299 lines
35 KiB
XML
299 lines
35 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E7825FFE2FF013CFDAE66E839" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Colossobolus aculeatus Wesener & Enghoff & Sierwald 2009, sp. n." docType="treatment" docUuid="FA93B3D8-7AD7-4E8F-93E9-CA4B329D4BB9" docUuidSource="ZooBank" docVersion="11" lastPageNumber="37" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="35" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>128</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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||
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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||
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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||
</mods:mods>
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||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791403" ID-GBIF-Taxon="163747653" ID-Zenodo-Dep="3791403" LSID="urn:lsid:zoobank.org:act:FA93B3D8-7AD7-4E8F-93E9-CA4B329D4BB9" httpUri="http://treatment.plazi.org/id/1772122E7825FFE2FF013CFDAE66E839" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35">
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||
<subSubSection box="[140,593,905,932]" pageId="34" pageNumber="35" type="nomenclature">
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||
<paragraph blockId="34.[140,988,905,967]" box="[140,593,905,932]" pageId="34" pageNumber="35">
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||
<heading box="[140,593,905,932]" fontSize="11" level="2" pageId="34" pageNumber="35" reason="3">
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<emphasis bold="true" box="[140,593,905,932]" pageId="34" pageNumber="35">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,516,905,932]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis bold="true" box="[140,408,905,932]" italics="true" pageId="34" pageNumber="35">Colossobolus aculeatus</emphasis>
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Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel box="[528,593,906,932]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,988,941,967]" pageId="34" pageNumber="35" type="description">
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||
<paragraph blockId="34.[140,988,905,967]" box="[140,988,941,967]" pageId="34" pageNumber="35">
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<uri box="[140,988,941,967]" pageId="34" pageNumber="35">
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||
urn:lsid:zoobank.org:act:
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<uuid box="[423,988,941,967]" pageId="34" pageNumber="35">FA93B3D8-7AD7-4E8F-93E9-CA4B329D4BB9</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="materials_examined">
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<paragraph blockId="34.[140,1108,1011,1672]" pageId="34" pageNumber="35">
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<emphasis bold="true" box="[140,371,1011,1037]" pageId="34" pageNumber="35">
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<emphasis box="[140,241,1011,1037]" inLineHeading="true" pageId="34" pageNumber="35" reason="1">Material</emphasis>
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examined:
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</emphasis>
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<specimenCount box="[378,422,1012,1038]" pageId="34" pageNumber="35" type="male">2 ♂</specimenCount>
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<materialsCitation ID-GBIF-Occurrence="2620669307" collectingDate="2003-12-01" collectorName="de Binara & B. L. Fisher" country="Madagascar" elevation="375" latitude="-13.255" location="Foret de Binara" longLatPrecision="21" longitude="49.616665" municipality="Province" pageId="34" pageNumber="35" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana" typeStatus="holotype">
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<emphasis box="[429,532,1011,1037]" italics="true" pageId="34" pageNumber="35">
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<typeStatus box="[429,526,1011,1037]" pageId="34" pageNumber="35">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCount box="[539,583,1012,1038]" pageId="34" pageNumber="35" type="male">1 ♂</specimenCount>
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(
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<quantity box="[599,697,1012,1038]" metricMagnitude="-1" metricUnit="m" metricValue="1.22" pageId="34" pageNumber="35" unit="mm" value="122.0">122 mm</quantity>
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long), CAS BLF 9555,
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<collectingCountry box="[969,1103,1011,1037]" name="Madagascar" pageId="34" pageNumber="35">Madagascar</collectingCountry>
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,
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<collectingMunicipality box="[140,238,1047,1073]" pageId="34" pageNumber="35">Province</collectingMunicipality>
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<collectingRegion box="[245,384,1047,1073]" country="Madagascar" name="Antsiranana" pageId="34" pageNumber="35">Antsiranana</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:1772122E7825FFE2FF013CFDAE66E839:9A04F5E37825FFE4FE063B63AD65EE60" box="[395,572,1047,1073]" country="Madagascar" latitude="-13.255" longLatPrecision="21" longitude="49.616665" municipality="Province" name="Foret de Binara" pageId="34" pageNumber="35" stateProvince="Antsiranana">
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Forêt
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<collectorName box="[461,572,1047,1073]" pageId="34" pageNumber="35">de Binara</collectorName>
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</location>
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,
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<locationDeviation box="[583,876,1046,1073]" location="Daraina" pageId="34" pageNumber="35">
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<quantity box="[583,664,1046,1073]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="34" pageNumber="35" unit="km" value="7.5">7.5 km</quantity>
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230° SW Daraina
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</locationDeviation>
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,
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<quantity box="[887,964,1046,1073]" metricMagnitude="2" metricUnit="m" metricValue="3.75" pageId="34" pageNumber="35" unit="m" value="375.0">
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<elevation box="[887,964,1046,1073]" metricMagnitude="2" metricUnit="m" metricValue="3.75" pageId="34" pageNumber="35" unit="m" value="375.0">375 m</elevation>
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</quantity>
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, tropical dry forest,
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<geoCoordinate box="[220,352,1082,1109]" degrees="13" direction="south" minutes="15" orientation="latitude" pageId="34" pageNumber="35" precision="15" seconds="18" value="-13.255">13°15’18”S</geoCoordinate>
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,
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<geoCoordinate box="[367,517,1082,1108]" degrees="049" direction="east" minutes="37" orientation="longitude" pageId="34" pageNumber="35" precision="15" seconds="00" value="49.616665">049°37’00”E</geoCoordinate>
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, leg.
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<collectorName box="[581,720,1082,1108]" pageId="34" pageNumber="35">B. L. Fisher</collectorName>
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,
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<date box="[734,864,1082,1108]" pageId="34" pageNumber="35" value="2003-12-01">
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<collectingDate box="[734,864,1082,1108]" pageId="34" pageNumber="35" value="2003-12-01">1.XII.2003</collectingDate>
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</date>
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="2620669301" collectingDate="2003-12-01" collectorName="de Binara & B. L. Fisher" country="Madagascar" elevation="375" latitude="-13.255" location="Foret de Binara" longLatPrecision="21" longitude="49.616665" municipality="Province" pageId="34" pageNumber="35" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana" typeStatus="paratype">
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<emphasis box="[878,978,1083,1108]" italics="true" pageId="34" pageNumber="35">
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<typeStatus box="[878,972,1083,1108]" pageId="34" pageNumber="35">Paratype</typeStatus>
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:
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</emphasis>
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<specimenCount box="[988,1036,1082,1108]" pageId="34" pageNumber="35" type="male">1 ♂</specimenCount>
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, same data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="diagnosis">
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<paragraph blockId="34.[140,1108,1011,1672]" pageId="34" pageNumber="35">
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<emphasis bold="true" box="[185,376,1152,1178]" pageId="34" pageNumber="35">
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<emphasis box="[185,275,1152,1178]" inLineHeading="true" pageId="34" pageNumber="35" reason="1">Similar</emphasis>
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species:
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</emphasis>
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the special U-shaped posterior gonopods are unique for
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="34" pageNumber="35">Colossobolus</emphasis>
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</taxonomicName>
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except for
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[348,559,1187,1214]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[348,559,1187,1214]" italics="true" pageId="34" pageNumber="35">C. pseudoaculeatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[566,631,1188,1214]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[637,775,1187,1214]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species">
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<emphasis box="[637,775,1187,1214]" italics="true" pageId="34" pageNumber="35">C. aculeatus</emphasis>
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</taxonomicName>
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differs from the latter in the presence of a sharp-ending preanal process, which is well-rounded in
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis italics="true" pageId="34" pageNumber="35">C. pseudoaculeatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[226,290,1258,1284]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
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(compare
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<figureCitation box="[414,517,1258,1284]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="34" pageNumber="35">Fig. 20C</figureCitation>
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with
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<figureCitation box="[584,689,1258,1284]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="34" pageNumber="35">Fig. 21B</figureCitation>
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). The lateral branch of the posterior gonopod telopodite is of equal width all along in
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[708,844,1293,1320]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species">
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<emphasis box="[708,844,1293,1320]" italics="true" pageId="34" pageNumber="35">C. aculeatus</emphasis>
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</taxonomicName>
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while it is basally slen- der with a swollen tip in
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[425,636,1328,1355]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[425,636,1328,1355]" italics="true" pageId="34" pageNumber="35">C. pseudoaculeatus</emphasis>
|
||
</taxonomicName>
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<taxonomicNameLabel box="[642,705,1329,1355]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
|
||
(compare
|
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<figureCitation box="[827,926,1328,1354]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="34" pageNumber="35">Fig. 20E</figureCitation>
|
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with
|
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<figureCitation box="[991,1095,1328,1354]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="34" pageNumber="35">Fig. 21E</figureCitation>
|
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).
|
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</paragraph>
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</subSubSection>
|
||
<subSubSection lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35" type="description">
|
||
<paragraph blockId="34.[140,1108,1011,1672]" pageId="34" pageNumber="35">
|
||
<emphasis bold="true" box="[185,335,1363,1389]" pageId="34" pageNumber="35">
|
||
<emphasis box="[185,330,1363,1389]" inLineHeading="true" pageId="34" pageNumber="35" reason="1">Description</emphasis>
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.
|
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</emphasis>
|
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<emphasis box="[345,497,1364,1389]" italics="true" pageId="34" pageNumber="35">Measurements</emphasis>
|
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: males with 51 body rings, circa
|
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<quantity box="[892,993,1364,1390]" metricMagnitude="-1" metricUnit="m" metricValue="1.22" pageId="34" pageNumber="35" unit="mm" value="122.0">122 mm</quantity>
|
||
long,
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="34" pageNumber="35" unit="mm" value="9.5">9.5 mm</quantity>
|
||
wide. Female unknown.
|
||
</paragraph>
|
||
<paragraph blockId="34.[140,1108,1011,1672]" pageId="34" pageNumber="35">
|
||
<emphasis box="[185,300,1434,1460]" italics="true" pageId="34" pageNumber="35">Coloration</emphasis>
|
||
affected by alcohol. Head, antennae, legs and telson red (
|
||
<figureCitation box="[953,1038,1434,1460]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="34" pageNumber="35">Figs 20</figureCitation>
|
||
A–C). Median part of collum dark, margins red (
|
||
<figureCitation box="[625,729,1469,1495]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="34" pageNumber="35">Fig. 20A</figureCitation>
|
||
). Pro- and anterior part of mesozonites dark brown, posterior part of meso- and metazonites red.
|
||
<emphasis box="[874,977,1505,1531]" italics="true" pageId="34" pageNumber="35">Antennae</emphasis>
|
||
protruding back to ring 3.
|
||
<emphasis box="[309,520,1539,1565]" italics="true" pageId="34" pageNumber="35">Male coxal processes</emphasis>
|
||
on coxae 3 and 4 barely visible, on coxae 5–7 absent (
|
||
<figureCitation box="[148,250,1575,1601]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="34" pageNumber="35">Fig. 20B</figureCitation>
|
||
).
|
||
<emphasis box="[270,430,1575,1601]" italics="true" pageId="34" pageNumber="35">Preanal process</emphasis>
|
||
sharp-edged, but not projecting over anal valves (
|
||
<figureCitation box="[990,1095,1575,1601]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="34" pageNumber="35">Fig. 20C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="34.[140,1108,1011,1672]" pageId="34" pageNumber="35">
|
||
<emphasis box="[185,372,1610,1636]" italics="true" pageId="34" pageNumber="35">Anterior gonopod</emphasis>
|
||
sternite elongated into a wide, broadly rounded lobe (
|
||
<figureCitation box="[987,1095,1610,1636]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="34" pageNumber="35">Fig. 20D</figureCitation>
|
||
). Mesal process of coxite weakly developed, protruding into a very short wide lobe with
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3820814" ID-Zenodo-Dep="3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="35" pageNumber="36" startId="35.[140,212,1526,1548]" targetBox="[140,1108,165,1506]" targetPageId="35">
|
||
<paragraph blockId="35.[140,1108,1526,1676]" pageId="35" pageNumber="36">
|
||
<emphasis bold="true" box="[140,250,1526,1548]" pageId="35" pageNumber="36">Figure 21.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[256,508,1526,1547]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[256,508,1526,1547]" italics="true" pageId="35" pageNumber="36">Colossobolus pseudoaculeatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[513,564,1527,1548]" pageId="35" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male
|
||
<typeStatus box="[624,708,1526,1548]" pageId="35" pageNumber="36">holotype</typeStatus>
|
||
.
|
||
<emphasis bold="true" box="[716,735,1526,1548]" pageId="35" pageNumber="36">A</emphasis>
|
||
head, lateral
|
||
<emphasis bold="true" box="[857,874,1526,1548]" pageId="35" pageNumber="36">B</emphasis>
|
||
telson
|
||
<emphasis bold="true" box="[939,958,1526,1548]" pageId="35" pageNumber="36">C</emphasis>
|
||
anterior gonopods, posterior view
|
||
<emphasis bold="true" box="[319,338,1558,1580]" pageId="35" pageNumber="36">D</emphasis>
|
||
anterior gonopods, anterior view
|
||
<emphasis bold="true" box="[652,667,1558,1580]" pageId="35" pageNumber="36">E</emphasis>
|
||
right posterior gonopod, posterior view
|
||
<emphasis bold="true" box="[1042,1056,1558,1580]" pageId="35" pageNumber="36">F</emphasis>
|
||
right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="36.[140,1108,167,616]" pageId="36" pageNumber="37">
|
||
a well-rounded tip. Lobe only slightly longer than sternite (
|
||
<figureCitation box="[797,903,167,193]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="36" pageNumber="37">Fig. 20D</figureCitation>
|
||
). Telopodite process long, becoming apically thicker than basally. Mesal margin laterally sharp, not projecting (
|
||
<figureCitation box="[231,332,237,263]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="36" pageNumber="37">Fig. 20F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="36.[140,1108,167,616]" pageId="36" pageNumber="37">
|
||
<emphasis box="[185,388,272,298]" italics="true" pageId="36" pageNumber="37">Posterior gonopods</emphasis>
|
||
telopodites positioned parallel to one another, generally Ushaped (
|
||
<figureCitation box="[237,387,307,334]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="36" pageNumber="37">Figs 20E, G</figureCitation>
|
||
). Coxite branch absent (
|
||
<figureCitation box="[683,790,307,334]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="36" pageNumber="37">Fig. 20E</figureCitation>
|
||
). Lateral branch of regular width, tip wide, well-rounded, not thicker than median or basal parts (
|
||
<figureCitation box="[930,1031,343,369]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="36" pageNumber="37">Fig. 20E</figureCitation>
|
||
). Central membranous area present as a wide, well-rounded knob (
|
||
<figureCitation box="[833,940,378,404]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="36" pageNumber="37">Fig. 20G</figureCitation>
|
||
). Main branch slightly longer than lateral branch, latero-apically towards lateral branch with a large membranous area (Fig, 20G).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="36" pageNumber="37" type="distribution">
|
||
<paragraph blockId="36.[140,1108,167,616]" pageId="36" pageNumber="37">
|
||
<emphasis bold="true" box="[185,486,483,509]" pageId="36" pageNumber="37">
|
||
<emphasis box="[185,336,483,509]" inLineHeading="true" pageId="36" pageNumber="37" reason="1">Distribution</emphasis>
|
||
and ecology
|
||
</emphasis>
|
||
:
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[500,631,483,510]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[500,631,483,510]" italics="true" pageId="36" pageNumber="37">C. aculeatus</emphasis>
|
||
</taxonomicName>
|
||
is only known from the Forêt Binara (
|
||
<figureCitation captionStart="Figure 10" captionStartId="18.[140,212,1654,1676]" captionTargetBox="[160,1106,1078,1634]" captionTargetId="figure@18.[140,1109,1071,1635]" captionTargetPageId="18" captionText="Figure 10. Distribution of the genus Colossobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820794" httpUri="https://zenodo.org/record/3820794/files/figure.png" pageId="36" pageNumber="37">Fig. 10</figureCitation>
|
||
) where it occurs in close proximity to
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[605,745,519,545]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[605,745,519,545]" italics="true" pageId="36" pageNumber="37">C. semicyclus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="36" pageNumber="37" type="etymology">
|
||
<paragraph blockId="36.[140,1108,167,616]" pageId="36" pageNumber="37">
|
||
<emphasis bold="true" box="[185,323,554,580]" pageId="36" pageNumber="37">
|
||
<emphasis box="[185,318,554,580]" inLineHeading="true" pageId="36" pageNumber="37" reason="1">Etymology</emphasis>
|
||
:
|
||
</emphasis>
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[330,428,554,580]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[330,428,554,580]" italics="true" pageId="36" pageNumber="37">aculeatus</emphasis>
|
||
</taxonomicName>
|
||
, adjective, refers to the sharp-edged preanal process, unique for
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[180,312,589,615]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[180,312,589,615]" italics="true" pageId="36" pageNumber="37">Colossobolus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |