DFF64248D03DBFD26" blockId="12.[151,1437,150,789]" box="[199,418,727,753]" pageId="12" pageNumber="565">
<emphasis id="36E7EADDC504833DFF64248D035CFD26" bold="true" box="[199,293,727,753]" pageId="12" pageNumber="565">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="4C896544C504833DFF6424A00020FCC2" box="[199,601,762,789]" pageId="12" pageNumber="565" type="distribution">
<paragraph id="042C36CFC504833DFF6424A00020FCC2" blockId="12.[151,1437,150,789]" box="[199,601,762,789]" pageId="12" pageNumber="565">
<emphasis id="36E7EADDC504833DFF6424A00318FCC3" bold="true" box="[199,353,762,788]" pageId="12" pageNumber="565">Distribution.</emphasis>
<collectingCountry id="7C84765FC504833DFEC424A103D2FCC2" box="[359,427,763,789]" name="Brazil" pageId="12" pageNumber="565">Brazil</collectingCountry>
(
<collectingRegion id="C657F82DC504833DFE1824A10389FCC2" box="[443,496,763,789]" country="Brazil" name="Para" pageId="12" pageNumber="565">Pará</collectingRegion>
) (
<figureCitation id="9CA82A4AC504833DFDA424A10035FCC2" box="[519,588,763,789]" captionStart="FIGURE 2" captionStartId="4.[152,255,1100,1124]" captionTargetBox="[158,1429,189,1070]" captionTargetId="figure-20@4.[151,1436,182,1077]" captionTargetPageId="4" captionText="FIGURE 2. Geographic distribution of Paropsocolis Sampaio &amp; Campos, gen. nov.." figureDoi="http://doi.org/10.5281/zenodo.8212472" httpUri="https://zenodo.org/record/8212472/files/figure.png" pageId="12" pageNumber="565">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>