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<document id="C3F1E68DB045DDEBAD7F851954652530" ID-DOI="10.5281/zenodo.5170505" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170505" ID-ZooBank="B39DE284-15C2-4ADE-B1B9-9146D3DFE764" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628285093964" checkinUser="felipe" docAuthor="Ferro, Michael L." docDate="2016" docId="CB3843409160FFB16CA2B86628D35203" docLanguage="en" docName="InsectaMundi.0472.1-57.pdf" docOrigin="Insecta Mundi 2016 (472)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Sonoma colberti Ferro 2016, new species" docType="treatment" docVersion="4" lastPageNumber="13" masterDocId="37013B38916EFFBF6C62B9072D725479" masterDocTitle="Fourteen new species of Sonoma Casey (Coleoptera: Staphylinidae: Pselaphinae) with a key to species from western North America" masterLastPageNumber="57" masterPageNumber="1" pageNumber="13" updateTime="1699040133542" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="9FB13250B834AA01180EBF73B68DFD22">Fourteen new species of Sonoma Casey (Coleoptera: Staphylinidae: Pselaphinae) with a key to species from western North America</mods:title>
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<mods:namePart id="6D8D7E3C8D4A277A86D99235358F991A">Ferro, Michael L.</mods:namePart>
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<mods:title id="800F387A47D9E90E5BBB4CF2E99B34B0">Insecta Mundi</mods:title>
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<mods:date id="5B49F2F51A6E035FA6A6699B64E8D9FC">2016</mods:date>
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<mods:number id="A987899E7DA58CE1611B1A93D65AFA62">2016-03-11</mods:number>
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<mods:number id="191A240F8F4BFAB39463EF29A5D58D86">472</mods:number>
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<subSubSection id="0B8BA1DD9160FFB16CA2B8662F185500" box="[192,618,353,378]" pageId="14" pageNumber="13" type="nomenclature">
<paragraph id="432EF2569160FFB16CA2B8662F185500" blockId="14.[192,618,353,410]" box="[192,618,353,378]" pageId="14" pageNumber="13">
<heading id="1866453A9160FFB16CA2B8662F185500" bold="true" box="[192,618,353,378]" fontSize="10" level="2" pageId="14" pageNumber="13" reason="3">
<emphasis id="71E52E449160FFB16CA2B8662F185500" bold="true" box="[192,618,353,378]" pageId="14" pageNumber="13">
8.
<taxonomicName id="849189D59160FFB16CBCB8652CCF5500" ID-CoL="6Z724" authority="Ferro, 2016" authorityName="Ferro" authorityYear="2016" box="[222,445,353,378]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="colberti" status="sp. nov.">
<emphasis id="71E52E449160FFB16CBCB8652CCF5500" bold="true" box="[222,445,353,378]" italics="true" pageId="14" pageNumber="13">Sonoma colberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AD6933F9160FFB16DA7B8652F185500" box="[453,618,353,378]" pageId="14" pageNumber="13" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0B8BA1DD9160FFB16CA2B8852F6257C3" pageId="14" pageNumber="13" type="description">
<paragraph id="432EF2569160FFB16CA2B8852CD455E3" blockId="14.[192,618,353,410]" box="[192,422,386,410]" pageId="14" pageNumber="13">
<figureCitation id="DBAAEED39160FFB16CA2B8852C7555E3" box="[192,263,386,410]" captionStart="Figures 711" captionStartId="23.[192,288,1808,1830]" captionTargetBox="[240,1378,224,1735]" captionTargetId="figure-13@23.[240,1378,192,1841]" captionTargetPageId="23" captionText="Figures 711. Sonoma spp. holotype habiti. 7) S. cobra. 8) S. colberti. 9) S. maryae. 10) S. quellazaire. 11) S. rossellinae." figureDoi="http://doi.org/10.5281/zenodo.5170509" httpUri="https://zenodo.org/record/5170509/files/figure.png" pageId="14" pageNumber="13">Fig. 8</figureCitation>
,
<figureCitation id="DBAAEED39160FFB16D77B8852C4755E3" box="[277,309,386,410]" captionStart="Figures 1518" captionStartId="25.[192,288,1821,1843]" captionTargetBox="[217,1410,194,1796]" captionTargetId="figure-13@25.[192,1421,192,1801]" captionTargetPageId="25" captionText="Figures 1518. Aedeagi of Sonoma spp. 15) S. stewarti new species. 16) S. wintuorum Chandler. Modified from Chandler (2003). 17) S. twaini new species. 18) S. colberti new species. Right side of figures is anatomical left." figureDoi="http://doi.org/10.5281/zenodo.5170513" httpUri="https://zenodo.org/record/5170513/files/figure.png" pageId="14" pageNumber="13">18</figureCitation>
; Map 11.
</paragraph>
<paragraph id="432EF2569160FFB16CA2B8C629435663" blockId="14.[192,1441,449,954]" pageId="14" pageNumber="13">
<emphasis id="71E52E449160FFB16CA2B8C62C1855A0" bold="true" box="[192,362,449,473]" pageId="14" pageNumber="13">Description.</emphasis>
<typeStatus id="9C2A4CF49160FFB16D0DB8C52C9255A3" box="[367,480,450,474]" pageId="14" pageNumber="13" type="holotype">Holotype</typeStatus>
, male. Measurements: head 0.23 long, 0.27 wide; pronotum 0.27 long, 0.28 wide; elytra 0.51 long, 0.26 wide; antennomeres 111 total 0.62; total length 1.90. Body uniformly brown. Body setae covered in elongate (greater than half the length of the eye).
</paragraph>
<paragraph id="432EF2569160FFB16C92BB2529BC5623" blockId="14.[192,1441,449,954]" pageId="14" pageNumber="13">
<emphasis id="71E52E449160FFB16C92BB252C435643" box="[240,305,546,570]" italics="true" pageId="14" pageNumber="13">Head</emphasis>
. Eyes prominent, maximum length in dorsal view 0.83x length of first antennal segment, with approximately 40 facets. Antennomere 2 approximately equal width of 1; 3 smallest.
</paragraph>
<paragraph id="432EF2569160FFB16C92BB65291D56E3" blockId="14.[192,1441,449,954]" pageId="14" pageNumber="13">
<emphasis id="71E52E449160FFB16C92BB652C3A5603" box="[240,328,610,634]" italics="true" pageId="14" pageNumber="13">Thorax</emphasis>
. Elytra with 3 indistinct sutural foveae; 2 foveae lateral to sutural fovea; central row of 24 foveae in basal 1/3. Winged. Metatrochanter elongate; metatibia unmodified.
</paragraph>
<paragraph id="432EF2569160FFB16C92BBA529EA56A0" blockId="14.[192,1441,449,954]" pageId="14" pageNumber="13">
<emphasis id="71E52E449160FFB16C92BBA52C1156C3" box="[240,355,674,698]" italics="true" pageId="14" pageNumber="13">Abdomen</emphasis>
. Tergite IV with transverse patch of microtrichia narrowly interrupted at midline. Basal lateral foveae on ventrites IVVI. Basal pubescence present on ventrites IIIVI.
</paragraph>
<paragraph id="432EF2569160FFB16C92BBE52F6257C3" blockId="14.[192,1441,449,954]" pageId="14" pageNumber="13">
<emphasis id="71E52E449160FFB16C92BBE52C165683" box="[240,356,738,762]" italics="true" pageId="14" pageNumber="13">Aedeagus</emphasis>
. Elongate. Left paramere: subequal in length to endophallus; subequal in width in basal 4/5; apical 1/5 widened, paddle shaped with 710 stout setae directed mesad. Endophallus: widest at basal 1/3; stout subapical projection 3/5 from base on right lateral margin; left apical process with wide posteriorly directed triangular projection, tip bent anteriorly; median apical process thin, elongate, shaped like a shepherds hook; right apical process a stout pointed projection; Right paramere: 2/3 length left paramere; subequal in width in basal 3/4; apical 1/4 widened, paddle shapped, with 710 stout setae directed mesad.
</paragraph>
</subSubSection>
<subSubSection id="0B8BA1DD9160FFB16CA2BAE52C3650A3" pageId="14" pageNumber="13" type="materials_examined">
<paragraph id="432EF2569160FFB16CA2BAE52C615063" blockId="14.[192,1440,993,1146]" pageId="14" pageNumber="13">
<emphasis id="71E52E449160FFB16CA2BAE52CF85780" bold="true" box="[192,394,993,1018]" pageId="14" pageNumber="13">Type Material.</emphasis>
<materialsCitation id="F3F9F80B9160FFB16DF3BAE52C615063" collectionCode="T, R, CSAC" collectorName="Haig Whiskeytown Shasta Co. Cal." county="Haig Whiskeytown Shasta Co." location="Haig Whiskeytown Shasta Co." pageId="14" pageNumber="13" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="9C2A4CF49160FFB16DF3BAE52F715783" box="[401,515,994,1018]" pageId="14" pageNumber="13" type="holotype">Holotype</typeStatus>
, male:
<collectionCode id="25806A939160FFB16E3BBAE52F1E5783" box="[601,620,994,1018]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="13">T</collectionCode>
.
<collectionCode id="25806A939160FFB16E12BAE52FF45783" box="[624,646,994,1018]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="13">R</collectionCode>
.
<collectorName id="EE6497809160FFB16EEBBAE529495783" box="[649,1083,994,1018]" pageId="14" pageNumber="13">
<collectingCounty id="AA4F8ADA9160FFB16EEBBAE529715783" box="[649,1027,994,1018]" pageId="14" pageNumber="13">Haig Whiskeytown Shasta Co.</collectingCounty>
Cal.
</collectorName>
III.9.1973 (
<specimenCount id="559739DF9160FFB168ACBAE5299D5783" box="[1230,1263,993,1018]" pageId="14" pageNumber="13" type="male">1♂</specimenCount>
). Deposited in
<collectionCode id="25806A939160FFB16CA2BD052C7D5063" box="[192,271,1026,1050]" pageId="14" pageNumber="13">CSAC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="432EF2569160FFB16C92BD252C9F5003" blockId="14.[192,1440,993,1146]" pageId="14" pageNumber="13">
<emphasis id="71E52E449160FFB16C92BD252C0E5043" bold="true" box="[240,380,1058,1082]" pageId="14" pageNumber="13">
<typeStatus id="9C2A4CF49160FFB16C92BD252C0E5043" box="[240,380,1058,1082]" pageId="14" pageNumber="13" type="paratype">Paratypes</typeStatus>
</emphasis>
(n=5).
<emphasis id="71E52E449160FFB16DACBD2529555040" bold="true" box="[462,1063,1057,1082]" pageId="14" pageNumber="13">
<collectingCountry id="3B86B2C69160FFB16DACBD252FCF5040" box="[462,701,1057,1082]" name="United States of America" pageId="14" pageNumber="13">UNITED STATES</collectingCountry>
:
<collectingRegion id="81553CB49160FFB16EAABD262EF55040" box="[712,903,1057,1081]" country="United States of America" name="California" pageId="14" pageNumber="13">CALIFORNIA</collectingRegion>
:
<collectingCounty id="AA4F8ADA9160FFB16FF3BD2629535040" box="[913,1057,1057,1081]" pageId="14" pageNumber="13">Shasta Co.</collectingCounty>
:
</emphasis>
<collectionCode id="25806A939160FFB1684EBD25294D5043" box="[1068,1087,1058,1082]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="13">T</collectionCode>
.
<collectionCode id="25806A939160FFB16821BD25292A5043" box="[1091,1112,1058,1082]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="13">R</collectionCode>
.Haig Whiskeytown Shasta Co. Cal. III.9.1973 (
<collectionCode id="25806A939160FFB16DCEBD452C8A5023" box="[428,504,1090,1114]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="14" pageNumber="13" type="Museum">CSCA</collectionCode>
<specimenCount id="559739DF9160FFB16D9CBD452F535023" box="[510,545,1089,1114]" pageId="14" pageNumber="13" type="male">4♂</specimenCount>
, one aedeagus only, one
<collectionCode id="25806A939160FFB16F28BD452ED65023" box="[842,932,1090,1114]" pageId="14" pageNumber="13">SLIDE</collectionCode>
). *
<collectionCode id="25806A939160FFB16FA3BD452EA75023" box="[961,981,1090,1114]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="13">T</collectionCode>
.
<collectionCode id="25806A939160FFB16FBBBD452E9D5023" box="[985,1007,1090,1114]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="13">R</collectionCode>
.Haig Whiskeytown Shasta Co. Cal.
<date id="372FD4969160FFB16CA2BD652C4B5003" box="[192,313,1122,1146]" pageId="14" pageNumber="13" value="1973-01-31">1-31-1973</date>
(
<collectionCode id="25806A939160FFB16D2BBD652CD05003" box="[329,418,1122,1146]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="14" pageNumber="13" type="Museum">FMNH</collectionCode>
) (
<specimenCount id="559739DF9160FFB16DDEBD652CAF5003" box="[444,477,1121,1146]" pageId="14" pageNumber="13" type="male">1♂</specimenCount>
).
</paragraph>
<paragraph id="432EF2569160FFB16CA2BDA62C3650A3" blockId="14.[192,1441,1185,1242]" pageId="14" pageNumber="13">
<emphasis id="71E52E449160FFB16CA2BDA62F4850C0" bold="true" box="[192,570,1185,1209]" pageId="14" pageNumber="13">Geographical Distribution.</emphasis>
<taxonomicName id="849189D59160FFB16E26BDA52E7C50C3" authorityName="Ferro" authorityYear="2016" box="[580,782,1186,1210]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="colberti">
<emphasis id="71E52E449160FFB16E26BDA52E7C50C3" box="[580,782,1186,1210]" italics="true" pageId="14" pageNumber="13">Sonoma colberti</emphasis>
</taxonomicName>
is only known from one location in Shasta County,
<collectingRegion id="81553CB49160FFB16CA2BDC52C3250A3" box="[192,320,1218,1242]" country="United States of America" name="California" pageId="14" pageNumber="13">California</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="0B8BA1DD9160FFB16CA2BC062F485263" pageId="14" pageNumber="13" type="discussion">
<paragraph id="432EF2569160FFB16CA2BC06295C5163" blockId="14.[192,1440,1281,1562]" box="[192,1070,1281,1306]" pageId="14" pageNumber="13">
<emphasis id="71E52E449160FFB16CA2BC062C295160" bold="true" box="[192,347,1281,1305]" pageId="14" pageNumber="13">Comments.</emphasis>
<taxonomicName id="849189D59160FFB16D00BC052F5A5163" authorityName="Ferro" authorityYear="2016" box="[354,552,1282,1306]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="colberti">
<emphasis id="71E52E449160FFB16D00BC052F5A5163" box="[354,552,1282,1306]" italics="true" pageId="14" pageNumber="13">Sonoma colberti</emphasis>
</taxonomicName>
was collected during January and March.
</paragraph>
<paragraph id="432EF2569160FFB16C92BC252F485263" blockId="14.[192,1440,1281,1562]" pageId="14" pageNumber="13">
Aedeagal characters of
<taxonomicName id="849189D59160FFB16E7EBC252FEE5143" authorityName="Ferro" authorityYear="2016" box="[540,668,1314,1338]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="colberti">
<emphasis id="71E52E449160FFB16E7EBC252FEE5143" box="[540,668,1314,1338]" italics="true" pageId="14" pageNumber="13">S. colberti</emphasis>
</taxonomicName>
are superficially similar to
<taxonomicName id="849189D59160FFB16867BC2529C15140" authorityName="Chandler. Modified" authorityYear="2003" box="[1029,1203,1314,1337]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="wintuorum">
<emphasis id="71E52E449160FFB16867BC2529C15140" box="[1029,1203,1314,1337]" italics="true" pageId="14" pageNumber="13">S. wintuorum</emphasis>
</taxonomicName>
,
<taxonomicName id="849189D59160FFB168A4BC2528485140" authorityName="Ferro" authorityYear="2016" box="[1222,1338,1314,1337]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="twaini">
<emphasis id="71E52E449160FFB168A4BC2528485140" box="[1222,1338,1314,1337]" italics="true" pageId="14" pageNumber="13">S. twaini</emphasis>
</taxonomicName>
, and
<taxonomicName id="849189D59160FFB169E5BC252C515120" authorityName="Ferro" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="stewarti">
<emphasis id="71E52E449160FFB169E5BC252C515120" italics="true" pageId="14" pageNumber="13">S. stewarti</emphasis>
</taxonomicName>
all of which share a similar gestalt comprised of narrow, elongate parameres and an elongate endophallus with elaborate apical hooks or processes. The folowing combination of characters will separate
<taxonomicName id="849189D59160FFB16D57BC852CC651E3" authorityName="Ferro" authorityYear="2016" box="[309,436,1410,1434]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="colberti">
<emphasis id="71E52E449160FFB16D57BC852CC651E3" box="[309,436,1410,1434]" italics="true" pageId="14" pageNumber="13">S. colberti</emphasis>
</taxonomicName>
from the others: endophallus with three projections [
<taxonomicName id="849189D59160FFB16803BC8529A651E0" authorityName="Ferro" authorityYear="2016" box="[1121,1236,1410,1433]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="twaini">
<emphasis id="71E52E449160FFB16803BC8529A651E0" box="[1121,1236,1410,1433]" italics="true" pageId="14" pageNumber="13">S. twaini</emphasis>
</taxonomicName>
and
<taxonomicName id="849189D59160FFB1697BBC8528D251E0" authorityName="Ferro" authorityYear="2016" box="[1305,1440,1410,1433]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="stewarti">
<emphasis id="71E52E449160FFB1697BBC8528D251E0" box="[1305,1440,1410,1433]" italics="true" pageId="14" pageNumber="13">S. stewarti</emphasis>
</taxonomicName>
with one,
<taxonomicName id="849189D59160FFB16D5ABCA52C9351C0" authorityName="Chandler. Modified" authorityYear="2003" box="[312,481,1442,1465]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="wintuorum">
<emphasis id="71E52E449160FFB16D5ABCA52C9351C0" box="[312,481,1442,1465]" italics="true" pageId="14" pageNumber="13">S. wintuorum</emphasis>
</taxonomicName>
with two]; subapical lateral projection of endophallus directed posteriorly and short, approximately 1/10 length of endophallus [
<taxonomicName id="849189D59160FFB16F79BCC52EEC51A0" authorityName="Ferro" authorityYear="2016" box="[795,926,1474,1497]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="stewarti">
<emphasis id="71E52E449160FFB16F79BCC52EEC51A0" box="[795,926,1474,1497]" italics="true" pageId="14" pageNumber="13">S. stewarti</emphasis>
</taxonomicName>
with projection 1/5 length of endophallus,
<taxonomicName id="849189D59160FFB16CA2BCE52C195180" authorityName="Chandler. Modified" authorityYear="2003" box="[192,363,1506,1529]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="wintuorum">
<emphasis id="71E52E449160FFB16CA2BCE52C195180" box="[192,363,1506,1529]" italics="true" pageId="14" pageNumber="13">S. wintuorum</emphasis>
</taxonomicName>
with projection greater than 1/3 length of endophallus,
<taxonomicName id="849189D59160FFB1684BBCE529E85180" authorityName="Ferro" authorityYear="2016" box="[1065,1178,1506,1529]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="twaini">
<emphasis id="71E52E449160FFB1684BBCE529E85180" box="[1065,1178,1506,1529]" italics="true" pageId="14" pageNumber="13">S. twaini</emphasis>
</taxonomicName>
with two projections perpendicular to endophallus].
</paragraph>
</subSubSection>
<subSubSection id="0B8BA1DD9160FFB16CA2BF4528D35203" pageId="14" pageNumber="13" type="etymology">
<paragraph id="432EF2569160FFB16CA2BF4528D35203" blockId="14.[192,1441,1602,1658]" pageId="14" pageNumber="13">
<emphasis id="71E52E449160FFB16CA2BF452C2E5223" bold="true" box="[192,348,1602,1626]" pageId="14" pageNumber="13">Etymology.</emphasis>
<taxonomicName id="849189D59160FFB16D01BF452F5B5223" authorityName="Ferro" authorityYear="2016" box="[355,553,1602,1626]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="colberti">
<emphasis id="71E52E449160FFB16D01BF452F5B5223" box="[355,553,1602,1626]" italics="true" pageId="14" pageNumber="13">Sonoma colberti</emphasis>
</taxonomicName>
is named for Stephen Colbert (silent “t”) a character created by Stephen Tyrone Colbert (sounded “t”), an author, philosopher, humanitarian, champion of science, and humorist.
</paragraph>
</subSubSection>
</treatment>
</document>