286 lines
32 KiB
XML
286 lines
32 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E780AFFC9FF013C87AB6EEC35" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Sanguinobolus maculosus Wesener & Enghoff & Sierwald 2009, sp. n." docType="treatment" docUuid="CD92BDAF-F533-40B5-A50B-DA179DE3B2CC" docUuidSource="ZooBank" docVersion="8" lastPageNumber="16" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="14" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>128</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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||
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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||
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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||
</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791378" ID-GBIF-Taxon="164088190" ID-Zenodo-Dep="3791378" LSID="urn:lsid:zoobank.org:act:CD92BDAF-F533-40B5-A50B-DA179DE3B2CC" httpUri="http://treatment.plazi.org/id/1772122E780AFFC9FF013C87AB6EEC35" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
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||
<subSubSection box="[140,629,1011,1038]" pageId="13" pageNumber="14" type="nomenclature">
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||
<paragraph blockId="13.[140,1006,1011,1073]" box="[140,629,1011,1038]" pageId="13" pageNumber="14">
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<heading box="[140,629,1011,1038]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="3">
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<emphasis bold="true" box="[140,629,1011,1038]" pageId="13" pageNumber="14">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,552,1011,1038]" class="Diplopoda" family="Pachybolidae" genus="Sanguinobolus" kingdom="Animalia" order="Spirobolida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis bold="true" box="[140,443,1011,1038]" italics="true" pageId="13" pageNumber="14">Sanguinobolus maculosus</emphasis>
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Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel box="[564,629,1011,1037]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,1006,1046,1073]" pageId="13" pageNumber="14" type="description">
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||
<paragraph blockId="13.[140,1006,1011,1073]" box="[140,1006,1046,1073]" pageId="13" pageNumber="14">
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<uri box="[140,1006,1046,1073]" pageId="13" pageNumber="14">
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||
urn:lsid:zoobank.org:act:
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<uuid box="[418,1006,1046,1073]" pageId="13" pageNumber="14">CD92BDAF-F533-40B5-A50B-DA179DE3B2CC</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="materials_examined">
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<paragraph blockId="13.[140,1108,1117,1672]" pageId="13" pageNumber="14">
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<emphasis bold="true" box="[140,373,1117,1143]" pageId="13" pageNumber="14">
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<emphasis box="[140,241,1117,1143]" inLineHeading="true" pageId="13" pageNumber="14" reason="1">Material</emphasis>
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examined:
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</emphasis>
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<specimenCount box="[382,429,1117,1144]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
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,
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<specimenCount box="[444,488,1117,1144]" pageId="13" pageNumber="14" type="generic">1 F.</specimenCount>
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<materialsCitation ID-GBIF-Occurrence="2620669309" collectingDate="2001-01-20" collectingDateMax="2001-01-26" collectingDateMin="2001-01-20" collectorName="Fisher & Griswold" country="Madagascar" elevation="925" latitude="-12.534445" location="Parc National Montagne d'Ambre" longLatPrecision="21" longitude="49.179447" municipality="Province" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana" typeStatus="holotype">
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<emphasis box="[496,599,1117,1143]" italics="true" pageId="13" pageNumber="14">
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<typeStatus box="[496,593,1117,1143]" pageId="13" pageNumber="14">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCount box="[608,655,1117,1144]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
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(
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<quantity box="[673,773,1117,1144]" metricMagnitude="-1" metricUnit="m" metricValue="1.18" pageId="13" pageNumber="14" unit="mm" value="118.0">118 mm</quantity>
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long), FMMC 3918.
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<collectingCountry name="Madagascar" pageId="13" pageNumber="14">Madagascar</collectingCountry>
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,
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<collectingMunicipality box="[220,318,1152,1178]" pageId="13" pageNumber="14">Province</collectingMunicipality>
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<collectingRegion box="[325,464,1152,1178]" country="Madagascar" name="Antsiranana" pageId="13" pageNumber="14">Antsiranana</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:1772122E780AFFC9FF013C87AB6EEC35:9A04F5E3780AFFCBFE513BF5AC39EECB" box="[476,864,1152,1179]" country="Madagascar" latitude="-12.534445" longLatPrecision="21" longitude="49.179447" municipality="Province" name="Parc National Montagne d'Ambre" pageId="13" pageNumber="14" stateProvince="Antsiranana">Parc National Montagne d’Ambre</location>
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,
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<locationDeviation location="Joffreville" pageId="13" pageNumber="14">
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<quantity box="[875,956,1152,1178]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="13" pageNumber="14" unit="km" value="3.6">3.6 km</quantity>
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235°SW Joffreville
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</locationDeviation>
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,
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<quantity box="[230,306,1187,1214]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="13" pageNumber="14" unit="m" value="925.0">
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<elevation box="[230,306,1187,1214]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="13" pageNumber="14" unit="m" value="925.0">925 m</elevation>
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</quantity>
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, montane rainforest,
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<geoCoordinate box="[546,670,1188,1214]" degrees="12" direction="south" minutes="32" orientation="latitude" pageId="13" pageNumber="14" precision="15" seconds="4" value="-12.534445">12°32’4” S</geoCoordinate>
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,
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<geoCoordinate box="[682,839,1187,1214]" degrees="049" direction="east" minutes="10" orientation="longitude" pageId="13" pageNumber="14" precision="15" seconds="46" value="49.179447">049°10’46” E</geoCoordinate>
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, leg.
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<collectorName box="[896,966,1187,1213]" pageId="13" pageNumber="14">Fisher</collectorName>
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,
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<collectorName box="[976,1080,1187,1213]" pageId="13" pageNumber="14">Griswold</collectorName>
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et al.,
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<date box="[181,342,1223,1249]" pageId="13" pageNumber="14" value="2001-01-20" valueMax="2001-01-26" valueMin="2001-01-20">
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<collectingDate box="[181,342,1223,1249]" pageId="13" pageNumber="14" value="2001-01-20" valueMax="2001-01-26" valueMin="2001-01-20">20–26.I.2001</collectingDate>
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</date>
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="2620669313" box="[352,937,1223,1249]" collectingDate="2001-01-20" collectingDateMax="2001-01-26" collectingDateMin="2001-01-20" collectorName="Fisher & Griswold" country="Madagascar" elevation="925" latitude="-12.534445" location="Parc National Montagne d'Ambre" longLatPrecision="21" longitude="49.179447" municipality="Province" pageId="13" pageNumber="14" specimenCount="1" specimenCount-female="1" stateProvince="Antsiranana" typeStatus="paratype">
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<emphasis box="[352,452,1223,1248]" italics="true" pageId="13" pageNumber="14">
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<typeStatus box="[352,446,1223,1248]" pageId="13" pageNumber="14">Paratype</typeStatus>
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:
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</emphasis>
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<specimenCount box="[459,502,1223,1249]" pageId="13" pageNumber="14" type="female">1 ♀</specimenCount>
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, FMMC 3918, same data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
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<paragraph blockId="13.[140,1108,1117,1672]" pageId="13" pageNumber="14">
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<emphasis bold="true" box="[185,335,1257,1283]" pageId="13" pageNumber="14">
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<emphasis box="[185,330,1257,1283]" inLineHeading="true" pageId="13" pageNumber="14" reason="1">Description</emphasis>
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.
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</emphasis>
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<emphasis box="[341,493,1259,1284]" italics="true" pageId="13" pageNumber="14">Measurements</emphasis>
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: male
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<typeStatus box="[565,664,1258,1284]" pageId="13" pageNumber="14">holotype</typeStatus>
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with 51 body rings, circa
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<quantity box="[949,1044,1258,1284]" metricMagnitude="-1" metricUnit="m" metricValue="1.18" pageId="13" pageNumber="14" unit="mm" value="118.0">118 mm</quantity>
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long,
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<quantity box="[140,230,1293,1320]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="13" pageNumber="14" unit="mm" value="9.8">9.8 mm</quantity>
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wide. Female
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<typeStatus box="[393,490,1294,1320]" pageId="13" pageNumber="14">paratype</typeStatus>
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with 52 body rings,
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<quantity box="[726,823,1293,1320]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="14" unit="mm" value="120.0">120 mm</quantity>
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long,
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<quantity box="[895,1000,1293,1320]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="13" pageNumber="14" unit="mm" value="10.5">10.5 mm</quantity>
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wide.
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</paragraph>
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<paragraph blockId="13.[140,1108,1117,1672]" pageId="13" pageNumber="14">
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<emphasis box="[185,300,1328,1354]" italics="true" pageId="13" pageNumber="14">Coloration</emphasis>
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faded in alcohol. Head, antennae, legs and telson red. Body rings dirty red-brownish.
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</paragraph>
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<paragraph blockId="13.[140,1108,1117,1672]" pageId="13" pageNumber="14">
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<emphasis box="[185,244,1399,1425]" italics="true" pageId="13" pageNumber="14">Head</emphasis>
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: each eye with circa 35–40 ocelli arranged in 7 or 8 vertical rows. Labrum with standard three irregular teeth and a single row of 10–12 stout marginal setae. Clypeus with two setiferous foveolae on each side. Antennal cavity present, not extending below eye. Antennae of medium length, protruding back to ring 4. Relative lengths of antennomeres: 1<2>3=4=5=6, antennomere 2 longest. Terminal antennomere with four large sensory cones located together inside a membranous area (
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<figureCitation box="[148,239,1610,1636]" captionStart="Figure 7" captionStartId="14.[140,212,1263,1285]" captionTargetBox="[141,1107,165,1244]" captionTargetId="figure@14.[140,1107,165,1247]" captionTargetPageId="14" captionText="Figure 7. Sanguinobolus maculosus sp. n., male paratype, SEM. A antenna, apical view, arrow indicates area shown in 7B B antennomere 6 with sensilla basiconica C male left midbody leg D detail of tarsus E left mandible, mesal view. 3iT = 3-combed internal teeth; eT = external tooth; LT = lateral tooth; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3767292" httpUri="https://zenodo.org/record/3767292/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
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). Antennomere 5 latero-apically with four rows, antennomere 6 with three rows (
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<figureCitation box="[208,292,1645,1672]" captionStart="Figure 7" captionStartId="14.[140,212,1263,1285]" captionTargetBox="[141,1107,165,1244]" captionTargetId="figure@14.[140,1107,165,1247]" captionTargetPageId="14" captionText="Figure 7. Sanguinobolus maculosus sp. n., male paratype, SEM. A antenna, apical view, arrow indicates area shown in 7B B antennomere 6 with sensilla basiconica C male left midbody leg D detail of tarsus E left mandible, mesal view. 3iT = 3-combed internal teeth; eT = external tooth; LT = lateral tooth; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3767292" httpUri="https://zenodo.org/record/3767292/files/figure.png" pageId="13" pageNumber="14">Fig. 7B</figureCitation>
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) of sensilla basiconica.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3767292" ID-Zenodo-Dep="3767292" httpUri="https://zenodo.org/record/3767292/files/figure.png" pageId="14" pageNumber="15" startId="14.[140,212,1263,1285]" targetBox="[141,1107,165,1244]" targetPageId="14">
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<paragraph blockId="14.[140,1107,1226,1381]" pageId="14" pageNumber="15">
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<emphasis bold="true" box="[140,239,1263,1285]" pageId="14" pageNumber="15">Figure 7.</emphasis>
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[245,468,1264,1285]" class="Diplopoda" family="Pachybolidae" genus="Sanguinobolus" kingdom="Animalia" order="Spirobolida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[245,468,1264,1285]" italics="true" pageId="14" pageNumber="15">Sanguinobolus maculosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[475,528,1264,1285]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
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, male paratype, SEM.
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<emphasis bold="true" box="[742,761,1263,1285]" pageId="14" pageNumber="15">A</emphasis>
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antenna, apical view, arrow indicates area shown in 7B
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<emphasis bold="true" box="[306,323,1295,1317]" pageId="14" pageNumber="15">B</emphasis>
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antennomere 6 with sensilla basiconica
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<emphasis bold="true" box="[694,713,1295,1317]" pageId="14" pageNumber="15">C</emphasis>
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male left midbody leg
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<emphasis bold="true" box="[927,946,1295,1317]" pageId="14" pageNumber="15">D</emphasis>
|
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detail of tarsus
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<emphasis bold="true" box="[1092,1107,1295,1317]" pageId="14" pageNumber="15">E</emphasis>
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left mandible, mesal view. 3iT = 3-combed internal teeth; eT = external tooth; LT = lateral tooth; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica.
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</paragraph>
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</caption>
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<paragraph blockId="14.[140,1108,1469,1672]" pageId="14" pageNumber="15">
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<emphasis box="[185,373,1469,1495]" italics="true" pageId="14" pageNumber="15">Gnathochilarium</emphasis>
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: of usual spirobolidan shape. Lamellae linguales each with two standard setae located behind one another. Stipites each with three apical setae. Palpi of gnathochilarium with numerous sensilla. Hypopharyngeal crest with a field of spinelike structures. Central pads of endochilarium separated into two levels, group of 10– 12 sensilla located apically in recessed area. Medial palps with a large group of sensilla each, located towards hypopharyngeal crest.
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</paragraph>
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<paragraph blockId="15.[140,1108,167,1636]" pageId="15" pageNumber="16">
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<emphasis box="[185,291,167,193]" italics="true" pageId="15" pageNumber="16">Mandible</emphasis>
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: external tooth simple, rounded; mesal tooth with three cusps (
|
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<figureCitation box="[1008,1095,167,193]" captionStart="Figure 7" captionStartId="14.[140,212,1263,1285]" captionTargetBox="[141,1107,165,1244]" captionTargetId="figure@14.[140,1107,165,1247]" captionTargetPageId="14" captionText="Figure 7. Sanguinobolus maculosus sp. n., male paratype, SEM. A antenna, apical view, arrow indicates area shown in 7B B antennomere 6 with sensilla basiconica C male left midbody leg D detail of tarsus E left mandible, mesal view. 3iT = 3-combed internal teeth; eT = external tooth; LT = lateral tooth; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3767292" httpUri="https://zenodo.org/record/3767292/files/figure.png" pageId="15" pageNumber="16">Fig. 7E</figureCitation>
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). Five or six pectinate lamellae. Mesal margin of pectinate area with circa two rows of small slender spines. Molar plate with numerous (8–10) transverse furrows.
|
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</paragraph>
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<paragraph blockId="15.[140,1108,167,1636]" box="[185,1088,272,299]" pageId="15" pageNumber="16">
|
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<emphasis box="[185,274,272,298]" italics="true" pageId="15" pageNumber="16">Collum:</emphasis>
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smooth, laterally not protruding as far as ventral surface of body ring 2.
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</paragraph>
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<paragraph blockId="15.[140,1108,167,1636]" pageId="15" pageNumber="16">
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<emphasis box="[185,308,307,333]" italics="true" pageId="15" pageNumber="16">Body rings:</emphasis>
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mostly smooth, with some irregular punctation and some transverse impressions on mesozona. Metazona and mesozona ventrally with deep longitudinal impressions. Ozopores starting at body ring 6, located closely in front of, but not touching suture between mesozona and metazona. Ozopore on ring 6 located much more ventrally than subsequent ozopores.
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</paragraph>
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<paragraph blockId="15.[140,1108,167,1636]" pageId="15" pageNumber="16">
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||
<emphasis box="[185,251,483,509]" italics="true" pageId="15" pageNumber="16">Telson</emphasis>
|
||
: anal valves with well-developed lips and micropunctation but with neither grooves nor setae. Preanal ring and subanal process well-rounded, not extending beyond anal valves. Preanal process inconspicuous.
|
||
</paragraph>
|
||
<paragraph blockId="15.[140,1108,167,1636]" pageId="15" pageNumber="16">
|
||
<emphasis box="[185,230,590,615]" italics="true" pageId="15" pageNumber="16">Legs</emphasis>
|
||
: coxae 1 and 2 elongated and fused with sternum, podomeres from prefemur to tarsus in both sexes each with 4–10 ventral/mesal setae. Length of midbody legs circa 1.2 times body diameter in males, circa 1.0 times body diameter in females. Each podomere with an apical ventral seta. Coxae 3 and beyond of rectangular shape. Tarsus with a stout dorso-apical seta and a pair of stout ventro-apical ones in males, in females with three pairs of setae, of which apical pair stout, others successively smaller.
|
||
</paragraph>
|
||
<paragraph blockId="15.[140,1108,167,1636]" pageId="15" pageNumber="16">
|
||
<emphasis box="[185,429,800,826]" italics="true" pageId="15" pageNumber="16">Male sexual characters</emphasis>
|
||
: tarsi 3 and beyond with a small tarsal pad not protruding past base of claw. Mesal margin of tarsus towards tarsal pad carrying five grooves with short sensory setae (
|
||
<figureCitation box="[369,502,871,897]" captionStart="Figure 7" captionStartId="14.[140,212,1263,1285]" captionTargetBox="[141,1107,165,1244]" captionTargetId="figure@14.[140,1107,165,1247]" captionTargetPageId="14" captionText="Figure 7. Sanguinobolus maculosus sp. n., male paratype, SEM. A antenna, apical view, arrow indicates area shown in 7B B antennomere 6 with sensilla basiconica C male left midbody leg D detail of tarsus E left mandible, mesal view. 3iT = 3-combed internal teeth; eT = external tooth; LT = lateral tooth; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3767292" httpUri="https://zenodo.org/record/3767292/files/figure.png" pageId="15" pageNumber="16">Figs 7C, D</figureCitation>
|
||
). An additional row of numerous short setae present on outer tarsal margin (
|
||
<figureCitation box="[408,500,906,932]" captionStart="Figure 7" captionStartId="14.[140,212,1263,1285]" captionTargetBox="[141,1107,165,1244]" captionTargetId="figure@14.[140,1107,165,1247]" captionTargetPageId="14" captionText="Figure 7. Sanguinobolus maculosus sp. n., male paratype, SEM. A antenna, apical view, arrow indicates area shown in 7B B antennomere 6 with sensilla basiconica C male left midbody leg D detail of tarsus E left mandible, mesal view. 3iT = 3-combed internal teeth; eT = external tooth; LT = lateral tooth; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3767292" httpUri="https://zenodo.org/record/3767292/files/figure.png" pageId="15" pageNumber="16">Fig. 7D</figureCitation>
|
||
). Coxae 3 and 4 each with a short process protruding posteriorly, but never overlapping adjacent coxa.
|
||
</paragraph>
|
||
<paragraph blockId="15.[140,1108,167,1636]" pageId="15" pageNumber="16">
|
||
<emphasis box="[185,378,976,1003]" italics="true" pageId="15" pageNumber="16">Anterior gonopod</emphasis>
|
||
sternite elongated into a wide, broadly-rounded lobe with a slightly narrowed tip (
|
||
<figureCitation box="[391,481,1011,1038]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="15" pageNumber="16">Fig. 6A</figureCitation>
|
||
). Mesal process of coxite weakly developed, protruding into a wide, very short process, shorter than sternite (
|
||
<figureCitation box="[736,825,1047,1074]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="15" pageNumber="16">Fig. 6A</figureCitation>
|
||
). Telopodite on posterior side basally with a circular groove (
|
||
<figureCitation box="[542,631,1082,1109]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="15" pageNumber="16">Fig. 6B</figureCitation>
|
||
). Telopodite process long, basally slender, proximally greatly enlarged into a characteristic swollen knob (
|
||
<figureCitation box="[846,933,1117,1144]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="15" pageNumber="16">Fig. 6B</figureCitation>
|
||
). Mesal margin not projecting (
|
||
<figureCitation box="[316,404,1152,1179]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="15" pageNumber="16">Fig. 6B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="15.[140,1108,167,1636]" pageId="15" pageNumber="16">
|
||
<emphasis box="[185,382,1187,1213]" italics="true" pageId="15" pageNumber="16">Posterior gonopods</emphasis>
|
||
unique (
|
||
<figureCitation box="[483,613,1187,1214]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="15" pageNumber="16">Figs 6C, D</figureCitation>
|
||
). Gonopod pair connected by a well-sclerotized and visible sternite. Telopodites positioned face-to-face with one another. Coxite and telopodite separated by a membranous area. Branch of coxite long and stout, almost as long as, but more slender than lateral main branch of telopodite (
|
||
<figureCitation captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="15" pageNumber="16">Figs 6C, D</figureCitation>
|
||
), latter curved towards opposite gonopod, not tapering (
|
||
<figureCitation box="[820,911,1328,1355]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="15" pageNumber="16">Fig. 6C</figureCitation>
|
||
). Main branch 2 times longer than wide, completely sclerotized. Mesal margin towards coxite branch with some membranous folds, which slightly protrude above mesal margin (
|
||
<figureCitation box="[1002,1094,1399,1426]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="15" pageNumber="16">Fig. 6D</figureCitation>
|
||
). Sperm canal discharging apically at mesal margin of lateral main branch.
|
||
</paragraph>
|
||
<paragraph blockId="15.[140,1108,167,1636]" pageId="15" pageNumber="16">
|
||
<emphasis box="[185,449,1469,1495]" italics="true" pageId="15" pageNumber="16">Female sexual characters</emphasis>
|
||
: vulva simple, consisting of two simple, subequally-sized, sclerotized plates, bivalve-like (
|
||
<figureCitation box="[496,566,1504,1530]" captionStart="Figure 8" captionStartId="16.[140,212,799,821]" captionTargetBox="[140,1108,165,780]" captionTargetId="figure@16.[140,1108,165,780]" captionTargetPageId="16" captionText="Figure 8. Sanguinobolus maculosus sp. n., female paratype, SEM, right vulva. Abbreviations: av = ante- rior valve; O = operculum; pv = posterior valve." figureDoi="http://doi.org/10.5281/zenodo.3820792" httpUri="https://zenodo.org/record/3820792/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
). Both plates only basally with 3 or 4 rows of setae located towards opening. Anterior plate slightly larger than posterior one (
|
||
<figureCitation box="[1026,1092,1539,1566]" captionStart="Figure 8" captionStartId="16.[140,212,799,821]" captionTargetBox="[140,1108,165,780]" captionTargetId="figure@16.[140,1108,165,780]" captionTargetPageId="16" captionText="Figure 8. Sanguinobolus maculosus sp. n., female paratype, SEM, right vulva. Abbreviations: av = ante- rior valve; O = operculum; pv = posterior valve." figureDoi="http://doi.org/10.5281/zenodo.3820792" httpUri="https://zenodo.org/record/3820792/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="15.[140,1108,167,1636]" box="[185,1101,1574,1601]" pageId="15" pageNumber="16">
|
||
<emphasis bold="true" box="[185,456,1574,1600]" pageId="15" pageNumber="16">
|
||
<emphasis box="[185,333,1574,1600]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">Intraspecific</emphasis>
|
||
variation:
|
||
</emphasis>
|
||
only known from
|
||
<typeStatus box="[668,767,1575,1601]" pageId="15" pageNumber="16">holotype</typeStatus>
|
||
and a single female
|
||
<typeStatus box="[997,1098,1575,1601]" pageId="15" pageNumber="16">paratype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[185,1079,1610,1636]" pageId="15" pageNumber="16" type="etymology">
|
||
<paragraph blockId="15.[140,1108,167,1636]" box="[185,1079,1610,1636]" pageId="15" pageNumber="16">
|
||
<emphasis bold="true" box="[185,323,1610,1636]" pageId="15" pageNumber="16">
|
||
<emphasis box="[185,318,1610,1636]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">Etymology</emphasis>
|
||
:
|
||
</emphasis>
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[330,438,1610,1636]" class="Diplopoda" family="Pachybolidae" genus="Sanguinobolus" kingdom="Animalia" order="Spirobolida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[330,438,1610,1636]" italics="true" pageId="15" pageNumber="16">maculosus</emphasis>
|
||
</taxonomicName>
|
||
, adjective, refers to the spotted black-red colour pattern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |