422 lines
49 KiB
XML
422 lines
49 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E781EFFDDFF013DE7ACB7E836" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Colossobolus oblongopedus Wesener & Enghoff & Sierwald 2009, sp. n." docType="treatment" docUuid="50DBF062-18CB-4A0A-AD61-10066B0A8604" docUuidSource="ZooBank" docVersion="11" lastPageNumber="28" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="26" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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||
<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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||
<mods:detail type="issue">
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||
<mods:number>19</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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||
<mods:start>1</mods:start>
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||
<mods:end>128</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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||
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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||
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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||
</mods:mods>
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||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791393" ID-GBIF-Taxon="163747655" ID-Zenodo-Dep="3791393" LSID="urn:lsid:zoobank.org:act:50DBF062-18CB-4A0A-AD61-10066B0A8604" httpUri="http://treatment.plazi.org/id/1772122E781EFFDDFF013DE7ACB7E836" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
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||
<subSubSection box="[140,642,659,686]" pageId="25" pageNumber="26" type="nomenclature">
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||
<paragraph blockId="25.[140,975,659,721]" box="[140,642,659,686]" pageId="25" pageNumber="26">
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||
<heading box="[140,642,659,686]" fontSize="11" level="2" pageId="25" pageNumber="26" reason="3">
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<emphasis bold="true" box="[140,642,659,686]" pageId="25" pageNumber="26">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,565,659,686]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis bold="true" box="[140,457,659,686]" italics="true" pageId="25" pageNumber="26">Colossobolus oblongopedus</emphasis>
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Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel box="[577,642,659,685]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,975,694,721]" pageId="25" pageNumber="26" type="description">
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||
<paragraph blockId="25.[140,975,659,721]" box="[140,975,694,721]" pageId="25" pageNumber="26">
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||
<uri box="[140,975,694,721]" pageId="25" pageNumber="26">
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||
urn:lsid:zoobank.org:act:
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<uuid box="[417,975,694,721]" pageId="25" pageNumber="26">50DBF062-18CB-4A0A-AD61-10066B0A8604</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="26" type="materials_examined">
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<paragraph blockId="25.[140,1108,765,1672]" pageId="25" pageNumber="26">
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<emphasis bold="true" box="[140,382,765,791]" pageId="25" pageNumber="26">
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<emphasis box="[140,245,765,791]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Material</emphasis>
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examined:
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="2608818313" collectingDate="2003-11-27" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" county="Province" elevation="30" latitude="-13.083333" location="Daraina" longLatPrecision="21" longitude="49.908337" municipality="Foret d'Analabe" pageId="25" pageNumber="26" specimenCode="BLF 9429" specimenCount="3" specimenCount-female="3" stateProvince="Antsiranana" typeStatus="holotype">
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9 ♂,
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<specimenCount box="[457,505,765,792]" pageId="25" pageNumber="26" type="female">3 ♀</specimenCount>
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<emphasis box="[516,622,765,791]" italics="true" pageId="25" pageNumber="26">
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<typeStatus box="[516,617,765,791]" pageId="25" pageNumber="26">Holotype</typeStatus>
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:
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</emphasis>
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1 ♂ (
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<quantity box="[700,804,765,792]" metricMagnitude="-1" metricUnit="m" metricValue="1.26" pageId="25" pageNumber="26" unit="mm" value="126.0">126 mm</quantity>
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long),
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<collectionCode box="[895,949,765,791]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="25" pageNumber="26" type="Museum">CAS</collectionCode>
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<specimenCode box="[960,1084,765,792]" collectionCode="BLF" pageId="25" pageNumber="26">BLF 9429</specimenCode>
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B,
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<collectingCountry box="[140,279,800,826]" name="Madagascar" pageId="25" pageNumber="26">Madagascar</collectingCountry>
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,
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<collectingCounty box="[295,396,800,826]" pageId="25" pageNumber="26">Province</collectingCounty>
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<collectingRegion box="[409,553,800,826]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
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,
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<collectingMunicipality box="[570,763,800,826]" pageId="25" pageNumber="26">Forêt d’Analabe</collectingMunicipality>
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, 30.0 km 72°ENE
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<location LSID="urn:lsid:plazi:treatment:1772122E781EFFDDFF013DE7ACB7E836:9A04F5E3781EFFDFFC633C54AB16E96B" box="[1006,1103,800,826]" country="Madagascar" county="Province" latitude="-13.083333" longLatPrecision="21" longitude="49.908337" municipality="Foret d'Analabe" name="Daraina" pageId="25" pageNumber="26" stateProvince="Antsiranana">Daraina</location>
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,
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<quantity box="[140,204,836,862]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="25" pageNumber="26" unit="m" value="30.0">
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<elevation box="[140,204,836,862]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="25" pageNumber="26" unit="m" value="30.0">30 m</elevation>
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</quantity>
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, littoral rainforest,
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<geoCoordinate box="[431,575,835,862]" degrees="13" direction="south" minutes="05" orientation="latitude" pageId="25" pageNumber="26" precision="15" seconds="00" value="-13.083333">13°05’00” S</geoCoordinate>
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,
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<geoCoordinate box="[589,751,835,862]" degrees="049" direction="east" minutes="54" orientation="longitude" pageId="25" pageNumber="26" precision="15" seconds="30" value="49.908337">049°54’30” E</geoCoordinate>
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, leg.
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<collectorName box="[813,952,835,862]" pageId="25" pageNumber="26">B. L. Fisher</collectorName>
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,
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<date box="[964,1103,836,862]" pageId="25" pageNumber="26" value="2003-11-27">
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<collectingDate box="[964,1103,836,862]" pageId="25" pageNumber="26" value="2003-11-27">27.XI.2003</collectingDate>
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</date>
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="2608818306" box="[140,859,871,897]" collectingDate="2003-11-27" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" county="Province" elevation="30" latitude="-13.083333" location="Daraina" longLatPrecision="21" longitude="49.908337" municipality="Foret d'Analabe" pageId="25" pageNumber="26" specimenCode="BLF 9429" specimenCount="1" specimenCount-female="1" stateProvince="Antsiranana" typeStatus="paratype">
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<emphasis box="[140,252,871,896]" italics="true" pageId="25" pageNumber="26">
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<typeStatus box="[140,246,871,896]" pageId="25" pageNumber="26">Paratypes</typeStatus>
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:
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</emphasis>
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1 ♂,
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<specimenCount box="[321,366,871,897]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
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,
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<collectionCode box="[382,436,871,897]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="25" pageNumber="26" type="Museum">CAS</collectionCode>
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<specimenCode box="[445,567,871,897]" collectionCode="BLF" pageId="25" pageNumber="26">BLF 9429</specimenCode>
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B, same data as holotype
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2608818239" collectingDate="2003-12-12" collectionCode="CAS, BLF" collectorName="B. L. Fisher" country="Madagascar" elevation="550" latitude="-13.211666" location="Foret d'Antsahabe" longLatPrecision="21" longitude="49.556667" municipality="Province" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Antsiranana" typeStatus="paratype">
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||
1 ♂,
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<specimenCount box="[933,978,871,897]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
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,
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<collectionCode box="[994,1048,871,897]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="25" pageNumber="26" type="Museum">CAS</collectionCode>
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/BLF10114" pageId="25" pageNumber="26">
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<collectionCode box="[1057,1108,871,897]" pageId="25" pageNumber="26">BLF</collectionCode>
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10114
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</accessionNumber>
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,
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<collectingMunicipality box="[234,335,906,932]" pageId="25" pageNumber="26">Province</collectingMunicipality>
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<collectingRegion box="[347,491,906,932]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
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,
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||
<location LSID="urn:lsid:plazi:treatment:1772122E781EFFDDFF013DE7ACB7E836:9A04F5E3781EFFDFFE773CFEAD8FE9F5" box="[506,726,906,932]" country="Madagascar" latitude="-13.211666" longLatPrecision="21" longitude="49.556667" municipality="Province" name="Foret d'Antsahabe" pageId="25" pageNumber="26" stateProvince="Antsiranana">Forêt d’Antsahabe</location>
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,
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<locationDeviation box="[741,1047,906,933]" location="Daraina" pageId="25" pageNumber="26">
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<quantity box="[741,844,906,932]" metricMagnitude="4" metricUnit="m" metricValue="1.14" pageId="25" pageNumber="26" unit="km" value="11.4">11.4 km</quantity>
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275°W Daraina
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</locationDeviation>
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,
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<quantity metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="25" pageNumber="26" unit="m" value="550.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="25" pageNumber="26" unit="m" value="550.0">550 m</elevation>
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</quantity>
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, tropical dry forest,
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<geoCoordinate box="[410,555,941,967]" degrees="13" direction="south" minutes="12" orientation="latitude" pageId="25" pageNumber="26" precision="15" seconds="42" value="-13.211666">13°12’42” S</geoCoordinate>
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,
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<geoCoordinate box="[570,734,941,968]" degrees="049" direction="east" minutes="33" orientation="longitude" pageId="25" pageNumber="26" precision="15" seconds="24" value="49.556667">049°33’24” E</geoCoordinate>
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, leg.
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<collectorName box="[799,940,941,968]" pageId="25" pageNumber="26">B. L.Fisher</collectorName>
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,
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<date box="[954,1102,941,967]" pageId="25" pageNumber="26" value="2003-12-12">
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<collectingDate box="[954,1102,941,967]" pageId="25" pageNumber="26" value="2003-12-12">12.XII.2003</collectingDate>
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</date>
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, general collecting
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="25.[140,1108,765,1672]" pageId="25" pageNumber="26">
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||
<emphasis bold="true" box="[185,484,1011,1037]" pageId="25" pageNumber="26">
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<emphasis box="[185,257,1011,1037]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Other</emphasis>
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material examined
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</emphasis>
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||
:
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<materialsCitation ID-GBIF-Occurrence="2608818280" collectingDate="1999-02-28" collectingDateMax="1999-03-06" collectingDateMin="1999-02-28" collectionCode="FMMC" collectorName="S. M. Goodman" country="Madagascar" elevation="1785" latitude="-13.9772215" location="Reserve Speciale de Manongarivo" longLatPrecision="21" longitude="48.42278" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Antsiranana">
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4 ♂,
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<specimenCount box="[555,641,1011,1038]" pageId="25" pageNumber="26" type="generic">1 imm.</specimenCount>
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||
,
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||
<collectionCode box="[652,745,1012,1038]" pageId="25" pageNumber="26">FMMC</collectionCode>
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||
,
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||
<collectingCountry box="[756,890,1011,1037]" name="Madagascar" pageId="25" pageNumber="26">Madagascar</collectingCountry>
|
||
, Prov.
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||
<collectingRegion box="[964,1103,1011,1037]" country="Madagascar" name="Antsiranana" pageId="25" pageNumber="26">Antsiranana</collectingRegion>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:1772122E781EFFDDFF013DE7ACB7E836:9A04F5E3781EFFDFFF013B63AD53EE60" box="[140,522,1047,1073]" country="Madagascar" latitude="-13.9772215" longLatPrecision="21" longitude="48.42278" name="Reserve Speciale de Manongarivo" pageId="25" pageNumber="26" stateProvince="Antsiranana">Réserve Spéciale de Manongarivo</location>
|
||
, camp
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||
<quantity box="[605,714,1046,1073]" metricMagnitude="3" metricUnit="m" metricValue="1.785" pageId="25" pageNumber="26" unit="m" value="1785.0">
|
||
<elevation box="[605,714,1046,1073]" metricMagnitude="3" metricUnit="m" metricValue="1.785" pageId="25" pageNumber="26" unit="m" value="1785.0">1, 785 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate box="[727,868,1046,1073]" degrees="13" direction="south" minutes="58" orientation="latitude" pageId="25" pageNumber="26" precision="15" seconds="38" value="-13.9772215">13°58‘38“ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[881,1025,1046,1073]" degrees="48" direction="east" minutes="25" orientation="longitude" pageId="25" pageNumber="26" precision="15" seconds="22" value="48.42278">48°25‘22“ E</geoCoordinate>
|
||
, leg.
|
||
<collectorName pageId="25" pageNumber="26">S. M. Goodman</collectorName>
|
||
,
|
||
<date box="[315,506,1082,1108]" pageId="25" pageNumber="26" value="1999-02-28" valueMax="1999-03-06" valueMin="1999-02-28">
|
||
<collectingDate box="[315,506,1082,1108]" pageId="25" pageNumber="26" value="1999-02-28" valueMax="1999-03-06" valueMin="1999-02-28">28.II–6.III.1999</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2608818241" box="[520,936,1082,1108]" collectingDate="1999-02-28" collectingDateMax="1999-03-06" collectingDateMin="1999-02-28" collectionCode="FMMC" collectorName="S. M. Goodman" country="Madagascar" elevation="1785" latitude="-13.9772215" location="Reserve Speciale de Manongarivo" longLatPrecision="21" longitude="48.42278" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Antsiranana">
|
||
<specimenCount box="[520,564,1082,1108]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectionCode box="[581,674,1082,1108]" pageId="25" pageNumber="26">FMMC</collectionCode>
|
||
, same data as previous
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2608818211" collectingDate="1999-02-28" collectingDateMax="1999-03-06" collectingDateMin="1999-02-28" collectionCode="FMMC" collectorName="S. M. Goodman" country="Madagascar" elevation="1785" latitude="-13.9772215" location="Reserve Speciale de Manongarivo" longLatPrecision="21" longitude="48.42278" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Antsiranana">
|
||
2 ♂,
|
||
<collectionCode box="[1010,1103,1082,1108]" pageId="25" pageNumber="26">FMMC</collectionCode>
|
||
, same data as previous
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="25" pageNumber="26" type="diagnosis">
|
||
<paragraph blockId="25.[140,1108,765,1672]" pageId="25" pageNumber="26">
|
||
<emphasis bold="true" box="[185,460,1152,1178]" pageId="25" pageNumber="26">
|
||
<emphasis box="[185,327,1152,1178]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Differential</emphasis>
|
||
diagnosis:
|
||
</emphasis>
|
||
the general shape of the posterior gonopods of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species">
|
||
<emphasis italics="true" pageId="25" pageNumber="26">Colossobolus oblongopedus</emphasis>
|
||
</taxonomicName>
|
||
is similar to those of
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[621,757,1188,1214]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[621,757,1188,1214]" italics="true" pageId="25" pageNumber="26">C. giganteus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[765,828,1188,1214]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
|
||
The coxite branch is far shorter in
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[264,445,1223,1249]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[264,445,1223,1249]" italics="true" pageId="25" pageNumber="26">C. oblongopedus</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[550,689,1223,1249]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[550,689,1223,1249]" italics="true" pageId="25" pageNumber="26">C. giganteus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[698,762,1223,1249]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
|
||
(compare
|
||
<figureCitation box="[891,993,1223,1249]" captionStart="Figure 14" captionStartId="26.[140,212,1050,1072]" captionTargetBox="[140,1108,165,1030]" captionTargetId="figure@26.[140,1108,165,1031]" captionTargetPageId="26" captionText="Figure 14. Colossobolus oblongopedus sp. n., male holotype, right posterior gonopod. A anterior view B posterior view. Cx = coxite; Cx-br = coxite branch; ma = membranous area; S-Gr = spermatic groove; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820802" httpUri="https://zenodo.org/record/3820802/files/figure.png" pageId="25" pageNumber="26">Fig. 14A</figureCitation>
|
||
with
|
||
<figureCitation captionStart="Figure 15" captionStartId="28.[140,212,1494,1516]" captionTargetBox="[140,1108,165,1409]" captionTargetId="figure@28.[140,1108,528,1439]" captionTargetPageId="28" captionText="Figure 15. Colossobolus giganteus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. Apo = apodeme; Co = collum; Cx = coxite; Cx-br = coxite branch; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; St = sternite; ST = stipites; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820804" httpUri="https://zenodo.org/record/3820804/files/figure.png" pageId="25" pageNumber="26">Fig. 15E</figureCitation>
|
||
), while the telopodite main branch is more regularly shaped in the latter than in
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[172,352,1293,1320]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[172,352,1293,1320]" italics="true" pageId="25" pageNumber="26">C. oblongopedus</emphasis>
|
||
</taxonomicName>
|
||
(compare
|
||
<figureCitation box="[478,578,1293,1319]" captionStart="Figure 14" captionStartId="26.[140,212,1050,1072]" captionTargetBox="[140,1108,165,1030]" captionTargetId="figure@26.[140,1108,165,1031]" captionTargetPageId="26" captionText="Figure 14. Colossobolus oblongopedus sp. n., male holotype, right posterior gonopod. A anterior view B posterior view. Cx = coxite; Cx-br = coxite branch; ma = membranous area; S-Gr = spermatic groove; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820802" httpUri="https://zenodo.org/record/3820802/files/figure.png" pageId="25" pageNumber="26">Fig. 14B</figureCitation>
|
||
with
|
||
<figureCitation box="[647,749,1293,1320]" captionStart="Figure 15" captionStartId="28.[140,212,1494,1516]" captionTargetBox="[140,1108,165,1409]" captionTargetId="figure@28.[140,1108,528,1439]" captionTargetPageId="28" captionText="Figure 15. Colossobolus giganteus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. Apo = apodeme; Co = collum; Cx = coxite; Cx-br = coxite branch; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; St = sternite; ST = stipites; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820804" httpUri="https://zenodo.org/record/3820804/files/figure.png" pageId="25" pageNumber="26">Fig. 15F</figureCitation>
|
||
). The telopodite process of the anterior gonopod features a short, pointed, retrorse process in
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[885,1025,1329,1355]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[885,1025,1329,1355]" italics="true" pageId="25" pageNumber="26">C. giganteus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1034,1102,1329,1355]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
|
||
, which is entirely absent in
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[454,634,1363,1390]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[454,634,1363,1390]" italics="true" pageId="25" pageNumber="26">C. oblongopedus</emphasis>
|
||
</taxonomicName>
|
||
(compare
|
||
<figureCitation box="[759,865,1363,1390]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="25" pageNumber="26">Fig. 13D</figureCitation>
|
||
with
|
||
<figureCitation box="[933,1042,1363,1390]" captionStart="Figure 15" captionStartId="28.[140,212,1494,1516]" captionTargetBox="[140,1108,165,1409]" captionTargetId="figure@28.[140,1108,528,1439]" captionTargetPageId="28" captionText="Figure 15. Colossobolus giganteus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. Apo = apodeme; Co = collum; Cx = coxite; Cx-br = coxite branch; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; St = sternite; ST = stipites; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820804" httpUri="https://zenodo.org/record/3820804/files/figure.png" pageId="25" pageNumber="26">Fig. 15D</figureCitation>
|
||
). The collum is completely red in
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[467,648,1399,1425]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[467,648,1399,1425]" italics="true" pageId="25" pageNumber="26">C. oblongopedus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[665,771,1399,1425]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="25" pageNumber="26">Fig. 13A</figureCitation>
|
||
), while it is centrally dark in
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,276,1434,1460]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[140,276,1434,1460]" italics="true" pageId="25" pageNumber="26">C. giganteus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[284,346,1434,1460]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation box="[363,468,1434,1461]" captionStart="Figure 15" captionStartId="28.[140,212,1494,1516]" captionTargetBox="[140,1108,165,1409]" captionTargetId="figure@28.[140,1108,528,1439]" captionTargetPageId="28" captionText="Figure 15. Colossobolus giganteus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. Apo = apodeme; Co = collum; Cx = coxite; Cx-br = coxite branch; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; St = sternite; ST = stipites; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820804" httpUri="https://zenodo.org/record/3820804/files/figure.png" pageId="25" pageNumber="26">Fig. 15A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="description">
|
||
<paragraph blockId="25.[140,1108,765,1672]" pageId="25" pageNumber="26">
|
||
<emphasis bold="true" box="[185,335,1469,1495]" pageId="25" pageNumber="26">
|
||
<emphasis box="[185,330,1469,1495]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis box="[346,498,1470,1495]" italics="true" pageId="25" pageNumber="26">Measurements</emphasis>
|
||
: males with 50–52 body rings,
|
||
<quantity box="[879,1039,1469,1496]" metricMagnitude="-1" metricUnit="m" metricValue="1.34" metricValueMax="1.48" metricValueMin="1.2" pageId="25" pageNumber="26" unit="mm" value="134.0" valueMax="148.0" valueMin="120.0">120–148 mm</quantity>
|
||
long,
|
||
<quantity box="[140,311,1504,1531]" metricMagnitude="-2" metricUnit="m" metricValue="1.095" metricValueMax="1.14" metricValueMin="1.05" pageId="25" pageNumber="26" unit="mm" value="10.95" valueMax="11.4" valueMin="10.5">10.5–11.4 mm</quantity>
|
||
wide. Females with 50–52 body rings,
|
||
<quantity box="[760,917,1504,1531]" metricMagnitude="-1" metricUnit="m" metricValue="1.2349999999999999" metricValueMax="1.25" metricValueMin="1.22" pageId="25" pageNumber="26" unit="mm" value="123.5" valueMax="125.0" valueMin="122.0">122–125 mm</quantity>
|
||
long,
|
||
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.155" metricValueMax="1.26" metricValueMin="1.05" pageId="25" pageNumber="26" unit="mm" value="11.55" valueMax="12.6" valueMin="10.5">10.5–12.6 mm</quantity>
|
||
wide.
|
||
</paragraph>
|
||
<paragraph blockId="25.[140,1108,765,1672]" pageId="25" pageNumber="26">
|
||
<emphasis box="[185,300,1575,1601]" italics="true" pageId="25" pageNumber="26">Coloration</emphasis>
|
||
affected by alcohol. Head, collum, antennae and telson red (
|
||
<figureCitation box="[991,1095,1575,1601]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="25" pageNumber="26">Fig. 13A</figureCitation>
|
||
). Legs also red, but tarsi whitish (
|
||
<figureCitation box="[505,608,1610,1636]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="25" pageNumber="26">Fig. 13B</figureCitation>
|
||
). Meso- and metazonites of body rings reddish, posterior margin with a thin, dark brown line.
|
||
<emphasis box="[731,834,1645,1671]" italics="true" pageId="25" pageNumber="26">Antennae</emphasis>
|
||
protruding back to ring
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3820802" ID-Zenodo-Dep="3820802" httpUri="https://zenodo.org/record/3820802/files/figure.png" pageId="26" pageNumber="27" startId="26.[140,212,1050,1072]" targetBox="[140,1108,165,1030]" targetPageId="26">
|
||
<paragraph blockId="26.[140,1108,1050,1136]" pageId="26" pageNumber="27">
|
||
<emphasis bold="true" box="[140,252,1050,1072]" pageId="26" pageNumber="27">Figure 14.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[258,489,1050,1071]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[258,489,1050,1071]" italics="true" pageId="26" pageNumber="27">Colossobolus oblongopedus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[495,547,1051,1072]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male
|
||
<typeStatus box="[609,693,1050,1072]" pageId="26" pageNumber="27">holotype</typeStatus>
|
||
, right posterior gonopod.
|
||
<emphasis bold="true" box="[938,957,1050,1072]" pageId="26" pageNumber="27">A</emphasis>
|
||
anterior view
|
||
<emphasis bold="true" box="[1091,1108,1050,1072]" pageId="26" pageNumber="27">B</emphasis>
|
||
posterior view. Cx = coxite; Cx-br = coxite branch; ma = membranous area; S-Gr = spermatic groove; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="26.[140,1108,1187,1672]" pageId="26" pageNumber="27">
|
||
4.
|
||
<emphasis box="[169,382,1187,1213]" italics="true" pageId="26" pageNumber="27">Male coxal processes</emphasis>
|
||
on coxae 3 and 4 weakly developed, on coxae 5–7 barely visible (
|
||
<figureCitation box="[148,250,1223,1249]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="26" pageNumber="27">Fig. 13B</figureCitation>
|
||
).
|
||
<emphasis box="[270,431,1223,1249]" italics="true" pageId="26" pageNumber="27">Preanal process</emphasis>
|
||
well-rounded, not projecting.
|
||
</paragraph>
|
||
<paragraph blockId="26.[140,1108,1187,1672]" pageId="26" pageNumber="27">
|
||
<emphasis box="[185,372,1258,1284]" italics="true" pageId="26" pageNumber="27">Anterior gonopod</emphasis>
|
||
sternite elongated into a wide, broadly rounded lobe (
|
||
<figureCitation box="[989,1093,1258,1284]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="26" pageNumber="27">Fig. 13C</figureCitation>
|
||
). Mesal process of coxite weakly developed, protruding into a short, wide lobe with a pointed tip (
|
||
<figureCitation box="[280,384,1328,1354]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="26" pageNumber="27">Fig. 13C</figureCitation>
|
||
). Telopodite on posterior side basally with a circular groove (
|
||
<figureCitation captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="26" pageNumber="27">Fig. 13D</figureCitation>
|
||
). Telopodite process long and well-rounded (
|
||
<figureCitation box="[721,829,1363,1390]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="26" pageNumber="27">Fig. 13D</figureCitation>
|
||
), mesal margin laterally sharp, slightly protruding in basal half (
|
||
<figureCitation box="[586,693,1399,1425]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="26" pageNumber="27">Fig. 13D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="26.[140,1108,1187,1672]" lastBlockId="27.[140,1108,167,616]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
|
||
<emphasis box="[185,385,1434,1460]" italics="true" pageId="26" pageNumber="27">Posterior gonopods</emphasis>
|
||
telopodites positioned parallel to one another. Coxite branch relatively short, wide, stout, less than half as long as lateral branch of telopodite (
|
||
<figureCitation captionStart="Figure 14" captionStartId="26.[140,212,1050,1072]" captionTargetBox="[140,1108,165,1030]" captionTargetId="figure@26.[140,1108,165,1031]" captionTargetPageId="26" captionText="Figure 14. Colossobolus oblongopedus sp. n., male holotype, right posterior gonopod. A anterior view B posterior view. Cx = coxite; Cx-br = coxite branch; ma = membranous area; S-Gr = spermatic groove; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820802" httpUri="https://zenodo.org/record/3820802/files/figure.png" pageId="26" pageNumber="27">Figs 14A, B</figureCitation>
|
||
). Latter slightly curved, basally wide, at midpoint tapering, tip again wide and swollen (
|
||
<figureCitation box="[238,340,1539,1566]" captionStart="Figure 14" captionStartId="26.[140,212,1050,1072]" captionTargetBox="[140,1108,165,1030]" captionTargetId="figure@26.[140,1108,165,1031]" captionTargetPageId="26" captionText="Figure 14. Colossobolus oblongopedus sp. n., male holotype, right posterior gonopod. A anterior view B posterior view. Cx = coxite; Cx-br = coxite branch; ma = membranous area; S-Gr = spermatic groove; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820802" httpUri="https://zenodo.org/record/3820802/files/figure.png" pageId="26" pageNumber="27">Fig. 14A</figureCitation>
|
||
). Central membranous area developed as a short, wide, well-rounded lobe (
|
||
<figureCitation box="[202,304,1575,1601]" captionStart="Figure 14" captionStartId="26.[140,212,1050,1072]" captionTargetBox="[140,1108,165,1030]" captionTargetId="figure@26.[140,1108,165,1031]" captionTargetPageId="26" captionText="Figure 14. Colossobolus oblongopedus sp. n., male holotype, right posterior gonopod. A anterior view B posterior view. Cx = coxite; Cx-br = coxite branch; ma = membranous area; S-Gr = spermatic groove; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820802" httpUri="https://zenodo.org/record/3820802/files/figure.png" pageId="26" pageNumber="27">Fig. 14A</figureCitation>
|
||
). Mesal main branch short, basal part running parallel to lateral branch. Apical part of main branch extending towards and touching lateral branch (
|
||
<figureCitation box="[992,1094,1610,1636]" captionStart="Figure 14" captionStartId="26.[140,212,1050,1072]" captionTargetBox="[140,1108,165,1030]" captionTargetId="figure@26.[140,1108,165,1031]" captionTargetPageId="26" captionText="Figure 14. Colossobolus oblongopedus sp. n., male holotype, right posterior gonopod. A anterior view B posterior view. Cx = coxite; Cx-br = coxite branch; ma = membranous area; S-Gr = spermatic groove; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820802" httpUri="https://zenodo.org/record/3820802/files/figure.png" pageId="26" pageNumber="27">Fig. 14B</figureCitation>
|
||
). Mesal margin of main branch with two tiny, well-rounded, overlapping membranous folds (
|
||
<figureCitation box="[211,315,167,193]" captionStart="Figure 14" captionStartId="26.[140,212,1050,1072]" captionTargetBox="[140,1108,165,1030]" captionTargetId="figure@26.[140,1108,165,1031]" captionTargetPageId="26" captionText="Figure 14. Colossobolus oblongopedus sp. n., male holotype, right posterior gonopod. A anterior view B posterior view. Cx = coxite; Cx-br = coxite branch; ma = membranous area; S-Gr = spermatic groove; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820802" httpUri="https://zenodo.org/record/3820802/files/figure.png" pageId="27" pageNumber="28">Fig. 14A</figureCitation>
|
||
). Main branch laterally with a very large, irregular-shaped membrane (
|
||
<figureCitation box="[147,285,202,228]" captionStart="Figure 14" captionStartId="26.[140,212,1050,1072]" captionTargetBox="[140,1108,165,1030]" captionTargetId="figure@26.[140,1108,165,1031]" captionTargetPageId="26" captionText="Figure 14. Colossobolus oblongopedus sp. n., male holotype, right posterior gonopod. A anterior view B posterior view. Cx = coxite; Cx-br = coxite branch; ma = membranous area; S-Gr = spermatic groove; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820802" httpUri="https://zenodo.org/record/3820802/files/figure.png" pageId="27" pageNumber="28">Figs 14A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="27.[140,1108,167,616]" pageId="27" pageNumber="28">
|
||
<emphasis bold="true" box="[185,479,237,263]" pageId="27" pageNumber="28">
|
||
<emphasis box="[185,344,237,263]" inLineHeading="true" pageId="27" pageNumber="28" reason="1">Intraspecific</emphasis>
|
||
variation:
|
||
</emphasis>
|
||
the specimens from the dry forest Antsahabe (52 body rings) and Manangarivo (53 body rings) are slightly larger and more voluminous than the
|
||
<typeStatus box="[347,398,308,334]" pageId="27" pageNumber="28">type</typeStatus>
|
||
series specimens from the littoral forest of Analabe, which possess 50 or 51 body rings. The gonopods from both populations are almost identical.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="27" pageNumber="28" type="distribution">
|
||
<paragraph blockId="27.[140,1108,167,616]" pageId="27" pageNumber="28">
|
||
<emphasis bold="true" box="[185,500,413,439]" pageId="27" pageNumber="28">
|
||
<emphasis box="[185,336,413,439]" inLineHeading="true" pageId="27" pageNumber="28" reason="1">Distribution</emphasis>
|
||
and ecology:
|
||
</emphasis>
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[511,689,413,440]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[511,689,413,440]" italics="true" pageId="27" pageNumber="28">C. oblongopedus</emphasis>
|
||
</taxonomicName>
|
||
occurs in the dry forest of Manangarivo, around Daraina and the tiny littoral forest fragment of Analabe (
|
||
<figureCitation box="[948,1030,448,474]" captionStart="Figure 10" captionStartId="18.[140,212,1654,1676]" captionTargetBox="[160,1106,1078,1634]" captionTargetId="figure@18.[140,1109,1071,1635]" captionTargetPageId="18" captionText="Figure 10. Distribution of the genus Colossobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820794" httpUri="https://zenodo.org/record/3820794/files/figure.png" pageId="27" pageNumber="28">Fig. 10</figureCitation>
|
||
). Both areas are now isolated from one another by vast areas of pseudosteppe (
|
||
<bibRefCitation author="Moat and Smith" firstAuthor="Moat" pageId="27" pageNumber="28" refId="ref57536" refString="Moat J, Smith P (2007) Atlas of the Vegetation of Madagascar. Kew Publishing, London, 124 pp." type="book" year="2007">Moat and Smith 2007</bibRefCitation>
|
||
).
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[222,396,519,545]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[222,396,519,545]" italics="true" pageId="27" pageNumber="28">C. oblongopedus</emphasis>
|
||
</taxonomicName>
|
||
occurs in Analabe sympatric with
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[782,880,519,545]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[782,880,519,545]" italics="true" pageId="27" pageNumber="28">C. minor</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[887,949,519,545]" pageId="27" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
|
||
, in Antsahabe together with
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[299,440,554,580]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[299,440,554,580]" italics="true" pageId="27" pageNumber="28">C. semicyclus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[447,508,554,580]" pageId="27" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[185,1006,589,616]" pageId="27" pageNumber="28" type="etymology">
|
||
<paragraph blockId="27.[140,1108,167,616]" box="[185,1006,589,616]" pageId="27" pageNumber="28">
|
||
<emphasis bold="true" box="[185,323,589,615]" pageId="27" pageNumber="28">
|
||
<emphasis box="[185,318,589,615]" inLineHeading="true" pageId="27" pageNumber="28" reason="1">Etymology</emphasis>
|
||
:
|
||
</emphasis>
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[330,472,589,615]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[330,472,589,615]" italics="true" pageId="27" pageNumber="28">oblongopedus</emphasis>
|
||
</taxonomicName>
|
||
, adjective, refers to the extraordinary long legs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |