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<document id="92829CFE12B906B7F85DD28B260D32CD" ID-DOI="10.5281/zenodo.5398985" ID-ISSN="1638-9387" ID-Zenodo-Dep="5398985" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1630329734525" checkinUser="marcus" docAuthor="Vallès, Yvonne, Valdés, Ángel &amp; Ortea, Jesús" docDate="2000" docId="096B7348DD6BFF80FD2CFC33FC9AF959" docLanguage="en" docName="Zoosystema.2000.22.1.15-31.pdf" docOrigin="Zoosystema 22 (1)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Okenia amoenula" docType="treatment" docVersion="1" lastPageNumber="17" masterDocId="F5520B30DD6AFF82FFC2FFA2FFC3FFF1" masterDocTitle="On the phanerobranch dorids of Angola (Mollusca, Nudibranchia): a crossroads of temperate and tropical species" masterLastPageNumber="31" masterPageNumber="15" pageNumber="16" updateTime="1740157300531" updateUser="juliana" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="839F6727570C5CCC74091F6A46299FE5">On the phanerobranch dorids of Angola (Mollusca, Nudibranchia): a crossroads of temperate and tropical species</mods:title>
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<mods:namePart id="545FBCA820563610C7BDF8D6790F8479">Vallès, Yvonne</mods:namePart>
<mods:affiliation id="53368E62FF474946B80DB9BF73BD6662">P. O. Box 15868, Petion Ville, W. I. (Haiti) valles @ acn 2. net</mods:affiliation>
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<mods:namePart id="C924983B8FC485341CCD07AACF154AB3">Valdés, Ángel</mods:namePart>
<mods:affiliation id="AF73610D22024D14D20D4CFC998EB75C">Department of Invertebrate Zoology and Geology, California Academy of Sciences Golden Gate Park, San Francisco, CA 94118 (USA) avaldes @ calacademy. org</mods:affiliation>
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<mods:namePart id="68D008D5EFDC6190A7425E2152897B54">Ortea, Jesús</mods:namePart>
<mods:affiliation id="1228A641BC1BF763191A63504668471D">Departamento de Biología de Organismos y Sistemas, Laboratorio de Zoología Universidad de Oviedo, 33071 Oviedo (Spain) jortea @ sci. cpd. uniovi. es</mods:affiliation>
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<mods:date id="52BE21399BA11645D71646DF84CA16F6">2000</mods:date>
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<treatment id="096B7348DD6BFF80FD2CFC33FC9AF959" LSID="urn:lsid:plazi:treatment:096B7348DD6BFF80FD2CFC33FC9AF959" httpUri="http://treatment.plazi.org/id/096B7348DD6BFF80FD2CFC33FC9AF959" lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16">
<subSubSection id="C9D891D5DD6BFF83FD2CFC33FBB3FC5A" box="[750,1136,912,939]" pageId="1" pageNumber="16" type="nomenclature">
<paragraph id="817DC25EDD6BFF83FD2CFC33FBB3FC5A" blockId="1.[750,1136,912,969]" box="[750,1136,912,939]" pageId="1" pageNumber="16">
<heading id="DA357532DD6BFF83FD2CFC33FBB3FC5A" box="[750,1136,912,939]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="16" reason="2">
<taxonomicName id="46C2B9DDDD6BFF83FD2CFC33FBB3FC5A" ID-CoL="9G5LY" authority="(Bergh, 1907)" baseAuthorityName="Bergh" baseAuthorityYear="1907" box="[750,1136,912,939]" class="Gastropoda" family="Goniodorididae" genus="Okenia" isUncertain="true" kingdom="Animalia" order="Nudibranchia" pageId="1" pageNumber="16" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD6BFF83FD2CFC33FCFDFC5A" bold="true" box="[750,830,913,939]" italics="true" pageId="1" pageNumber="16">Okenia</emphasis>
cf.
<emphasis id="B3B61E4CDD6BFF83FCA4FC33FC12FC5A" bold="true" box="[870,977,913,939]" italics="true" pageId="1" pageNumber="16">amoenula</emphasis>
(
<bibRefCitation id="E553BFAFDD6BFF83FC1DFC33FBABFC5A" author="Bergh R." bookContentInfo="144 p." box="[991,1128,913,939]" journalOrPublisher="Transactions of the South African Philosophical Society" pageId="1" pageNumber="16" part="17" refId="ref8210" refString="Bergh R. 1907. - The Opisthobranchiata of South Africa. Transactions of the South African Philosophical Society 17: 144 p., 14 pls." title="The Opisthobranchiata of South Africa" type="book" year="1907">Bergh, 1907</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD6BFF83FC45FC0DFC14FC38" box="[903,983,943,969]" pageId="1" pageNumber="16" type="description">
<paragraph id="817DC25EDD6BFF83FC45FC0DFC14FC38" blockId="1.[750,1136,912,969]" box="[903,983,943,969]" pageId="1" pageNumber="16">
(
<figureCitation id="19F9DEDBDD6BFF83FC4DFC0DFC0FFC38" box="[911,972,943,969]" captionStart="FIG" captionStartId="2.[114,125,758,775]" captionTargetBox="[120,1207,205,709]" captionTargetId="figure-509@2.[111,1218,196,722]" captionTargetPageId="2" captionText="FIG. 1. — Okenia cf. amoenula (MNHN), dorsal view of the living animal. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5398989" httpUri="https://zenodo.org/record/5398989/files/figure.png" pageId="1" pageNumber="16">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD6BFF83FD67FC43FB79FC08" box="[677,1210,993,1017]" pageId="1" pageNumber="16" type="reference_group">
<paragraph id="817DC25EDD6BFF83FD67FC43FB79FC08" blockId="1.[677,1210,993,1192]" box="[677,1210,993,1017]" pageId="1" pageNumber="16">
<taxonomicName id="46C2B9DDDD6BFF83FD67FC43FBDFFC08" authority="Bergh, 1907: 80 - 82" authorityName="Bergh" authorityPageNumber="80 - 82" authorityYear="1907" box="[677,1052,993,1017]" class="Gastropoda" family="Goniodorididae" genus="Idaliella" kingdom="Animalia" order="Nudibranchia" pageId="1" pageNumber="16" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD6BFF83FD67FC43FC90FC08" box="[677,851,993,1017]" italics="true" pageId="1" pageNumber="16">Idaliella amoenula</emphasis>
<bibRefCitation id="E553BFAFDD6BFF83FC9BFC43FBDFFC08" author="Bergh R." bookContentInfo="144 p." box="[857,1052,993,1017]" journalOrPublisher="Transactions of the South African Philosophical Society" pageId="1" pageNumber="16" part="17" refId="ref8210" refString="Bergh R. 1907. - The Opisthobranchiata of South Africa. Transactions of the South African Philosophical Society 17: 144 p., 14 pls." title="The Opisthobranchiata of South Africa" type="book" year="1907">Bergh, 1907: 80-82</bibRefCitation>
</taxonomicName>
, pl. 8, figs 6-11.
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD6BFF83FD67FBA7FC4EFBA1" pageId="1" pageNumber="16" type="materials_examined">
<paragraph id="817DC25EDD6BFF83FD67FBA7FC4EFBA1" blockId="1.[677,1210,993,1192]" pageId="1" pageNumber="16">
MATERIAL EXAMINED. —
<materialsCitation id="31AAC803DD6BFF83FC63FBA6FC4EFBA1" pageId="1" pageNumber="16">
<emphasis id="B3B61E4CDD6BFF83FC63FBA6FB91FBEF" bold="true" box="[929,1106,1028,1054]" pageId="1" pageNumber="16">
<collectingRegion id="43060CBCDD6BFF83FC63FBA6FB8EFBEF" box="[929,1101,1028,1054]" country="Angola" name="Luanda" pageId="1" pageNumber="16">Luanda Province</collectingRegion>
.
</emphasis>
Punta das Palmeirinhas,
<geoCoordinate id="E4F6A499DD6BFF83FCF5FBBDFCBFFBC6" box="[823,892,1055,1079]" degrees="9" direction="south" minutes="09" orientation="latitude" pageId="1" pageNumber="16" precision="925" value="-9.15">9°09S</geoCoordinate>
,
<geoCoordinate id="E4F6A499DD6BFF83FC4BFBBDFC1DFBC4" box="[905,990,1054,1079]" degrees="12" direction="east" minutes="58" orientation="longitude" pageId="1" pageNumber="16" precision="925" value="12.966666">12°58E</geoCoordinate>
,
<date id="F57CE49EDD6BFF83FC29FBBDFBFAFBC6" box="[1003,1081,1055,1079]" pageId="1" pageNumber="16" value="1987-02">II.1987</date>
,
<specimenCount id="97C409D7DD6BFF83FB85FBBDFB79FBC6" box="[1095,1210,1055,1079]" pageId="1" pageNumber="16" type="generic">1 specimen</specimenCount>
<quantity id="463A6FBBDD6BFF83FD67FB9AFD23FBBF" box="[677,736,1079,1104]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="16" unit="mm" value="6.0">6 mm</quantity>
preserved length.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD6BFF83FD67FBFEFB61FB59" pageId="1" pageNumber="16" type="distribution">
<paragraph id="817DC25EDD6BFF83FD67FBFEFB61FB59" blockId="1.[677,1210,993,1192]" pageId="1" pageNumber="16">
DISTRIBUTION. —
<taxonomicName id="46C2B9DDDD6BFF83FCAFFBFFFBE3FB84" baseAuthorityName="Bergh" baseAuthorityYear="1907" box="[877,1056,1117,1141]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="1" pageNumber="16" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD6BFF83FCAFFBFFFBE3FB84" box="[877,1056,1117,1141]" italics="true" pageId="1" pageNumber="16">Okenia amoenula</emphasis>
</taxonomicName>
has only been reported from
<collectingCountry id="F9D582CEDD6BFF83FCFDFBD4FC02FB7F" box="[831,961,1142,1166]" name="South Africa" pageId="1" pageNumber="16">South Africa</collectingCountry>
(
<bibRefCitation id="E553BFAFDD6BFF83FC13FBD4FBB4FB7F" author="Gosliner T. M." bookContentInfo="Sea Challengers, Monterey, 136 p." box="[977,1143,1142,1166]" pageId="1" pageNumber="16" refId="ref8424" refString="Gosliner T. M. 1987 a. - Nudibranchs of Southern Africa. A Guide to Opisthobranch Mollusks of Southern Africa. Sea Challengers, Monterey, 136 p." title="Nudibranchs of Southern Africa. A Guide to Opisthobranch Mollusks of Southern Africa" type="book" year="1987">Gosliner 1987a</bibRefCitation>
). This paper is the first reference for this species to
<collectingCountry id="F9D582CEDD6BFF83FB97FB32FB5DFB59" box="[1109,1182,1168,1192]" name="Angola" pageId="1" pageNumber="16">Angola</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD6BFF83FD67FB42FBB9F956" pageId="1" pageNumber="16" type="description">
<paragraph id="817DC25EDD6BFF83FD67FB42FCF6FB09" blockId="1.[677,1210,1248,1704]" box="[677,821,1248,1274]" pageId="1" pageNumber="16">DESCRIPTION</paragraph>
<paragraph id="817DC25EDD6BFF83FD67FB5DFBB9F956" blockId="1.[677,1210,1248,1704]" pageId="1" pageNumber="16">
The background body colour is opaque white in the living animal. In the centre of the dorsum there are a few reddish spots and short lines of the same colour (
<figureCitation id="19F9DEDBDD6BFF83FCF4FAF9FCB6FA84" box="[822,885,1371,1397]" captionStart="FIG" captionStartId="2.[114,125,758,775]" captionTargetBox="[120,1207,205,709]" captionTargetId="figure-509@2.[111,1218,196,722]" captionTargetPageId="2" captionText="FIG. 1. — Okenia cf. amoenula (MNHN), dorsal view of the living animal. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5398989" httpUri="https://zenodo.org/record/5398989/files/figure.png" pageId="1" pageNumber="16">Fig. 1</figureCitation>
). The oral tentacles are white and have four yellow tips each. The dorsum has, on each lateral margin, ten short papillae with yellow pigmentation on the apex. All over the dorsum there is a network of spicules. The rhinophores are white, non-retractile, and ringed with 18 lamellae. There are 11 unipinnate (the largest one is bipinnate) branchial leaves white in colour, having the rachis yellow (in the distal region edged with two red lines). The tail has a yellow line in the centre with few red spots.
</paragraph>
</subSubSection>
<caption id="D5BD92D6DD68FF80FFB0FD54FC95FCF6" ID-DOI="http://doi.org/10.5281/zenodo.5398989" ID-Zenodo-Dep="5398989" box="[114,854,758,775]" httpUri="https://zenodo.org/record/5398989/files/figure.png" pageId="2" pageNumber="17" startId="2.[114,125,758,775]" targetBox="[120,1207,205,709]" targetPageId="2">
<paragraph id="817DC25EDD68FF80FFB0FD54FC95FCF6" blockId="2.[114,854,758,775]" box="[114,854,758,775]" pageId="2" pageNumber="17">
FIG. 1. —
<taxonomicName id="46C2B9DDDD68FF80FF03FD54FE75FCF6" authority="(MNHN)" baseAuthorityName="MNHN" box="[193,438,758,775]" class="Gastropoda" family="Goniodorididae" genus="Okenia" isUncertain="true" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FF03FD54FF39FCF6" box="[193,250,758,775]" italics="true" pageId="2" pageNumber="17">Okenia</emphasis>
cf.
<emphasis id="B3B61E4CDD68FF80FED8FD54FEA8FCF6" box="[282,363,758,775]" italics="true" pageId="2" pageNumber="17">amoenula</emphasis>
(MNHN)
</taxonomicName>
, dorsal view of the living animal. Scale bar: 1 mm.
</paragraph>
</caption>
<subSubSection id="C9D891D5DD68FF80FFB0FCD2FCF5FB6E" pageId="2" pageNumber="17" type="discussion">
<paragraph id="817DC25EDD68FF80FFB0FCD2FF15FC79" blockId="2.[114,648,880,1704]" box="[114,214,880,906]" pageId="2" pageNumber="17">REMARKS</paragraph>
<paragraph id="817DC25EDD68FF80FFB0FC2DFEFDFBB2" blockId="2.[114,648,880,1704]" pageId="2" pageNumber="17">The single specimen collected was badly preserved, making dissection and investigation of the internal anatomy impossible. In the process of removing the foregut the radula was lost. For that reason a positive identification of the specimen could not be done.</paragraph>
<paragraph id="817DC25EDD68FF80FFB0FBE5FDAFF959" blockId="2.[114,648,880,1704]" pageId="2" pageNumber="17">
<bibRefCitation id="E553BFAFDD68FF80FFB0FBE5FEC1FB90" author="Bergh R." bookContentInfo="144 p." box="[114,258,1095,1121]" journalOrPublisher="Transactions of the South African Philosophical Society" pageId="2" pageNumber="17" part="17" refId="ref8210" refString="Bergh R. 1907. - The Opisthobranchiata of South Africa. Transactions of the South African Philosophical Society 17: 144 p., 14 pls." title="The Opisthobranchiata of South Africa" type="book" year="1907">Bergh (1907)</bibRefCitation>
described
<taxonomicName id="46C2B9DDDD68FF80FEBAFBE5FF70FB71" authority="(Bergh, 1907)" baseAuthorityName="Bergh" baseAuthorityYear="1907" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FEBAFBE5FDF3FB90" box="[376,560,1095,1121]" italics="true" pageId="2" pageNumber="17">Okenia amoenula</emphasis>
(
<bibRefCitation id="E553BFAFDD68FF80FDFDFBE5FF68FB71" author="Bergh R." bookContentInfo="144 p." journalOrPublisher="Transactions of the South African Philosophical Society" pageId="2" pageNumber="17" part="17" refId="ref8210" refString="Bergh R. 1907. - The Opisthobranchiata of South Africa. Transactions of the South African Philosophical Society 17: 144 p., 14 pls." title="The Opisthobranchiata of South Africa" type="book" year="1907">Bergh, 1907</bibRefCitation>
)
</taxonomicName>
in the binomen
<taxonomicName id="46C2B9DDDD68FF80FEB1FBC4FF6FFB6E" authority="Bergh, 1907" authorityName="Bergh" authorityYear="1907" class="Gastropoda" family="Goniodorididae" genus="Idaliella" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FEB1FBC4FDFBFB71" box="[371,568,1126,1152]" italics="true" pageId="2" pageNumber="17">Idaliella amoenula</emphasis>
<bibRefCitation id="E553BFAFDD68FF80FD83FBC4FF6FFB6E" author="Bergh R." bookContentInfo="144 p." journalOrPublisher="Transactions of the South African Philosophical Society" pageId="2" pageNumber="17" part="17" refId="ref8210" refString="Bergh R. 1907. - The Opisthobranchiata of South Africa. Transactions of the South African Philosophical Society 17: 144 p., 14 pls." title="The Opisthobranchiata of South Africa" type="book" year="1907">Bergh, 1907</bibRefCitation>
</taxonomicName>
, as being a white animal with three crimson lines on the dorsum, which joined together in the anterior part, and a crimson patch on the anal area. The dorsal papillae were reddish and yellow, the branchial leaves had a crimson colour, and the rhinophores were yellow. Our specimen, as well as Berghs, has a white background colour, red and yellow pigmentation on the lateral papillae, and the branchial leaves are also reddish (with two red lines). However, our specimen shows a few differences in colour pattern from Berghs material (white rhinophores instead of yellow, white branchial leaves instead of crimson), and with that described by
<bibRefCitation id="E553BFAFDD68FF80FEC3F9B1FE7AF9DC" author="Gosliner T. M." bookContentInfo="Sea Challengers, Monterey, 136 p." box="[257,441,1555,1581]" pageId="2" pageNumber="17" refId="ref8424" refString="Gosliner T. M. 1987 a. - Nudibranchs of Southern Africa. A Guide to Opisthobranch Mollusks of Southern Africa. Sea Challengers, Monterey, 136 p." title="Nudibranchs of Southern Africa. A Guide to Opisthobranch Mollusks of Southern Africa" type="book" year="1987">Gosliner (1987a)</bibRefCitation>
from
<collectingCountry id="F9D582CEDD68FF80FE3CF9B1FD44F9DC" box="[510,647,1555,1581]" name="South Africa" pageId="2" pageNumber="17">South Africa</collectingCountry>
(dorsum with more red pigmentation in Gosliners specimen and yellow lateral papillae, but also with red spots), which could be caused by differences in size. Otherwise they are identical.
</paragraph>
<paragraph id="817DC25EDD68FF80FD67FCD2FCF5FB6E" blockId="2.[677,1212,880,1704]" pageId="2" pageNumber="17">
<taxonomicName id="46C2B9DDDD68FF80FD67FCD2FC43FC7A" baseAuthorityName="Bergh" baseAuthorityYear="1907" box="[677,896,880,907]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FD67FCD2FC43FC7A" box="[677,896,880,907]" italics="true" pageId="2" pageNumber="17">Okenia amoenula</emphasis>
</taxonomicName>
has been considered by
<bibRefCitation id="E553BFAFDD68FF80FD67FC2DFCBEFC58" author="Schmekel L." box="[677,893,911,937]" journalOrPublisher="Veliger" pageId="2" pageNumber="17" pagination="355 - 360" part="21" refId="ref8798" refString="Schmekel L. 1979. - First record of Okenia impexa Marcus, 1957 from the Western Atlantic in the Mediterranean. Veliger 21: 355 - 360." title="First record of Okenia impexa Marcus, 1957 from the Western Atlantic in the Mediterranean" type="journal article" year="1979">Schmekel (1979)</bibRefCitation>
as a synonym of
<taxonomicName id="46C2B9DDDD68FF80FBA0FC2DFC35FC39" authority="(Ihering, 1886)" baseAuthorityName="Ihering" baseAuthorityYear="1886" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="mediterranea">
<emphasis id="B3B61E4CDD68FF80FBA0FC2DFCF6FC39" italics="true" pageId="2" pageNumber="17">Okenia mediterranea</emphasis>
(Ihering, 1886)
</taxonomicName>
.
<taxonomicName id="46C2B9DDDD68FF80FBC1FC0CFD22FC14" baseAuthorityName="Ihering" baseAuthorityYear="1886" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="mediterranea">
<emphasis id="B3B61E4CDD68FF80FBC1FC0CFD22FC14" italics="true" pageId="2" pageNumber="17">Okenia mediterranea</emphasis>
</taxonomicName>
, like our material of
<taxonomicName id="46C2B9DDDD68FF80FC1CFC6EFBADFC16" baseAuthorityName="Bergh" baseAuthorityYear="1907" box="[990,1134,972,999]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FC1CFC6EFBADFC16" box="[990,1134,972,999]" italics="true" pageId="2" pageNumber="17">O. amoenula</emphasis>
</taxonomicName>
, has a yellow line in the centre of the tail with some red spots, and the rhinophores are white. However, our specimen lacks tubercles in the centre of the dorsum, which is a definitive character that separates the two species (
<bibRefCitation id="E553BFAFDD68FF80FB8FFBC4FCEAFB6E" author="Valdes A. &amp; Ortea J." journalOrPublisher="Veliger" pageId="2" pageNumber="17" pagination="223 - 234" part="38" refId="ref8858" refString="Valdes A. &amp; Ortea J. 1995. - Revised taxonomy of some species of the genus Okenia Menke, 1830 (Mollusca: Nudibranchia) from the Atlantic Ocean, with the description of a new species. Veliger 38: 223 - 234." title="Revised taxonomy of some species of the genus Okenia Menke, 1830 (Mollusca: Nudibranchia) from the Atlantic Ocean, with the description of a new species" type="journal article" year="1995">Valdés &amp; Ortea 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD68FF80FD67FB01FC9AF959" pageId="2" pageNumber="17" type="description">
<paragraph id="817DC25EDD68FF80FD67FB01FC9AF959" blockId="2.[677,1212,880,1704]" pageId="2" pageNumber="17">
<taxonomicName id="46C2B9DDDD68FF80FD67FB01FCA7FB4C" baseAuthorityName="Bergh" baseAuthorityYear="1907" box="[677,868,1187,1213]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FD67FB01FCA7FB4C" box="[677,868,1187,1213]" italics="true" pageId="2" pageNumber="17">Okenia amoenula</emphasis>
</taxonomicName>
is similar in colour to
<taxonomicName id="46C2B9DDDD68FF80FBAFFB01FBDFFB2D" authority="Valdes &amp; Ortea, 1995" authorityName="Valdes &amp; Ortea" authorityYear="1995" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="hispanica">
<emphasis id="B3B61E4CDD68FF80FBAFFB01FCCEFB2D" italics="true" pageId="2" pageNumber="17">Okenia hispanica</emphasis>
<bibRefCitation id="E553BFAFDD68FF80FCD4FB60FBDFFB2D" author="Valdes A. &amp; Ortea J." box="[790,1052,1218,1244]" journalOrPublisher="Veliger" pageId="2" pageNumber="17" pagination="223 - 234" part="38" refId="ref8858" refString="Valdes A. &amp; Ortea J. 1995. - Revised taxonomy of some species of the genus Okenia Menke, 1830 (Mollusca: Nudibranchia) from the Atlantic Ocean, with the description of a new species. Veliger 38: 223 - 234." title="Revised taxonomy of some species of the genus Okenia Menke, 1830 (Mollusca: Nudibranchia) from the Atlantic Ocean, with the description of a new species" type="journal article" year="1995">Valdés &amp; Ortea, 1995</bibRefCitation>
</taxonomicName>
. Both species have a white background colour, a white tail with a yellow line, and yellow pigmentation on the gill. However, they are clearly distinguishable by several external features. The dorsum of
<taxonomicName id="46C2B9DDDD68FF80FBB2FA9EFD2EFA85" authorityName="Valdes &amp; Ortea" authorityYear="1995" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="hispanica">
<emphasis id="B3B61E4CDD68FF80FBB2FA9EFD2EFA85" italics="true" pageId="2" pageNumber="17">O. hispanica</emphasis>
</taxonomicName>
has two large pink patches, just behind each rhinophore, and another one on the anal area, the rhinophores are pink, and the oral tentacles are white, whereas in
<taxonomicName id="46C2B9DDDD68FF80FC0EFA14FB9BFA20" baseAuthorityName="Bergh" baseAuthorityYear="1907" box="[972,1112,1462,1489]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FC0EFA14FB9BFA20" box="[972,1112,1462,1489]" italics="true" pageId="2" pageNumber="17">O. amoenula</emphasis>
</taxonomicName>
the dorsum is white with red spots or lines (lacking large patches), the rhinophores are white or yellow and the oral tentacles are white with four small yellow spots. Other differences include the absence of tubercles on the dorsum of
<taxonomicName id="46C2B9DDDD68FF80FC18F9F2FB9CF99A" authorityName="Valdes &amp; Ortea" authorityYear="1995" box="[986,1119,1616,1643]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="hispanica">
<emphasis id="B3B61E4CDD68FF80FC18F9F2FB9CF99A" box="[986,1119,1616,1643]" italics="true" pageId="2" pageNumber="17">O. hispanica</emphasis>
</taxonomicName>
and the radular morphology, already compared by
<bibRefCitation id="E553BFAFDD68FF80FBB7F9CDFC96F959" author="Valdes A. &amp; Ortea J." journalOrPublisher="Veliger" pageId="2" pageNumber="17" pagination="223 - 234" part="38" refId="ref8858" refString="Valdes A. &amp; Ortea J. 1995. - Revised taxonomy of some species of the genus Okenia Menke, 1830 (Mollusca: Nudibranchia) from the Atlantic Ocean, with the description of a new species. Veliger 38: 223 - 234." title="Revised taxonomy of some species of the genus Okenia Menke, 1830 (Mollusca: Nudibranchia) from the Atlantic Ocean, with the description of a new species" type="journal article" year="1995">Valdés &amp; Ortea (1995)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>