416 lines
31 KiB
XML
416 lines
31 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.410.6955" ID-GBIF-Dataset="c00cebd9-2243-47b2-a4e1-5c61f05e4126" ID-PMC="PMC4042707" ID-Pensoft-Pub="1313-2970-410-41" ID-PubMed="24899849" ID-ZBK="501921E5E350470FA4726D65E737B133" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-410-41" ModsDocOrigin="ZooKeys 410" ModsDocTitle="Calcaridorylaimus castaneae sp. n. (Nematoda, Dorylaimidae) from Bulgaria with an identification key to the species of the genus" checkinTime="1451245874272" checkinUser="pensoft" docAuthor="Nedelchev, Sevdan, Elshishka, Milka, Lazarova, Stela, Radoslavov, Georgi, Hristov, Peter & Peneva, Vlada" docDate="2014" docId="C2BEF1EC124001E34AC1F1B8BF86F888" docLanguage="en" docName="ZooKeys 410: 41-61" docOrigin="ZooKeys 410" docSource="http://dx.doi.org/10.3897/zookeys.410.6955" docTitle="Calcaridorylaimus castaneae Nedelchev, Elshishka, Lazarova, Radoslavov, Hristov & Peneva, 2014, sp. n." docType="treatment" docUuid="9BF1D302-1986-47C5-B0D8-2F5DC75BD6D2" docUuidSource="ZooBank" docVersion="4" lastPageNumber="46" masterDocId="FFC7FB640361FFC48766EA3E113EFF85" masterDocTitle="Calcaridorylaimus castaneae sp. n. (Nematoda, Dorylaimidae) from Bulgaria with an identification key to the species of the genus" masterLastPageNumber="61" masterPageNumber="41" pageNumber="44" updateTime="1668158520999" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Calcaridorylaimus castaneae sp. n. (Nematoda, Dorylaimidae) from Bulgaria with an identification key to the species of the genus</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nedelchev, Sevdan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Elshishka, Milka</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lazarova, Stela</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Radoslavov, Georgi</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hristov, Peter</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Peneva, Vlada</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>410</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>41</mods:start>
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<mods:end>61</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.410.6955</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.410.6955</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-410-41</mods:identifier>
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<mods:identifier type="ZBK">501921E5E350470FA4726D65E737B133</mods:identifier>
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<mods:identifier type="ZooBank">501921E5E350470FA4726D65E737B133</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152053209" LSID="urn:lsid:zoobank.org:act:9BF1D302-1986-47C5-B0D8-2F5DC75BD6D2" httpUri="http://treatment.plazi.org/id/C2BEF1EC124001E34AC1F1B8BF86F888" lastPageId="5" lastPageNumber="46" pageId="3" pageNumber="44">
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<subSubSection pageId="3" pageNumber="44" type="nomenclature">
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<paragraph pageId="3" pageNumber="44">
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<taxonomicName LSID="http://zoobank.org/9BF1D302-1986-47C5-B0D8-2F5DC75BD6D2" class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus castaneae" order="Dorylaimida" pageId="3" pageNumber="44" phylum="Nematoda" rank="species" species="castaneae">Calcaridorylaimus castaneae</taxonomicName>
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<taxonomicNameLabel pageId="3" pageNumber="44">sp. n.</taxonomicNameLabel>
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Figs 2-6
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="44" type="measurements">
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<paragraph pageId="3" pageNumber="44">Measurements.</paragraph>
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<paragraph pageId="3" pageNumber="44">See Table 1.</paragraph>
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<caption pageId="3" pageNumber="44">
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<paragraph pageId="3" pageNumber="44">
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Table 1. Morphometrics of
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<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus castaneae" order="Dorylaimida" pageId="3" pageNumber="44" phylum="Nematoda" rank="species" species="castaneae">Calcaridorylaimus castaneae</taxonomicName>
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sp. n., females and males, from Belasitsa Mountain. All measurements, unless indicated otherwise, in
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<normalizedToken originalValue="μm">μm</normalizedToken>
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and in the form: mean
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<normalizedToken originalValue="±">+/-</normalizedToken>
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SD (range).
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</paragraph>
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</caption>
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<paragraph pageId="3" pageNumber="44">
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<table pageId="3" pageNumber="44">
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<tr pageId="3" pageNumber="44">
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">Habitat</th>
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<th colspan="5" pageId="3" pageNumber="44" rowspan="1">Mixed broadleaf forest</th>
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<th colspan="2" pageId="3" pageNumber="44" rowspan="1">Chestnut forest</th>
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</tr>
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<tr pageId="3" pageNumber="44">
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">Locality and year of collection</th>
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<th colspan="3" pageId="3" pageNumber="44" rowspan="1">140b 2003</th>
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<th colspan="2" pageId="3" pageNumber="44" rowspan="1">140b 2012</th>
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<th colspan="2" pageId="3" pageNumber="44" rowspan="1">104g 2003</th>
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</tr>
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<tr pageId="3" pageNumber="44">
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">Characters</th>
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">Holotype</th>
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<th colspan="6" pageId="3" pageNumber="44" rowspan="1">Paratypes</th>
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</tr>
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<tr pageId="3" pageNumber="44">
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">Females</th>
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">Males</th>
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">Females</th>
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">Males</th>
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">Females</th>
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">Males</th>
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</tr>
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<tr pageId="3" pageNumber="44">
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">n</th>
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">40</th>
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">20</th>
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">10</th>
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">10</th>
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">18</th>
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<th colspan="1" pageId="3" pageNumber="44" rowspan="1">12</th>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="46" pageId="3" pageNumber="44" type="description">
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<paragraph pageId="3" pageNumber="44">Description.</paragraph>
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<paragraph pageId="3" pageNumber="44">
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Female. Body slender, more or less curved ventrally. Cuticle ca 2
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<normalizedToken originalValue="μm">μm</normalizedToken>
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at anterior part of neck, 3
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<normalizedToken originalValue="μm">μm</normalizedToken>
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thick at midbody, 4.5-5.5
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<normalizedToken originalValue="μm">μm</normalizedToken>
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at postanal region; outer layer with fine transverse striae. Lateral chord ca 1/4 of body width. Three dorsal, two ventral and three lateral pores are observed in the spear area. Lip region practically not offset, lip region width ca 20% of its height. Lips partly fused, labial papilae slightly protruding. Body at proximal end of pharynx 3-4 times the width of the lip region diameter. Amphidial aperture 5-6
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<normalizedToken originalValue="μm">μm</normalizedToken>
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wide or about half the lip region width. Odontostyle 1.2-1.3 times the lip region diameter, aperture occupying 35-40% of its length. Odontophore simple 1.3-1.7 times odontostyle length. Guiding ring at 8.5-9
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<normalizedToken originalValue="μm">μm</normalizedToken>
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from anterior end. Nerve ring surrounding the pharynx at 36-39% of neck length from head end. Hemizonid and conspicuous excretory pore observed in the nerve ring region. Pharyngeal characters (five females and five males): pharynx beginning to widen at 56-60% and attaining its full width at 61-64% of neck length from anterior end. DO (for terminology see
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<bibRefCitation pageId="3" pageNumber="44">Loof and Coomans 1970</bibRefCitation>
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) lying near the point where the pharynx attains its full width;
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<normalizedToken originalValue="DO–DN">DO-DN</normalizedToken>
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8-13
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<normalizedToken originalValue="μm">μm</normalizedToken>
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. The two S1N lying at a small distance behind the middle of the distance
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<normalizedToken originalValue="DN–S">DN-S</normalizedToken>
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2N, the anterior one (S1N1) smaller, ca 2
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<normalizedToken originalValue="μm">μm</normalizedToken>
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diam; S1N2 comparatively large and distinct, 3-4
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<normalizedToken originalValue="μm">μm</normalizedToken>
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diam. DN nucleolus 4-4.5
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<normalizedToken originalValue="μm">μm</normalizedToken>
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diam.; S2N 2-3
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<normalizedToken originalValue="μm">μm</normalizedToken>
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diam. (DN>S1N2>S2N). Locations (%):
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</paragraph>
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<paragraph pageId="3" pageNumber="44">
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<table pageId="3" pageNumber="44">
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<tr pageId="3" pageNumber="44">
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<td colspan="1" pageId="3" pageNumber="44" rowspan="1">11</td>
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<td colspan="1" pageId="3" pageNumber="44" rowspan="1">2</td>
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</tr>
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<tr pageId="3" pageNumber="44">
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<td colspan="1" pageId="3" pageNumber="44" rowspan="1">12</td>
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<td colspan="1" pageId="3" pageNumber="44" rowspan="1">2</td>
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</tr>
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<tr pageId="3" pageNumber="44">
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<td colspan="1" pageId="3" pageNumber="44" rowspan="1">1112</td>
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</tr>
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</table>
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</paragraph>
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<paragraph pageId="4" pageNumber="45">
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<pageBreakToken pageId="4" pageNumber="45" start="start">Cardia</pageBreakToken>
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conoid, variable in length, microvilli visible only here. Genital system didelphic-amphidelphic, ovaries reflexed, reaching rarely the vulva level. Oviducts and ovaries very long compared to uteri. Uteri not differentiated, short, anterior 96-125
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<normalizedToken originalValue="μm">μm</normalizedToken>
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and posterior 95-135
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<normalizedToken originalValue="μm">μm</normalizedToken>
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long, in one female with no sperm inside the genital system uteri shorter, 83
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<normalizedToken originalValue="μm">μm</normalizedToken>
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each, sphincter between uterus and pars dilatata oviductus well developed. Sperm present in uteri and pars dilatata oviductus. Synchronous uterine eggs 1-3, measuring 72-78
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<normalizedToken originalValue="×">x</normalizedToken>
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32-44
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<normalizedToken originalValue="μm">μm</normalizedToken>
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. Vulva transverse. Vagina extending to 60-70% inwards: pars proximalis 9-13
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<normalizedToken originalValue="μm">μm</normalizedToken>
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wide, 14.5-19.5
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<normalizedToken originalValue="μm">μm</normalizedToken>
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long, pars refringens more or less rounded trapezoid, 10-12.5
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<normalizedToken originalValue="μm">μm</normalizedToken>
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wide, 4-6
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<normalizedToken originalValue="μm">μm</normalizedToken>
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deep; pars distalis approx. 1.5
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(terminology following
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<bibRefCitation author="De Ley, P" journalOrPublisher="Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Biologie" pageId="9" pageNumber="50" pagination="13 - 34" title="Terrestrial nematodes from the Galapagos Archipelago 2: Redescription of Aporcelaimellus obtusicaudatus (Bastian, 1865) Altherr, 1968, with review of similar species and a nomenclature for the vagina in Dorylaimida (Nematoda)." volume="63" year="1993">De Ley et al. 1993</bibRefCitation>
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). Peculiar tongue-like valve present at intestine-prerectum junction. Rectum 1.3-1.5, prerectum 3-4 times anal body width long, respectively. Tail first conoid, then more or less uniformly tapering to a narrowly rounded terminus. Posterior part of tail usually slightly curved dorsally. Two pairs of caudal pores, one subventral, other subdorsal.
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</paragraph>
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<paragraph pageId="5" pageNumber="46">
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<pageBreakToken pageId="5" pageNumber="46" start="start">Male</pageBreakToken>
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. General morphology similar to that of female, body curved ventrally in J-shape when fixed. Genital system diorchic, testes opposed, well developed. Spicules dorylaimoid, with double contour on dorsal arm, 1.3-1.6 times the corresponding body diameter long; ventral arm smaller than dorsal. A spur present dorsally before the distal tip, distinctly visible in extruded spicules. Lateral guiding pieces 9-11
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<normalizedToken originalValue="μm">μm</normalizedToken>
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long or ca. 23 % spicule length. In addition to adcloacal pair seven to twelve (mostly nine or ten), regularly spaced ventromedian supplements present (9 supplements in 8 specimens; 10 suppl. - in 7, and 7, 8 and 12 each in one specimen). Prerectum 4.5-7.5 times the corresponding body diameter long, extending 0.7-1.8 body widths anterior to the supplement series. Tail dorsally conoid and broadly rounded. One subdorsal and one subterminal pair of caudal pores.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="46" type="differential diagnosis and relationships">
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<paragraph pageId="5" pageNumber="46">Differential diagnosis and relationships.</paragraph>
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<paragraph pageId="5" pageNumber="46">
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<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus castaneae" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="castaneae">Calcaridorylaimus castaneae</taxonomicName>
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sp. n. differs from all species in the genus by a combination of the following characters: long body (1.4-2.1 mm), lip region practically not offset, short odontostyle (14.5-16
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<normalizedToken originalValue="μm">μm</normalizedToken>
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in females and 14-16
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<normalizedToken originalValue="μm">μm</normalizedToken>
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in males) and short female tail (75.5-110.5
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, c=14.7-23.6;
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<normalizedToken originalValue="c’=2.9–">c'=2.9-</normalizedToken>
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4.4); vulva transverse, 38-46
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<normalizedToken originalValue="μm">μm</normalizedToken>
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long spicules with spur before its distal end. The new species is most similar to
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<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus andrassyi" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="andrassyi">Calcaridorylaimus andrassyi</taxonomicName>
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from which it can be differentiated by having transverse vulva vs pore-like and shorter spicules (38-46
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<normalizedToken originalValue="μm">μm</normalizedToken>
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vs 52-57
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<normalizedToken originalValue="μm">μm</normalizedToken>
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). Further,
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<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus castaneae" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="castaneae">Calcaridorylaimus castaneae</taxonomicName>
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differs from
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<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus ruwenzorii" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="ruwenzorii">Calcaridorylaimus ruwenzorii</taxonomicName>
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by having shorter odontostyle (14.5-16
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<normalizedToken originalValue="μm">μm</normalizedToken>
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vs 19.5-25
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<normalizedToken originalValue="μm">μm</normalizedToken>
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), and tail (75.5-110.5
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<normalizedToken originalValue="μm">μm</normalizedToken>
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vs 160
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, higher c (c=14.7-23.6 vs c=10) and lower
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<normalizedToken originalValue="c’">c'</normalizedToken>
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(
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<normalizedToken originalValue="c’=2.9–">c'=2.9-</normalizedToken>
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4.4 vs 7) values in females. It can be differentiated from
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<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus arcticus" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="arcticus">Calcaridorylaimus arcticus</taxonomicName>
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,
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<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus beatus" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="beatus">Calcaridorylaimus beatus</taxonomicName>
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,
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<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus calcarifer" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="calcarifer">Calcaridorylaimus calcarifer</taxonomicName>
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and
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<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus signatus" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="signatus">Calcaridorylaimus signatus</taxonomicName>
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by having different vulva shape (transverse vs longitudinal) and shorter spicules (38-46
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<normalizedToken originalValue="μm">μm</normalizedToken>
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vs 57-67
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<normalizedToken originalValue="μm">μm</normalizedToken>
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; 48-55
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<normalizedToken originalValue="μm">μm</normalizedToken>
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; 52-54
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<normalizedToken originalValue="μm">μm</normalizedToken>
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and 72
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<normalizedToken originalValue="μm">μm</normalizedToken>
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). From
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<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus sirgeli" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="sirgeli">Calcaridorylaimus sirgeli</taxonomicName>
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it differs by having transverse vulva vs pore-like, higher c (c=14.7-23.6 vs c=9.7-11.3) and lower
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<normalizedToken originalValue="c’">c'</normalizedToken>
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(
|
||
<normalizedToken originalValue="c’=2.9–">c'=2.9-</normalizedToken>
|
||
4.4 vs
|
||
<normalizedToken originalValue="c’=5.3–">c'=5.3-</normalizedToken>
|
||
6.7) values. Finally,
|
||
<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus castaneae" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="castaneae">Calcaridorylaimus castaneae</taxonomicName>
|
||
differs from
|
||
<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus promissus" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="promissus">Calcaridorylaimus promissus</taxonomicName>
|
||
and
|
||
<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus simillimus" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="simillimus">Calcaridorylaimus simillimus</taxonomicName>
|
||
by having longer odontostyle (14.5-16
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
vs 13
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
and 11
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
) and shorter tail (75.5-110.5
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
vs 158-178
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
and 175
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="5" pageNumber="46">
|
||
Presence of the conspicuous excretory pore observed in
|
||
<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus castaneae" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="castaneae">Calcaridorylaimus castaneae</taxonomicName>
|
||
is unusual for members of Dorylaimida, however, similar structure has been mentioned only for two
|
||
<taxonomicName class="Adenophorea" family="Longidoridae" genus="Longidorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Longidorus" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="genus">Longidorus</taxonomicName>
|
||
species, namely
|
||
<taxonomicName class="Adenophorea" family="Longidoridae" genus="Longidorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Longidorus macrosoma" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="macrosoma">Longidorus macrosoma</taxonomicName>
|
||
Hooper, 1961 and
|
||
<taxonomicName class="Adenophorea" family="Longidoridae" genus="Longidorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Longidorus carniolensis" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="carniolensis">Longidorus carniolensis</taxonomicName>
|
||
<normalizedToken originalValue="Širca">Sirca</normalizedToken>
|
||
et al., 2011 (
|
||
<bibRefCitation author="Aboul-Eid, HZ" journalOrPublisher="Nematologica" pageId="9" pageNumber="50" pagination="437 - 450" title="Histological anatomy of the excretory and reproductive systems of Longidorus macrosoma." url="10.1163/187529269X00777" volume="15" year="1969">Aboul-Eid 1969</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Sirca, S" journalOrPublisher="ZooKeys" pageId="10" pageNumber="51" pagination="1 - 27" title="Longidorus carniolensis sp. n. (Nematoda, Longidoridae) from vineyard soil in Slovenia." url="10.3897/zookeys.141.1906" volume="141" year="2011">
|
||
<normalizedToken originalValue="Širca">Sirca</normalizedToken>
|
||
et al. 2011
|
||
</bibRefCitation>
|
||
) and for several
|
||
<taxonomicName class="Adenophorea" family="Dorylaimidae" genus="Mesodorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesodorylaimus" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="genus">Mesodorylaimus</taxonomicName>
|
||
species origination from Antarctica -
|
||
<taxonomicName class="Adenophorea" family="Dorylaimidae" genus="Mesodorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesodorylaimus chipevi" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="chipevi">Mesodorylaimus chipevi</taxonomicName>
|
||
Nedelchev & Peneva, 2000,
|
||
<taxonomicName class="Adenophorea" family="Dorylaimidae" genus="Mesodorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesodorylaimus antarcticus" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="antarcticus">Mesodorylaimus antarcticus</taxonomicName>
|
||
Nedelchev & Peneva, 2000,
|
||
<taxonomicName class="Adenophorea" family="Dorylaimidae" genus="Mesodorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesodorylaimus masleni" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="masleni">Mesodorylaimus masleni</taxonomicName>
|
||
Nedelchev & Peneva, 2000,
|
||
<taxonomicName class="Adenophorea" family="Dorylaimidae" genus="Mesodorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesodorylaimus imperator" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="imperator">Mesodorylaimus imperator</taxonomicName>
|
||
Loof, 1975 (
|
||
<bibRefCitation author="Nedelchev, S" journalOrPublisher="Russian Journal of Nematology" pageId="10" pageNumber="51" pagination="161 - 172" title="Description of three new species of the genus Mesodorylaimus Andrassy, 1959 (Nematoda: Dorylaimidae) from Livingston Island, Antarctica, with notes on M. imperator Loof, 1975." volume="8" year="2000">Nedelchev and Peneva 2000</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="46" type="type locality and plant association">
|
||
<paragraph pageId="5" pageNumber="46">Type locality and plant association.</paragraph>
|
||
<paragraph pageId="5" pageNumber="46">
|
||
Belasitsa Mountain, south-western Bulgaria, litter from old broadleaf forest (100-140 years) dominated by
|
||
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Castanea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Castanea sativa" order="Fagales" pageId="5" pageNumber="46" phylum="Tracheophyta" rank="species" species="sativa">Castanea sativa</taxonomicName>
|
||
, mixed with
|
||
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus daleshampii" order="Fagales" pageId="5" pageNumber="46" phylum="Tracheophyta" rank="species" species="daleshampii">Quercus daleshampii</taxonomicName>
|
||
and
|
||
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus sylvatica" order="Fagales" pageId="5" pageNumber="46" phylum="Tracheophyta" rank="species" species="sylvatica">Fagus sylvatica</taxonomicName>
|
||
. Site is located in the vicinity of Belasitsa hut,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="41.37">41°22'12"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="15" value="23.186666">23°11'12"E</geoCoordinate>
|
||
(sub-compartment 140b). Second locality: young sweet chestnut forest (30-40 years old) near Belasitsa village (sub-compartment 104g).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="46" type="type material">
|
||
<paragraph pageId="5" pageNumber="46">Type material.</paragraph>
|
||
<paragraph pageId="5" pageNumber="46">Holotype and 80 paratype females and 47 males deposited in the nematode collection of the Institute of Biodiversity and Ecosystem Research, Sofia, Bulgaria. Other paratypes deposited as follows: four females and two males in the Nematode Collection of the Foodland Environment Research Agency, Sand Hutton, UK (former Rothamsted Nematode Collection); three females and three males in the USDA Nematode Collection, Beltsville, Maryland, USA; two females and two males in the Riverside Nematode Collection, University of California, Riverside, USA; four females, and four males in the Wageningen Nematode Collection (WANECO), Wageningen, the Netherlands; four females and three males in the Nematode Collection of the Zoology Museum of the Ghent University, Belgium.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="46" type="etymology">
|
||
<paragraph pageId="5" pageNumber="46">Etymology.</paragraph>
|
||
<paragraph pageId="5" pageNumber="46">
|
||
The scientific name is derived from the generic name of dominant tree species, the sweet chestnut tree (
|
||
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Castanea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Castanea" order="Fagales" pageId="5" pageNumber="46" phylum="Tracheophyta" rank="genus">Castanea</taxonomicName>
|
||
) in the forest where this nematode was found.
|
||
</paragraph>
|
||
<caption pageId="5" pageNumber="46">
|
||
<paragraph pageId="5" pageNumber="46">
|
||
Figure 2.
|
||
<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus castaneae" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="castaneae">Calcaridorylaimus castaneae</taxonomicName>
|
||
sp. n. Female: A Pharyngeal gland nuclei B Anterior region C Pharyngeal region D Genital system E Vulval region F Sperm cells in uterus. Scale bars: A, B, E, F - 30
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
; C, D - 50
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="5" pageNumber="46">
|
||
<paragraph pageId="5" pageNumber="46">
|
||
Figure 3.
|
||
<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus castaneae" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="castaneae">Calcaridorylaimus castaneae</taxonomicName>
|
||
sp. n. Female: A Posterior region. Male: B, C Extruded spicules with supplements D Posterior region. Scale bar:
|
||
<normalizedToken originalValue="A–D">A-D</normalizedToken>
|
||
– 30
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="5" pageNumber="46">
|
||
<paragraph pageId="5" pageNumber="46">
|
||
Figure 4.
|
||
<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus castaneae" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="castaneae">Calcaridorylaimus castaneae</taxonomicName>
|
||
sp. n. Female: A Anterior end C Amphid
|
||
<normalizedToken originalValue="F–I">F-I</normalizedToken>
|
||
Tail shapes MaleB Anterior end D Posterior end with extruded spicules, arrow indicating the spur E Posterior end. Scale bars: A, B,
|
||
<normalizedToken originalValue="D–I">D-I</normalizedToken>
|
||
– 20
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
; C - 6
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="5" pageNumber="46">
|
||
<paragraph pageId="5" pageNumber="46">
|
||
Figure 5.
|
||
<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus castaneae" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="castaneae">Calcaridorylaimus castaneae</taxonomicName>
|
||
sp. n. Female: A Entire body B Lip region D Prerectum, arrow pointing tongue-like valve E Pharyngeal bulb F Vulval region with posterior uterus G Vulval region with egg in posterior uterus H, I Vulval region J Cardia L Lateral field. Male: A Entire body C Lip region K Cardia M Supplements. Scale bars: A - 200
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
; B, C, M - 6
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
;
|
||
<normalizedToken originalValue="E–L">E-L</normalizedToken>
|
||
– 20
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="5" pageNumber="46">
|
||
<paragraph pageId="5" pageNumber="46">
|
||
Figure 6.
|
||
<taxonomicName class="Enoplea" family="Dorylaimidae" genus="Calcaridorylaimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calcaridorylaimus castaneae" order="Dorylaimida" pageId="5" pageNumber="46" phylum="Nematoda" rank="species" species="castaneae">Calcaridorylaimus castaneae</taxonomicName>
|
||
sp. n. Male: A Posterior end B Sperm cells in testis
|
||
<normalizedToken originalValue="C–F">C-F</normalizedToken>
|
||
Spicular region C Lateral piece of spicules D, E Extruded spicules, arrows pointing the spur F Spicules in the body. Scale bars: A, B - 20
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
;
|
||
<normalizedToken originalValue="C–F">C-F</normalizedToken>
|
||
– 18
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |