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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E7856FF97FF013FD3AB0DE880" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Alluviobolus Wesener &amp; Enghoff &amp; Sierwald 2009" docType="key" docVersion="10" lastPageNumber="82" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="82" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesener, Thomas</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sierwald, Petra</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>19</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>128</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791451" ID-GBIF-Taxon="163747616" ID-Zenodo-Dep="3791451" LSID="urn:lsid:plazi:treatment:1772122E7856FF97FF013FD3AB0DE880" httpUri="http://treatment.plazi.org/id/1772122E7856FF97FF013FD3AB0DE880" lastPageNumber="82" pageId="81" pageNumber="82">
<subSubSection box="[140,515,166,193]" pageId="81" pageNumber="82" type="nomenclature">
<paragraph blockId="81.[140,515,166,193]" box="[140,515,166,193]" pageId="81" pageNumber="82">
<heading bold="true" box="[140,515,166,193]" fontSize="11" level="1" pageId="81" pageNumber="82" reason="1">
<emphasis bold="true" box="[140,515,166,193]" pageId="81" pageNumber="82">
Key to the
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[270,418,166,193]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[270,418,166,193]" italics="true" pageId="81" pageNumber="82">Alluviobolus</emphasis>
</taxonomicName>
species:
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="81" pageNumber="82" type="key">
<keyStep pageId="81" pageNumber="82">
<paragraph blockId="81.[140,1108,237,722]" pageId="81" pageNumber="82">
<keyLead pageId="81" pageNumber="82">
<specimenCount box="[140,303,237,263]" pageId="81" pageNumber="82" type="male">1 Male</specimenCount>
coxae 3–7 with processes (
<figureCitation box="[612,717,237,263]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="81" pageNumber="82">Figs 44B</figureCitation>
, 45B). Telopodite of posterior gonopods with a sclerite of elongated triangular shape (
<emphasis box="[869,881,273,298]" italics="true" pageId="81" pageNumber="82">x</emphasis>
in
<figureCitation box="[925,1036,272,298]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="81" pageNumber="82">Figs 44H</figureCitation>
, 45I). Body with 47–49 body rings......................................................................
<emphasis bold="true" box="[1092,1107,307,333]" pageId="81" pageNumber="82">2</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="81.[140,1108,237,722]" pageId="81" pageNumber="82">
<keyLead pageId="81" pageNumber="82">
– Male coxal processes absent (
<figureCitation box="[588,696,343,369]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="81" pageNumber="82">Fig. 47B</figureCitation>
). Telopodite of posterior gonopods with a sclerite of wide rectangular shape (
<emphasis box="[724,736,379,404]" italics="true" pageId="81" pageNumber="82">x</emphasis>
in
<figureCitation box="[777,884,378,404]" captionStart="Figure 47" captionStartId="85.[140,212,825,847]" captionTargetBox="[140,1108,165,794]" captionTargetId="figure@85.[140,1108,465,804]" captionTargetPageId="85" captionText="Figure 47. Alluviobolus antanosy sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767336" httpUri="https://zenodo.org/record/3767336/files/figure.png" pageId="81" pageNumber="82">Fig. 47G</figureCitation>
). Body with 42–44 body rings........................................................................
<emphasis bold="true" box="[897,1108,413,439]" pageId="81" pageNumber="82">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[897,1036,413,439]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="antanosy" status="sp. n.">
<emphasis bold="true" box="[897,1036,413,439]" italics="true" pageId="81" pageNumber="82">A. antanosy</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,413,439]" pageId="81" pageNumber="82" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="81" pageNumber="82">
<paragraph blockId="81.[140,1108,237,722]" pageId="81" pageNumber="82">
<keyLead pageId="81" pageNumber="82">
2 Colour on mesozonites grey, metazonites black, dorsally with a wide red stripe. Male coxae 3 and 4 with slender processes (
<figureCitation box="[829,934,483,510]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="81" pageNumber="82">Fig. 44B</figureCitation>
). Telopodite of posterior gonopod without a lateral membranous fringe (
<figureCitation box="[890,995,519,545]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="81" pageNumber="82">Fig. 44H</figureCitation>
)............. .....................................................................................
<emphasis bold="true" box="[880,1108,553,580]" pageId="81" pageNumber="82">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[880,1035,553,580]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="laticlavius" status="sp. n.">
<emphasis bold="true" box="[880,1035,553,580]" italics="true" pageId="81" pageNumber="82">A. laticlavius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,554,580]" pageId="81" pageNumber="82" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="81.[140,1108,237,722]" pageId="81" pageNumber="82">
<keyLead pageId="81" pageNumber="82">
– Colour on mesozonites dark green, metazonites red, dorsally on mesozonites with a slender orange stripe (
<figureCitation box="[580,686,624,650]" captionStart="Figure 42" captionStartId="77.[139,211,599,621]" captionTargetBox="[139,1107,165,580]" captionTargetId="figure@77.[139,1108,165,580]" captionTargetPageId="77" captionText="Figure 42. Alluviobolus tsimelahy sp. n., A living male from Tsimelahy B male and female specimen from Ebosika." figureDoi="http://doi.org/10.5281/zenodo.3767330" httpUri="https://zenodo.org/record/3767330/files/figure.png" pageId="81" pageNumber="82">Fig. 42A</figureCitation>
). Male coxae 3–5 with a large, swollen process (
<figureCitation box="[383,486,659,686]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. A–C, F–I male holotype, D, E female paratype. A head, lateral view B legs 1–7, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="81" pageNumber="82">Figs 45B</figureCitation>
,
<figureCitation box="[498,549,659,686]" captionStart="Figure 46" captionStartId="83.[140,212,1014,1036]" captionTargetBox="[140,1108,165,990]" captionTargetId="figure@83.[140,1108,165,667]" captionTargetPageId="83" captionText="Figure 46. Alluviobolus tsimelahy sp. n. specimens from Ebosika, A–G, I–M male, H female. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E gnathochilarium, mentum F gnathochilarium, dorsal view on endochilarium G, H left midbody legs I anterior gonopod, anterior view Į right posterior gonopod, anterior view K anterior gonopod, posterior view L right posterior gonopod, telopodite, posterior view M right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="81" pageNumber="82">46B</figureCitation>
). Telopodite of posterior gonopod laterally with a membranous fringe (
<figureCitation box="[482,578,694,721]" captionStart="Figure 45" captionStartId="79.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1295]" captionTargetId="figure@79.[140,1108,165,887]" captionTargetPageId="79" captionText="Figure 45. Alluviobolus tsimelahy sp. n. A–C, F–I male holotype, D, E female paratype. A head, lateral view B legs 1–7, ventral view C telson D vulva and sterno-coxite of second leg, posterior view E left mandible, mesal view F anterior gonopod, anterior view G right posterior gonopod, anterior view H anterior gonopod, posterior view I right posterior gonopod, posterior view. 3iT = 3-combed internal tooth; Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; LT = lateral tooth; Md = mandible; mp = molar plate; O = operculum; pL = pectinate lamella; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = coxosternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767332" httpUri="https://zenodo.org/record/3767332/files/figure.png" pageId="81" pageNumber="82">Figs 45I</figureCitation>
,
<figureCitation box="[590,637,695,722]" captionStart="Figure 46" captionStartId="83.[140,212,1014,1036]" captionTargetBox="[140,1108,165,990]" captionTargetId="figure@83.[140,1108,165,667]" captionTargetPageId="83" captionText="Figure 46. Alluviobolus tsimelahy sp. n. specimens from Ebosika, A–G, I–M male, H female. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E gnathochilarium, mentum F gnathochilarium, dorsal view on endochilarium G, H left midbody legs I anterior gonopod, anterior view Į right posterior gonopod, anterior view K anterior gonopod, posterior view L right posterior gonopod, telopodite, posterior view M right posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; cp = central pads; Cx = coxite; Endo = endochilarium; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767334" httpUri="https://zenodo.org/record/3767334/files/figure.png" pageId="81" pageNumber="82">46L</figureCitation>
).................................
<emphasis bold="true" box="[890,1108,694,721]" pageId="81" pageNumber="82">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[890,1035,694,721]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="tsimelahy" status="sp. n.">
<emphasis bold="true" box="[890,1035,694,721]" italics="true" pageId="81" pageNumber="82">A. tsimelahy</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,695,721]" pageId="81" pageNumber="82" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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