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<document id="326345BDEC333AD1AA3906AFCB51DD22" ID-DOI="10.11646/zootaxa.5566.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14612921" ID-ZooBank="C39ECCDA-23CC-478C-90BC-E6CFE1595E7E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736289794118" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2025" docId="863EA81AFFA75F0BFF117D30E6F6FE93" docLanguage="en" docName="zootaxa.5566.1.2.pdf" docOrigin="Zootaxa 5566 (1)" docSource="https://doi.org/10.11646/zootaxa.5566.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nesamblyops nunni Sokolov, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="56" masterDocId="7A07D062FFA65F0FFF867B07E556FFD2" masterDocTitle="Three new blind species of the genus Nesamblyops Jeannel, 1937 (Anillini: Carabidae: Coleoptera) from New Zealand" masterLastPageNumber="60" masterPageNumber="52" pageNumber="53" updateTime="1736291255819" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D568E3586989C9ECEEABEA4D007C05EC">Three new blind species of the genus Nesamblyops Jeannel, 1937 (Anillini: Carabidae: Coleoptera) from New Zealand</mods:title>
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<mods:date id="935157769FCDA04444D12A5942F17D3A">2025</mods:date>
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<treatment id="863EA81AFFA75F0BFF117D30E6F6FE93" ID-DOI="http://doi.org/10.5281/zenodo.14612819" ID-Zenodo-Dep="14612819" LSID="urn:lsid:plazi:treatment:863EA81AFFA75F0BFF117D30E6F6FE93" httpUri="http://treatment.plazi.org/id/863EA81AFFA75F0BFF117D30E6F6FE93" lastPageId="4" lastPageNumber="56" pageId="1" pageNumber="53">
<subSubSection id="468D4A87FFA75F0EFF117D30E4A2F983" box="[151,500,1591,1618]" pageId="1" pageNumber="53" type="nomenclature">
<paragraph id="0E28190CFFA75F0EFF117D30E4A2F983" blockId="1.[151,500,1591,1653]" box="[151,500,1591,1618]" pageId="1" pageNumber="53">
<heading id="5560AE60FFA75F0EFF117D30E4A2F983" bold="true" box="[151,500,1591,1618]" fontSize="11" level="1" pageId="1" pageNumber="53" reason="1">
<emphasis id="3CE3C51EFFA75F0EFF117D30E4A2F983" bold="true" box="[151,500,1591,1618]" pageId="1" pageNumber="53">
<taxonomicName id="C997628FFFA75F0EFF117D30E4D1F983" authorityName="Sokolov" authorityYear="2025" box="[151,391,1591,1618]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="nunni" status="sp. nov.">
<emphasis id="3CE3C51EFFA75F0EFF117D30E4D1F983" bold="true" box="[151,391,1591,1618]" italics="true" pageId="1" pageNumber="53">Nesamblyops nunni</emphasis>
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,
<taxonomicNameLabel id="27D07865FFA75F0EFE107D30E4A2F983" box="[406,500,1591,1617]" pageId="1" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="468D4A87FFA75F0EFF117D5CE4EFF9A7" box="[151,441,1627,1653]" pageId="1" pageNumber="53" type="description">
<paragraph id="0E28190CFFA75F0EFF117D5CE4EFF9A7" blockId="1.[151,500,1591,1653]" box="[151,441,1627,1653]" pageId="1" pageNumber="53">
<figureCitation id="96AC0589FFA75F0EFF117D5CE448F9A7" box="[151,286,1627,1653]" captionStart="FIGURE 1" captionStartId="2.[151,250,996,1020]" captionTargetBox="[154,1411,194,961]" captionTargetId="figure-15@2.[151,1436,181,972]" captionTargetPageId="2" captionText="FIGURE 1. Digital habitus images of blind Nesamblyops species, dorsal aspect. A—N. nunni (Old Man Range, Central Otago, South Island), B—N. larochellei (Lewis Pass, Buller, South Island), C—N. pygmaeus (Waiotauru Road, Tararua Forest Park, Wellington, North Island). Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14612923" httpUri="https://zenodo.org/record/14612923/files/figure.png" pageId="1" pageNumber="53">Figures. 1A</figureCitation>
,
<figureCitation id="96AC0589FFA75F0EFEAF7D5CE418F9A7" box="[297,334,1627,1653]" captionStart="FIGURE 2" captionStartId="2.[151,250,1501,1525]" captionTargetBox="[156,1432,1118,1473]" captionTargetId="figure-76@2.[151,1436,1116,1477]" captionTargetPageId="2" captionText="FIGURE 2. Digital images of pronota of blind Nesamblyops species, dorsal aspect. A—N. nunni (Old Man Range, Central Otago, South Island), B—N. larochellei (Lewis Pass, Buller, South Island), C—N. pygmaeus (Waiotauru Road, Tararua Forest Park, Wellington, North Island). Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14612925" httpUri="https://zenodo.org/record/14612925/files/figure.png" pageId="1" pageNumber="53">2A</figureCitation>
,
<figureCitation id="96AC0589FFA75F0EFEDC7D5CE4C9F9A7" box="[346,415,1627,1653]" captionStart="FIGURE 3" captionStartId="3.[151,250,916,940]" captionTargetBox="[157,1430,187,886]" captionTargetId="figure-37@3.[151,1435,181,892]" captionTargetPageId="3" captionText="FIGURE 3. Line drawings of male genitalia of blind Nesamblyops species. N. nunni (Old Man Range, Central Otago, South Island): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. larochellei (Lewis Pass, Buller, South Island): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. pygmaeus (Waiotauru Road, Tararua Forest Park, Wellington, North Island): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14612927" httpUri="https://zenodo.org/record/14612927/files/figure.png" pageId="1" pageNumber="53">3AC</figureCitation>
,
<figureCitation id="96AC0589FFA75F0EFE2D7D5CE4EFF9A7" box="[427,441,1627,1653]" captionStart="FIGURE 4" captionStartId="5.[151,250,1648,1672]" captionTargetBox="[151,1436,181,1624]" captionTargetId="figure-37@5.[151,1436,181,1624]" captionTargetPageId="5" captionText="FIGURE 4. Distribution records for the blind Nesamblyops species and images of the male median lobe obtained from the collected specimens (inset illustrates specimen localities). Black circle—N. pygmaeus; red circle—N. larochellei; orange circles—N. nunni. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14612929" httpUri="https://zenodo.org/record/14612929/files/figure.png" pageId="1" pageNumber="53">4</figureCitation>
</paragraph>
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<paragraph id="0E28190CFFA75F0EFF117D9BE499F92D" blockId="1.[151,1437,1692,2043]" pageId="1" pageNumber="53">
<materialsCitation id="BEFF1351FFA75F0EFF117D9BE499F92D" collectingDate="1975-03-15" collectionCode="NZAC, R" collectorName="CO Shingle Rd. Old Man Range &amp; Entomology Division D. S. I. &amp; Arthropod Collection" country="New Zealand" determinerName="B. P. Moore" location="Shingle Rd." municipality="Rd. Old Man Range" pageId="1" pageNumber="53" specimenCode="NZAC04002740" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="3CE3C51EFFA75F0EFF117D9BE460F964" bold="true" box="[151,310,1692,1718]" pageId="1" pageNumber="53">
<typeStatus id="D12CA7AEFFA75F0EFF117D9BE460F964" box="[151,310,1692,1718]" pageId="1" pageNumber="53" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
,
<specimenCount id="1891D285FFA75F0EFEC07D9AE4C1F964" box="[326,407,1692,1719]" count="1" pageId="1" pageNumber="53" type="male">1 male</specimenCount>
, in
<collectionCode id="688681C9FFA75F0EFE427D9AE745F965" box="[452,531,1693,1719]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="1" pageNumber="53" type="Other institutional type">NZAC</collectionCode>
, labeled: \
<collectingCountry id="7680599CFFA75F0EFD127D9AE67AF964" box="[660,812,1692,1719]" name="New Zealand" pageId="1" pageNumber="53">New Zealand</collectingCountry>
<collectorName id="A3627CDAFFA75F0EFCB07D9AE1F8F965" box="[822,1198,1692,1719]" pageId="1" pageNumber="53">
CO
<location id="0B484FD7FFA75F0EFCE07D9BE6BBF964" LSID="urn:lsid:plazi:treatment:863EA81AFFA75F0BFF117D30E6F6FE93:0B484FD7FFA75F0EFCE07D9BE6BBF964" box="[870,1005,1692,1718]" country="New Zealand" municipality="Rd. Old Man Range" name="Shingle Rd." pageId="1" pageNumber="53">
Shingle
<collectingMunicipality id="EE4C8376FFA75F0EFC437D9BE6BBF964" box="[965,1005,1692,1718]" pageId="1" pageNumber="53">Rd.</collectingMunicipality>
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Old Man Range
</collectorName>
<date id="7A293FCCFFA75F0EFB317D9BE01BF965" box="[1207,1357,1692,1719]" pageId="1" pageNumber="53" value="1975-03-15">
<collectingDate id="6A6DC624FFA75F0EFB317D9BE01BF965" box="[1207,1357,1692,1719]" pageId="1" pageNumber="53" value="1975-03-15">15 Mar 1975</collectingDate>
</date>
\ litter 75/127 \
<collectorName id="A3627CDAFFA75F0EFF7F7DC7E764F909" box="[249,562,1728,1755]" pageId="1" pageNumber="53">Entomology Division D.S.I.</collectorName>
<collectionCode id="688681C9FFA75F0EFDB57DC6E710F909" box="[563,582,1729,1755]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="53">R</collectionCode>
.
<collectingCountry id="7680599CFFA75F0EFDD47DC6E7B1F908" box="[594,743,1728,1755]" name="New Zealand" pageId="1" pageNumber="53">New Zealand</collectingCountry>
\
<taxonomicName id="C997628FFFA75F0EFD7A7DC7E147F909" authority="Broun" authorityName="Broun" box="[764,1041,1728,1755]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="3CE3C51EFFA75F0EFD7A7DC7E692F908" box="[764,964,1728,1754]" italics="true" pageId="1" pageNumber="53">Zeanillus pallidus</emphasis>
Broun
</taxonomicName>
det.
<determinerName id="FBF4F5C8FFA75F0EFBC07DC6E1E8F909" box="[1094,1214,1729,1755]" pageId="1" pageNumber="53">B.P.Moore</determinerName>
74 \ NZ
<collectorName id="A3627CDAFFA75F0EFAAE7DC7E45AF92C" pageId="1" pageNumber="53">Arthropod Collection</collectorName>
<specimenCode id="5E31B177FFA75F0EFE957DE2E499F92D" box="[275,463,1765,1791]" collectionCode="NZAC" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="1" pageNumber="53" type="Other institutional type">NZAC04002740</specimenCode>
</materialsCitation>
</paragraph>
<paragraph id="0E28190CFFA75F0EFF417C0FE140F8B9" blockId="1.[151,1437,1692,2043]" pageId="1" pageNumber="53">
<emphasis id="3CE3C51EFFA75F0EFF417C0FE43DF8F0" bold="true" box="[199,363,1800,1826]" pageId="1" pageNumber="53">
<typeStatus id="D12CA7AEFFA75F0EFF417C0FE43DF8F0" box="[199,363,1800,1826]" pageId="1" pageNumber="53" type="paratype">PARATYPES</typeStatus>
</emphasis>
(
<specimenCount id="1891D285FFA75F0EFEFC7C0EE75EF8F0" box="[378,520,1800,1827]" count="2" pageId="1" pageNumber="53" type="generic">2 specimens</specimenCount>
),
<specimenCount id="1891D285FFA75F0EFD9C7C0EE732F8F0" box="[538,612,1800,1827]" count="1" pageId="1" pageNumber="53" type="male">1 male</specimenCount>
<materialsCitation id="BEFF1351FFA75F0EFDEC7C0FE024F8F1" box="[618,1394,1800,1827]" collectorName="CO Old Man Rng &amp; Dec" country="New Zealand" location="New Zealand" pageId="1" pageNumber="53" specimenCount="1" typeStatus="paratype">
labeled: \
<collectingCountry id="7680599CFFA75F0EFD517C0EE63AF8F0" box="[727,876,1800,1827]" name="New Zealand" pageId="1" pageNumber="53">New Zealand</collectingCountry>
<collectorName id="A3627CDAFFA75F0EFCF47C0EE16EF8F1" box="[882,1080,1800,1827]" pageId="1" pageNumber="53">CO Old Man Rng</collectorName>
nr summit 31
<collectorName id="A3627CDAFFA75F0EFB517C0EE052F8F1" box="[1239,1284,1801,1827]" pageId="1" pageNumber="53">Dec</collectorName>
98 \ 2289
</materialsCitation>
\;
<specimenCount id="1891D285FFA75F0EFA087C0EE59BF894" count="1" pageId="1" pageNumber="53" type="male">1 male</specimenCount>
<materialsCitation id="BEFF1351FFA75F0EFF537C2BE157F8B9" collectingDate="2008-01-17" collectorName="Snow Farm &amp; Pisa Range &amp; J. Nunn &amp; Sokolov I. M." country="New Zealand" elevation="1520" location="Molecular" pageId="1" pageNumber="53" specimenCount="1" typeStatus="paratype">
labeled: \
<collectingCountry id="7680599CFFA75F0EFEC57C2AE488F894" box="[323,478,1836,1863]" name="New Zealand" pageId="1" pageNumber="53">New Zealand</collectingCountry>
: CO,
<collectorName id="A3627CDAFFA75F0EFD9B7C2AE7F4F895" box="[541,674,1837,1863]" pageId="1" pageNumber="53">Snow Farm</collectorName>
,
<collectorName id="A3627CDAFFA75F0EFD2B7C2BE67AF895" box="[685,812,1836,1863]" pageId="1" pageNumber="53">Pisa Range</collectorName>
,
<quantity id="C96FB4E9FFA75F0EFCBE7C2BE6D9F895" box="[824,911,1836,1863]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="1" pageNumber="53" unit="m" value="1520.0">
<elevation id="85BAFE3FFFA75F0EFCBE7C2BE6D9F895" box="[824,911,1836,1863]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="1" pageNumber="53" unit="m" value="1520.0">1520 m</elevation>
</quantity>
, moss and soil sample, subalpine tussock land, F41/001852
<collectorName id="A3627CDAFFA75F0EFEA07C56E420F8B9" box="[294,374,1873,1899]" pageId="1" pageNumber="53">J.Nunn</collectorName>
<date id="7A293FCCFFA75F0EFEFB7C56E752F8B9" box="[381,516,1873,1899]" pageId="1" pageNumber="53" value="2008-01-17">
<collectingDate id="6A6DC624FFA75F0EFEFB7C56E752F8B9" box="[381,516,1873,1899]" pageId="1" pageNumber="53" value="2008-01-17">17 Jan 2008</collectingDate>
</date>
\
<location id="0B484FD7FFA75F0EFD9C7C57E7DBF8B8" LSID="urn:lsid:plazi:treatment:863EA81AFFA75F0BFF117D30E6F6FE93:0B484FD7FFA75F0EFD9C7C57E7DBF8B8" box="[538,653,1872,1898]" country="New Zealand" name="Molecular" pageId="1" pageNumber="53">Molecular</location>
voucher # 90
<collectorName id="A3627CDAFFA75F0EFCAB7C57E694F8B9" box="[813,962,1872,1899]" pageId="1" pageNumber="53">Sokolov I.M.</collectorName>
2009
</materialsCitation>
\.
</paragraph>
<paragraph id="0E28190CFFA75F0EFF417C73E004F829" blockId="1.[151,1437,1692,2043]" pageId="1" pageNumber="53">
<emphasis id="3CE3C51EFFA75F0EFF417C73E4E9F85C" bold="true" box="[199,447,1908,1934]" pageId="1" pageNumber="53">Additional material:</emphasis>
<materialsCitation id="BEFF1351FFA75F0EFE4E7C72E617F85C" box="[456,833,1908,1935]" country="New Zealand" location="New Zealand" pageId="1" pageNumber="53" specimenCount="1" specimenCount-female="1">
<specimenCount id="1891D285FFA75F0EFE4E7C72E77AF85C" box="[456,556,1908,1935]" count="1" pageId="1" pageNumber="53" type="female">1 female</specimenCount>
labeled: \
<collectingCountry id="7680599CFFA75F0EFD2F7C72E617F85C" box="[681,833,1908,1935]" name="New Zealand" pageId="1" pageNumber="53">New Zealand</collectingCountry>
</materialsCitation>
<materialsCitation id="BEFF1351FFA75F0EFCCD7C72E652F861" collectorName="CO Waikaia Forest Whitecoomb Rd" country="New Zealand" county="Beech Forest" location="Washed" municipality="Apr" pageId="1" pageNumber="53" specimenCount="1">
<collectorName id="A3627CDAFFA75F0EFCCD7C72E1BFF85C" box="[843,1257,1908,1935]" pageId="1" pageNumber="53">CO Waikaia Forest Whitecoomb Rd</collectorName>
<locationDeviation id="1F9A002EFFA75F0EFB747C73E450F860" location="Post Office Ck" pageId="1" pageNumber="53">
<quantity id="C96FB4E9FFA75F0EFB747C73E072F85C" box="[1266,1316,1908,1934]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="53" unit="km" value="4.0">4km</quantity>
N of Post Office Ck
</locationDeviation>
/
<location id="0B484FD7FFA75F0EFE9F7C9FE427F860" LSID="urn:lsid:plazi:treatment:863EA81AFFA75F0BFF117D30E6F6FE93:0B484FD7FFA75F0EFE9F7C9FE427F860" box="[281,369,1944,1970]" country="New Zealand" county="Beech Forest" municipality="Apr" name="Washed" pageId="1" pageNumber="53">Washed</location>
soil sample 25
<collectingMunicipality id="EE4C8376FFA75F0EFD9B7C9EE71FF861" box="[541,585,1945,1971]" pageId="1" pageNumber="53">Apr</collectingMunicipality>
07
<collectingCounty id="E7496180FFA75F0EFDF77C9FE652F861" box="[625,772,1944,1971]" pageId="1" pageNumber="53">Beech Forest</collectingCounty>
</materialsCitation>
\;
<materialsCitation id="BEFF1351FFA75F0EFCA67C9EE1C2F860" box="[800,1172,1944,1971]" country="New Zealand" location="New Zealand" pageId="1" pageNumber="53" specimenCount="3" specimenCount-female="3">
<specimenCount id="1891D285FFA75F0EFCA67C9EE6DDF860" box="[800,907,1944,1971]" count="3" pageId="1" pageNumber="53" type="female">3 females</specimenCount>
labeled: \
<collectingCountry id="7680599CFFA75F0EFC797C9EE1C2F860" box="[1023,1172,1944,1971]" name="New Zealand" pageId="1" pageNumber="53">New Zealand</collectingCountry>
</materialsCitation>
<materialsCitation id="BEFF1351FFA75F0EFB1D7C9EE1EDF804" collectingDate="1974-11-20" collectionCode="R" collectorName="CO Old Woman Range &amp; Dugdale &amp; Entomology Division D. S. I." country="New Zealand" elevation="1389" location="New Zealand" pageId="1" pageNumber="53" specimenCount="1">
<collectorName id="A3627CDAFFA75F0EFB1D7C9EE0CBF861" box="[1179,1437,1944,1971]" pageId="1" pageNumber="53">CO Old Woman Range</collectorName>
<quantity id="C96FB4E9FFA75F0EFF117CBAE5B3F805" box="[151,229,1981,2007]" metricMagnitude="3" metricUnit="m" metricValue="1.389" pageId="1" pageNumber="53" unit="m" value="1389.0">
<elevation id="85BAFE3FFFA75F0EFF117CBAE5B3F805" box="[151,229,1981,2007]" metricMagnitude="3" metricUnit="m" metricValue="1.389" pageId="1" pageNumber="53" unit="m" value="1389.0">1389m</elevation>
</quantity>
<date id="7A293FCCFFA75F0EFF6B7CBAE4D6F805" box="[237,384,1981,2007]" pageId="1" pageNumber="53" value="1974-11-20">
<collectingDate id="6A6DC624FFA75F0EFF6B7CBAE4D6F805" box="[237,384,1981,2007]" pageId="1" pageNumber="53" value="1974-11-20">20 Nov 1974</collectingDate>
</date>
J.S.
<collectorName id="A3627CDAFFA75F0EFE2E7CBBE759F804" box="[424,527,1980,2006]" pageId="1" pageNumber="53">Dugdale</collectorName>
\ litter 74/100 \
<collectorName id="A3627CDAFFA75F0EFD417CBBE155F805" box="[711,1027,1980,2007]" pageId="1" pageNumber="53">Entomology Division D.S.I.</collectorName>
<collectionCode id="688681C9FFA75F0EFB857CBAE140F805" box="[1027,1046,1981,2007]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="53">R</collectionCode>
.
<collectingCountry id="7680599CFFA75F0EFBA37CBAE1EDF804" box="[1061,1211,1980,2007]" name="New Zealand" pageId="1" pageNumber="53">New Zealand</collectingCountry>
</materialsCitation>
\
<taxonomicName id="C997628FFFA75F0EFB547CBBE58BF829" authority="Broun" authorityName="Broun" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="3CE3C51EFFA75F0EFB547CBBE0CAF804" box="[1234,1436,1980,2006]" italics="true" pageId="1" pageNumber="53">Zeanillus pallidus</emphasis>
Broun
</taxonomicName>
det.
<determinerName id="FBF4F5C8FFA75F0EFE927CE6E4C2F829" box="[276,404,2017,2043]" pageId="1" pageNumber="53">B.P. Moore</determinerName>
\
<collectingCountry id="7680599CFFA75F0EFE2F7CE6E498F829" box="[425,462,2017,2043]" name="New Zealand" pageId="1" pageNumber="53">NZ</collectingCountry>
<materialsCitation id="BEFF1351FFA75F0EFE527CE7E06AF828" box="[468,1340,2016,2043]" collectionCode="NZAC" collectorName="Arthropod Collection" country="New Zealand" location="Private Bag" pageId="1" pageNumber="53" specimenCount="1" stateProvince="Auckland">
<collectorName id="A3627CDAFFA75F0EFE527CE7E791F828" box="[468,711,2016,2042]" pageId="1" pageNumber="53">Arthropod Collection</collectorName>
,
<collectionCode id="688681C9FFA75F0EFD547CE6E677F829" box="[722,801,2017,2043]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="1" pageNumber="53" type="Other institutional type">NZAC</collectionCode>
,
<location id="0B484FD7FFA75F0EFCAA7CE7E6E6F829" LSID="urn:lsid:plazi:treatment:863EA81AFFA75F0BFF117D30E6F6FE93:0B484FD7FFA75F0EFCAA7CE7E6E6F829" box="[812,944,2016,2043]" country="New Zealand" name="Private Bag" pageId="1" pageNumber="53" stateProvince="Auckland">Private Bag</location>
92170
<collectingRegion id="CC53D7EEFFA75F0EFB847CE6E1C9F829" box="[1026,1183,2017,2043]" country="New Zealand" name="Auckland" pageId="1" pageNumber="53">AUCKLAND</collectingRegion>
<collectingCountry id="7680599CFFA75F0EFB207CE6E06AF828" box="[1190,1340,2016,2043]" name="New Zealand" pageId="1" pageNumber="53">New Zealand</collectingCountry>
</materialsCitation>
\.
</paragraph>
</subSubSection>
<caption id="5AE84984FFA45F0DFF1178E3E737FB96" ID-DOI="http://doi.org/10.5281/zenodo.14612923" ID-Zenodo-Dep="14612923" httpUri="https://zenodo.org/record/14612923/files/figure.png" pageId="2" pageNumber="54" startId="2.[151,250,996,1020]" targetBox="[154,1411,194,961]" targetPageId="2" targetType="figure">
<paragraph id="0E28190CFFA45F0DFF1178E3E737FB96" blockId="2.[151,1437,996,1092]" pageId="2" pageNumber="54">
<emphasis id="3CE3C51EFFA45F0DFF1178E3E45BFC2E" bold="true" box="[151,269,996,1020]" pageId="2" pageNumber="54">FIGURE 1</emphasis>
. Digital habitus images of blind
<taxonomicName id="C997628FFFA45F0DFDD178E3E666FC2E" box="[599,816,996,1020]" pageId="2" pageNumber="54">
<emphasis id="3CE3C51EFFA45F0DFDD178E3E788FC2E" box="[599,734,996,1020]" italics="true" pageId="2" pageNumber="54">Nesamblyops</emphasis>
species
</taxonomicName>
, dorsal aspect.
<emphasis id="3CE3C51EFFA45F0DFC4F78E3E68DFC2E" bold="true" box="[969,987,996,1020]" pageId="2" pageNumber="54">A</emphasis>
<taxonomicName id="C997628FFFA45F0DFC7378E2E11AFC2E" authorityName="Sokolov" authorityYear="2025" box="[1013,1100,997,1020]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="3CE3C51EFFA45F0DFC7378E2E11AFC2E" box="[1013,1100,997,1020]" italics="true" pageId="2" pageNumber="54">N. nunni</emphasis>
</taxonomicName>
(Old Man Range, Central Otago, South Island),
<emphasis id="3CE3C51EFFA45F0DFEA87F0FE469FBF2" bold="true" box="[302,319,1032,1056]" pageId="2" pageNumber="54">B</emphasis>
<taxonomicName id="C997628FFFA45F0DFEDF7F0EE4B2FBF2" authorityName="Sokolov" authorityYear="2025" box="[345,484,1032,1056]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="larochellei">
<emphasis id="3CE3C51EFFA45F0DFEDF7F0EE4B2FBF2" box="[345,484,1032,1056]" italics="true" pageId="2" pageNumber="54">N. larochellei</emphasis>
</taxonomicName>
(Lewis Pass, Buller, South Island),
<emphasis id="3CE3C51EFFA45F0DFCDE7F0FE63CFBF2" bold="true" box="[856,874,1032,1056]" pageId="2" pageNumber="54">C</emphasis>
<taxonomicName id="C997628FFFA45F0DFC027F0EE15EFBF2" authorityName="Sokolov" authorityYear="2025" box="[900,1032,1033,1056]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="3CE3C51EFFA45F0DFC027F0EE15EFBF2" box="[900,1032,1033,1056]" italics="true" pageId="2" pageNumber="54">N. pygmaeus</emphasis>
</taxonomicName>
(Waiotauru Road, Tararua Forest Park, Wellington, North Island). Scale bar 1.0 mm.
</paragraph>
</caption>
<caption id="5AE84984FFA45F0DFF117EDAE7CDF9EF" ID-DOI="http://doi.org/10.5281/zenodo.14612925" ID-Zenodo-Dep="14612925" httpUri="https://zenodo.org/record/14612925/files/figure.png" pageId="2" pageNumber="54" startId="2.[151,250,1501,1525]" targetBox="[156,1432,1118,1473]" targetPageId="2" targetType="figure">
<paragraph id="0E28190CFFA45F0DFF117EDAE7CDF9EF" blockId="2.[151,1437,1501,1597]" pageId="2" pageNumber="54">
<emphasis id="3CE3C51EFFA45F0DFF117EDAE443FA27" bold="true" box="[151,277,1501,1525]" pageId="2" pageNumber="54">FIGURE 2.</emphasis>
Digital images of pronota of blind
<taxonomicName id="C997628FFFA45F0DFD0D7EDAE630FA27" box="[651,870,1501,1525]" pageId="2" pageNumber="54">
<emphasis id="3CE3C51EFFA45F0DFD0D7EDAE644FA27" box="[651,786,1501,1525]" italics="true" pageId="2" pageNumber="54">Nesamblyops</emphasis>
species
</taxonomicName>
, dorsal aspect.
<emphasis id="3CE3C51EFFA45F0DFB817EDAE14FFA27" bold="true" box="[1031,1049,1501,1525]" pageId="2" pageNumber="54">A</emphasis>
<taxonomicName id="C997628FFFA45F0DFBB47ED9E1DAFA27" authorityName="Sokolov" authorityYear="2025" box="[1074,1164,1502,1525]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="3CE3C51EFFA45F0DFBB47ED9E1DAFA27" box="[1074,1164,1502,1525]" italics="true" pageId="2" pageNumber="54">N. nunni</emphasis>
</taxonomicName>
(Old Man Range, Central Otago, South Island),
<emphasis id="3CE3C51EFFA45F0DFEF17D06E4DEF9CB" bold="true" box="[375,392,1537,1561]" pageId="2" pageNumber="54">B</emphasis>
<taxonomicName id="C997628FFFA45F0DFE277D05E77AF9CB" authorityName="Sokolov" authorityYear="2025" box="[417,556,1537,1561]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="larochellei">
<emphasis id="3CE3C51EFFA45F0DFE277D05E77AF9CB" box="[417,556,1537,1561]" italics="true" pageId="2" pageNumber="54">N. larochellei</emphasis>
</taxonomicName>
(Lewis Pass, Buller, South Island),
<emphasis id="3CE3C51EFFA45F0DFC1C7D06E6FAF9CB" bold="true" box="[922,940,1537,1561]" pageId="2" pageNumber="54">C</emphasis>
<taxonomicName id="C997628FFFA45F0DFC407D05E11EF9CB" authorityName="Sokolov" authorityYear="2025" box="[966,1096,1538,1561]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="3CE3C51EFFA45F0DFC407D05E11EF9CB" box="[966,1096,1538,1561]" italics="true" pageId="2" pageNumber="54">N. pygmaeus</emphasis>
</taxonomicName>
(Waiotauru Road, Tararua Forest Park, Wellington, North Island). Scale bar 0.1 mm.
</paragraph>
</caption>
<subSubSection id="468D4A87FFA45F0DFF417D88E7C4F91F" pageId="2" pageNumber="54" type="etymology">
<paragraph id="0E28190CFFA45F0DFF417D88E7C4F91F" blockId="2.[151,1436,1679,2030]" pageId="2" pageNumber="54">
<emphasis id="3CE3C51EFFA45F0DFF417D88E418F97B" bold="true" box="[199,334,1679,1705]" pageId="2" pageNumber="54">Etymology.</emphasis>
The specific epithet is a Latinized eponym in the genitive case and is based on the surname of John T. Nunn, one of the collectors of this species.
</paragraph>
</subSubSection>
<subSubSection id="468D4A87FFA45F0DFF417DD0E73FF88F" pageId="2" pageNumber="54" type="materials_examined">
<paragraph id="0E28190CFFA45F0DFF417DD0E110F920" blockId="2.[151,1436,1679,2030]" box="[199,1094,1751,1778]" pageId="2" pageNumber="54">
<emphasis id="3CE3C51EFFA45F0DFF417DD0E430F923" bold="true" box="[199,358,1751,1777]" pageId="2" pageNumber="54">Type locality.</emphasis>
<collectingCountry id="7680599CFFA45F0DFEEA7DDFE753F923" box="[364,517,1751,1778]" name="New Zealand" pageId="2" pageNumber="54">New Zealand</collectingCountry>
, South Island, CO, Central
<collectingRegion id="CC53D7EEFFA45F0DFCBF7DD0E6D6F923" box="[825,896,1751,1777]" country="New Zealand" name="Otago" pageId="2" pageNumber="54">Otago</collectingRegion>
, Old Man Range.
</paragraph>
<paragraph id="0E28190CFFA45F0DFF417DFCE73FF88F" blockId="2.[151,1436,1679,2030]" pageId="2" pageNumber="54">
<emphasis id="3CE3C51EFFA45F0DFF417DFCE40BF8C7" bold="true" box="[199,349,1787,1813]" pageId="2" pageNumber="54">Recognition.</emphasis>
Adults of this species (
<figureCitation id="96AC0589FFA45F0DFDE57DFCE7ECF8C7" box="[611,698,1787,1813]" captionStart="FIGURE 1" captionStartId="2.[151,250,996,1020]" captionTargetBox="[154,1411,194,961]" captionTargetId="figure-15@2.[151,1436,181,972]" captionTargetPageId="2" captionText="FIGURE 1. Digital habitus images of blind Nesamblyops species, dorsal aspect. A—N. nunni (Old Man Range, Central Otago, South Island), B—N. larochellei (Lewis Pass, Buller, South Island), C—N. pygmaeus (Waiotauru Road, Tararua Forest Park, Wellington, North Island). Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14612923" httpUri="https://zenodo.org/record/14612923/files/figure.png" pageId="2" pageNumber="54">Fig. 1A</figureCitation>
) can be distinguished from the adults of other blind
<taxonomicName id="C997628FFFA45F0DFA817DFCE5BEF8EB" pageId="2" pageNumber="54">
<emphasis id="3CE3C51EFFA45F0DFA817DFCE0CAF8C7" box="[1287,1436,1787,1813]" italics="true" pageId="2" pageNumber="54">Nesamblyops</emphasis>
species
</taxonomicName>
described in this revision by the absence of microsculpture on the frons of the head and disc of the pronotum, and by the structure of the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="468D4A87FFA45F0BFF417C60E6F6FE93" lastPageId="4" lastPageNumber="56" pageId="2" pageNumber="54" type="description">
<paragraph id="0E28190CFFA45F0DFF417C60E1D7F853" blockId="2.[151,1436,1679,2030]" box="[199,1153,1895,1922]" pageId="2" pageNumber="54">
<emphasis id="3CE3C51EFFA45F0DFF417C60E40FF853" bold="true" box="[199,345,1895,1921]" pageId="2" pageNumber="54">Description.</emphasis>
Large for genus (SBL range
<quantity id="C96FB4E9FFA45F0DFD197C60E612F850" box="[671,836,1895,1922]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.99" metricValueMin="1.71" pageId="2" pageNumber="54" unit="mm" value="1.85" valueMax="1.99" valueMin="1.71">1.711.99 mm</quantity>
, mean 1.86±
<quantity id="C96FB4E9FFA45F0DFC557C60E16FF850" box="[979,1081,1895,1922]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="54" unit="mm" value="0.11">0.11 mm</quantity>
, n=6).
</paragraph>
<paragraph id="0E28190CFFA45F0DFF417C8CE10CF83F" blockId="2.[151,1436,1679,2030]" pageId="2" pageNumber="54">
<emphasis id="3CE3C51EFFA45F0DFF417C8CE471F877" box="[199,295,1931,1957]" italics="true" pageId="2" pageNumber="54">Habitus.</emphasis>
Body form (
<figureCitation id="96AC0589FFA45F0DFE3B7C8CE743F877" box="[445,533,1931,1957]" captionStart="FIGURE 1" captionStartId="2.[151,250,996,1020]" captionTargetBox="[154,1411,194,961]" captionTargetId="figure-15@2.[151,1436,181,972]" captionTargetPageId="2" captionText="FIGURE 1. Digital habitus images of blind Nesamblyops species, dorsal aspect. A—N. nunni (Old Man Range, Central Otago, South Island), B—N. larochellei (Lewis Pass, Buller, South Island), C—N. pygmaeus (Waiotauru Road, Tararua Forest Park, Wellington, North Island). Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14612923" httpUri="https://zenodo.org/record/14612923/files/figure.png" pageId="2" pageNumber="54">Fig. 1A</figureCitation>
) slightly convex, elongate ovoid, general proportions moderately narrow (WE/ SBL 0.38±0.011), head moderately wide relative to pronotum (WH/WPm 0.76±0.017), elytra slightly wide relative to pronotum in comparison with other blind species of genus (WPm/WE 0.73±0.021).
</paragraph>
<caption id="5AE84984FFA55F0CFF117893E028FBEE" ID-DOI="http://doi.org/10.5281/zenodo.14612927" ID-Zenodo-Dep="14612927" httpUri="https://zenodo.org/record/14612927/files/figure.png" pageId="3" pageNumber="55" startId="3.[151,250,916,940]" targetBox="[157,1430,187,886]" targetPageId="3" targetType="figure">
<paragraph id="0E28190CFFA55F0CFF117893E028FBEE" blockId="3.[151,1437,916,1084]" pageId="3" pageNumber="55">
<emphasis id="3CE3C51EFFA55F0CFF117893E442FC7E" bold="true" box="[151,276,916,940]" pageId="3" pageNumber="55">FIGURE 3.</emphasis>
Line drawings of male genitalia of blind
<taxonomicName id="C997628FFFA55F0CFD457893E6F4FC7E" box="[707,930,916,940]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="55" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="3CE3C51EFFA55F0CFD457893E61CFC7E" box="[707,842,916,940]" italics="true" pageId="3" pageNumber="55">Nesamblyops</emphasis>
species.
</taxonomicName>
<taxonomicName id="C997628FFFA55F0CFC2E7892E157FC7E" authorityName="Sokolov" authorityYear="2025" box="[936,1025,917,940]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="55" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="3CE3C51EFFA55F0CFC2E7892E157FC7E" box="[936,1025,917,940]" italics="true" pageId="3" pageNumber="55">N. nunni</emphasis>
</taxonomicName>
(Old Man Range, Central Otago, South Island):
<emphasis id="3CE3C51EFFA55F0CFF6A78BFE5A8FC02" bold="true" box="[236,254,952,976]" pageId="3" pageNumber="55">A</emphasis>
—left paramere, left lateral aspect,
<emphasis id="3CE3C51EFFA55F0CFDEC78BFE72DFC02" bold="true" box="[618,635,952,976]" pageId="3" pageNumber="55">B</emphasis>
—right paramere, right lateral aspect,
<emphasis id="3CE3C51EFFA55F0CFB8578BFE143FC02" bold="true" box="[1027,1045,952,976]" pageId="3" pageNumber="55">C</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="C997628FFFA55F0CFA0378BEE452FC26" authorityName="Sokolov" authorityYear="2025" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="55" phylum="Arthropoda" rank="species" species="larochellei">
<emphasis id="3CE3C51EFFA55F0CFA0378BEE452FC26" italics="true" pageId="3" pageNumber="55">N. larochellei</emphasis>
</taxonomicName>
(Lewis Pass, Buller, South Island):
<emphasis id="3CE3C51EFFA55F0CFD0678DBE7C4FC26" bold="true" box="[640,658,988,1012]" pageId="3" pageNumber="55">D</emphasis>
—left paramere, left lateral aspect,
<emphasis id="3CE3C51EFFA55F0CFB8378DBE140FC26" bold="true" box="[1029,1046,988,1012]" pageId="3" pageNumber="55">E</emphasis>
—right paramere, right lateral aspect,
<emphasis id="3CE3C51EFFA55F0CFF117F07E5F0FBCA" bold="true" box="[151,166,1024,1048]" pageId="3" pageNumber="55">F</emphasis>
—median lobe, right lateral aspect.
<taxonomicName id="C997628FFFA55F0CFD9E7F06E7CAFBCA" authorityName="Sokolov" authorityYear="2025" box="[536,668,1025,1048]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="55" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="3CE3C51EFFA55F0CFD9E7F06E7CAFBCA" box="[536,668,1025,1048]" italics="true" pageId="3" pageNumber="55">N. pygmaeus</emphasis>
</taxonomicName>
(Waiotauru Road, Tararua Forest Park, Wellington, North Island):
<emphasis id="3CE3C51EFFA55F0CFACB7F07E037FBCA" bold="true" box="[1357,1377,1024,1048]" pageId="3" pageNumber="55">G</emphasis>
—left paramere, left lateral aspect,
<emphasis id="3CE3C51EFFA55F0CFE3A7F23E486FBEE" bold="true" box="[444,464,1060,1084]" pageId="3" pageNumber="55">H</emphasis>
—right paramere, right lateral aspect,
<emphasis id="3CE3C51EFFA55F0CFCD47F23E60AFBEE" bold="true" box="[850,860,1060,1084]" pageId="3" pageNumber="55">I</emphasis>
—median lobe, right lateral aspect. Scale bar 0.1 mm.
</paragraph>
</caption>
<paragraph id="0E28190CFFA55F0CFF417F89E61BFB7B" blockId="3.[151,1437,1166,2021]" box="[199,845,1166,1193]" pageId="3" pageNumber="55">
<emphasis id="3CE3C51EFFA55F0CFF417F89E45BFB7A" box="[199,269,1166,1192]" italics="true" pageId="3" pageNumber="55">Color.</emphasis>
Body color brunneorufous, appendages testaceous.
</paragraph>
<paragraph id="0E28190CFFA55F0CFF417FB5E038FB22" blockId="3.[151,1437,1166,2021]" pageId="3" pageNumber="55">
<emphasis id="3CE3C51EFFA55F0CFF417FB5E42FFB1E" box="[199,377,1202,1228]" italics="true" pageId="3" pageNumber="55">Microsculpture.</emphasis>
Mesh pattern of irregularly isodiametric sculpticells distinctly present over dorsal surface of labrum, clypeal area and vertex of head. Frons and disc of pronotum smooth (without evident microsculpture).
</paragraph>
<paragraph id="0E28190CFFA55F0CFF417FFDE459FA76" blockId="3.[151,1437,1166,2021]" pageId="3" pageNumber="55">
<emphasis id="3CE3C51EFFA55F0CFF417FFDE46FFAC6" box="[199,313,1274,1300]" italics="true" pageId="3" pageNumber="55">Prothorax</emphasis>
. Pronotum (
<figureCitation id="96AC0589FFA55F0CFE4C7FFDE772FAC6" box="[458,548,1274,1300]" captionStart="FIGURE 2" captionStartId="2.[151,250,1501,1525]" captionTargetBox="[156,1432,1118,1473]" captionTargetId="figure-76@2.[151,1436,1116,1477]" captionTargetPageId="2" captionText="FIGURE 2. Digital images of pronota of blind Nesamblyops species, dorsal aspect. A—N. nunni (Old Man Range, Central Otago, South Island), B—N. larochellei (Lewis Pass, Buller, South Island), C—N. pygmaeus (Waiotauru Road, Tararua Forest Park, Wellington, North Island). Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14612925" httpUri="https://zenodo.org/record/14612925/files/figure.png" pageId="3" pageNumber="55">Fig. 2A</figureCitation>
) moderately long in comparison to elytra (LP/LE 0.40±0.01) and moderately transverse (WPm/LP 1.19±0.03), with lateral margins rectilinear and markedly constricted posteriorly (WPm/WPp 1.45±0.08). Anterior angles indistinct, posterior angles strongly obtuse (136144°), almost completely rounded. Width between anterior angles slightly greater than between posterior angles (WPa/WPp 1.09±0.07). Basal margin rectilinear.
</paragraph>
<paragraph id="0E28190CFFA55F0CFF417EA9E63FF9C2" blockId="3.[151,1437,1166,2021]" pageId="3" pageNumber="55">
<emphasis id="3CE3C51EFFA55F0CFF417EA9E443FA1A" box="[199,277,1454,1480]" italics="true" pageId="3" pageNumber="55">Elytra.</emphasis>
Slightly ovoid, widely depressed along suture, comparatively long (LE/SBL 0.59±0.01) and moderately narrow (WE/LE 0.64±0.02). Humeri completely rounded. Lateral margins slightly divergent at basal fourth, subparallel at middle and evenly rounded to apex in apical third.
</paragraph>
<paragraph id="0E28190CFFA55F0CFF417D1DE1AEF8E2" blockId="3.[151,1437,1166,2021]" pageId="3" pageNumber="55">
<emphasis id="3CE3C51EFFA55F0CFF417D1DE422F9E6" box="[199,372,1562,1588]" italics="true" pageId="3" pageNumber="55">Male genitalia.</emphasis>
Median lobe of aedeagus (
<figureCitation id="96AC0589FFA55F0CFD207D1DE7AAF9E6" box="[678,764,1562,1588]" captionStart="FIGURE 3" captionStartId="3.[151,250,916,940]" captionTargetBox="[157,1430,187,886]" captionTargetId="figure-37@3.[151,1435,181,892]" captionTargetPageId="3" captionText="FIGURE 3. Line drawings of male genitalia of blind Nesamblyops species. N. nunni (Old Man Range, Central Otago, South Island): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. larochellei (Lewis Pass, Buller, South Island): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. pygmaeus (Waiotauru Road, Tararua Forest Park, Wellington, North Island): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14612927" httpUri="https://zenodo.org/record/14612927/files/figure.png" pageId="3" pageNumber="55">Fig. 3C</figureCitation>
) moderately arcuate and slightly twisted. Shaft subparallel, abruptly tapering in apically. Apex small, triangular, with narrowly rounded tip. Apical orifice of moderate length, occupies apical third of the shaft length. Ventral margin of median lobe almost straight. Walls of shaft without of poriferous canals. Copulatory sclerites united in one moderately sclerotized plate, occupying basal third of the shaft length. Presumably, sclerotized parts of dorsal V-contour and medial rC-sclerite merged together to form a sclerotized contour bordering the plate basally and slightly protruding from basal orifice. Left paramere (
<figureCitation id="96AC0589FFA55F0CFABD7DC9E0C2F93A" box="[1339,1428,1742,1768]" captionStart="FIGURE 3" captionStartId="3.[151,250,916,940]" captionTargetBox="[157,1430,187,886]" captionTargetId="figure-37@3.[151,1435,181,892]" captionTargetPageId="3" captionText="FIGURE 3. Line drawings of male genitalia of blind Nesamblyops species. N. nunni (Old Man Range, Central Otago, South Island): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. larochellei (Lewis Pass, Buller, South Island): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. pygmaeus (Waiotauru Road, Tararua Forest Park, Wellington, North Island): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14612927" httpUri="https://zenodo.org/record/14612927/files/figure.png" pageId="3" pageNumber="55">Fig. 3A</figureCitation>
) short, with non-attenuated apex, bearing one long and one short setae. Right paramere (
<figureCitation id="96AC0589FFA55F0CFBEC7DF5E196F8DE" box="[1130,1216,1778,1804]" captionStart="FIGURE 3" captionStartId="3.[151,250,916,940]" captionTargetBox="[157,1430,187,886]" captionTargetId="figure-37@3.[151,1435,181,892]" captionTargetPageId="3" captionText="FIGURE 3. Line drawings of male genitalia of blind Nesamblyops species. N. nunni (Old Man Range, Central Otago, South Island): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. larochellei (Lewis Pass, Buller, South Island): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. pygmaeus (Waiotauru Road, Tararua Forest Park, Wellington, North Island): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14612927" httpUri="https://zenodo.org/record/14612927/files/figure.png" pageId="3" pageNumber="55">Fig. 3B</figureCitation>
) short, of moderate width, bearing two long setae, which are shorter the length of paramere. Ring sclerite not examined.
</paragraph>
<paragraph id="0E28190CFFA55F0CFF417C3DE67CF886" blockId="3.[151,1437,1166,2021]" box="[199,810,1850,1877]" pageId="3" pageNumber="55">
<emphasis id="3CE3C51EFFA55F0CFF417C3DE4BBF886" box="[199,493,1850,1876]" italics="true" pageId="3" pageNumber="55">Female internal genitalia.</emphasis>
Spermatheca not examined.
</paragraph>
<paragraph id="0E28190CFFA55F0CFF417C59E5A7F812" blockId="3.[151,1437,1166,2021]" pageId="3" pageNumber="55">
<emphasis id="3CE3C51EFFA55F0CFF417C59E75FF8AA" bold="true" box="[199,521,1886,1912]" pageId="3" pageNumber="55">Geographical distribution.</emphasis>
The range of the species is the southern part of the South Island, where it is restricted to the ridges of Central
<collectingRegion id="CC53D7EEFFA55F0CFD8B7C85E707F84E" box="[525,593,1922,1948]" country="New Zealand" name="Otago" pageId="3" pageNumber="55">Otago</collectingRegion>
adjacent to the west of the Clutha River in its middle course (
<figureCitation id="96AC0589FFA55F0CFA877C85E013F84E" box="[1281,1349,1922,1948]" captionStart="FIGURE 4" captionStartId="5.[151,250,1648,1672]" captionTargetBox="[151,1436,181,1624]" captionTargetId="figure-37@5.[151,1436,181,1624]" captionTargetPageId="5" captionText="FIGURE 4. Distribution records for the blind Nesamblyops species and images of the male median lobe obtained from the collected specimens (inset illustrates specimen localities). Black circle—N. pygmaeus; red circle—N. larochellei; orange circles—N. nunni. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14612929" httpUri="https://zenodo.org/record/14612929/files/figure.png" pageId="3" pageNumber="55">Fig. 4</figureCitation>
, orange circles).
</paragraph>
<paragraph id="0E28190CFFA55F0BFF417CCDE0D5FF63" blockId="3.[151,1437,1166,2021]" lastBlockId="4.[151,1437,151,321]" lastPageId="4" lastPageNumber="56" pageId="3" pageNumber="55">
<emphasis id="3CE3C51EFFA55F0CFF417CCDE47CF836" bold="true" box="[199,298,1994,2020]" pageId="3" pageNumber="55">Habitat.</emphasis>
Specimens were collected using a washing technique from moss and soil samples, some specimens bear label “litter” without details (as plant communities labels mention beech forest and subalpine tussock land).
</paragraph>
<paragraph id="0E28190CFFA25F0BFF417BBDE6F6FE93" blockId="4.[151,1437,151,321]" pageId="4" pageNumber="56">
<emphasis id="3CE3C51EFFA25F0BFF417BBDE426FF06" bold="true" box="[199,368,186,212]" pageId="4" pageNumber="56">Relationships.</emphasis>
The merged copulatory sclerites of peculiar shape of the male genitalia suggest an isolated position of
<taxonomicName id="C997628FFFA25F0BFE9F7BD8E42CFF2A" authorityName="Sokolov" authorityYear="2025" box="[281,378,223,248]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="3CE3C51EFFA25F0BFE9F7BD8E42CFF2A" box="[281,378,223,248]" italics="true" pageId="4" pageNumber="56">N. nunni</emphasis>
</taxonomicName>
within the genus and only remote relatedness to the other species of
<taxonomicName id="C997628FFFA25F0BFB0D7BD8E076FF2B" authorityName=", Jeannel" authorityYear="1937" box="[1163,1312,223,249]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="3CE3C51EFFA25F0BFB0D7BD8E076FF2B" box="[1163,1312,223,249]" italics="true" pageId="4" pageNumber="56">Nesamblyops</emphasis>
</taxonomicName>
. The same details of the configuration of copulative sclerites indicate that
<taxonomicName id="C997628FFFA25F0BFCD57A04E6E5FECE" authorityName="Sokolov" authorityYear="2025" box="[851,947,259,284]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="nunni">
<emphasis id="3CE3C51EFFA25F0BFCD57A04E6E5FECE" box="[851,947,259,284]" italics="true" pageId="4" pageNumber="56">N. nunni</emphasis>
</taxonomicName>
belongs to a separate lineage in comparison with the other blind representatives of
<taxonomicName id="C997628FFFA25F0BFDC17A20E78AFE93" authorityName=", Jeannel" authorityYear="1937" box="[583,732,295,321]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="3CE3C51EFFA25F0BFDC17A20E78AFE93" box="[583,732,295,321]" italics="true" pageId="4" pageNumber="56">Nesamblyops</emphasis>
</taxonomicName>
described below.
</paragraph>
</subSubSection>
</treatment>
</document>