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<document id="D745494B0DB90A183BDAD93BECB5F731" ENCODING="UTF-8" ID-GBIF-Dataset="55732097-d5aa-4063-a6b4-e4e75f27c917" ModsDocAuthor="Kohout, R. J." ModsDocDate="2007" ModsDocID="21282" ModsDocOrigin="http://hdl.handle.net/10199/15380" ModsDocTitle="Revision of the subgenus Aulacomyrma Emery of the genus Polyrhachis F. Smith, with descriptions of new species." checkinTime="1243372153277" checkinUser="christiana" docAuthor="Kohout, R. J." docDate="2007" docId="14E18B9A55A37CF2DD9C0D388C02DC37" docLanguage="en" docName="21282" docOrigin="Advances in ant systematics (Hymenoptera: Formicidae): Homage to E. O. Wilson - 50 years of contributions. (Memoirs of the American Entomological Institute 80)" docSource="http://hdl.handle.net/10199/15380" docTitle="Polyrhachis (Aulacomyrma) kokoda Kohout, 2007, new species" docType="treatment" docVersion="12" lastPageNumber="227" masterDocId="D65858900E89C21BBC0C1FCAE0C75724" masterDocTitle="Revision of the subgenus Aulacomyrma Emery of the genus Polyrhachis F. Smith, with descriptions of new species." masterLastPageNumber="253" masterPageNumber="186" pageNumber="226" updateTime="1701311414827" updateUser="plazi">
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<mods:title id="2D4E35AC24B16FA6934F62EC49F20930">Revision of the subgenus Aulacomyrma Emery of the genus Polyrhachis F. Smith, with descriptions of new species.</mods:title>
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<treatment id="14E18B9A55A37CF2DD9C0D388C02DC37" ID-DOI="http://doi.org/10.5281/zenodo.6240921" ID-GBIF-Taxon="100134647" ID-Zenodo-Dep="6240921" LSID="urn:lsid:plazi:treatment:14E18B9A55A37CF2DD9C0D388C02DC37" httpUri="http://treatment.plazi.org/id/14E18B9A55A37CF2DD9C0D388C02DC37" lastPageNumber="227" pageNumber="226">
<subSubSection id="2B312AF9D5748898B515C8898170CB47" type="nomenclature">
<paragraph id="6E60413B1E1230B68876508B20171961" pageNumber="226">
<taxonomicName id="437FA4FC6C9DBDC4BEA8DB9B2FDC1DCF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:238317" class="Insecta" family="Formicidae" genus="Polyrhachis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Polyrhachis (Aulacomyrma) kokoda Kohout" order="Hymenoptera" pageNumber="226" phylum="Arthropoda" rank="species" species="kokoda" subGenus="Aulacomyrma">Polyrhachis (Aulacomyrma) kokoda</taxonomicName>
,
<taxonomicNameLabel id="C758363B3F3FA045EFCCB8660C157BFC" rank="species">new species</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="5063B1C3B269431F39E09C875E721026" type="description">
<paragraph id="FBA56ED59A2F7185A621A1D90B35427A" pageNumber="226">Figures 74, 77, 80</paragraph>
</subSubSection>
<subSubSection id="28618DD25274FD08D3D1266C8254A19F" type="nomenclature">
<paragraph id="461B553F8083441672A9AD9EBA0E5D64" pageNumber="226">TYPE MATERIAL</paragraph>
<paragraph id="91E580F743636D44B76965E8DFB9683C" pageNumber="226">
HOLOTYPE:
<materialsCitation id="4352E5C6D3CC8D0E6351B8E3EE461EBF" ID-GBIF-Occurrence="923893380" collectingDate="1972-03-09" collectorName="P. M. Room" country="PAPUA NEW GUINEA" location="Kokoda" specimenCode=" 250" stateProvince="Northern Prov" typeStatus="PARATYPES">
PAPUA NEW GUINEA,
<collectingRegion id="9770B79EB411E9F9649FD125E82485F0">Northern Prov</collectingRegion>
.,
<location id="0C94D76732F7DD360A4E493A64D151DE" LSID="plazi:4A1779D05E59716394E5C9157C1A492D:17AAE93791676B2CF22001D4038BDDA9" country="PAPUA NEW GUINEA" name="Kokoda" stateProvince="Northern Prov">Kokoda</location>
,
<collectingDate id="E01894FFC6A4E9ABE01DE0476192B2AA">9. iii. 1972</collectingDate>
,
<collectorName id="E8E3CB05399D5F13273A1734D42F6D64">P. M. Room</collectorName>
<specimenCode id="CF7B981A076B0FFC68164E7CFB7D6DBD"># 250</specimenCode>
(worker).
<typeStatus id="AFDB217B7B51C36B627935E02E7363DB">PARATYPES</typeStatus>
: data as for
<typeStatus id="1E0F800E19FB932074E3CEFCD2BFBF4F">holotype</typeStatus>
(1 worker)
</materialsCitation>
;
<materialsCitation id="C68E38C51F3AA782D42A83C963DE9835" ID-GBIF-Occurrence="923893381" collectingDate="1971-01-18" collectionCode="ANIC, MCZC, QMBA" collectorName="B. B. Lowery" country="Papua New Guinea" location="Oivi Ridge" stateProvince="Northern Prov" typeStatus="holotype">
<location id="2362E16E18BB022B2C6311F2B3B9576E" LSID="plazi:4A1779D05E59716394E5C9157C1A492D:3B3CF8A1A94B0B5D571FB4298985ADBF" country="Papua New Guinea" name="Oivi Ridge" stateProvince="Northern Prov">Oivi Ridge</location>
,
<location id="FF0ED2E1E6CA481C84024C1FCDCC2F0C" LSID="plazi:4A1779D05E59716394E5C9157C1A492D:CC930FA16837D2FC41F43FC0425D7AD8" country="Papua New Guinea" name="Kokoda" stateProvince="Northern Prov">Kokoda</location>
Rd, 1200 ft, rf.,
<collectingDate id="BE34099CAC13BC917A4FF21B25C2B3D2">18. i. 1971</collectingDate>
,
<collectorName id="A90895B2756CDD7E3A1420D9FAAA28E6">B. B. Lowery</collectorName>
(1 worker). Type distribution:
<typeStatus id="AD813B31A29F52ABB98E1B6B17994E9E">holotype</typeStatus>
in
<collectionCode id="534F23DF5939207520AB6DD05AE5C575" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection">
<collectionCode id="DA97C35F8A614F376E83AAADACD55623" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection">
<collectionCode id="BE704EBA54A97851B71A44E84EDEAFDA" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection">
<collectionCode id="1428684ECAF1E9D992624627AC5D6DA3" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection">
<collectionCode id="8047E11D694351EB9589E33115DC73E7" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection">ANIC</collectionCode>
</collectionCode>
</collectionCode>
</collectionCode>
</collectionCode>
; 1
<typeStatus id="BA5CB082C0B522A1EF812F390FD5F711">paratype</typeStatus>
each in
<collectionCode id="D88B834E23521FA981D06F051DFCBC61" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">
<collectionCode id="DBE2833F5909C6880EA9CD01340E65D4" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">
<collectionCode id="366837E4A697827955406F2F3523DA41" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">
<collectionCode id="4402A1E3AF7CDCF5356A2FFE2A811A8A" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">
<collectionCode id="EF09F1A2FBF61BDB337A7B081AF514F9" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">MCZC</collectionCode>
</collectionCode>
</collectionCode>
</collectionCode>
</collectionCode>
and
<collectionCode id="F9B17A522BF9FE40F56B0EAEABB647C6" collectionName="Australia, Queensland, South Brisbane, Queensland Museum">
<collectionCode id="A52EC260203088DE9242A39DF3C143B2" collectionName="Australia, Queensland, South Brisbane, Queensland Museum">
<collectionCode id="AFA0D24FA2001330CCC0BB352AC05FD8" collectionName="Australia, Queensland, South Brisbane, Queensland Museum">
<collectionCode id="E1BF22EA538F0624924D15D11C742F91" collectionName="Australia, Queensland, South Brisbane, Queensland Museum">
<collectionCode id="A550CBE6F0D1FD81573491FB89A58B8A" collectionName="Australia, Queensland, South Brisbane, Queensland Museum">QMBA</collectionCode>
</collectionCode>
</collectionCode>
</collectionCode>
</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B9564CBE6D421C1899AF343F924FC530" type="description">
<paragraph id="7D9AB1FD023F5CA30C7CC9672F465ACF" pageNumber="226">WORKER</paragraph>
<paragraph id="974F9065EAB2FAF25CB6DB6D7E0B2049" pageNumber="226">Dimensions: TL c. 4.64 - 5.09 (4.79); HL 1.22 - 1.31 (1.31); HW 1.00 - 1.06 (1.03); CI 79 - 82 (79); SL 1.36 - 1.43 (1.43); SI 135 - 139 (139); PW 0.97 - 1.03 (1.00); MTL 1.31 - 1.40 (1.37) (3 measured).</paragraph>
<paragraph id="1F88B0386537D47BD8B9C02088181AFA" pageNumber="226">Clypeus with anterior margin medially truncate or very shallowly emarginate; median longitudinal carina weakly elevated posteriorly before descending towards shallowly impressed basal margin. Frontal carinae sinuate, with rather short, laminate, anteriorly truncate lobes. Sides of head in front of eyes convex; narrowed behind into convex preoccipital margin. Eyes strongly convex, clearly breaking cephalic outline in full face view. Mesosoma immarginate, dorsum sloping onto sides and propodeal declivity in unbroken curve. Pronotal dorsum very strongly raised anteriorly, convex in profile. Pronotal humeri armed with acute, rather long, downward curved spines; bases broad, margins raised. Promesonotal suture distinct; metanotal groove lacking. Petiole with both faces convex and dorsal margin somewhat angular; lateral spines short, curved backwards. Anterior face of first gastral segment convex.</paragraph>
<paragraph id="9FCFA761402D6E71CD90C8E19C5A5F74" pageNumber="226">Mandibles distinctly, longitudinally striate. Sculpture of head, mesosoma and petiole consisting of regularly spaced, rather smooth and glossy striae. Head with longitudinal, mostly anteriorly converging striae; clypeal striae terminating just short of anterior margin. Striae on mesosomal dorsum forming an imperfect elliptical pattern, divided in half by promesonotal suture; outermost striae continued obliquely onto sides and posteriorly extending onto propodeal declivity. Petiole with tranverse, dorsally arched striae on anterior face; inversely V-shaped on posterior face and converging upwards towards dorsal margin. Gaster finely, microscopically, shagreened.</paragraph>
<paragraph id="48D9F7706F24D99BFFFC07E2983650C4" lastPageNumber="227" pageNumber="226">Abundant, long, mostly erect or curved, golden hairs on most of body and appendages, hairs reaching greatest diameter of eyes in length, distinctly shorter on antennal scapes and along dorsal margin of petiole. Propodeal declivity with patch of very short, somewhat dorsally curved, off-white hairs. Appressed, golden pubescence very sparse on dorsum of head and mesosoma; more distinct on pronotal dorsum, forming V-shaped patch running from pronotal spines across the segment, and posteriorly towards propodeal declivity. Thin covering of mostly white or silvery pubescence on sides of mesosoma, coxae, appendages and posterior margins of gastral segments; pubescence most abundant and distinctly reddish on gastral dorsum, somewhat obscuring underlying sculpture.</paragraph>
<paragraph id="E7BAB1EFD6D52ED630E98B85FE8314FA" pageNumber="227">Black, masticatory margins of mandibles, antennal scapes and most of legs medium to dark reddish-brown. Trochanters and basal femora distinctly yellow.</paragraph>
<paragraph id="8E387CBAB917D6C09C2EB8C4B52DA648" pageNumber="227">Sexuals and immature stages unknown.</paragraph>
</subSubSection>
<subSubSection id="FF10AD1885DB924CB01898D3146AAE78" type="materials_examined">
<paragraph id="585EB1310AD8368EE493089F4CCBBE83" pageNumber="227">ADDITIONAL MATERIAL EXAMINED</paragraph>
<paragraph id="985A7ECA954F6B12AC7C0A86976598BE" pageNumber="227">
<materialsCitation id="7AECD79FDFC3B8E69BAD718E8B14E9C2" ID-GBIF-Occurrence="923893382" collectingDate="1959-09-03" collectorName="T. C. Maa" country="PAPUA NEW GUINEA" latitude="-9.45" location="Port Moresby" longitude="147.15" stateProvince="Catalina Estate">
PAPUA NEW GUINEA:
<collectingRegion id="6B7282B018DD0800A84E4A133BAE58F2">Catalina Estate</collectingRegion>
,
<locationDeviation id="7339051866248792CA8E1134677C7697">48 km N of</locationDeviation>
<location id="929556C29B2F72F8E51D9943A3BE877A" LSID="plazi:4A1779D05E59716394E5C9157C1A492D:FA5535E9E518460B45BED4F8C0E466A2" country="PAPUA NEW GUINEA" latitude="-9.45" longitude="147.15" name="Port Moresby" stateProvince="Catalina Estate">Port Moresby</location>
,
<geoCoordinate id="B177B3BBBD257FE7CE6E39F60FA68117" direction="south" orientation="latitude" value="-9.45">09 º 27 S</geoCoordinate>
,
<geoCoordinate id="800271F20FE3F8E91B041AB7E5690FCC" direction="east" orientation="longitude" value="147.15">147 º 09 E</geoCoordinate>
, 500 m,
<collectingDate id="E0DD00560533B3816610EEDAA9CCC92D">3. ix. 1959</collectingDate>
(
<collectorName id="D654306196FB12EE314A64B1A33EDB7C">T. C. Maa</collectorName>
) (w)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="55B25B4D03EE3178C1EE213970A9C28B" type="etymology">
<paragraph id="4883AEC312E155A2CFC277DDA182C2C5" pageNumber="227">ETYMOLOGY</paragraph>
<paragraph id="8A3D41B028A4E4D8336F6291EB997DC6" pageNumber="227">Named after the type locality, Kokoda village, situated on the northern rim of the Owen Stanley Ranges in the Northern Province of Papua New Guinea.</paragraph>
</subSubSection>
<subSubSection id="362ADFE3771F6287BF160412D8C28F63" type="discussion">
<paragraph id="9592526BB9B58FCA50542E8CC768A895" pageNumber="227">REMARKS</paragraph>
<paragraph id="85D55AD5B3FBD2F38A5A3FB331958FAC" pageNumber="227">
<taxonomicName id="DEFA025E0C4E90A4EF564C1CE1032E51" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:238317" class="Insecta" family="Formicidae" genus="Polyrhachis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Polyrhachis (Aulacomyrma) kokoda Kohout" order="Hymenoptera" pageNumber="227" phylum="Arthropoda" rank="species" species="kokoda">P. kokoda</taxonomicName>
is superficially similar to
<taxonomicName id="5CD161EE0158C7065361D0BC778433C4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:238319" class="Insecta" family="Formicidae" genus="Polyrhachis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Polyrhachis (Aulacomyrma) mamba Kohout" order="Hymenoptera" pageNumber="227" phylum="Arthropoda" rank="species" species="mamba">P. mamba</taxonomicName>
described below, with both species evidently sympatric. The paratype of kokoda was collected together with a mamba queen at the same locality at Oivi Ridge and they were subsequently mounted on the same pin. Both species share the somewhat elliptical striation of the mesosomal dorsum but there the similarity ends. They differ in numerous characters, including the shape of the eyes, which in kokoda are strongly convex and clearly project beyond the lateral outline of the head. In contrast the eyes in mamba are virtually flat and sunk into a shallow concavity in the cephalic sculpture and they do not break the cephalic outline in full face view. In lateral view the outline of the mesosoma in kokoda features a strongly convex pronotal dorsum and the propodeal dorsum descends into the declivity in an uninterrupted curve. In mamba the pronotum is flatter and the propodeal dorsum descends abruptly into a rather short, vertical declivity. The dorsum of the petiole has an acute margin in kokoda, while it is bluntly rounded in mamba. The first gastral segment of kokoda is finely shagreened in contrast to mamba, where its sides are distinctly, longitudinally striate. The pubescence on the head and body is golden in kokoda, with a distinct reddish tint on the gastral dorsum, while in mamba the pubescence is uniformly greyish or white.
</paragraph>
</subSubSection>
</treatment>
</document>