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<document id="9C4AE4F6853421043A8CDB34B06E835E" ID-DOI="10.35463/j.apr.2019.02.01" ID-ISSN="1842-371x" ID-Zenodo-Dep="10520652" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1705449792158" checkinUser="felipe" docAuthor="SCHLAGINTWEIT, FELIX, RASHIDI, KOOROSH, YARAHMADZAHI, HAMED, HABIBIMOOD, SHARAM, AMIRSHAHKARMI, MAHNAZ, AHMADI, HOSSAIN &amp; KHOKAN, HOSSAIN" docDate="2019" docId="E10087A34B61D056FF26EB09FE7CFBB1" docLanguage="en" docName="ActaPalaeontolRom.15.2.3-13.pdf" docOrigin="Acta Palaeontologica Romaniae 15 (2)" docSource="http://dx.doi.org/10.35463/j.apr.2019.02.01" docStyle="DocumentStyle:CA474AAD92E97C533D63E0C5C0F159F1.1:ActaPalaeontolRom.2019-.journal_article" docStyleId="CA474AAD92E97C533D63E0C5C0F159F1" docStyleName="ActaPalaeontolRom.2019-.journal_article" docStyleVersion="1" docTitle="Dissocladella chahtorshiana Rashidi &amp; Schlagintweit" docType="treatment" docVersion="3" lastPageNumber="9" masterDocId="1D39FFDB4B65D051FFB1EF1EFFA0FF9C" masterDocTitle="Dissocladella? Chahtorshiana Rashidi &amp; Schlagintweit N. Sp., A New Dasycladale (Green Algae) From The Paleocene Of Iran" masterLastPageNumber="13" masterPageNumber="3" pageNumber="7" updateTime="1706815965324" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9F226AD84DA940133CEBDF30C18AAB10">Dissocladella? Chahtorshiana Rashidi &amp; Schlagintweit N. Sp., A New Dasycladale (Green Algae) From The Paleocene Of Iran</mods:title>
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<mods:namePart id="42550A041695AEA83E4DF0D17B3D811C">SCHLAGINTWEIT, FELIX</mods:namePart>
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<mods:namePart id="644D58F56BFF0D4C762ABCA52A1DCC67">RASHIDI, KOOROSH</mods:namePart>
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<mods:namePart id="1183041727A110D29D050FED5DFC03CC">YARAHMADZAHI, HAMED</mods:namePart>
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<mods:namePart id="8E8C055B0616A1946F7F96F88BE6564F">AMIRSHAHKARMI, MAHNAZ</mods:namePart>
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<mods:namePart id="5350B29D802BA2D830419244892F9B86">AHMADI, HOSSAIN</mods:namePart>
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<taxonomicName id="AEA94D364B61D055FF26EB09FE76FBB3" ID-CoL="46XC" authority="Rashidi &amp; Schlagintweit" authorityName="Rashidi &amp; Schlagintweit" box="[151,470,1047,1071]" class="Ulvophyceae" family="Triploporellaceae" genus="Dissocladella" kingdom="Plantae" order="Dasycladales" pageId="4" pageNumber="7" phylum="Chlorophyta" rank="species" species="chahtorshiana">
<emphasis id="5BDDEAA74B61D055FF26EB09FE89FBB3" box="[151,297,1047,1071]" italics="true" pageId="4" pageNumber="7">Dissocladella</emphasis>
? 
<emphasis id="5BDDEAA74B61D055FE8DEB09FE76FBB3" box="[316,470,1047,1071]" italics="true" pageId="4" pageNumber="7">chahtorshiana</emphasis>
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n. sp.
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<paragraph id="691636B54B61D055FF26EB28FEDCFBD2" blockId="4.[151,772,1078,2053]" box="[151,380,1078,1102]" pageId="4" pageNumber="7">
<figureCitation id="F1922A304B61D055FF26EB28FEB3FBD1" box="[151,275,1078,1102]" captionStart="Fig" captionStartId="5.[243,278,1748,1769]" captionTargetBox="[243,1419,154,1711]" captionTargetId="figure-112@5.[243,1419,154,1711]" captionTargetPageId="5" captionText="Fig. 5 Typical microfacies of samples with Dissocladella? chahtorshiana n. sp. (D), from the Paleocene of the Kuh-e- Chah Torsh (a–c) and Kuh-e-Patorgi (d-e) sections. a–b Bioclastic packstone with dasycladalean (e.g., Acroporella? anceps Segonzac, A), halimedacean (ha) algae, gastropods, and porcelaneous benthic foraminifera (e.g., soritoidids). c Bioclastic packstone with dasycladalean algae (e.g., Clypeina elliotti Beckmann &amp; Beckmann, Cl). d Bioclastic packstone with dasycladaleans (e.g., Rostroporella oviformis Segonzac, R). Thin-sections: Ah 185 (a–b), CT 4 (c), 2pz29 (d). e Weathered rock surface with thallus fragments of Dissocladella? chahtorshiana n. sp. The maximum observed length amounts for ~ 17.5 mm (arrow). Kuh-e-Patorgi area." figureDoi="http://doi.org/10.5281/zenodo.10520662" httpUri="https://zenodo.org/record/10520662/files/figure.png" pageId="4" pageNumber="7">Figs. 5A–D</figureCitation>
pars, 6–8
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<emphasis id="5BDDEAA74B61D055FF26EB4BFEDAFBF1" bold="true" box="[151,378,1109,1133]" pageId="4" pageNumber="7">Origin of the name:</emphasis>
The name refers to the type-locality Mount Chah Torsh, Central 
<collectingCountry id="11BE76254B61D055FE75EB6AFE4FFB17" box="[452,495,1140,1163]" name="Iran" pageId="4" pageNumber="7">Iran</collectingCountry>
(
<figureCitation id="F1922A304B61D055FE4FEB6DFDC5FB17" box="[510,613,1139,1163]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[347,382,824,845]" captionStartId-1="1.[258,293,1578,1599]" captionStartId-2="3.[227,262,1906,1927]" captionTargetBox-0="[350,1240,167,788]" captionTargetBox-1="[258,1329,945,1541]" captionTargetBox-2="[227,1360,210,1869]" captionTargetId-0="figure-516@1.[347,1240,167,790]" captionTargetId-1="figure-614@1.[258,1329,945,1541]" captionTargetId-2="figure-52@3.[227,1360,210,1869]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionTargetPageId-2="3" captionText-0="Fig. 1 Tentative location of the Kuh-e-Chah Torsh (1) and Kuh-e-Patorgi (2) sections plot- ted on the tectonic map of Iran (modified from Zanchi et al., 2009). Abbreviations: AA = Anatolian–Armenian Block, AMC = Anarak Metamorphic Complex, KDF = Kopeh Dagh Foredeep, MZT = Main Zagros Trust)." captionText-1="Fig. 2 Location (1) and field view (2) of Kuh-e-Chah Torsh, the type-locality of Dissocladella? chahtorshiana n. sp. For the position within the territory of Iran see figure 1. Larger Benthic Foraminifera in (2): Dictyoconus? pachymarginalis Schroeder (?late Bedoulian-Gargasian?, a), Orbitoides gensacicus (Leymerie) (late Maastrichtian, b), Sistanites iranicus Rahaghi (Selandian, c), Orduella sphaerica Sirel (Selandian, d)." captionText-2="Fig. 3 Simplified lithostratigraphic column of the Kuh-e-Chah Torsh section with position of samples and distribution of benthic foraminifera and calcareous algae. Inferred SBZ zones after Serra-Kiel et al. (1998), Sirel (2012), and Hottinger (2014) (modified from Schlagintweit &amp; Rashidi, 2019, in press.). ." figureDoi-0="http://doi.org/10.5281/zenodo.10520654" figureDoi-1="http://doi.org/10.5281/zenodo.10520656" figureDoi-2="http://doi.org/10.5281/zenodo.10520658" httpUri-0="https://zenodo.org/record/10520654/files/figure.png" httpUri-1="https://zenodo.org/record/10520656/files/figure.png" httpUri-2="https://zenodo.org/record/10520658/files/figure.png" pageId="4" pageNumber="7">Figs. 1–3</figureCitation>
).
</paragraph>
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<paragraph id="691636B54B61D055FF26EB8CFE86FB54" blockId="4.[151,772,1078,2053]" pageId="4" pageNumber="7">
<materialsCitation id="D9C13CE84B61D055FF26EB8CFE86FB54" collectionCode="CT" pageId="4" pageNumber="7" specimenCode="CT 4" specimenCount="1" typeStatus="holotype">
<emphasis id="5BDDEAA74B61D055FF26EB8CFEA8FB36" bold="true" box="[151,264,1170,1194]" pageId="4" pageNumber="7">
<typeStatus id="B61288174B61D055FF26EB8CFEA3FB36" box="[151,259,1170,1194]" pageId="4" pageNumber="7" type="holotype">Holotype</typeStatus>
:
</emphasis>
Oblique section shown in 
<figureCitation id="F1922A304B61D055FDF8EB8CFD1FFB36" box="[585,703,1170,1194]" captionStart="Fig" captionStartId="9.[226,261,1669,1690]" captionTargetBox="[226,1361,186,1633]" captionTargetId="figure-296@9.[226,1361,186,1633]" captionTargetPageId="9" captionText="Fig. 8 Dissocladella? chahtorshiana n. sp. from Selandian carbonates of Kuh-e-Chah Torsh, Central Iran. a–f, variously oriented oblique sections. Note the claviform thallus in d, and the presence of short secondaries visible in a (right side) or f (upper side). The holotype is shown in c. Thin-sections: CT 4 (a–d, f), Ah 185 (e)." figureDoi="http://doi.org/10.5281/zenodo.10520668" httpUri="https://zenodo.org/record/10520668/files/figure.png" pageId="4" pageNumber="7">Figure 8C</figureCitation>
, thin-section 
<specimenCode id="390F9ECE4B61D055FF58EBAEFE82FB54" box="[233,290,1200,1224]" collectionCode="CT" country="South Africa" lsid="urn:lsid:biocol.org:col:14492" name="University of Cape Town" pageId="4" pageNumber="7" type="Herbarium">CT 4</specimenCode>
.
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</paragraph>
<paragraph id="691636B54B61D055FF26EBD1FE34FAB9" blockId="4.[151,772,1078,2053]" pageId="4" pageNumber="7">
<emphasis id="5BDDEAA74B61D055FF26EBD1FE2CFB7B" bold="true" box="[151,396,1231,1255]" pageId="4" pageNumber="7">Horizon and locality:</emphasis>
Selandian carbonates from Mount Chah Torsh, lithostratigraphically not defined (= no formation name assigned).
</paragraph>
</subSubSection>
<subSubSection id="21B3653E4B61D056FF26EA32FE7CFBB1" lastPageId="7" lastPageNumber="10" pageId="4" pageNumber="7" type="description">
<paragraph id="691636B54B61D055FF26EA32FB5CFB17" blockId="4.[151,772,1078,2053]" lastBlockId="4.[816,1437,1047,2053]" pageId="4" pageNumber="7">
<emphasis id="5BDDEAA74B61D055FF26EA32FE84FAD8" bold="true" box="[151,292,1324,1348]" pageId="4" pageNumber="7">Description:</emphasis>
<collectionCode id="0FB8AE704B61D055FE9AEA35FE9EFADF" box="[299,318,1323,1347]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="7" type="Herbarium">A</collectionCode>
medium-sized, well calcified Dasycladale with a cylindrical to slightly clavate thallus (
<figureCitation id="F1922A304B61D055FD39EA54FD5EFAFE" box="[648,766,1354,1378]" captionStart="Fig" captionStartId="8.[205,240,1742,1763]" captionTargetBox="[205,1383,162,1706]" captionTargetId="figure-56@8.[205,1383,162,1706]" captionTargetPageId="8" captionText="Fig. 7 Dissocladella? chahtorshiana n. sp. from Selandian carbonates of Kuh-e-Chah Torsh, Central Iran. a–b, longitudinal (upper part) and tangential (lower part) sections. Primaries are arranged in alternating position between successive verticils. In shallow tangential sections they are close-set and display rounded transverse outline. In deep tangential section (b, middle part) the transverse sections are rounded triangular. c–h, detailed views of the primaries, showing short secondaries. Thin-sections: Ah 185 (a–b), CT 4 (c–h)." figureDoi="http://doi.org/10.5281/zenodo.10520666" httpUri="https://zenodo.org/record/10520666/files/figure.png" pageId="4" pageNumber="7">Fig. 7A–B</figureCitation>
, 
<figureCitation id="F1922A304B61D055FF26EA76FF4FFA1C" box="[151,239,1384,1408]" captionStart="Fig" captionStartId="9.[226,261,1669,1690]" captionTargetBox="[226,1361,186,1633]" captionTargetId="figure-296@9.[226,1361,186,1633]" captionTargetPageId="9" captionText="Fig. 8 Dissocladella? chahtorshiana n. sp. from Selandian carbonates of Kuh-e-Chah Torsh, Central Iran. a–f, variously oriented oblique sections. Note the claviform thallus in d, and the presence of short secondaries visible in a (right side) or f (upper side). The holotype is shown in c. Thin-sections: CT 4 (a–d, f), Ah 185 (e)." figureDoi="http://doi.org/10.5281/zenodo.10520668" httpUri="https://zenodo.org/record/10520668/files/figure.png" pageId="4" pageNumber="7">Fig. 8D</figureCitation>
). The large main axis of roughly two third of diameter is bordered by a rather smooth surface except common widenings of the proximal part of the laterals, assumed of secondary origin (
<figureCitation id="F1922A304B61D055FE6DEADBFDEFFA41" box="[476,591,1477,1501]" captionStart="Fig" captionStartId="6.[213,248,1585,1606]" captionTargetBox="[213,1402,154,1549]" captionTargetId="figure-338@6.[213,1402,154,1549]" captionTargetPageId="6" captionText="Fig. 6 Dissocladella? chahtorshiana n. sp. from Selandian carbonates of Kuh-e-Chah Torsh, Central Iran. a–f, transverse sections, partly slightly oblique. Note the different morphologies of the primaries, only slightly widening (a–b), or clubshaped (e–f). Secondary laterals partly visible in c, e, and f. g, oblique section. Thin-sections: Ah 185 (a, c-d), CT 4 (b, e-g)." figureDoi="http://doi.org/10.5281/zenodo.10520664" httpUri="https://zenodo.org/record/10520664/files/figure.png" pageId="4" pageNumber="7">Fig. 6A, F</figureCitation>
) (compare Pl. 1, 
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Fig. 
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<bibRefCitation id="0D384B444B61D055FF4CEAFDFE40FA67" author="Radoicic, R. &amp; Conrad, M. A. &amp; Carras, N." box="[253,480,1507,1531]" pageId="4" pageNumber="7" pagination="311 - 317" refId="ref5543" refString="Radoicic, R., Conrad, M. A., Carras, N., 2005. Observations on Neomeris cretacea var. undulata Raineri followed by Dissocladella bonardii, n. sp. (Dasycladales, green algae). Revue de Paleobiologie, 24 (1): 311 - 317." type="journal article" year="2005">Radoičic et al., 2005</bibRefCitation>
). The primariy laterals are numerous and regularly alternating between consecutive verticils. In shallow tangential sections they are close-set and of round outline (
<figureCitation id="F1922A304B61D055FE24E921FDAFF9CB" box="[405,527,1599,1623]" captionStart="Fig" captionStartId="8.[205,240,1742,1763]" captionTargetBox="[205,1383,162,1706]" captionTargetId="figure-56@8.[205,1383,162,1706]" captionTargetPageId="8" captionText="Fig. 7 Dissocladella? chahtorshiana n. sp. from Selandian carbonates of Kuh-e-Chah Torsh, Central Iran. a–b, longitudinal (upper part) and tangential (lower part) sections. Primaries are arranged in alternating position between successive verticils. In shallow tangential sections they are close-set and display rounded transverse outline. In deep tangential section (b, middle part) the transverse sections are rounded triangular. c–h, detailed views of the primaries, showing short secondaries. Thin-sections: Ah 185 (a–b), CT 4 (c–h)." figureDoi="http://doi.org/10.5281/zenodo.10520666" httpUri="https://zenodo.org/record/10520666/files/figure.png" pageId="4" pageNumber="7">Fig. 7A–B</figureCitation>
, lower part). In deep tangential sections they appear of triangular outline (with rounded edges) (
<figureCitation id="F1922A304B61D055FEFCE963FE02F908" box="[333,418,1661,1685]" captionStart="Fig" captionStartId="8.[205,240,1742,1763]" captionTargetBox="[205,1383,162,1706]" captionTargetId="figure-56@8.[205,1383,162,1706]" captionTargetPageId="8" captionText="Fig. 7 Dissocladella? chahtorshiana n. sp. from Selandian carbonates of Kuh-e-Chah Torsh, Central Iran. a–b, longitudinal (upper part) and tangential (lower part) sections. Primaries are arranged in alternating position between successive verticils. In shallow tangential sections they are close-set and display rounded transverse outline. In deep tangential section (b, middle part) the transverse sections are rounded triangular. c–h, detailed views of the primaries, showing short secondaries. Thin-sections: Ah 185 (a–b), CT 4 (c–h)." figureDoi="http://doi.org/10.5281/zenodo.10520666" httpUri="https://zenodo.org/record/10520666/files/figure.png" pageId="4" pageNumber="7">Fig. 7B</figureCitation>
, middle part). The primaries are usually arranged perpendicular to the axis in cylindrical morphologies (
<figureCitation id="F1922A304B61D055FE8AE9A4FE31F94E" box="[315,401,1722,1746]" captionStart="Fig" captionStartId="8.[205,240,1742,1763]" captionTargetBox="[205,1383,162,1706]" captionTargetId="figure-56@8.[205,1383,162,1706]" captionTargetPageId="8" captionText="Fig. 7 Dissocladella? chahtorshiana n. sp. from Selandian carbonates of Kuh-e-Chah Torsh, Central Iran. a–b, longitudinal (upper part) and tangential (lower part) sections. Primaries are arranged in alternating position between successive verticils. In shallow tangential sections they are close-set and display rounded transverse outline. In deep tangential section (b, middle part) the transverse sections are rounded triangular. c–h, detailed views of the primaries, showing short secondaries. Thin-sections: Ah 185 (a–b), CT 4 (c–h)." figureDoi="http://doi.org/10.5281/zenodo.10520666" httpUri="https://zenodo.org/record/10520666/files/figure.png" pageId="4" pageNumber="7">Fig. 7B</figureCitation>
); in slightly claviform morphologies they are slightly inclined upwards (
<figureCitation id="F1922A304B61D055FDD4E9C6FD61F96C" box="[613,705,1752,1776]" captionStart="Fig" captionStartId="9.[226,261,1669,1690]" captionTargetBox="[226,1361,186,1633]" captionTargetId="figure-296@9.[226,1361,186,1633]" captionTargetPageId="9" captionText="Fig. 8 Dissocladella? chahtorshiana n. sp. from Selandian carbonates of Kuh-e-Chah Torsh, Central Iran. a–f, variously oriented oblique sections. Note the claviform thallus in d, and the presence of short secondaries visible in a (right side) or f (upper side). The holotype is shown in c. Thin-sections: CT 4 (a–d, f), Ah 185 (e)." figureDoi="http://doi.org/10.5281/zenodo.10520668" httpUri="https://zenodo.org/record/10520668/files/figure.png" pageId="4" pageNumber="7">Fig. 8D</figureCitation>
). The shape of the primaries is variable: slightly widening outwards and without detectable secondaries (
<figureCitation id="F1922A304B61D055FDD8E808FD7CF8B2" box="[617,732,1814,1838]" captionStart="Fig" captionStartId="6.[213,248,1585,1606]" captionTargetBox="[213,1402,154,1549]" captionTargetId="figure-338@6.[213,1402,154,1549]" captionTargetPageId="6" captionText="Fig. 6 Dissocladella? chahtorshiana n. sp. from Selandian carbonates of Kuh-e-Chah Torsh, Central Iran. a–f, transverse sections, partly slightly oblique. Note the different morphologies of the primaries, only slightly widening (a–b), or clubshaped (e–f). Secondary laterals partly visible in c, e, and f. g, oblique section. Thin-sections: Ah 185 (a, c-d), CT 4 (b, e-g)." figureDoi="http://doi.org/10.5281/zenodo.10520664" httpUri="https://zenodo.org/record/10520664/files/figure.png" pageId="4" pageNumber="7">Fig. 6A–B</figureCitation>
) or with a distinct swelling (club-shaped morphology) and secondaries (
<figureCitation id="F1922A304B61D055FE97E84DFE39F8F7" box="[294,409,1875,1899]" captionStart="Fig" captionStartId="6.[213,248,1585,1606]" captionTargetBox="[213,1402,154,1549]" captionTargetId="figure-338@6.[213,1402,154,1549]" captionTargetPageId="6" captionText="Fig. 6 Dissocladella? chahtorshiana n. sp. from Selandian carbonates of Kuh-e-Chah Torsh, Central Iran. a–f, transverse sections, partly slightly oblique. Note the different morphologies of the primaries, only slightly widening (a–b), or clubshaped (e–f). Secondary laterals partly visible in c, e, and f. g, oblique section. Thin-sections: Ah 185 (a, c-d), CT 4 (b, e-g)." figureDoi="http://doi.org/10.5281/zenodo.10520664" httpUri="https://zenodo.org/record/10520664/files/figure.png" pageId="4" pageNumber="7">Fig. 6E–F</figureCitation>
). Sometimes the swollen part of the primaries are almost touching (or do so) with neighbouring ones. The different morphologies might refer to different parts of the thallus (sterile? fertile?). In transverse or longitudinal sections, two secondaries distinctly shorter than the primaries and widening distally are detectable (
<figureCitation id="F1922A304B61D055FC25EB09FBE7FBB3" box="[916,1095,1047,1071]" captionStart="Fig" captionStartId="8.[205,240,1742,1763]" captionTargetBox="[205,1383,162,1706]" captionTargetId="figure-56@8.[205,1383,162,1706]" captionTargetPageId="8" captionText="Fig. 7 Dissocladella? chahtorshiana n. sp. from Selandian carbonates of Kuh-e-Chah Torsh, Central Iran. a–b, longitudinal (upper part) and tangential (lower part) sections. Primaries are arranged in alternating position between successive verticils. In shallow tangential sections they are close-set and display rounded transverse outline. In deep tangential section (b, middle part) the transverse sections are rounded triangular. c–h, detailed views of the primaries, showing short secondaries. Thin-sections: Ah 185 (a–b), CT 4 (c–h)." figureDoi="http://doi.org/10.5281/zenodo.10520666" httpUri="https://zenodo.org/record/10520666/files/figure.png" pageId="4" pageNumber="7">Fig. 7C–D, E–G</figureCitation>
). The exact number of secondaries per primary however lateral is unknown. Often they appear as a united micritic diverging masse upon the distal surface of the primaries (
<figureCitation id="F1922A304B61D055FBECEB6DFB4CFB17" box="[1117,1260,1139,1163]" captionStart="Fig" captionStartId="8.[205,240,1742,1763]" captionTargetBox="[205,1383,162,1706]" captionTargetId="figure-56@8.[205,1383,162,1706]" captionTargetPageId="8" captionText="Fig. 7 Dissocladella? chahtorshiana n. sp. from Selandian carbonates of Kuh-e-Chah Torsh, Central Iran. a–b, longitudinal (upper part) and tangential (lower part) sections. Primaries are arranged in alternating position between successive verticils. In shallow tangential sections they are close-set and display rounded transverse outline. In deep tangential section (b, middle part) the transverse sections are rounded triangular. c–h, detailed views of the primaries, showing short secondaries. Thin-sections: Ah 185 (a–b), CT 4 (c–h)." figureDoi="http://doi.org/10.5281/zenodo.10520666" httpUri="https://zenodo.org/record/10520666/files/figure.png" pageId="4" pageNumber="7">Fig. 7C, E, G</figureCitation>
).
</paragraph>
<paragraph id="691636B54B61D055FC81EB8CFC47FB7B" blockId="4.[816,1437,1047,2053]" pageId="4" pageNumber="7">
<emphasis id="5BDDEAA74B61D055FC81EB8CFC71FB36" bold="true" box="[816,977,1170,1194]" pageId="4" pageNumber="7">Comparisons:</emphasis>
From the Paleogene, the following species of 
<taxonomicName id="AEA94D364B61D055FC3FEBAEFB80FB54" box="[910,1056,1200,1224]" class="Ulvophyceae" family="Triploporellaceae" genus="Dissocladella" kingdom="Plantae" order="Dasycladales" pageId="4" pageNumber="7" phylum="Chlorophyta" rank="genus">
<emphasis id="5BDDEAA74B61D055FC3FEBAEFB80FB54" box="[910,1056,1200,1224]" italics="true" pageId="4" pageNumber="7">Dissocladella</emphasis>
</taxonomicName>
have been described (data from 
<bibRefCitation id="0D384B444B61D055FC81EBD1FC7CFB7B" author="Barattolo, F." box="[816,988,1231,1255]" pageId="4" pageNumber="7" pagination="17 - 40" refId="ref4454" refString="Barattolo, F., 2002. Late Cretaceous-Paleogene Dasycladaleans and the K / T boundary problem. In: Bucur, I. I., Filipescu, S. (eds.), Research advances in calcareous algae and microbial carbonates, Proceedings of the 4 th IFAA Regional Meeting, Cluj Napoca, Romania, 29 August- 5 September 2001, pp. 17 - 40." type="proceedings paper" year="2002">Barattolo, 2002</bibRefCitation>
):
</paragraph>
<paragraph id="691636B54B61D055FC81EBF1FB8CFAFE" blockId="4.[816,1437,1047,2053]" pageId="4" pageNumber="7">
<taxonomicName id="AEA94D364B61D055FC81EBF1FC4FFA9A" authority="Elliott" authorityName="Elliott" box="[816,1007,1262,1286]" class="Ulvophyceae" family="Triploporellaceae" genus="Dissocladella" kingdom="Plantae" order="Dasycladales" pageId="4" pageNumber="7" phylum="Chlorophyta" rank="species" species="deserta">
<emphasis id="5BDDEAA74B61D055FC81EBF1FC02FA9A" box="[816,930,1262,1286]" italics="true" pageId="4" pageNumber="7">D. deserta</emphasis>
Elliott
</taxonomicName>
(Selandian-Ypresian): 
<taxonomicName id="AEA94D364B61D055FB5CEBF1FAFFFA9A" box="[1261,1375,1262,1286]" class="Ulvophyceae" family="Triploporellaceae" genus="Dissocladella" kingdom="Plantae" order="Dasycladales" pageId="4" pageNumber="7" phylum="Chlorophyta" rank="species" species="deserta">
<emphasis id="5BDDEAA74B61D055FB5CEBF1FAFFFA9A" box="[1261,1375,1262,1286]" italics="true" pageId="4" pageNumber="7">D. deserta</emphasis>
</taxonomicName>
from the Paleocene of 
<collectingCountry id="11BE76254B61D055FC59EA13FBB5FAB8" box="[1000,1045,1293,1316]" name="Iraq" pageId="4" pageNumber="7">Iraq</collectingCountry>
, and lower Eocene of 
<collectingCountry id="11BE76254B61D055FAB2EA13FAE3FAB9" box="[1283,1347,1293,1317]" name="Egypt" pageId="4" pageNumber="7">Egypt</collectingCountry>
(
<bibRefCitation id="0D384B444B61D055FAE0EA13FCC7FADF" author="Elliott, G. F." pageId="4" pageNumber="7" pagination="1 - 111" refId="ref4740" refString="Elliott, G. F., 1968. Permian to Palaeocene calcareous algae (Dasycladaceae) of the Middle East. Bulletin of the British Museum Natural History (Geology) Supplement, 4, 1 - 111." type="journal article" year="1968">Elliott, 1968</bibRefCitation>
) is a very small species (D = 
<quantity id="AE519B504B61D055FB73EA35FAFFFADF" box="[1218,1375,1323,1347]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" metricValueMax="4.699999999999999" metricValueMin="3.9" pageId="4" pageNumber="7" unit="mm" value="0.43" valueMax="0.47" valueMin="0.39">0.39-0.47 mm</quantity>
), and about twelve primaries.
</paragraph>
<paragraph id="691636B54B61D055FC81EA77FCDFFA41" blockId="4.[816,1437,1047,2053]" pageId="4" pageNumber="7">
<taxonomicName id="AEA94D364B61D055FC81EA77FBB5FA1C" authority="Radoicic" authorityName="Radoicic" box="[816,1045,1384,1408]" class="Ulvophyceae" family="Triploporellaceae" genus="Dissocladella" kingdom="Plantae" order="Dasycladales" pageId="4" pageNumber="7" phylum="Chlorophyta" rank="species" species="gracilis">
<emphasis id="5BDDEAA74B61D055FC81EA77FC08FA1C" box="[816,936,1384,1408]" italics="true" pageId="4" pageNumber="7">D. gracilis</emphasis>
Radoičić
</taxonomicName>
(Thanetian of 
<collectingCountry id="11BE76254B61D055FB77EA76FA86FA1C" box="[1222,1318,1384,1408]" name="Slovenia" pageId="4" pageNumber="7">Slovenia</collectingCountry>
, 
<bibRefCitation id="0D384B444B61D055FA86EA76FCC5FA03" author="Radoicic, R." pageId="4" pageNumber="7" pagination="309 - 313" refId="ref5477" refString="Radoicic, R., 1991. Dissocladella gracilis n. sp. (Dasycladales) from the Thanetian of Majevica mountain (NE Bosnia, Dinarides). Revue de Paleobiologie, 10: 309 - 313." type="journal article" year="1991">Radoičić, 1991</bibRefCitation>
): as name already implies, it represents a very small alga (D: 
<quantity id="AE519B504B61D055FC22EAB8FB9CFA22" box="[915,1084,1446,1470]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" metricValueMax="2.9" metricValueMin="1.6" pageId="4" pageNumber="7" unit="mm" value="0.22499999999999998" valueMax="0.29" valueMin="0.16">0.16-0.290 mm</quantity>
, max. 
<quantity id="AE519B504B61D055FB36EAB8FB57FA22" box="[1159,1271,1446,1470]" metricMagnitude="-4" metricUnit="m" metricValue="3.6799999999999997" pageId="4" pageNumber="7" unit="mm" value="0.368">0.368 mm</quantity>
) with four primaries.
</paragraph>
<paragraph id="691636B54B61D055FC81EAFAFC07F9EA" blockId="4.[816,1437,1047,2053]" pageId="4" pageNumber="7">
<taxonomicName id="AEA94D364B61D055FC81EAFAFB29FA60" authority="Mu &amp; Wang" authorityName="Mu &amp; Wang" authorityYear="1985" box="[816,1161,1507,1532]" class="Ulvophyceae" family="Triploporellaceae" genus="Dissocladella" kingdom="Plantae" order="Dasycladales" pageId="4" pageNumber="7" phylum="Chlorophyta" rank="species" species="longjangensis">
<emphasis id="5BDDEAA74B61D055FC81EAFAFC4BFA67" box="[816,1003,1507,1531]" italics="true" pageId="4" pageNumber="7">D. longjangensis</emphasis>
Mu &amp; Wang
</taxonomicName>
(Lutetian of 
<collectingCountry id="11BE76254B61D055FA94EAFDFAC7FA67" box="[1317,1383,1507,1531]" name="China" pageId="4" pageNumber="7">China</collectingCountry>
, 
<bibRefCitation id="0D384B444B61D055FAC9EAFAFC49F986" author="Mu, X. &amp; Wang, Y. - J." pageId="4" pageNumber="7" pagination="289 - 296" refId="ref5357" refString="Mu, X., Wang, Y. - J., 1985. Some calcareous algae from the Eocene of Tingri, Xizang, China. Acta Micropaleontologica Sinica, 2: 289 - 296." type="journal article" year="1985">Mu and Wang, 1985</bibRefCitation>
): medium-sized species (D up to 
<quantity id="AE519B504B61D055FADFE91CFCFCF9A4" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="7" unit="mm" value="0.65">0.65 mm</quantity>
, see 
<bibRefCitation id="0D384B444B61D055FC2BE93FFB02F9A4" author="Kuss, J. &amp; Herbig, H. - G." box="[922,1186,1568,1592]" pageId="4" pageNumber="7" pagination="249 - 280" refId="ref5222" refString="Kuss, J., Herbig, H. - G., 1993. Biogeograpgy, facies and taxonomy of Early Tertiary green algae from Egypt and Morocco. Bolletino della Societa Paleontologica Italiana, Spec. vol. 1: 249 - 280." type="journal article" year="1993">Kuss and Herbig, 1993</bibRefCitation>
, Paleogene of 
<collectingCountry id="11BE76254B61D055FAFCE93EFA30F9A4" box="[1357,1424,1568,1592]" name="Egypt" pageId="4" pageNumber="7">Egypt</collectingCountry>
), and a main axis occupying about half the diameter (d/D 0.50-0.55).
</paragraph>
<paragraph id="691636B54B61D055FC81E963FBB1F94E" blockId="4.[816,1437,1047,2053]" pageId="4" pageNumber="7">
<taxonomicName id="AEA94D364B61D055FC81E963FBB6F909" authority="Segonzac" authorityName="Segonzac" box="[816,1046,1660,1685]" class="Ulvophyceae" family="Triploporellaceae" genus="Dissocladella" kingdom="Plantae" order="Dasycladales" pageId="4" pageNumber="7" phylum="Chlorophyta" rank="species" species="lunata">
<emphasis id="5BDDEAA74B61D055FC81E963FC3FF908" box="[816,927,1660,1684]" italics="true" pageId="4" pageNumber="7">D. lunata</emphasis>
Segonzac
</taxonomicName>
(
<location id="6C76606E4B61D055FB9CE963FB39F909" LSID="urn:lsid:plazi:treatment:E10087A34B61D056FF26EB09FE7CFBB1:6C76606E4B61D055FB9CE963FB39F909" box="[1069,1177,1661,1685]" country="France" latitude="0.282" longLatPrecision="78" longitude="-0.415" name="Thanetian" pageId="4" pageNumber="7">Thanetian</location>
of 
<collectingCountry id="11BE76254B61D055FB61E963FABBF908" box="[1232,1307,1661,1684]" name="France" pageId="4" pageNumber="7">France</collectingCountry>
, 
<bibRefCitation id="0D384B444B61D055FA81E963FCC5F92F" author="Segonzac, G." pageId="4" pageNumber="7" pagination="439 - 463" refId="ref6020" refString="Segonzac, G., 1979. Algues calacires du Thanetien d'Eseranza (Aude) (Dasyckadacees, Corallinacees). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 115 (3 - 4): 439 - 463." type="journal article" year="1979">Segonzac, 1979</bibRefCitation>
): small sized species (D: 
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<geoCoordinate id="0C9D50724B61D055FB10E985FB7DF92F" box="[1185,1245,1691,1715]" degrees="0.282" direction="north" orientation="latitude" pageId="4" pageNumber="7" precision="55" value="0.282">0.282</geoCoordinate>
<geoCoordinate id="0C9D50724B61D055FB6CE982FA82F92F" box="[1245,1314,1691,1715]" degrees="0.415" direction="west" orientation="longitude" pageId="4" pageNumber="7" precision="55" value="-0.415">-0.415</geoCoordinate>
mm
</quantity>
) with four to six primaries.
</paragraph>
<paragraph id="691636B54B61D055FC81E9C7FA27F8B2" blockId="4.[816,1437,1047,2053]" pageId="4" pageNumber="7">
<taxonomicName id="AEA94D364B61D055FC81E9C7FC7DF96C" authority="Pia" authorityName="Pia" authorityYear="1936" box="[816,989,1752,1776]" class="Ulvophyceae" family="Triploporellaceae" genus="Dissocladella" kingdom="Plantae" order="Dasycladales" pageId="4" pageNumber="7" phylum="Chlorophyta" rank="species" species="savitriae">
<emphasis id="5BDDEAA74B61D055FC81E9C7FC11F96C" box="[816,945,1753,1776]" italics="true" pageId="4" pageNumber="7">D. savitriae</emphasis>
Pia
</taxonomicName>
(Danian-Thanetian): large-sized and annulated representative of the genus (D: 1.5-2.0 mm) with more than 40 primaries per verticil (
<bibRefCitation id="0D384B444B61D055FB00E808FADBF8B2" author="Rao, L. R. &amp; Pia, J." box="[1201,1403,1814,1838]" pageId="4" pageNumber="7" pagination="1 - 49" refId="ref5692" refString="Rao, L. R., Pia, J., 1936. Fossil algae from the uppermost Cretaceous beds (the Niniyur Group) of the Trichinopoly district, S. India. Mem Geol Surv India, Pal Indica, N. S. 21 (4): 1 - 49." type="journal article" year="1936">Rao and Pia, 1936</bibRefCitation>
).
</paragraph>
<paragraph id="691636B54B61D055FC81E828FB31F834" blockId="4.[816,1437,1047,2053]" pageId="4" pageNumber="7">
<taxonomicName id="AEA94D364B61D055FC81E828FB8EF8D1" authority="Radoicic" authorityName="Radoicic" box="[816,1070,1845,1869]" class="Ulvophyceae" family="Triploporellaceae" genus="Dissocladella" kingdom="Plantae" order="Dasycladales" pageId="4" pageNumber="7" phylum="Chlorophyta" rank="species" species="turnsekae">
<emphasis id="5BDDEAA74B61D055FC81E828FC61F8D1" box="[816,961,1845,1869]" italics="true" pageId="4" pageNumber="7">D. turnsekae</emphasis>
Radoičić
</taxonomicName>
(Danian-Selandian of 
<collectingCountry id="11BE76254B61D055FA88E82BFA39F8D1" box="[1337,1433,1845,1869]" name="Slovenia" pageId="4" pageNumber="7">Slovenia</collectingCountry>
, 
<bibRefCitation id="0D384B444B61D055FC81E84DFC76F8F7" author="Radoicic, R." box="[816,982,1875,1899]" pageId="4" pageNumber="7" pagination="103 - 121" refId="ref5516" refString="Radoicic, R., 1998. Two new Paleocene Dasycladales from Slovenia. Annales Geologique de la Peninsule Balkanique, 62: 103 - 121." type="journal article" year="1998">Radoičić, 1998</bibRefCitation>
): it represents a very small species (D = 
<quantity id="AE519B504B61D055FC81E86CFC6CF816" box="[816,972,1906,1930]" metricMagnitude="-4" metricUnit="m" metricValue="2.9499999999999997" metricValueMax="3.5" metricValueMin="2.4" pageId="4" pageNumber="7" unit="mm" value="0.295" valueMax="0.35" valueMin="0.24">0.24-0.35 mm</quantity>
) with about eight primaries aligned vertically between successive whorls.
</paragraph>
<paragraph id="691636B54B61D057FC81E8AEFEF8F82C" blockId="4.[816,1437,1047,2053]" lastBlockId="6.[151,771,1730,2060]" lastPageId="6" lastPageNumber="9" pageId="4" pageNumber="7">
We may also note some resemblances of the lateral morphology with the poorly known 
<taxonomicName id="AEA94D364B61D055FB2FE8D0FACFF87A" box="[1182,1391,1998,2022]" class="Ulvophyceae" family="Dasycladaceae" genus="Jodotella" kingdom="Plantae" order="Dasycladales" pageId="4" pageNumber="7" phylum="Chlorophyta" rank="species" species="volpensis">
<emphasis id="5BDDEAA74B61D055FB2FE8D0FACFF87A" box="[1182,1391,1998,2022]" italics="true" pageId="4" pageNumber="7">Jodotella volpensis</emphasis>
</taxonomicName>
described by 
<bibRefCitation id="0D384B444B61D055FC02E8F3FBCEF799" author="Segonzac, G." box="[947,1134,2029,2053]" pageId="4" pageNumber="7" pagination="123 - 136" refId="ref5983" refString="Segonzac, G., 1976. Dasycladacees nouvelles ou peu connues du Thanetian des Pyrenees. Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112 (1 - 2): 123 - 136." type="journal article" year="1976">Segonzac (1976)</bibRefCitation>
with two sections from the Thanetian of 
<collectingCountry id="11BE76254B63D057FE85E9DCFE20F945" box="[308,384,1730,1753]" name="France" pageId="6" pageNumber="9">France</collectingCountry>
. The transverse section shown in plate 3, fig. 1, shows primaries that are widened proximally, displaying distinct swelling in the middle part, and two secondaries. This alga is distinctly smaller, displays a much lower d/D ratio (about 0.38), a reduced number of laterals (w = 16). 
<taxonomicName id="AEA94D364B63D057FEEFE845FE61F8EF" authorityName="L. Morellet &amp; J. Morellet" authorityYear="1913" box="[350,449,1883,1907]" class="Ulvophyceae" family="Dasycladaceae" genus="Jodotella" kingdom="Plantae" order="Dasycladales" pageId="6" pageNumber="9" phylum="Chlorophyta" rank="genus">
<emphasis id="5BDDEAA74B63D057FEEFE845FE61F8EF" box="[350,449,1883,1907]" italics="true" pageId="6" pageNumber="9">Jodotella</emphasis>
</taxonomicName>
is different by its fertile ampullae set in clusters laterally to the primaries (
<bibRefCitation id="0D384B444B63D057FD17E864FEEDF82C" author="Deloffre, R. &amp; Genot, P." pageId="6" pageNumber="9" pagination="1 - 247" refId="ref4615" refString="Deloffre, R., Genot, P., 1982. Les Algues Dasycladales du Cenozoique. Bulletin des Centres de Recherches Exploration-Production Elf-Aqutitaine, Mem. 4: 1 - 247." type="journal article" year="1982">Deloffre and Génot, 1982</bibRefCitation>
).
</paragraph>
<caption id="3DD6663D4B60D054FF42E9CAFD4DF813" ID-DOI="http://doi.org/10.5281/zenodo.10520662" ID-Zenodo-Dep="10520662" httpUri="https://zenodo.org/record/10520662/files/figure.png" pageId="5" pageNumber="8" startId="5.[243,278,1748,1769]" targetBox="[243,1419,154,1711]" targetPageId="5" targetType="figure">
<paragraph id="691636B54B60D054FF42E9CAFD4DF813" blockId="5.[243,1421,1748,1935]" pageId="5" pageNumber="8">
<emphasis id="5BDDEAA74B60D054FF42E9CAFE8FF975" bold="true" box="[243,303,1748,1769]" pageId="5" pageNumber="8">Fig. 5</emphasis>
Typical microfacies of samples with 
<taxonomicName id="AEA94D364B60D054FD16E9CAFC8BF976" box="[679,811,1748,1770]" class="Ulvophyceae" family="Triploporellaceae" genus="Dissocladella" kingdom="Plantae" order="Dasycladales" pageId="5" pageNumber="8" phylum="Chlorophyta" rank="genus">
<emphasis id="5BDDEAA74B60D054FD16E9CAFC8BF976" box="[679,811,1748,1770]" italics="true" pageId="5" pageNumber="8">Dissocladella</emphasis>
</taxonomicName>
? 
<emphasis id="5BDDEAA74B60D054FC8FE9CAFC69F976" box="[830,969,1748,1770]" italics="true" pageId="5" pageNumber="8">chahtorshiana</emphasis>
n. sp. (
<emphasis id="5BDDEAA74B60D054FBA9E9CBFB89F976" box="[1048,1065,1749,1770]" italics="true" pageId="5" pageNumber="8">D</emphasis>
), from the Paleocene of the Kuh-e- Chah Torsh (
<emphasis id="5BDDEAA74B60D054FEC4E9EEFE38F899" bold="true" box="[373,408,1776,1797]" pageId="5" pageNumber="8">a–c</emphasis>
) and Kuh-e-Patorgi (
<emphasis id="5BDDEAA74B60D054FDD8E9EEFD29F899" bold="true" box="[617,649,1776,1797]" pageId="5" pageNumber="8">d-e</emphasis>
) sections. 
<emphasis id="5BDDEAA74B60D054FD42E9EEFCB8F899" bold="true" box="[755,792,1776,1797]" pageId="5" pageNumber="8">a–b</emphasis>
Bioclastic packstone with dasycladalean (e.g., 
<taxonomicName id="AEA94D364B60D054FB57E9F1FE3EF8BD" authority="Segonzac, A" authorityName="Segonzac, A" class="Ulvophyceae" family="Triploporellaceae" genus="Acroporella" kingdom="Plantae" order="Dasycladales" pageId="5" pageNumber="8" phylum="Chlorophyta" rank="species" species="anceps">
<emphasis id="5BDDEAA74B60D054FB57E9F1FAFAF899" box="[1254,1370,1775,1797]" italics="true" pageId="5" pageNumber="8">Acroporella</emphasis>
? 
<emphasis id="5BDDEAA74B60D054FADDE9EEFEBEF8BD" italics="true" pageId="5" pageNumber="8">anceps</emphasis>
Segonzac, 
<emphasis id="5BDDEAA74B60D054FE3EE815FE3EF8BD" box="[399,414,1803,1825]" italics="true" pageId="5" pageNumber="8">A</emphasis>
</taxonomicName>
), halimedacean (ha) algae, gastropods, and porcelaneous benthic foraminifera (e.g., soritoidids). 
<emphasis id="5BDDEAA74B60D054FAFEE815FAFAF8BC" bold="true" box="[1359,1370,1803,1824]" pageId="5" pageNumber="8">c</emphasis>
Bioclastic packstone with dasycladalean algae (e.g., 
<taxonomicName id="AEA94D364B60D054FD68E839FB37F8A1" authority="Beckmann &amp; Beckmann, Cl" authorityName="Beckmann &amp; Beckmann, Cl" box="[729,1175,1831,1853]" class="Ulvophyceae" family="Dasycladaceae" genus="Clypeina" kingdom="Plantae" order="Dasycladales" pageId="5" pageNumber="8" phylum="Chlorophyta" rank="species" species="elliotti">
<emphasis id="5BDDEAA74B60D054FD68E839FCD8F8A1" box="[729,888,1831,1853]" italics="true" pageId="5" pageNumber="8">Clypeina elliotti</emphasis>
Beckmann &amp; Beckmann, 
<emphasis id="5BDDEAA74B60D054FB31E839FB37F8A1" box="[1152,1175,1831,1853]" italics="true" pageId="5" pageNumber="8">Cl</emphasis>
</taxonomicName>
). 
<emphasis id="5BDDEAA74B60D054FB1CE839FB1AF8A0" bold="true" box="[1197,1210,1831,1852]" pageId="5" pageNumber="8">d</emphasis>
Bioclastic packstone with dasycladaleans (e.g., 
<taxonomicName id="AEA94D364B60D054FE4AE85CFCC8F8C4" authority="Segonzac, R" authorityName="Segonzac, R" authorityYear="1970" box="[507,872,1858,1880]" class="Ulvophyceae" family="Dasycladaceae" genus="Rostroporella" kingdom="Plantae" order="Dasycladales" pageId="5" pageNumber="8" phylum="Chlorophyta" rank="species" species="oviformis">
<emphasis id="5BDDEAA74B60D054FE4AE85CFD44F8C4" box="[507,740,1858,1880]" italics="true" pageId="5" pageNumber="8">Rostroporella oviformis</emphasis>
Segonzac, 
<emphasis id="5BDDEAA74B60D054FCE8E85DFCC8F8C4" box="[857,872,1859,1880]" italics="true" pageId="5" pageNumber="8">R</emphasis>
</taxonomicName>
). Thin-sections: Ah 185 (
<emphasis id="5BDDEAA74B60D054FBDCE85DFB32F8C4" bold="true" box="[1133,1170,1859,1880]" pageId="5" pageNumber="8">a–b</emphasis>
), CT 4 (
<emphasis id="5BDDEAA74B60D054FB5EE85DFB5AF8C4" bold="true" box="[1263,1274,1859,1880]" pageId="5" pageNumber="8">c</emphasis>
), 2pz29 (
<emphasis id="5BDDEAA74B60D054FAECE85DFACAF8C4" bold="true" box="[1373,1386,1859,1880]" pageId="5" pageNumber="8">d</emphasis>
). 
<emphasis id="5BDDEAA74B60D054FA33E85DFA2DF8C4" bold="true" box="[1410,1421,1859,1880]" pageId="5" pageNumber="8">e</emphasis>
Weathered rock surface with thallus fragments of 
<taxonomicName id="AEA94D364B60D054FD5AE840FCCFF8E8" box="[747,879,1886,1908]" class="Ulvophyceae" family="Triploporellaceae" genus="Dissocladella" kingdom="Plantae" order="Dasycladales" pageId="5" pageNumber="8" phylum="Chlorophyta" rank="genus">
<emphasis id="5BDDEAA74B60D054FD5AE840FCCFF8E8" box="[747,879,1886,1908]" italics="true" pageId="5" pageNumber="8">Dissocladella</emphasis>
</taxonomicName>
? 
<emphasis id="5BDDEAA74B60D054FC32E840FBAEF8E8" box="[899,1038,1886,1908]" italics="true" pageId="5" pageNumber="8">chahtorshiana</emphasis>
n. sp. The maximum observed length amounts for ~ 17.5 mm (arrow). Kuh-e-Patorgi area.
</paragraph>
</caption>
<caption id="3DD6663D4B63D057FF64E92FFADEF9E2" ID-DOI="http://doi.org/10.5281/zenodo.10520664" ID-Zenodo-Dep="10520664" httpUri="https://zenodo.org/record/10520664/files/figure.png" pageId="6" pageNumber="9" startId="6.[213,248,1585,1606]" targetBox="[213,1402,154,1549]" targetPageId="6" targetType="figure">
<paragraph id="691636B54B63D057FF64E92FFADEF9E2" blockId="6.[213,1427,1585,1662]" pageId="6" pageNumber="9">
<emphasis id="5BDDEAA74B63D057FF64E92FFEB2F9DA" bold="true" box="[213,274,1585,1606]" pageId="6" pageNumber="9">Fig. 6</emphasis>
<taxonomicName id="AEA94D364B63D057FEADE92FFE00F9DB" box="[284,416,1585,1607]" class="Ulvophyceae" family="Triploporellaceae" genus="Dissocladella" kingdom="Plantae" order="Dasycladales" pageId="6" pageNumber="9" phylum="Chlorophyta" rank="genus">
<emphasis id="5BDDEAA74B63D057FEADE92FFE00F9DB" box="[284,416,1585,1607]" italics="true" pageId="6" pageNumber="9">Dissocladella</emphasis>
</taxonomicName>
? 
<emphasis id="5BDDEAA74B63D057FE02E92FFD9EF9DB" box="[435,574,1585,1607]" italics="true" pageId="6" pageNumber="9">chahtorshiana</emphasis>
n. sp. from Selandian carbonates of Kuh-e-Chah Torsh, Central Iran. 
<emphasis id="5BDDEAA74B63D057FAB3E92FFA89F9DA" bold="true" box="[1282,1321,1585,1606]" pageId="6" pageNumber="9">a–f,</emphasis>
transverse sections, partly slightly oblique. Note the different morphologies of the primaries, only slightly widening (
<emphasis id="5BDDEAA74B63D057FAB6E953FA8CF9FE" bold="true" box="[1287,1324,1613,1634]" pageId="6" pageNumber="9">a–b</emphasis>
), or clubshaped (
<emphasis id="5BDDEAA74B63D057FE97E976FEE5F9E1" bold="true" box="[294,325,1640,1661]" pageId="6" pageNumber="9">e–f</emphasis>
). Secondary laterals partly visible in 
<emphasis id="5BDDEAA74B63D057FD1DE976FD17F9E1" bold="true" box="[684,695,1640,1661]" pageId="6" pageNumber="9">c</emphasis>
, 
<emphasis id="5BDDEAA74B63D057FD72E976FD6DF9E1" bold="true" box="[707,717,1640,1661]" pageId="6" pageNumber="9">e</emphasis>
, and 
<emphasis id="5BDDEAA74B63D057FCB3E976FCAAF9E1" bold="true" box="[770,778,1640,1661]" pageId="6" pageNumber="9">f</emphasis>
. 
<emphasis id="5BDDEAA74B63D057FCA7E976FC82F9E1" bold="true" box="[790,802,1640,1661]" pageId="6" pageNumber="9">g</emphasis>
, oblique section. Thin-sections: Ah 185 (a, c-d), CT 4 (b, e-g).
</paragraph>
</caption>
<paragraph id="691636B54B63D057FF26E8A9FDD2F790" blockId="6.[151,771,1730,2060]" pageId="6" pageNumber="9">
<emphasis id="5BDDEAA74B63D057FF26E8A9FE26F853" bold="true" box="[151,390,1975,1999]" pageId="6" pageNumber="9">Dimensions (in mm):</emphasis>
The main biometric parameters are compiled in 
<tableCitation id="242B030E4B63D057FEAEE8C8FED2F872" box="[287,370,2006,2030]" captionStart="Table 1" captionStartId="7.[231,296,191,215]" captionTargetId="graphics-873@7.[231,1402,338,978]" captionTargetPageId="7" captionText="Table 1 Dimensions of Dissocladella? chahtorshiana n. sp. from the Paleogene of Iran (in mm, except ratio d/D). D: outer diameter; d: inner diameter; pmax: maximum diameter of primary laterals; l: length of primary laterals; p’max: maximum diameter of secondary laterals; l’max: maximum length of secondary laterals; h: verticil spacing; w: number of primary laterals in a verticil." pageId="6" pageNumber="9">Table 1</tableCitation>
. The maximum observed length of a thallus fragment is about 
<quantity id="AE519B504B63D057FE17E8EAFDA5F790" box="[422,517,2036,2060]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="6" pageNumber="9" unit="mm" value="17.5">17.5 mm</quantity>
(
<figureCitation id="F1922A304B63D057FDA2E8EAFDC6F790" box="[531,614,2036,2060]" captionStart="Fig" captionStartId="5.[243,278,1748,1769]" captionTargetBox="[243,1419,154,1711]" captionTargetId="figure-112@5.[243,1419,154,1711]" captionTargetPageId="5" captionText="Fig. 5 Typical microfacies of samples with Dissocladella? chahtorshiana n. sp. (D), from the Paleocene of the Kuh-e- Chah Torsh (a–c) and Kuh-e-Patorgi (d-e) sections. a–b Bioclastic packstone with dasycladalean (e.g., Acroporella? anceps Segonzac, A), halimedacean (ha) algae, gastropods, and porcelaneous benthic foraminifera (e.g., soritoidids). c Bioclastic packstone with dasycladalean algae (e.g., Clypeina elliotti Beckmann &amp; Beckmann, Cl). d Bioclastic packstone with dasycladaleans (e.g., Rostroporella oviformis Segonzac, R). Thin-sections: Ah 185 (a–b), CT 4 (c), 2pz29 (d). e Weathered rock surface with thallus fragments of Dissocladella? chahtorshiana n. sp. The maximum observed length amounts for ~ 17.5 mm (arrow). Kuh-e-Patorgi area." figureDoi="http://doi.org/10.5281/zenodo.10520662" httpUri="https://zenodo.org/record/10520662/files/figure.png" pageId="6" pageNumber="9">Fig. 5E</figureCitation>
).
</paragraph>
<paragraph id="691636B54B63D056FC81E9DCFE7CFBB1" blockId="6.[816,1437,1730,2030]" lastBlockId="7.[151,770,1015,1069]" lastPageId="7" lastPageNumber="10" pageId="6" pageNumber="9">
<emphasis id="5BDDEAA74B63D057FC81E9DCFBB6F946" bold="true" box="[816,1046,1730,1754]" pageId="6" pageNumber="9">Palaeoenvironment:</emphasis>
<taxonomicName id="AEA94D364B63D057FB94E9DCFB17F946" box="[1061,1207,1730,1754]" class="Ulvophyceae" family="Triploporellaceae" genus="Dissocladella" kingdom="Plantae" order="Dasycladales" pageId="6" pageNumber="9" phylum="Chlorophyta" rank="genus">
<emphasis id="5BDDEAA74B63D057FB94E9DCFB17F946" box="[1061,1207,1730,1754]" italics="true" pageId="6" pageNumber="9">Dissocladella</emphasis>
</taxonomicName>
? 
<emphasis id="5BDDEAA74B63D057FB63E9DCFACCF946" box="[1234,1388,1730,1754]" italics="true" pageId="6" pageNumber="9">chahtorshiana</emphasis>
occurs in bioclastic packstones with various dasycladalean algae (e.g., 
<taxonomicName id="AEA94D364B63D057FC0AE9E1FBBBF88B" box="[955,1051,1791,1815]" class="Ulvophyceae" family="Dasycladaceae" genus="Clypeina" kingdom="Plantae" order="Dasycladales" pageId="6" pageNumber="9" phylum="Chlorophyta" rank="genus">
<emphasis id="5BDDEAA74B63D057FC0AE9E1FBBBF88B" box="[955,1051,1791,1815]" italics="true" pageId="6" pageNumber="9">Clypeina</emphasis>
</taxonomicName>
, 
<taxonomicName id="AEA94D364B63D057FB81E9E1FB05F88B" authorityName="J. V. F. Lamouroux" authorityYear="1816" box="[1072,1189,1791,1815]" class="Ulvophyceae" family="Dasycladaceae" genus="Cymopolia" kingdom="Plantae" order="Dasycladales" pageId="6" pageNumber="9" phylum="Chlorophyta" rank="genus">
<emphasis id="5BDDEAA74B63D057FB81E9E1FB05F88B" box="[1072,1189,1791,1815]" italics="true" pageId="6" pageNumber="9">Cymopolia</emphasis>
</taxonomicName>
), among some rather large taxa such as 
<taxonomicName id="AEA94D364B63D057FC47E800FAC2F8AA" authority="Segonzac" authorityName="Segonzac, R" authorityYear="1970" box="[1014,1378,1822,1846]" class="Ulvophyceae" family="Dasycladaceae" genus="Rostroporella" kingdom="Plantae" order="Dasycladales" pageId="6" pageNumber="9" phylum="Chlorophyta" rank="species" species="oviformis">
<emphasis id="5BDDEAA74B63D057FC47E800FB55F8AA" box="[1014,1269,1822,1846]" italics="true" pageId="6" pageNumber="9">Rostroporella oviformis</emphasis>
Segonzac
</taxonomicName>
(
<figureCitation id="F1922A304B63D057FAC0E800FCF4F8C8" captionStart="Fig" captionStartId="5.[243,278,1748,1769]" captionTargetBox="[243,1419,154,1711]" captionTargetId="figure-112@5.[243,1419,154,1711]" captionTargetPageId="5" captionText="Fig. 5 Typical microfacies of samples with Dissocladella? chahtorshiana n. sp. (D), from the Paleocene of the Kuh-e- Chah Torsh (a–c) and Kuh-e-Patorgi (d-e) sections. a–b Bioclastic packstone with dasycladalean (e.g., Acroporella? anceps Segonzac, A), halimedacean (ha) algae, gastropods, and porcelaneous benthic foraminifera (e.g., soritoidids). c Bioclastic packstone with dasycladalean algae (e.g., Clypeina elliotti Beckmann &amp; Beckmann, Cl). d Bioclastic packstone with dasycladaleans (e.g., Rostroporella oviformis Segonzac, R). Thin-sections: Ah 185 (a–b), CT 4 (c), 2pz29 (d). e Weathered rock surface with thallus fragments of Dissocladella? chahtorshiana n. sp. The maximum observed length amounts for ~ 17.5 mm (arrow). Kuh-e-Patorgi area." figureDoi="http://doi.org/10.5281/zenodo.10520662" httpUri="https://zenodo.org/record/10520662/files/figure.png" pageId="6" pageNumber="9">Fig. 5D</figureCitation>
), halimedacean algae (
<figureCitation id="F1922A304B63D057FBFBE822FB1DF8C8" box="[1098,1213,1852,1876]" captionStart="Fig" captionStartId="5.[243,278,1748,1769]" captionTargetBox="[243,1419,154,1711]" captionTargetId="figure-112@5.[243,1419,154,1711]" captionTargetPageId="5" captionText="Fig. 5 Typical microfacies of samples with Dissocladella? chahtorshiana n. sp. (D), from the Paleocene of the Kuh-e- Chah Torsh (a–c) and Kuh-e-Patorgi (d-e) sections. a–b Bioclastic packstone with dasycladalean (e.g., Acroporella? anceps Segonzac, A), halimedacean (ha) algae, gastropods, and porcelaneous benthic foraminifera (e.g., soritoidids). c Bioclastic packstone with dasycladalean algae (e.g., Clypeina elliotti Beckmann &amp; Beckmann, Cl). d Bioclastic packstone with dasycladaleans (e.g., Rostroporella oviformis Segonzac, R). Thin-sections: Ah 185 (a–b), CT 4 (c), 2pz29 (d). e Weathered rock surface with thallus fragments of Dissocladella? chahtorshiana n. sp. The maximum observed length amounts for ~ 17.5 mm (arrow). Kuh-e-Patorgi area." figureDoi="http://doi.org/10.5281/zenodo.10520662" httpUri="https://zenodo.org/record/10520662/files/figure.png" pageId="6" pageNumber="9">Fig. 5A–B</figureCitation>
) gastropods, benthic foraminifera, and rare fragments of corals. The association generally points to an open marine, outer platform environment of moderate water energy. 
<collectionCode id="0FB8AE704B63D057FB43E886FAA5F82C" box="[1266,1285,1944,1968]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="6" pageNumber="9" type="Herbarium">A</collectionCode>
similar association including 
<taxonomicName id="AEA94D364B63D057FBB3E8A9FB36F853" authorityName="G. Segonzac" authorityYear="1972" box="[1026,1174,1975,1999]" class="Ulvophyceae" family="Dasycladaceae" genus="Rostroporella" kingdom="Plantae" order="Dasycladales" pageId="6" pageNumber="9" phylum="Chlorophyta" rank="genus">
<emphasis id="5BDDEAA74B63D057FBB3E8A9FB36F853" box="[1026,1174,1975,1999]" italics="true" pageId="6" pageNumber="9">Rostroporella</emphasis>
</taxonomicName>
, 
<taxonomicName id="AEA94D364B63D057FB1EE8A9FAAFF853" box="[1199,1295,1975,1999]" class="Ulvophyceae" family="Dasycladaceae" genus="Clypeina" kingdom="Plantae" order="Dasycladales" pageId="6" pageNumber="9" phylum="Chlorophyta" rank="genus">
<emphasis id="5BDDEAA74B63D057FB1EE8A9FAAFF853" box="[1199,1295,1975,1999]" italics="true" pageId="6" pageNumber="9">Clypeina</emphasis>
</taxonomicName>
, 
<taxonomicName id="AEA94D364B63D057FA99E8A9FA3DF853" authorityName="J. V. F. Lamouroux" authorityYear="1816" box="[1320,1437,1975,1999]" class="Ulvophyceae" family="Dasycladaceae" genus="Cymopolia" kingdom="Plantae" order="Dasycladales" pageId="6" pageNumber="9" phylum="Chlorophyta" rank="genus">
<emphasis id="5BDDEAA74B63D057FA99E8A9FA3DF853" box="[1320,1437,1975,1999]" italics="true" pageId="6" pageNumber="9">Cymopolia</emphasis>
</taxonomicName>
div. sp., has been reported by 
<bibRefCitation id="0D384B444B63D056FB2EE8C8FF78FB93" author="Deloffre, R. &amp; Radoicic, R." lastPageId="7" lastPageNumber="10" pageId="6" pageNumber="9" pagination="61 - 95" refId="ref4649" refString="Deloffre, R., Radoicic, R., 1978. Algues calcaires (Dasycladales) du Paleocene de Slovenie (Yougoslavie). Bulletin de Centres Recherches Exploration-Production Elf-Aquitaine, 2 (1): 61 - 95." type="journal article" year="1978">Deloffre and Radoičić (1978</bibRefCitation>
, p. 63) from the Paleocene of 
<collectingCountry id="11BE76254B62D056FD9AECE9FD28FB93" box="[555,648,1015,1039]" name="Slovenia" pageId="7" pageNumber="10">Slovenia</collectingCountry>
and designated as &quot;reefal environment&quot;.
</paragraph>
</subSubSection>
</treatment>
</document>