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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFEBFFA95AB6F8DF4586CEF8" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus sapaensis V. S. Dang &amp; Naiki 2019" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571866" ID-Zenodo-Dep="7571866" LSID="urn:lsid:plazi:treatment:43284262FFEBFFA95AB6F8DF4586CEF8" httpUri="http://treatment.plazi.org/id/43284262FFEBFFA95AB6F8DF4586CEF8" lastPageId="119" pageId="116">
<subSubSection box="[136,1116,1913,1940]" pageId="116" type="nomenclature">
<paragraph blockId="116.[136,1452,1913,2048]" box="[136,1116,1913,1940]" pageId="116">
<emphasis bold="true" box="[136,171,1914,1940]" pageId="116">59.</emphasis>
<taxonomicName ID-CoL="8Y54Z" authority="V. S. Dang &amp; Naiki (2019: 39)" authorityName="V. S. Dang &amp; Naiki" authorityPageNumber="39" authorityYear="2019" box="[178,761,1913,1940]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="116" phylum="Tracheophyta" rank="species" species="sapaensis">
<emphasis bold="true" box="[178,424,1913,1940]" italics="true" pageId="116">Lasianthus sapaensis</emphasis>
V.S.
<bibRefCitation author="Dang, V. S. &amp; Naiki, A." box="[474,761,1914,1940]" pageId="116" pagination="33 - 40" refId="ref54157" refString="Dang, V. S. &amp; Naiki, A. (2019) Two new species of Lasianthus (Rubiaceae) from Hoang Lien National Park, northern Vietnam. Phytotaxa 423 (1): 33 - 40. https: // doi. org / 10.11646 / phytotaxa. 423.1.4" type="journal article" year="2019">Dang &amp; Naiki (2019: 39)</bibRefCitation>
</taxonomicName>
<emphasis bold="true" box="[761,789,1914,1940]" pageId="116">—</emphasis>
<figureCitation box="[789,868,1914,1940]" captionStart="FIGURE 88" captionStartId="117.[136,229,1763,1785]" captionTargetBox="[170,1414,194,1735]" captionTargetId="figure-18@117.[166,1418,193,1739]" captionTargetPageId="117" captionText="FIGURE 88. Lasianthus sapaensis V.S.Dang &amp; Naiki (from Dang 230, VNM and the herbarium of Iriomote Station (Japan)): A. Branch, B. Close-up of young branch and leaves, C. Stipules, D. Abaxial leaf surfaces, E–F. Inflorescence showing side view calyx, G. Mature fruit, H. Seeds." figureDoi="http://doi.org/10.5281/zenodo.7571824" httpUri="https://zenodo.org/record/7571824/files/figure.png" pageId="116">Fig. 88</figureCitation>
(plate),
<figureCitation box="[962,1042,1914,1940]" captionStart="FIGURE 89" captionStartId="118.[136,229,1814,1836]" captionTargetBox="[176,1405,203,1775]" captionTargetId="figure-24@118.[165,1418,193,1790]" captionTargetPageId="118" captionText="FIGURE 89. Distribution maps of Lasianthus sapaensis, L. saprosmoides, L. saxorum and L. schmidtii." figureDoi="http://doi.org/10.5281/zenodo.7571826" httpUri="https://zenodo.org/record/7571826/files/figure.png" pageId="116">Fig. 89</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection lastPageId="118" pageId="116" type="materials_examined">
<paragraph blockId="116.[136,1452,1913,2048]" pageId="116">
<emphasis bold="true" box="[136,194,1954,1975]" pageId="116">Type:</emphasis>
—
<materialsCitation collectingDate="2018-11-11" collectionCode="VNM, HN" collectorName="Dang V. S. &amp; Naiki A. &amp; Dang" country="Vietnam" county="Sa Pa District" elevation="1906" location="Fansipan" municipality="Hoang Lien National Park" pageId="116" specimenCode="VNM00042932, VNM00042933, VNM00042934" specimenCount="1" stateProvince="Lao Cai" typeStatus="holotype">
<collectingCountry box="[219,332,1954,1975]" name="Vietnam" pageId="116">VIETNAM</collectingCountry>
.
<collectingRegion box="[341,511,1954,1976]" country="Vietnam" name="Lao Cai" pageId="116">Lao Cai Province</collectingRegion>
,
<collectingCounty box="[521,655,1954,1976]" pageId="116">Sa Pa District</collectingCounty>
,
<collectingMunicipality box="[664,919,1954,1976]" pageId="116">Hoang Lien National Park</collectingMunicipality>
, on the way to
<location LSID="urn:lsid:plazi:treatment:43284262FFEBFFA95AB6F8DF4586CEF8:CE5EA5AFFFEBFFAA5E16F8074459CBF0" box="[1064,1151,1954,1976]" country="Vietnam" county="Sa Pa District" municipality="Hoang Lien National Park" name="Fansipan" pageId="116" stateProvince="Lao Cai">Fansipan</location>
peak from
<location LSID="urn:lsid:plazi:treatment:43284262FFEBFFA95AB6F8DF4586CEF8:CE5EA5AFFFEBFFAA5ED2F807457FCBF0" box="[1260,1369,1954,1976]" country="Vietnam" county="Sa Pa District" municipality="Hoang Lien National Park" name="Ton station" pageId="116" stateProvince="Lao Cai">Ton station</location>
,
<quantity box="[1379,1452,1953,1975]" metricMagnitude="3" metricUnit="m" metricValue="1.906" pageId="116" unit="m" value="1906.0">
<elevation box="[1379,1452,1953,1975]" metricMagnitude="3" metricUnit="m" metricValue="1.906" pageId="116" unit="m" value="1906.0">1906 m</elevation>
</quantity>
elev.,
<date box="[244,427,1990,2012]" pageId="116" value="2018-11-11">
<collectingDate box="[244,427,1990,2012]" pageId="116" value="2018-11-11">11 November 2018</collectingDate>
</date>
,
<emphasis box="[435,735,1990,2011]" italics="true" pageId="116">
<collectorName box="[435,529,1990,2011]" pageId="116">Dang V.S.</collectorName>
&amp;
<collectorName box="[555,634,1990,2011]" pageId="116">Naiki A.</collectorName>
,
<collectorName box="[642,695,1990,2011]" pageId="116">Dang</collectorName>
230
</emphasis>
(
<typeStatus box="[746,832,1990,2012]" pageId="116">holotype</typeStatus>
<collectionCode box="[835,896,1990,2011]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="116" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[909,1069,1990,2011]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="116" type="Herbarium">VNM00042932</specimenCode>
]; isotypes
<collectionCode box="[1165,1206,1990,2011]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="116" type="Herbarium">HN</collectionCode>
!,
<collectionCode box="[1217,1278,1990,2011]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="116" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[1291,1448,1990,2011]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="116" type="Herbarium">VNM00042933</specimenCode>
,
<specimenCode box="[189,347,2026,2047]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="116" type="Herbarium">VNM00042934</specimenCode>
], the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFEBFFA95AB6F8DF4586CEF8:CE5EA5AFFFEBFFAA5831F84F4288C448" box="[527,686,2026,2048]" country="Vietnam" county="Sa Pa District" municipality="Hoang Lien National Park" name="Iriomote Station" pageId="116" stateProvince="Lao Cai">Iriomote Station</location>
, (Japan))
</materialsCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571824" ID-Zenodo-Dep="7571824" httpUri="https://zenodo.org/record/7571824/files/figure.png" pageId="117" startId="117.[136,229,1763,1785]" targetBox="[170,1414,194,1735]" targetPageId="117">
<paragraph blockId="117.[136,1452,1763,1857]" pageId="117">
<emphasis bold="true" box="[136,265,1763,1785]" pageId="117">FIGURE 88.</emphasis>
<taxonomicName authority="V. S. Dang &amp; Naiki" authorityName="V. S. Dang &amp; Naiki" box="[271,655,1763,1785]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="117" phylum="Tracheophyta" rank="species" species="sapaensis">
<emphasis box="[271,475,1763,1785]" italics="true" pageId="117">Lasianthus sapaensis</emphasis>
V.S.Dang &amp; Naiki
</taxonomicName>
(from
<emphasis box="[721,816,1763,1785]" italics="true" pageId="117">Dang 230</emphasis>
, VNM and the herbarium of Iriomote Station (Japan)):
<emphasis bold="true" box="[1347,1371,1763,1784]" pageId="117">A.</emphasis>
Branch,
<emphasis bold="true" box="[136,158,1799,1820]" pageId="117">B.</emphasis>
Close-up of young branch and leaves,
<emphasis bold="true" box="[540,564,1799,1821]" pageId="117">C.</emphasis>
Stipules,
<emphasis bold="true" box="[661,685,1799,1820]" pageId="117">D.</emphasis>
Abaxial leaf surfaces,
<emphasis bold="true" box="[908,954,1799,1820]" pageId="117">E–F.</emphasis>
Inflorescence showing side view calyx,
<emphasis bold="true" box="[1350,1375,1799,1821]" pageId="117">G.</emphasis>
Mature fruit,
<emphasis bold="true" box="[189,214,1835,1856]" pageId="117">H.</emphasis>
Seeds.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571826" ID-Zenodo-Dep="7571826" box="[136,1136,1814,1836]" httpUri="https://zenodo.org/record/7571826/files/figure.png" pageId="118" startId="118.[136,229,1814,1836]" targetBox="[176,1405,203,1775]" targetPageId="118">
<paragraph blockId="118.[136,1136,1814,1836]" box="[136,1136,1814,1836]" pageId="118">
<emphasis bold="true" box="[136,265,1814,1836]" pageId="118">FIGURE 89.</emphasis>
Distribution maps of
<taxonomicName box="[476,680,1814,1836]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="118" phylum="Tracheophyta" rank="species" species="sapaensis">
<emphasis box="[476,680,1814,1836]" italics="true" pageId="118">Lasianthus sapaensis</emphasis>
</taxonomicName>
,
<taxonomicName box="[693,849,1814,1836]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="118" phylum="Tracheophyta" rank="species" species="saprosmoides">
<emphasis box="[693,849,1814,1836]" italics="true" pageId="118">L. saprosmoides</emphasis>
</taxonomicName>
,
<taxonomicName box="[861,970,1815,1836]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="118" phylum="Tracheophyta" rank="species" species="saxorum">
<emphasis box="[861,970,1815,1836]" italics="true" pageId="118">L. saxorum</emphasis>
</taxonomicName>
and
<taxonomicName box="[1016,1132,1814,1836]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="118" phylum="Tracheophyta" rank="species" species="schmidtii">L. schmidtii</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="118.[136,1451,1919,2053]" pageId="118">
<emphasis bold="true" box="[136,398,1919,1945]" pageId="118">
<collectingRegion box="[136,262,1919,1945]" pageId="118">Specimens</collectingRegion>
examined:
</emphasis>
—
<materialsCitation collectingDate="2018-11-11" collectionCode="VNM" collectorName="Dang V. S &amp; Naiki A. &amp; Dang &amp; Japan" country="Vietnam" elevation="1906" location="Iriomote Station" municipality="Hoang Lien National Park" pageId="118" specimenCount="1" stateProvince="Lao Cai">
<collectingCountry box="[427,559,1919,1945]" name="Vietnam" pageId="118">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[573,694,1919,1945]" pageId="118">Northern:</emphasis>
<collectingRegion box="[706,913,1919,1945]" country="Vietnam" name="Lao Cai" pageId="118">Lao Cai Province</collectingRegion>
,
<collectingMunicipality box="[928,1238,1919,1945]" pageId="118">Hoang Lien National Park</collectingMunicipality>
,
<quantity box="[1254,1343,1919,1945]" metricMagnitude="3" metricUnit="m" metricValue="1.906" pageId="118" unit="m" value="1906.0">
<elevation box="[1254,1343,1919,1945]" metricMagnitude="3" metricUnit="m" metricValue="1.906" pageId="118" unit="m" value="1906.0">1906 m</elevation>
</quantity>
elev.,
<date pageId="118" value="2018-11-11">
<collectingDate pageId="118" value="2018-11-11">11 November 2018</collectingDate>
</date>
,
<emphasis box="[331,638,1955,1981]" italics="true" pageId="118">
<collectorName box="[331,435,1955,1981]" pageId="118">Dang V.S</collectorName>
&amp;
<collectorName box="[471,564,1955,1981]" pageId="118">Naiki A.</collectorName>
,
<collectorName box="[576,638,1956,1981]" pageId="118">Dang</collectorName>
</emphasis>
230a (
<collectionCode box="[714,784,1955,1981]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="118" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFEBFFA95AB6F8DF4586CEF8:CE5EA5AFFFE9FFA859E3F80644B5CBF5" box="[989,1171,1955,1981]" country="Vietnam" municipality="Hoang Lien National Park" name="Iriomote Station" pageId="118" stateProvince="Lao Cai">Iriomote Station</location>
(
<collectorName box="[1187,1256,1955,1981]" pageId="118">Japan</collectorName>
))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="119" pageId="118" type="distribution">
<paragraph blockId="118.[136,1451,1919,2053]" pageId="118">
<emphasis bold="true" box="[189,345,1991,2017]" pageId="118">Distribution:</emphasis>
—Endemic to
<collectingCountry box="[523,618,1991,2017]" name="Vietnam" pageId="118">Vietnam</collectingCountry>
(currently known only from Hoang Lien National Park of
<collectingRegion country="Vietnam" name="Lao Cai" pageId="118">Lao Cai Province</collectingRegion>
).
</paragraph>
<paragraph blockId="119.[136,1452,158,689]" pageId="119">
<emphasis bold="true" box="[189,421,158,184]" pageId="119">Habitat &amp; Ecology:</emphasis>
—Growing along roadside, streams, and edges of the secondary forest at an elevation around
<quantity box="[136,294,194,221]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="119" unit="m" value="2000.0" valueMax="2100.0" valueMin="1900.0">1900–2100 m</quantity>
.
</paragraph>
<paragraph blockId="119.[136,1452,158,689]" box="[189,913,230,257]" pageId="119">
<emphasis bold="true" box="[189,323,230,256]" pageId="119">Phenology:</emphasis>
—Flowering and fruiting from August to November.
</paragraph>
<paragraph blockId="119.[136,1452,158,689]" box="[189,610,266,295]" pageId="119">
<emphasis bold="true" box="[189,405,266,292]" pageId="119">Vernacular name:</emphasis>
—Xú hưƠng sapa.
</paragraph>
<paragraph blockId="119.[136,1452,158,689]" pageId="119">
<emphasis bold="true" box="[189,265,302,328]" pageId="119">Notes:</emphasis>
—
<taxonomicName box="[294,532,302,328]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="119" phylum="Tracheophyta" rank="species" species="sapaensis">
<emphasis box="[294,532,302,328]" italics="true" pageId="119">Lasianthus sapaensis</emphasis>
</taxonomicName>
was described from a Vietnamese specimen (
<emphasis box="[1043,1154,302,328]" italics="true" pageId="119">Dang 230</emphasis>
) by
<bibRefCitation author="Dang, V. S. &amp; Naiki, A." box="[1205,1448,302,329]" pageId="119" pagination="33 - 40" refId="ref54157" refString="Dang, V. S. &amp; Naiki, A. (2019) Two new species of Lasianthus (Rubiaceae) from Hoang Lien National Park, northern Vietnam. Phytotaxa 423 (1): 33 - 40. https: // doi. org / 10.11646 / phytotaxa. 423.1.4" type="journal article" year="2019">Dang &amp; Naiki (2019)</bibRefCitation>
. It can easily be recognized from other species of
<taxonomicName box="[702,825,338,364]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="119" phylum="Tracheophyta" rank="genus">
<emphasis box="[702,825,338,364]" italics="true" pageId="119">Lasianthus</emphasis>
</taxonomicName>
in having densely appressed pubescent to sericious on branches, ovate or ovate-oblong to oblong-lanceolate leaves, secondary veins 6–7 on each side of midrib, peduncles
<quantity box="[136,229,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="119" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3–7 mm</quantity>
long, flower white, and fruits globose, ovoid-globose to depressed globose, blue when ripe.
</paragraph>
<paragraph blockId="119.[136,1452,158,689]" pageId="119">
<taxonomicName box="[189,428,446,472]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="119" phylum="Tracheophyta" rank="species" species="sapaensis">
<emphasis box="[189,428,446,472]" italics="true" pageId="119">Lasianthus sapaensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName authority="Hook. f. (1880: 190)" authorityName="Hook. f." authorityPageNumber="190" authorityYear="1880" box="[573,959,446,472]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="119" phylum="Tracheophyta" rank="species" species="micranthus">
<emphasis box="[573,729,446,472]" italics="true" pageId="119">L. micranthus</emphasis>
Hook.f. (1880: 190)
</taxonomicName>
but differs from the latter in having densely appressed pubescent to sericious on branches (vs. sparsely puberulent to glabrous), petioles
<quantity box="[1158,1250,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="119" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 mm</quantity>
long (vs.
<quantity box="[1359,1452,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" pageId="119" unit="mm" value="6.5" valueMax="9.0" valueMin="4.0">4–9 mm</quantity>
long), calyx tube
<quantity box="[331,424,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="119" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long (vs.
<quantity box="[535,600,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="119" unit="mm" value="1.0">1 mm</quantity>
long) and calyx lobes
<quantity box="[852,966,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="119" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
long (vs.
<quantity box="[1077,1191,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="119" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long), corolla tube
<quantity metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="119" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
long (vs.
<quantity box="[292,385,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="119" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
long) and corolla lobes
<quantity box="[656,749,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="119" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long (vs.
<quantity box="[860,974,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="119" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
long).
</paragraph>
<paragraph blockId="119.[136,1452,158,689]" pageId="119">
It is also similar to
<taxonomicName authority="C. Y. Wu &amp; H. Zhu (1994: 75)" authorityName="C. Y. Wu &amp; H. Zhu" authorityPageNumber="75" authorityYear="1994" box="[415,1212,590,617]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="119" phylum="Tracheophyta" rank="subSpecies" species="biermannii" subSpecies="crassipedunculatus">
<emphasis box="[415,569,590,616]" italics="true" pageId="119">L. biermannii</emphasis>
subsp.
<emphasis box="[658,874,590,616]" italics="true" pageId="119">crassipedunculatus</emphasis>
C.Y.Wu &amp; H.Zhu (1994: 75)
</taxonomicName>
which is endemic to
<collectingCountry box="[136,206,626,652]" name="China" pageId="119">China</collectingCountry>
, but differs from the latter in having shorter stipules (
<quantity box="[803,918,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="119" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
vs.
<quantity box="[963,1056,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="119" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
long), peduncles
<quantity box="[1254,1347,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="119" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3–7 mm</quantity>
long (vs.
<quantity box="[136,257,662,689]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="119" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 mm</quantity>
long), bracts inconspicuous (vs.
<quantity box="[626,733,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="119" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">6–10 mm</quantity>
long) and smaller calyx lobes (
<quantity box="[1084,1200,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="119" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
long vs.
<quantity box="[1302,1367,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="119" unit="mm" value="5.0">5 mm</quantity>
long).
</paragraph>
</subSubSection>
</treatment>
</document>
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