264 lines
30 KiB
XML
264 lines
30 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E781AFFD8FF013DADADEDECD9" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Colossobolus minor Wesener & Enghoff & Sierwald 2009, sp. n." docType="treatment" docUuid="BC5D77D7-9152-4DE3-94AA-CF2F1E83229C" docUuidSource="ZooBank" docVersion="11" lastPageNumber="31" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="30" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Wesener, Thomas</mods:namePart>
|
||
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
|
||
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Enghoff, Henrik</mods:namePart>
|
||
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
|
||
<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Sierwald, Petra</mods:namePart>
|
||
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
|
||
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2009</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2009-09-04</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>19</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>19</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>128</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
|
||
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
|
||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
|
||
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791388" ID-GBIF-Taxon="163747657" ID-Zenodo-Dep="3791388" LSID="urn:lsid:zoobank.org:act:BC5D77D7-9152-4DE3-94AA-CF2F1E83229C" httpUri="http://treatment.plazi.org/id/1772122E781AFFD8FF013DADADEDECD9" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||
<subSubSection box="[140,555,729,756]" pageId="29" pageNumber="30" type="nomenclature">
|
||
<paragraph blockId="29.[140,987,729,792]" box="[140,555,729,756]" pageId="29" pageNumber="30">
|
||
<heading box="[140,555,729,756]" fontSize="11" level="2" pageId="29" pageNumber="30" reason="3">
|
||
<emphasis bold="true" box="[140,555,729,756]" pageId="29" pageNumber="30">
|
||
<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,478,729,756]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis bold="true" box="[140,370,729,756]" italics="true" pageId="29" pageNumber="30">Colossobolus minor</emphasis>
|
||
Wesener
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[490,555,730,756]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[140,987,765,792]" pageId="29" pageNumber="30" type="description">
|
||
<paragraph blockId="29.[140,987,729,792]" box="[140,987,765,792]" pageId="29" pageNumber="30">
|
||
<uri box="[140,987,765,792]" pageId="29" pageNumber="30">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid box="[422,987,765,792]" pageId="29" pageNumber="30">BC5D77D7-9152-4DE3-94AA-CF2F1E83229C</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="29" pageNumber="30" type="materials_examined">
|
||
<paragraph blockId="29.[140,1108,835,1672]" pageId="29" pageNumber="30">
|
||
<emphasis bold="true" box="[140,372,835,861]" pageId="29" pageNumber="30">
|
||
<emphasis box="[140,241,835,861]" inLineHeading="true" pageId="29" pageNumber="30" reason="1">Material</emphasis>
|
||
examined:
|
||
</emphasis>
|
||
<specimenCount box="[380,426,835,862]" pageId="29" pageNumber="30" type="male">5 ♂</specimenCount>
|
||
<materialsCitation ID-GBIF-Occurrence="2620669311" collectingDate="2003-11-27" collectorName="B. L. Fisher" country="Madagascar" county="Province" elevation="30" latitude="-13.083333" location="Daraina" longLatPrecision="21" longitude="49.908337" municipality="Foret d'Analabe" pageId="29" pageNumber="30" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana" typeStatus="holotype">
|
||
<emphasis box="[434,537,835,861]" italics="true" pageId="29" pageNumber="30">
|
||
<typeStatus box="[434,531,835,861]" pageId="29" pageNumber="30">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount box="[545,591,836,862]" pageId="29" pageNumber="30" type="male">1 ♂</specimenCount>
|
||
(
|
||
<quantity box="[608,692,835,862]" metricMagnitude="-2" metricUnit="m" metricValue="8.6" pageId="29" pageNumber="30" unit="mm" value="86.0">86 mm</quantity>
|
||
long), CAS BLF 9429-A,
|
||
<collectingCountry name="Madagascar" pageId="29" pageNumber="30">Madagascar</collectingCountry>
|
||
,
|
||
<collectingCounty box="[186,284,871,897]" pageId="29" pageNumber="30">Province</collectingCounty>
|
||
<collectingRegion box="[292,431,871,897]" country="Madagascar" name="Antsiranana" pageId="29" pageNumber="30">Antsiranana</collectingRegion>
|
||
,
|
||
<collectingMunicipality box="[442,625,871,897]" pageId="29" pageNumber="30">Forêt d’Analabe</collectingMunicipality>
|
||
, 30.0 km 72° ENE
|
||
<location LSID="urn:lsid:plazi:treatment:1772122E781AFFD8FF013DADADEDECD9:9A04F5E3781AFFDBFCDE3C13ACE8E9D0" box="[851,945,871,897]" country="Madagascar" county="Province" latitude="-13.083333" longLatPrecision="21" longitude="49.908337" municipality="Foret d'Analabe" name="Daraina" pageId="29" pageNumber="30" stateProvince="Antsiranana">Daraina</location>
|
||
,
|
||
<quantity box="[957,1019,871,897]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="30" unit="m" value="30.0">
|
||
<elevation box="[957,1019,871,897]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="30" unit="m" value="30.0">30 m</elevation>
|
||
</quantity>
|
||
, littoral rainforest,
|
||
<geoCoordinate box="[263,405,906,933]" degrees="13" direction="south" minutes="05" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="00" value="-13.083333">13°05’00” S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[419,578,906,933]" degrees="049" direction="east" minutes="54" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="30" value="49.908337">049°54’30” E</geoCoordinate>
|
||
, leg.
|
||
<collectorName box="[641,778,906,932]" pageId="29" pageNumber="30">B. L. Fisher</collectorName>
|
||
,
|
||
<date box="[791,926,906,932]" pageId="29" pageNumber="30" value="2003-11-27">
|
||
<collectingDate box="[791,926,906,932]" pageId="29" pageNumber="30" value="2003-11-27">27.XI.2003</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation ID-GBIF-Occurrence="2620669325" collectingDate="2003-11-27" collectorName="B. L. Fisher" country="Madagascar" county="Province" elevation="30" latitude="-13.083333" location="Daraina" longLatPrecision="21" longitude="49.908337" municipality="Foret d'Analabe" pageId="29" pageNumber="30" specimenCount="4" specimenCount-male="4" stateProvince="Antsiranana" typeStatus="paratype">
|
||
<emphasis box="[939,1045,907,932]" italics="true" pageId="29" pageNumber="30">
|
||
<typeStatus box="[939,1041,907,932]" pageId="29" pageNumber="30">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount box="[1055,1102,906,932]" pageId="29" pageNumber="30" type="male">4 ♂</specimenCount>
|
||
, same data as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="29" pageNumber="30" type="diagnosis">
|
||
<paragraph blockId="29.[140,1108,835,1672]" pageId="29" pageNumber="30">
|
||
<emphasis bold="true" box="[185,449,976,1002]" pageId="29" pageNumber="30">
|
||
<emphasis box="[185,324,976,1002]" inLineHeading="true" pageId="29" pageNumber="30" reason="1">Differential</emphasis>
|
||
diagnosis:
|
||
</emphasis>
|
||
the small size, dark body ring colour and uniquely-shaped, overlapping posterior gonopods of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[535,635,1012,1038]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[535,635,1012,1038]" italics="true" pageId="29" pageNumber="30">C. minor</emphasis>
|
||
</taxonomicName>
|
||
identify this species unambiguously.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="29" pageNumber="30" type="description">
|
||
<paragraph blockId="29.[140,1108,835,1672]" pageId="29" pageNumber="30">
|
||
<emphasis bold="true" box="[185,335,1046,1072]" pageId="29" pageNumber="30">
|
||
<emphasis box="[185,330,1046,1072]" inLineHeading="true" pageId="29" pageNumber="30" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis box="[342,494,1047,1072]" italics="true" pageId="29" pageNumber="30">Measurements</emphasis>
|
||
: males with 47 body rings,
|
||
<quantity box="[807,948,1047,1073]" metricMagnitude="-2" metricUnit="m" metricValue="9.2" metricValueMax="10.0" metricValueMin="8.4" pageId="29" pageNumber="30" unit="mm" value="92.0" valueMax="100.0" valueMin="84.0">84–100 mm</quantity>
|
||
long,
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.4" metricValueMin="8.6" pageId="29" pageNumber="30" unit="mm" value="9.0" valueMax="9.4" valueMin="8.6">8.6–9.4 mm</quantity>
|
||
wide. Females unknown.
|
||
</paragraph>
|
||
<paragraph blockId="29.[140,1108,835,1672]" pageId="29" pageNumber="30">
|
||
<emphasis box="[185,300,1117,1143]" italics="true" pageId="29" pageNumber="30">Coloration</emphasis>
|
||
affected by alcohol. Head, antennae, legs and telson red (
|
||
<figureCitation box="[955,1094,1117,1144]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="29" pageNumber="30">Figs 16A, B</figureCitation>
|
||
). Pro- and mesozona dark brown, posterior half of metazona red.
|
||
<emphasis box="[873,976,1153,1179]" italics="true" pageId="29" pageNumber="30">Antennae</emphasis>
|
||
protruding back to ring 3.
|
||
<emphasis box="[316,530,1187,1213]" italics="true" pageId="29" pageNumber="30">Male coxal processes</emphasis>
|
||
on coxae 3–5 weakly developed, on coxae 6 and 7 barely visible (
|
||
<figureCitation box="[301,404,1223,1250]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="29" pageNumber="30">Fig. 16A</figureCitation>
|
||
).
|
||
<emphasis box="[424,584,1223,1249]" italics="true" pageId="29" pageNumber="30">Preanal process</emphasis>
|
||
well-rounded (
|
||
<figureCitation box="[758,860,1223,1250]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="29" pageNumber="30">Fig. 16B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="29.[140,1108,835,1672]" pageId="29" pageNumber="30">
|
||
<emphasis box="[185,372,1258,1284]" italics="true" pageId="29" pageNumber="30">Anterior gonopod</emphasis>
|
||
sternite elongated into a wide, broadly rounded lobe (
|
||
<figureCitation box="[989,1094,1258,1285]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="29" pageNumber="30">Fig. 16C</figureCitation>
|
||
). Mesal process of coxite weakly developed, protruding into a short, wide lobe with a well-rounded tip (
|
||
<figureCitation box="[347,452,1328,1355]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="29" pageNumber="30">Fig. 16C</figureCitation>
|
||
). Telopodite basally with a groove (
|
||
<figureCitation box="[858,965,1328,1355]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="29" pageNumber="30">Fig. 16D</figureCitation>
|
||
). Telopodite process long and well-rounded. Mesal margin laterally sharp-edged, at midpoint slightly protruding (
|
||
<figureCitation box="[306,413,1399,1426]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="29" pageNumber="30">Fig. 16D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="29.[140,1108,835,1672]" pageId="29" pageNumber="30">
|
||
<emphasis box="[185,371,1434,1460]" italics="true" pageId="29" pageNumber="30">Posterior gonopod</emphasis>
|
||
telopodites positioned face-to-face to one another. Coxite branch absent (
|
||
<figureCitation box="[230,334,1469,1496]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="29" pageNumber="30">Fig. 16E</figureCitation>
|
||
), lateral branch of telopodite straight, tapering (
|
||
<figureCitation box="[895,999,1469,1496]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="29" pageNumber="30">Fig. 16E</figureCitation>
|
||
). Central membranous area present as a short, wide, well-rounded lobe (
|
||
<figureCitation box="[825,924,1504,1531]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="29" pageNumber="30">Fig. 16E</figureCitation>
|
||
). Main branches of both posterior gonopods overlapping each other (
|
||
<figureCitation box="[722,856,1539,1566]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="29" pageNumber="30">Figs 16E, F</figureCitation>
|
||
). Main branch shorter than lateral branch, former at half of its length dividing into two parts, a mesal sclerotized finger-shaped process and a lateral wider membranous part (
|
||
<figureCitation box="[801,932,1610,1636]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="29" pageNumber="30">Figs 16E, F</figureCitation>
|
||
). Finger-shaped processes of both posterior gonopods overlap each others membranous parts (
|
||
<figureCitation box="[995,1095,1645,1672]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="29" pageNumber="30">Fig. 16E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3820806" ID-Zenodo-Dep="3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="30" pageNumber="31" startId="30.[140,212,1329,1351]" targetBox="[140,1108,165,1309]" targetPageId="30">
|
||
<paragraph blockId="30.[140,1108,1329,1511]" pageId="30" pageNumber="31">
|
||
<emphasis bold="true" box="[140,253,1329,1351]" pageId="30" pageNumber="31">Figure 16.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[260,430,1329,1351]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[260,430,1329,1351]" italics="true" pageId="30" pageNumber="31">Colossobolus minor</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[437,490,1330,1351]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male
|
||
<typeStatus box="[553,637,1329,1351]" pageId="30" pageNumber="31">holotype</typeStatus>
|
||
.
|
||
<emphasis bold="true" box="[648,667,1329,1351]" pageId="30" pageNumber="31">A</emphasis>
|
||
legs 1–7, ventral view
|
||
<emphasis bold="true" box="[885,902,1329,1351]" pageId="30" pageNumber="31">B</emphasis>
|
||
telson
|
||
<emphasis bold="true" box="[971,990,1329,1351]" pageId="30" pageNumber="31">C</emphasis>
|
||
anterior gonopods, anterior view
|
||
<emphasis bold="true" box="[347,366,1361,1383]" pageId="30" pageNumber="31">D</emphasis>
|
||
anterior gonopods, posterior view
|
||
<emphasis bold="true" box="[690,705,1361,1383]" pageId="30" pageNumber="31">E</emphasis>
|
||
left posterior gonopod, anterior view
|
||
<emphasis bold="true" box="[1057,1071,1361,1383]" pageId="30" pageNumber="31">F</emphasis>
|
||
left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="30" pageNumber="31" type="distribution">
|
||
<paragraph blockId="30.[140,1113,1539,1672]" pageId="30" pageNumber="31">
|
||
<emphasis bold="true" box="[185,485,1539,1565]" pageId="30" pageNumber="31">
|
||
<emphasis box="[185,336,1539,1565]" inLineHeading="true" pageId="30" pageNumber="31" reason="1">Distribution</emphasis>
|
||
and ecology
|
||
</emphasis>
|
||
:
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[498,597,1540,1566]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[498,597,1540,1566]" italics="true" pageId="30" pageNumber="31">C. minor</emphasis>
|
||
</taxonomicName>
|
||
is only known from the littoral forest of Analabe (
|
||
<figureCitation box="[193,277,1575,1601]" captionStart="Figure 10" captionStartId="18.[140,212,1654,1676]" captionTargetBox="[160,1106,1078,1634]" captionTargetId="figure@18.[140,1109,1071,1635]" captionTargetPageId="18" captionText="Figure 10. Distribution of the genus Colossobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820794" httpUri="https://zenodo.org/record/3820794/files/figure.png" pageId="30" pageNumber="31">Fig. 10</figureCitation>
|
||
), occurring sympatric with the larger
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[703,879,1575,1601]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[703,879,1575,1601]" italics="true" pageId="30" pageNumber="31">C. oblongopedus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="30" pageNumber="31" type="etymology">
|
||
<paragraph blockId="30.[140,1113,1539,1672]" pageId="30" pageNumber="31">
|
||
<emphasis bold="true" box="[185,323,1610,1636]" pageId="30" pageNumber="31">
|
||
<emphasis box="[185,318,1610,1636]" inLineHeading="true" pageId="30" pageNumber="31" reason="1">Etymology</emphasis>
|
||
:
|
||
</emphasis>
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[333,399,1611,1636]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[333,399,1611,1636]" italics="true" pageId="30" pageNumber="31">minor</emphasis>
|
||
</taxonomicName>
|
||
, adjective, refers to the small size of the species.
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species">
|
||
<emphasis italics="true" pageId="30" pageNumber="31">Colossobolus minor</emphasis>
|
||
</taxonomicName>
|
||
is the smallest known species of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |