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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFB4FFF05AB6F96C44C6CE68" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus chinensis Benth. 1861" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571616" ID-Zenodo-Dep="7571616" LSID="urn:lsid:plazi:treatment:43284262FFB4FFF05AB6F96C44C6CE68" httpUri="http://treatment.plazi.org/id/43284262FFB4FFF05AB6F96C44C6CE68" lastPageId="46" pageId="43">
<subSubSection box="[136,997,1736,1763]" pageId="43" type="nomenclature">
<paragraph blockId="43.[136,1452,1736,2087]" box="[136,997,1736,1763]" pageId="43">
<emphasis bold="true" box="[136,171,1737,1763]" pageId="43">13.</emphasis>
<taxonomicName ID-CoL="6NZ7H" ID-ENA="358833" authority="Benth. (1861: 160)" authorityName="Benth." authorityPageNumber="160" authorityYear="1861" box="[178,641,1736,1763]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="43" phylum="Tracheophyta" rank="species" species="chinensis">
<emphasis bold="true" box="[178,423,1736,1763]" italics="true" pageId="43">Lasianthus chinensis</emphasis>
Benth. (1861: 160)
</taxonomicName>
<figureCitation box="[672,750,1737,1763]" captionStart="FIGURE 31" captionStartId="44.[136,229,1775,1797]" captionTargetBox="[169,1414,212,1746]" captionTargetId="figure-25@44.[165,1418,209,1750]" captionTargetPageId="44" captionText="FIGURE 31. Lasianthus chinensis Benth. (from Dang 188, VNM): A. Branch, B. Stipules, C. Abaxial leaf surfaces, DG. Inflorescence showing side view flowers and young fruits, H. Young fruits, I. Cross section of fruit with pyrenes." figureDoi="http://doi.org/10.5281/zenodo.7571644" httpUri="https://zenodo.org/record/7571644/files/figure.png" pageId="43">Fig. 31</figureCitation>
(plate),
<figureCitation box="[844,923,1737,1763]" captionStart="FIGURE 32" captionStartId="45.[136,229,1781,1803]" captionTargetBox="[176,1410,204,1747]" captionTargetId="figure-17@45.[165,1418,193,1757]" captionTargetPageId="45" captionText="FIGURE 32. Distribution maps of Lasianthus chinensis, L. chrysoneurus, L. curtisii and L. dakglaiensis." figureDoi="http://doi.org/10.5281/zenodo.7571646" httpUri="https://zenodo.org/record/7571646/files/figure.png" pageId="43">Fig. 32</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection lastPageId="45" pageId="43" type="materials_examined">
<paragraph blockId="43.[136,1452,1736,2087]" box="[136,1235,1776,1799]" pageId="43">
<emphasis bold="true" box="[136,194,1777,1798]" pageId="43">Type:</emphasis>
<materialsCitation box="[219,1231,1776,1799]" country="China" county="Champion" location="Hong Kong" municipality="Festland" pageId="43" specimenCount="1" typeStatus="lectotype">
<collectingCountry box="[219,297,1777,1798]" name="China" pageId="43">CHINA</collectingCountry>
.
<location LSID="urn:lsid:plazi:treatment:43284262FFB4FFF05AB6F96C44C6CE68:CE5EA5AFFFB4FFF55B0CF9544180CB4E" box="[306,422,1777,1798]" country="China" county="Champion" municipality="Festland" name="Hong Kong" pageId="43">Hong Kong</location>
,
<collectingMunicipality box="[431,512,1777,1799]" pageId="43">Festland</collectingMunicipality>
bei,
<emphasis box="[560,699,1777,1798]" italics="true" pageId="43">
<collectingCounty box="[560,660,1777,1798]" pageId="43">Champion</collectingCounty>
s.n.
</emphasis>
(
<typeStatus box="[711,801,1777,1799]" pageId="43">lectotype</typeStatus>
K! [L000763825], designated by
<bibRefCitation author="Zhu, H." box="[1128,1224,1777,1799]" pageId="43" pagination="116 - 150" refId="ref55444" refString="Zhu, H. (2001) A taxonomic revision of the genus Lasianthus Jack (Rubiaceae) from Thailand. Acta Phytotaxonomica Sinica 39: 116 - 150." type="journal article" year="2001">Zhu 2001</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="43.[136,1452,1736,2087]" pageId="43">
<taxonomicName authority="Champ. ex Benth. (1852: 196)" authorityName="Benth." authorityPageNumber="196" authorityYear="1852" box="[136,622,1812,1835]" class="Magnoliopsida" family="Rubiaceae" genus="Mephitidia" kingdom="Plantae" order="Gentianales" pageId="43" phylum="Tracheophyta" rank="species" species="chinensis">
<emphasis box="[136,334,1813,1834]" italics="true" pageId="43">Mephitidia chinensis</emphasis>
<materialsCitation box="[338,622,1812,1835]" country="China" county="Champion" location="Type" municipality="Festland" pageId="43" specimenCount="1" typeStatus="lectotype">Champ.ex Benth.(1852:196)</materialsCitation>
</taxonomicName>
.
<location LSID="urn:lsid:plazi:treatment:43284262FFB4FFF05AB6F96C44C6CE68:CE5EA5AFFFB4FFF5584BF8B04281CB62" box="[629,679,1813,1834]" country="China" county="Champion" municipality="Festland" name="Type" pageId="43">Type</location>
:—
<collectingCountry box="[707,786,1813,1834]" name="China" pageId="43">CHINA</collectingCountry>
.
<location LSID="urn:lsid:plazi:treatment:43284262FFB4FFF05AB6F96C44C6CE68:CE5EA5AFFFB4FFF55926F8B043AFCB62" box="[792,905,1813,1834]" country="China" county="Champion" municipality="Festland" name="Hong Kong" pageId="43">Hong Kong</location>
,
<collectingMunicipality box="[912,993,1813,1835]" pageId="43">Festland</collectingMunicipality>
bei,
<emphasis box="[1036,1173,1813,1834]" italics="true" pageId="43">
<collectingCounty box="[1036,1136,1813,1834]" pageId="43">Champion</collectingCounty>
s.n.
</emphasis>
(
<typeStatus box="[1182,1272,1813,1835]" pageId="43">lectotype</typeStatus>
K! [K000763825], designated by
<bibRefCitation author="Zhu, H." box="[328,425,1849,1871]" pageId="43" pagination="116 - 150" refId="ref55444" refString="Zhu, H. (2001) A taxonomic revision of the genus Lasianthus Jack (Rubiaceae) from Thailand. Acta Phytotaxonomica Sinica 39: 116 - 150." type="journal article" year="2001">Zhu 2001</bibRefCitation>
).
</paragraph>
<paragraph blockId="43.[136,1452,1736,2087]" pageId="43">
<taxonomicName authority="Pierre ex Pit. (1924: 388)" authorityName="Pit." authorityPageNumber="388" authorityYear="1924" box="[136,586,1885,1907]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="43" phylum="Tracheophyta" rank="species" species="dinhensis">
<emphasis box="[136,338,1885,1906]" italics="true" pageId="43">Lasianthus dinhensis</emphasis>
<materialsCitation box="[343,586,1885,1907]" country="Cambodia" location="Poilane" municipality="Type" pageId="43" specimenCount="2" stateProvince="Ba Ria-Vung Tau" typeStatus="syntype">Pierre ex Pit. (1924: 388)</materialsCitation>
</taxonomicName>
.
<collectingMunicipality box="[594,642,1885,1906]" pageId="43">Type</collectingMunicipality>
:—
<collectingCountry box="[675,808,1885,1906]" name="Cambodia" pageId="43">CAMBODIA</collectingCountry>
. Kampot,
<emphasis box="[904,1020,1885,1906]" italics="true" pageId="43">
<location LSID="urn:lsid:plazi:treatment:43284262FFB4FFF05AB6F96C44C6CE68:CE5EA5AFFFB4FFF559B6F8F843F5CB3A" box="[904,979,1885,1906]" country="Cambodia" municipality="Type" name="Poilane" pageId="43" stateProvince="Ba Ria-Vung Tau">Poilane</location>
283
</emphasis>
(
<typeStatus box="[1032,1117,1885,1906]" pageId="43">syntypes</typeStatus>
P!);
<collectingCountry box="[1164,1278,1885,1906]" name="Vietnam" pageId="43">VIETNAM</collectingCountry>
.
<collectingRegion country="Vietnam" name="Ba Ria-Vung Tau" pageId="43">Ba Ria-Vung Tau Province</collectingRegion>
,
<emphasis box="[287,401,1921,1942]" italics="true" pageId="43">Pierre 3257</emphasis>
(
<typeStatus box="[414,499,1921,1942]" pageId="43">syntypes</typeStatus>
P!; isosyntypes K!, BM!).
</paragraph>
<paragraph blockId="43.[136,1452,1736,2087]" pageId="43">
<taxonomicName authority="Pit. (1924: 388)" authorityName="Pit." authorityPageNumber="388" authorityYear="1924" box="[136,664,1957,1979]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="43" phylum="Tracheophyta" rank="variety" species="dinhensis" variety="tonkinensis">
<emphasis box="[136,340,1957,1978]" italics="true" pageId="43">Lasianthus dinhensis</emphasis>
<materialsCitation box="[348,664,1957,1979]" collectingDate="1920-11-18" country="Laos" location="Type" pageId="43" specimenCount="1" stateProvince="Xiangkhouang" typeStatus="lectotype">
var.
<emphasis box="[391,499,1957,1978]" italics="true" pageId="43">tonkinensis</emphasis>
Pit. (1924: 388)
</materialsCitation>
</taxonomicName>
.
<location LSID="urn:lsid:plazi:treatment:43284262FFB4FFF05AB6F96C44C6CE68:CE5EA5AFFFB4FFF5589DF80042F5CBF2" box="[675,723,1957,1978]" country="Laos" name="Type" pageId="43" stateProvince="Xiangkhouang">Type</location>
:—
<collectingCountry box="[756,820,1957,1978]" name="Laos" pageId="43">LAOS</collectingCountry>
.
<collectingRegion box="[831,975,1957,1979]" country="Laos" name="Xiangkhouang" pageId="43">Xiangkhouang</collectingRegion>
,
<date box="[986,1176,1957,1979]" pageId="43" value="1920-11-18">
<collectingDate box="[986,1176,1957,1979]" pageId="43" value="1920-11-18">18 November 1920</collectingDate>
</date>
,
<emphasis box="[1188,1319,1957,1978]" italics="true" pageId="43">Poilane 2327</emphasis>
(
<typeStatus box="[1333,1423,1957,1979]" pageId="43">lectotype</typeStatus>
P! [P03980279]; isolectotypes P! [P03980278, P03980280]; designated by
<bibRefCitation author="Napiroon, T. &amp; Chayamarit, K. &amp; Dawson, S. &amp; Till, W. &amp; Balslev, H." box="[884,1087,1993,2015]" pageId="43" pagination="1 - 38" refId="ref54811" refString="Napiroon, T., Chayamarit, K., Dawson, S., Till, W. &amp; Balslev, H. (2020) A synopsis of Lasianthus (Lasiantheae, Rubiaceae) in Thailand and two additional new species. Phytotaxa 439 (1): 1 - 38. https: // doi. org / 10.11646 / phytotaxa. 439.1.1" type="journal article" year="2020">
Napiroon
<emphasis box="[981,1030,1993,2014]" italics="true" pageId="43">et al.</emphasis>
2020
</bibRefCitation>
).
</paragraph>
<paragraph blockId="43.[136,1452,1736,2087]" pageId="43">
<taxonomicName authority="Pierre ex Pit. (1924: 388)" authorityName="Pit." authorityPageNumber="388" authorityYear="1924" box="[136,752,2029,2051]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="43" phylum="Tracheophyta" rank="variety" species="dinhensis" variety="glabrescens">
<emphasis box="[136,338,2029,2050]" italics="true" pageId="43">Lasianthus dinhensis</emphasis>
<materialsCitation box="[344,752,2029,2051]" collectingDate="1877-03" country="Vietnam" location="Chua Chang mountain" municipality="Type" pageId="43" specimenCount="1" stateProvince="Dong Nai" typeStatus="holotype">
var.
<emphasis box="[386,500,2029,2050]" italics="true" pageId="43">glabrescens</emphasis>
Pierre ex Pit. (1924: 388)
</materialsCitation>
</taxonomicName>
.
<collectingMunicipality box="[761,809,2029,2050]" pageId="43">Type</collectingMunicipality>
:—
<collectingCountry box="[841,954,2029,2050]" name="Vietnam" pageId="43">VIETNAM</collectingCountry>
.
<collectingRegion box="[963,1151,2029,2051]" country="Vietnam" name="Dong Nai" pageId="43">Dong Nai Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFB4FFF05AB6F96C44C6CE68:CE5EA5AFFFB4FFF55EB7F8484545C44B" box="[1161,1379,2029,2051]" country="Vietnam" municipality="Type" name="Chua Chang mountain" pageId="43" stateProvince="Dong Nai">Chua Chang mountain</location>
,
<date pageId="43" value="1877-03">
<collectingDate pageId="43" value="1877-03">March 1877</collectingDate>
</date>
,
<emphasis box="[249,364,2065,2086]" italics="true" pageId="43">Pierre 3258</emphasis>
(
<typeStatus box="[376,462,2065,2087]" pageId="43">holotype</typeStatus>
P!; isotype P!).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571644" ID-Zenodo-Dep="7571644" httpUri="https://zenodo.org/record/7571644/files/figure.png" pageId="44" startId="44.[136,229,1775,1797]" targetBox="[169,1414,212,1746]" targetPageId="44">
<paragraph blockId="44.[136,1451,1775,1833]" pageId="44">
<emphasis bold="true" box="[136,265,1775,1797]" pageId="44">FIGURE 31.</emphasis>
<taxonomicName authority="Benth." authorityName="Benth." box="[271,543,1775,1797]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="44" phylum="Tracheophyta" rank="species" species="chinensis">
<emphasis box="[271,472,1775,1796]" italics="true" pageId="44">Lasianthus chinensis</emphasis>
Benth.
</taxonomicName>
(from
<emphasis box="[608,703,1775,1796]" italics="true" pageId="44">Dang 188</emphasis>
, VNM):
<emphasis bold="true" box="[792,815,1775,1796]" pageId="44">A.</emphasis>
Branch,
<emphasis bold="true" box="[902,924,1775,1796]" pageId="44">B.</emphasis>
Stipules,
<emphasis bold="true" box="[1020,1043,1775,1797]" pageId="44">C.</emphasis>
Abaxial leaf surfaces,
<emphasis bold="true" box="[1262,1316,1775,1797]" pageId="44">DG.</emphasis>
Inflorescence showing side view flowers and young fruits,
<emphasis bold="true" box="[568,593,1811,1832]" pageId="44">H.</emphasis>
Young fruits,
<emphasis bold="true" box="[729,745,1811,1832]" pageId="44">I.</emphasis>
Cross section of fruit with pyrenes.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571646" ID-Zenodo-Dep="7571646" box="[136,1148,1781,1803]" httpUri="https://zenodo.org/record/7571646/files/figure.png" pageId="45" startId="45.[136,229,1781,1803]" targetBox="[176,1410,204,1747]" targetPageId="45">
<paragraph blockId="45.[136,1148,1781,1803]" box="[136,1148,1781,1803]" pageId="45">
<emphasis bold="true" box="[136,265,1781,1803]" pageId="45">FIGURE 32.</emphasis>
Distribution maps of
<taxonomicName box="[476,676,1781,1802]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="45" phylum="Tracheophyta" rank="species" species="chinensis">
<emphasis box="[476,676,1781,1802]" italics="true" pageId="45">Lasianthus chinensis</emphasis>
</taxonomicName>
,
<taxonomicName box="[689,843,1781,1803]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="45" phylum="Tracheophyta" rank="species" species="chrysoneurus">
<emphasis box="[689,843,1781,1803]" italics="true" pageId="45">L. chrysoneurus</emphasis>
</taxonomicName>
,
<taxonomicName box="[855,949,1782,1803]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="45" phylum="Tracheophyta" rank="species" species="curtisii">
<emphasis box="[855,949,1782,1803]" italics="true" pageId="45">L. curtisii</emphasis>
</taxonomicName>
and
<taxonomicName box="[995,1144,1781,1803]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="45" phylum="Tracheophyta" rank="species" species="dakglaiensis">L. dakglaiensis</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="45.[136,1452,1849,2056]" pageId="45">
<emphasis bold="true" box="[136,396,1849,1875]" pageId="45">
<collectingRegion box="[136,262,1849,1875]" pageId="45">Specimens</collectingRegion>
examined:
</emphasis>
<materialsCitation collectingDate="1924-07" collectorName="Petelot" country="Vietnam" elevation="1500" location="Sapa District" pageId="45" specimenCount="1" stateProvince="Lao Cai">
<collectingCountry box="[425,557,1850,1876]" name="Vietnam" pageId="45">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[569,690,1850,1876]" pageId="45">Northern:</emphasis>
<collectingRegion box="[699,902,1850,1876]" country="Vietnam" name="Lao Cai" pageId="45">Lao Cai Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFB4FFF05AB6F96C44C6CE68:CE5EA5AFFFB2FFF359AAF89F440DCB1C" box="[916,1067,1850,1876]" country="Vietnam" name="Sapa District" pageId="45" stateProvince="Lao Cai">Sapa District</location>
,
<quantity box="[1080,1167,1849,1876]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="45" unit="m" value="1500.0">
<elevation box="[1080,1167,1849,1876]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="45" unit="m" value="1500.0">1500 m</elevation>
</quantity>
elev.,
<date box="[1244,1359,1850,1876]" pageId="45" value="1924-07">
<collectingDate box="[1244,1359,1850,1876]" pageId="45" value="1924-07">July 1924</collectingDate>
</date>
,
<emphasis italics="true" pageId="45">
<collectorName box="[1372,1451,1850,1876]" pageId="45">Petelot</collectorName>
1668
</emphasis>
(P)
</materialsCitation>
.
<materialsCitation collectingDate="2018-07-20" collectionCode="VNM" collectorName="Dang V. S. &amp; Hoang N. T. B. N. &amp; Dang" country="Vietnam" elevation="300" location="Hon Ba Nature Reserve" pageId="45" specimenCount="1" stateProvince="Khanh Hoa">
<emphasis bold="true" box="[252,353,1885,1911]" pageId="45">Central:</emphasis>
<collectingRegion box="[362,604,1886,1912]" country="Vietnam" name="Khanh Hoa" pageId="45">Khanh Hoa Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFB4FFF05AB6F96C44C6CE68:CE5EA5AFFFB2FFF35857F8FB435BCB30" box="[617,893,1886,1912]" country="Vietnam" name="Hon Ba Nature Reserve" pageId="45" stateProvince="Khanh Hoa">Hon Ba Nature Reserve</location>
,
<quantity box="[907,980,1886,1912]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="45" unit="m" value="300.0">
<elevation box="[907,980,1886,1912]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="45" unit="m" value="300.0">300 m</elevation>
</quantity>
elev.,
<date box="[1057,1209,1886,1912]" pageId="45" value="2018-07-20">
<collectingDate box="[1057,1209,1886,1912]" pageId="45" value="2018-07-20">20 July 2018</collectingDate>
</date>
,
<emphasis italics="true" pageId="45">
<collectorName box="[1222,1336,1886,1912]" pageId="45">Dang V.S.</collectorName>
&amp;
<collectorName pageId="45">Hoang N.T.B.N.</collectorName>
,
<collectorName box="[245,307,1922,1947]" pageId="45">Dang</collectorName>
188
</emphasis>
(
<collectionCode box="[369,440,1922,1948]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="45" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2014-07-13" collectionCode="FU, VNM" collectorName="Tagane &amp; Kanemitsu &amp; Dang V. S. &amp; Loi X. N. &amp; Thach N. D. &amp; Hieu P. N. H." country="Vietnam" elevation="900" location="Tran" pageId="45" specimenCount="1" stateProvince="Central">
in the same location,
<quantity box="[693,763,1922,1948]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="45" unit="m" value="900.0">
<elevation box="[693,763,1922,1948]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="45" unit="m" value="900.0">900 m</elevation>
</quantity>
elev.,
<date box="[833,979,1922,1948]" pageId="45" value="2014-07-13">
<collectingDate box="[833,979,1922,1948]" pageId="45" value="2014-07-13">13 July 2014</collectingDate>
</date>
,
<emphasis italics="true" pageId="45">
<collectorName box="[989,1070,1922,1947]" pageId="45">Tagane</collectorName>
S.,
<collectorName box="[1110,1230,1922,1947]" pageId="45">Kanemitsu</collectorName>
H.,
<collectorName box="[1276,1388,1922,1948]" pageId="45">Dang V.S.</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFB4FFF05AB6F96C44C6CE68:CE5EA5AFFFB2FFF35F48F827458DCBD3" box="[1398,1451,1922,1947]" country="Vietnam" name="Tran" pageId="45" stateProvince="Central">Tran</location>
H. with
<collectorName box="[225,322,1958,1983]" pageId="45">Loi X.N.</collectorName>
,
<collectorName box="[332,465,1958,1984]" pageId="45">Thach N.D.</collectorName>
,
<collectorName box="[476,611,1958,1983]" pageId="45">Hieu P.N.H.</collectorName>
<collectingDate box="[617,799,1957,1984]" isEnumeration="true" pageId="45">
<date box="[617,690,1957,1983]" pageId="45" value="1626-05">V1626</date>
&amp;
<date box="[725,799,1957,1983]" pageId="45" value="1678-05">V1678</date>
</collectingDate>
</emphasis>
(
<collectionCode box="[813,853,1958,1984]" country="China" name="Fudan University, Department of Biology" pageId="45">FU</collectionCode>
,
<collectionCode box="[864,934,1958,1984]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="45" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation collectingDate="1869-05" collectionCode="VNM" collectorName="Pierre" country="Vietnam" county="Ba Ria - Vung Tau Province" location="Nui Dinh" municipality="Tan Thanh District" pageId="45" specimenCount="1" stateProvince="Southern">
<emphasis bold="true" box="[952,1072,1957,1983]" pageId="45">
<collectingRegion box="[952,1067,1957,1983]" pageId="45">Southern</collectingRegion>
:
</emphasis>
<collectingCounty box="[1078,1399,1958,1984]" pageId="45">Ba Ria Vung Tau Province</collectingCounty>
,
<collectingMunicipality pageId="45">Tan Thanh District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFB4FFF05AB6F96C44C6CE68:CE5EA5AFFFB2FFF35B05F86F418FCBAC" box="[315,425,1994,2020]" country="Vietnam" county="Ba Ria - Vung Tau Province" municipality="Tan Thanh District" name="Nui Dinh" pageId="45" stateProvince="Southern">Nui Dinh</location>
,
<date box="[437,556,1993,2020]" pageId="45" value="1869-05">
<collectingDate box="[437,556,1993,2020]" pageId="45" value="1869-05">May 1869</collectingDate>
</date>
,
<emphasis box="[569,704,1994,2020]" italics="true" pageId="45">
<collectorName box="[569,640,1994,2019]" pageId="45">Pierre</collectorName>
3257
</emphasis>
(
<collectionCode box="[720,791,1994,2020]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="45" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1877-03" collectionCode="VNM" collectorName="Pierre" country="Vietnam" location="Nui Chua Chan" municipality="Xuan Loc District" pageId="45" specimenCount="1" stateProvince="Dong Nai">
<collectingRegion box="[812,1034,1994,2020]" country="Vietnam" name="Dong Nai" pageId="45">Dong Nai Province</collectingRegion>
,
<collectingMunicipality box="[1047,1255,1994,2020]" pageId="45">Xuan Loc District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFB4FFF05AB6F96C44C6CE68:CE5EA5AFFFB2FFF35ECDF86F4581CBAC" box="[1267,1447,1994,2020]" country="Vietnam" municipality="Xuan Loc District" name="Nui Chua Chan" pageId="45" stateProvince="Dong Nai">Nui Chua Chan</location>
,
<date box="[136,275,2030,2056]" pageId="45" value="1877-03">
<collectingDate box="[136,275,2030,2056]" pageId="45" value="1877-03">March 1877</collectingDate>
</date>
,
<emphasis box="[286,420,2030,2056]" italics="true" pageId="45">
<collectorName box="[286,357,2030,2055]" pageId="45">Pierre</collectorName>
3258
</emphasis>
(
<collectionCode box="[435,506,2030,2056]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="45" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="46" type="distribution">
<paragraph blockId="46.[136,1452,158,545]" pageId="46">
<emphasis bold="true" box="[189,345,158,184]" pageId="46">Distribution:</emphasis>
—Widespread in tropical and subtropical regions in Asia (
<collectingCountry box="[1043,1163,158,184]" name="Cambodia" pageId="46">Cambodia</collectingCountry>
,
<collectingCountry box="[1180,1250,158,184]" name="China" pageId="46">China</collectingCountry>
,
<collectingCountry box="[1267,1324,159,185]" name="Laos" pageId="46">Laos</collectingCountry>
,
<collectingCountry box="[1342,1447,158,184]" name="Malaysia" pageId="46">Malaysia</collectingCountry>
,
<collectingCountry box="[136,265,194,220]" name="Philippines" pageId="46">Philippines</collectingCountry>
,
<collectingCountry box="[277,382,194,220]" name="Thailand" pageId="46">Thailand</collectingCountry>
). In
<collectingCountry box="[432,530,194,220]" name="Vietnam" pageId="46">Vietnam</collectingCountry>
,
<taxonomicName box="[542,777,194,220]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="chinensis">
<emphasis box="[542,777,194,220]" italics="true" pageId="46">Lasianthus chinensis</emphasis>
</taxonomicName>
occurs in Ba Ria—Vung Tau,
<collectingRegion box="[1129,1242,194,221]" country="Vietnam" name="Dong Nai" pageId="46">Dong Nai</collectingRegion>
,
<collectingRegion box="[1255,1387,194,221]" country="Vietnam" name="Khanh Hoa" pageId="46">Khanh Hoa</collectingRegion>
,
<collectingRegion country="Vietnam" name="Lam Dong" pageId="46">Lam Dong</collectingRegion>
,
<collectingRegion box="[212,323,230,257]" country="Vietnam" name="Lang Son" pageId="46">Lang Son</collectingRegion>
, and
<collectingRegion box="[382,474,230,257]" country="Vietnam" name="Lao Cai" pageId="46">Lao Cai</collectingRegion>
.
</paragraph>
<paragraph blockId="46.[136,1452,158,545]" pageId="46">
<emphasis bold="true" box="[189,426,266,292]" pageId="46">Habitat &amp; Ecology:</emphasis>
—Growing under the shade of evergreen forest or on the edge of the forest at
<quantity box="[1323,1452,266,293]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="9.0" metricValueMin="3.0" pageId="46" unit="m" value="600.0" valueMax="900.0" valueMin="300.0">300900 m</quantity>
elev.
</paragraph>
<paragraph blockId="46.[136,1452,158,545]" box="[189,880,338,365]" pageId="46">
<emphasis bold="true" box="[189,323,338,364]" pageId="46">Phenology:</emphasis>
—Flowering and fruiting from July to November.
</paragraph>
<paragraph blockId="46.[136,1452,158,545]" box="[189,681,374,403]" pageId="46">
<emphasis bold="true" box="[189,405,374,400]" pageId="46">Vernacular name:</emphasis>
—Xú hưƠng trung quốc.
</paragraph>
<paragraph blockId="46.[136,1452,158,545]" pageId="46">
<emphasis bold="true" box="[189,265,410,436]" pageId="46">Notes:</emphasis>
—Three Vietnamese taxa,
<taxonomicName authority="Pierre ex Pit. (1924: 388)" authorityName="Pit." authorityPageNumber="388" authorityYear="1924" box="[565,995,410,437]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="dinhensis">
<emphasis box="[565,701,410,436]" italics="true" pageId="46">L. dinhensis</emphasis>
Pierre ex Pit. (1924: 388)
</taxonomicName>
,
<taxonomicName authority="Pit. (1924: 388)" authorityName="Pit." authorityPageNumber="388" authorityYear="1924" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="variety" species="dinhensis" variety="tonkinensis">
<emphasis box="[1008,1144,410,436]" italics="true" pageId="46">L. dinhensis</emphasis>
var.
<emphasis box="[1199,1325,410,436]" italics="true" pageId="46">tonkinensis</emphasis>
Pit. (1924: 388)
</taxonomicName>
and
<taxonomicName authority="Pierre ex Pit. (1924: 388)" authorityName="Pit." authorityPageNumber="388" authorityYear="1924" box="[239,843,446,473]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="variety" species="dinhensis" variety="glabrescens">
<emphasis box="[239,374,446,472]" italics="true" pageId="46">L. dinhensis</emphasis>
var.
<emphasis box="[426,559,446,472]" italics="true" pageId="46">glabrescens</emphasis>
Pierre ex Pit. (1924: 388)
</taxonomicName>
were treated as the synonyms of
<taxonomicName box="[1208,1341,446,472]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="chinensis">
<emphasis box="[1208,1341,446,472]" italics="true" pageId="46">L. chinensis</emphasis>
</taxonomicName>
by
<bibRefCitation author="Zhu, H. &amp; Roos, M. C. &amp; Ridsdale, C. E." pageId="46" pagination="1 - 102" refId="ref55608" refString="Zhu, H., Roos, M. C. &amp; Ridsdale, C. E. (2012) A taxonomic revision of the Malesian species of Lasianthus (Rubiaceae). Blumea 57: 1 - 102. https: // doi. org / 10.3767 / 000651912 X 652012" type="journal article" year="2012">
Zhu
<emphasis italics="true" pageId="46">et al.</emphasis>
(2012)
</bibRefCitation>
.
<taxonomicName box="[258,491,482,508]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="chinensis">
<emphasis box="[258,491,482,508]" italics="true" pageId="46">Lasianthus chinensis</emphasis>
</taxonomicName>
can easily be recognized in the field in having more than
<quantity box="[1126,1193,482,509]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="46" unit="cm" value="14.0">14 cm</quantity>
long leaves, secondary veins 1012 on each side of midrib, calyx elongated campanulate,
<quantity box="[883,976,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="46" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
long, and corolla white.
</paragraph>
</subSubSection>
</treatment>
</document>