264 lines
24 KiB
XML
264 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1158.100518" ID-Pensoft-Pub="1313-2970-1158-1" ID-Pensoft-UUID="299C29A6693A5054BA17D1663FBCCEE0" ID-ZooBank="047C3E08B3F244F484050AD39F23E9E8" ModsDocID="1313-2970-1158-1" checkinTime="1681371110153" checkinUser="pensoft" docAuthor="dela Cruz, Ian Niel & Ohara, Masahiro" docDate="2023" docId="75C9F8D83E375EE5A52275A7A0FB31A7" docLanguage="en" docName="ZooKeys 1158: 1-26" docOrigin="ZooKeys 1158" docPubDate="2023-04-12" docSource="http://dx.doi.org/10.3897/zookeys.1158.100518" docTitle="Atholus bakeri" docType="treatment" docVersion="1" id="299C29A6693A5054BA17D1663FBCCEE0" lastPageNumber="1" masterDocId="299C29A6693A5054BA17D1663FBCCEE0" masterDocTitle="Revision of the genus Atholus Thomson, 1859 (Coleoptera, Histeridae, Histerinae) from the Philippines with additional records" masterLastPageNumber="26" masterPageNumber="1" pageNumber="1" updateTime="1681371110153" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Atholus Thomson, 1859 (Coleoptera, Histeridae, Histerinae) from the Philippines with additional records</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>dela Cruz, Ian Niel</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2806-1337</mods:nameIdentifier>
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<mods:affiliation>Entomological Laboratory, Graduate School of Agriculture, Hokkaido University, N 9, W 9, Sapporo, 060 - 8589, Japan & Department of Biology, College of Mathematics and Natural Sciences, Caraga State University, Butuan City, 8600, Philippines</mods:affiliation>
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<mods:nameIdentifier type="email">histermushi@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ohara, Masahiro</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2806-1337</mods:nameIdentifier>
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<mods:affiliation>Entomological Laboratory, Graduate School of Agriculture, Hokkaido University, N 9, W 9, Sapporo, 060 - 8589, Japan</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1158</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>26</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1158.100518</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1158.100518</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1158-1</mods:identifier>
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<mods:identifier type="ZooBank">047C3E08B3F244F484050AD39F23E9E8</mods:identifier>
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<mods:identifier type="Pensoft-UUID">299C29A6693A5054BA17D1663FBCCEE0</mods:identifier>
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</mods:mods>
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<subSection pageId="0" pageNumber="1" type="systematics">
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<treatment LSID="urn:lsid:plazi:treatment:75C9F8D83E375EE5A52275A7A0FB31A7" httpUri="http://treatment.plazi.org/id/75C9F8D83E375EE5A52275A7A0FB31A7" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName ID-CoL="688GG" LSID="75C9F8D8-3E37-5EE5-A522-75A7A0FB31A7" authority="(Bickhardt, 1914)" baseAuthorityName="Bickhardt" baseAuthorityYear="1914" class="Insecta" family="Histeridae" genus="Atholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atholus bakeri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bakeri">Atholus bakeri (Bickhardt, 1914)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Philippine Atholus, dorsal habitus 1 A. philippinensis (Marseul, 1854) [IC- 21 - 18] 2 A. coelestis (Marseul, 1857) [IC- 21 - 20] 3 A. torquatus (Marseul, 1854) [IC- 21 - 49] 4 A. pirithous (Marseul, 1873) [IC- 21 - 47] 5 A. nitidissimus Desbordes, 1925 [syntype image] 6 A. bakeri (Bickhardt, 1914) [syntype image, No. 1639]. Scale bar: 1.00 mm." figureDoi="10.3897/zookeys.1158.100518.figures1-6" httpUri="https://binary.pensoft.net/fig/836675" pageId="0" pageNumber="1">Figs 6</figureCitation>
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<figureCitation captionStart="Figures 54–57" captionStartId="F12" captionText="Figures 54 - 57. Atholus bakeri (Bickhardt, 1914) 54 head and pronotum, dorsal view 55 head, frontal view 56 propygidium and pygidium 57 habitus, ventral view." figureDoi="10.3897/zookeys.1158.100518.figures54-57" httpUri="https://binary.pensoft.net/fig/836686" pageId="0" pageNumber="1">, 54-57</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Bickhardt" authorityYear="1914" class="Insecta" family="Histeridae" genus="Hister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hister bakeri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bakeri">Hister bakeri</taxonomicName>
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Bickhardt, 1914: 428 [Luzon Island].
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Bickhardt" authorityYear="1914" class="Insecta" family="Histeridae" genus="Hister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hister (Atholus) bakeri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bakeri" subGenus="Atholus">Hister (Atholus) bakeri</taxonomicName>
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:
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<bibRefCitation author="Bickhardt, H" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="1" refId="B7" refString="Bickhardt, H, 1917. Histeridae. In: Wytsman P (Ed.) Genera Insectorum, fascicules 166b: 113-302. [La Haye.]" title="Histeridae. In: Wytsman P (Ed.) Genera Insectorum, fascicules 166 b: 113 - 302. [La Haye.]" year="1917">Bickhardt 1917</bibRefCitation>
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: 193 [catalogued].
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Bickhardt" baseAuthorityYear="1914" class="Insecta" family="Histeridae" genus="Atholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atholus bakeri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bakeri">Atholus bakeri</taxonomicName>
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:
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<bibRefCitation author="Bickhardt, H" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="1" refId="B6" refString="Bickhardt, H, 1914. Philippinische Histeriden: I. Philippine Journal of Science 9(D): 423-429." title="Philippinische Histeriden: I. Philippine Journal of Science 9 (D): 423 - 429." year="1914">Bickhardt 1914</bibRefCitation>
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: 428;
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<bibRefCitation author="Mazur, S" journalOrPublisher="Polskie Pismo Entomologiczne" pageId="0" pageNumber="1" pagination="1 - 376" refId="B36" refString="Mazur, S, 1984. A world catalogue of Histeridae. Polskie Pismo Entomologiczne 54 (3-4): 1 - 376" title="A world catalogue of Histeridae." volume="54" year="1984">Mazur 1984</bibRefCitation>
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: 211;
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<bibRefCitation author="Mazur, S" journalOrPublisher="Polskie Pismo Entomologiczne" pageId="0" pageNumber="1" refId="B37" refString="Mazur, S, 1997. A world catalogue of Histeridae (Coleoptera: Histeroidea). Genus International Journal of Invertebrate Taxonomy (Supplement), Polskie Towarzystwo Taksonomiczne 1-373." title="A world catalogue of Histeridae (Coleoptera: Histeroidea). Genus International Journal of Invertebrate Taxonomy (Supplement), Polskie Towarzystwo Taksonomiczne 1 - 373." year="1997">1997</bibRefCitation>
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: 129;
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<bibRefCitation author="Mazur, S" journalOrPublisher="Warsaw University of Life Sciences SGGW Press, Warsaw" pageId="0" pageNumber="1" refId="B40" refString="Mazur, S, 2011. A Concise Catalogue of the Histeridae (Insecta: Coleoptera). Warsaw University of Life Sciences SGGW Press, Warsaw" title="A Concise Catalogue of the Histeridae (Insecta: Coleoptera)." year="2011">2011</bibRefCitation>
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: 103 [catalogued].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" location="Luzon Island" specimenCount="1" typeStatus="syntype">
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<specimenCount type="generic" typeStatus="syntype">1 syntype</specimenCount>
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[
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<emphasis bold="true" pageId="0" pageNumber="1">
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<location LSID="urn:lsid:plazi:treatment:75C9F8D83E375EE5A52275A7A0FB31A7:C70A27FA6B40BC7BF606F512447ADB8F" name="Luzon Island">Luzon Island</location>
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</emphasis>
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] based on images, "
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<taxonomicName baseAuthorityName="Bickhardt" baseAuthorityYear="1914" class="Insecta" family="Histeridae" genus="Atholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atholus bakeri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bakeri">
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<emphasis italics="true" pageId="0" pageNumber="1">Atholus bakeri</emphasis>
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</taxonomicName>
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n. sp. Bickh. / Los Banos, / P.I., Baker. / 1639" [sex undetermined, measurements not available] (MNHUB)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This species has lateral pronotal striae interrupted in the anterolateral angle, and strong punctations on its entire pygidium. According to the original description of
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<bibRefCitation author="Bickhardt, H" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="1" refId="B6" refString="Bickhardt, H, 1914. Philippinische Histeriden: I. Philippine Journal of Science 9(D): 423-429." title="Philippinische Histeriden: I. Philippine Journal of Science 9 (D): 423 - 429." year="1914">Bickhardt (1914)</bibRefCitation>
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,
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<taxonomicName baseAuthorityName="Bickhardt" baseAuthorityYear="1914" class="Insecta" family="Histeridae" genus="Atholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atholus bakeri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bakeri">
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<emphasis italics="true" pageId="0" pageNumber="1">Atholus bakeri</emphasis>
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</taxonomicName>
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(Bickhardt, 1914) is most similar to
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<taxonomicName lsidName="A. torquatus" pageId="0" pageNumber="1" rank="species" species="torquatus">
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<emphasis italics="true" pageId="0" pageNumber="1">A. torquatus</emphasis>
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</taxonomicName>
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(Marseul, 1854) except that the propygidium and pygidium of
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<taxonomicName lsidName="A. bakeri" pageId="0" pageNumber="1" rank="species" species="bakeri">
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<emphasis italics="true" pageId="0" pageNumber="1">A. bakeri</emphasis>
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</taxonomicName>
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(Bickhardt, 1914) are strongly punctate. However, based on the syntype observed, the punctation is not as prominent as described and, in fact quite similar to that of
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<taxonomicName lsidName="A. torquatus" pageId="0" pageNumber="1" rank="species" species="torquatus">
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<emphasis italics="true" pageId="0" pageNumber="1">A. torquatus</emphasis>
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</taxonomicName>
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(Marseul, 1854). The punctures are finer towards the apical end of the pygidium. Another distinguishable feature of
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<taxonomicName lsidName="A. bakeri" pageId="0" pageNumber="1" rank="species" species="bakeri">
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<emphasis italics="true" pageId="0" pageNumber="1">A. bakeri</emphasis>
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</taxonomicName>
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(Bickhardt, 1914) is the medially straight frontal stria, while in
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<taxonomicName lsidName="A. torquatus" pageId="0" pageNumber="1" rank="species" species="torquatus">
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<emphasis italics="true" pageId="0" pageNumber="1">A. torquatus</emphasis>
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</taxonomicName>
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(Marseul, 1854) it is weakly bent inwardly. However, several studied individuals of
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<taxonomicName lsidName="A. torquatus" pageId="0" pageNumber="1" rank="species" species="torquatus">
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<emphasis italics="true" pageId="0" pageNumber="1">A. torquatus</emphasis>
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</taxonomicName>
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(Marseul, 1854) likewise seem to have their frontal stria medially straight.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Re-description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Body</emphasis>
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(Fig.
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<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Philippine Atholus, dorsal habitus 1 A. philippinensis (Marseul, 1854) [IC- 21 - 18] 2 A. coelestis (Marseul, 1857) [IC- 21 - 20] 3 A. torquatus (Marseul, 1854) [IC- 21 - 49] 4 A. pirithous (Marseul, 1873) [IC- 21 - 47] 5 A. nitidissimus Desbordes, 1925 [syntype image] 6 A. bakeri (Bickhardt, 1914) [syntype image, No. 1639]. Scale bar: 1.00 mm." figureDoi="10.3897/zookeys.1158.100518.figures1-6" httpUri="https://binary.pensoft.net/fig/836675" pageId="0" pageNumber="1">6</figureCitation>
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) oval, moderately convex and black; tibia and antenna rufous.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
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: clypeus (Figs
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<figureCitation captionStart="Figures 54–57" captionStartId="F12" captionText="Figures 54 - 57. Atholus bakeri (Bickhardt, 1914) 54 head and pronotum, dorsal view 55 head, frontal view 56 propygidium and pygidium 57 habitus, ventral view." figureDoi="10.3897/zookeys.1158.100518.figures54-57" httpUri="https://binary.pensoft.net/fig/836686" pageId="0" pageNumber="1">54</figureCitation>
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,
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<figureCitation captionStart="Figures 54–57" captionStartId="F12" captionText="Figures 54 - 57. Atholus bakeri (Bickhardt, 1914) 54 head and pronotum, dorsal view 55 head, frontal view 56 propygidium and pygidium 57 habitus, ventral view." figureDoi="10.3897/zookeys.1158.100518.figures54-57" httpUri="https://binary.pensoft.net/fig/836686" pageId="0" pageNumber="1">55</figureCitation>
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) slightly crenate on anterolateral margin, apical margin slightly extended; frontal stria medially straight, complete, moderately impressed; eyes large, convex, clearly visible dorsally; mandibles with rounded outer margin curved inwardly; sub-apical tooth on left mandible large; mandibular apex acutely pointed.
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</paragraph>
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<caption doi="10.3897/zookeys.1158.100518.figures54-57" httpUri="https://binary.pensoft.net/fig/836686" pageId="0" pageNumber="1" start="Figures 54–57" startId="F12">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figures 54-57.</emphasis>
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<taxonomicName baseAuthorityName="Bickhardt" baseAuthorityYear="1914" class="Insecta" family="Histeridae" genus="Atholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atholus bakeri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bakeri">
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<emphasis italics="true" pageId="0" pageNumber="1">Atholus bakeri</emphasis>
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</taxonomicName>
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(Bickhardt, 1914)
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<emphasis bold="true" pageId="0" pageNumber="1">54</emphasis>
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head and pronotum, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="1">55</emphasis>
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head, frontal view
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<emphasis bold="true" pageId="0" pageNumber="1">56</emphasis>
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propygidium and pygidium
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<emphasis bold="true" pageId="0" pageNumber="1">57</emphasis>
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habitus, ventral view.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pronotum</emphasis>
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: marginal pronotal stria (Figs
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<figureCitation captionStart="Figures 54–57" captionStartId="F12" captionText="Figures 54 - 57. Atholus bakeri (Bickhardt, 1914) 54 head and pronotum, dorsal view 55 head, frontal view 56 propygidium and pygidium 57 habitus, ventral view." figureDoi="10.3897/zookeys.1158.100518.figures54-57" httpUri="https://binary.pensoft.net/fig/836686" pageId="0" pageNumber="1">54</figureCitation>
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,
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<figureCitation captionStart="Figures 54–57" captionStartId="F12" captionText="Figures 54 - 57. Atholus bakeri (Bickhardt, 1914) 54 head and pronotum, dorsal view 55 head, frontal view 56 propygidium and pygidium 57 habitus, ventral view." figureDoi="10.3897/zookeys.1158.100518.figures54-57" httpUri="https://binary.pensoft.net/fig/836686" pageId="0" pageNumber="1">55</figureCitation>
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) laterally complete, continuous onto apical angle and crenate behind head; lateral pronotal stria moderately impressed, slightly crenate; apical end shortened and bent inwardly in a curved hook; lateral portion rather distant from margin; its basal end abbreviated from basal fifth of pronotal length.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Elytra</emphasis>
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: elytral epipleuron (Fig.
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<figureCitation captionStart="Figures 54–57" captionStartId="F12" captionText="Figures 54 - 57. Atholus bakeri (Bickhardt, 1914) 54 head and pronotum, dorsal view 55 head, frontal view 56 propygidium and pygidium 57 habitus, ventral view." figureDoi="10.3897/zookeys.1158.100518.figures54-57" httpUri="https://binary.pensoft.net/fig/836686" pageId="0" pageNumber="1">57</figureCitation>
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) with few coarse punctures on apical half; marginal epipleural stria present on apical half; marginal elytral stria (Fig.
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<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Philippine Atholus, dorsal habitus 1 A. philippinensis (Marseul, 1854) [IC- 21 - 18] 2 A. coelestis (Marseul, 1857) [IC- 21 - 20] 3 A. torquatus (Marseul, 1854) [IC- 21 - 49] 4 A. pirithous (Marseul, 1873) [IC- 21 - 47] 5 A. nitidissimus Desbordes, 1925 [syntype image] 6 A. bakeri (Bickhardt, 1914) [syntype image, No. 1639]. Scale bar: 1.00 mm." figureDoi="10.3897/zookeys.1158.100518.figures1-6" httpUri="https://binary.pensoft.net/fig/836675" pageId="0" pageNumber="1">6</figureCitation>
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) complete, slightly impressed; external and internal subhumeral striae absent; oblique humeral stria lightly impressed on basal third; dorsal elytral striae 1-4 complete; dorsal elytral stria 5 and sutural elytral stria present on apical half; disk with fine punctures.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
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: propygidium (Fig.
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<figureCitation captionStart="Figures 54–57" captionStartId="F12" captionText="Figures 54 - 57. Atholus bakeri (Bickhardt, 1914) 54 head and pronotum, dorsal view 55 head, frontal view 56 propygidium and pygidium 57 habitus, ventral view." figureDoi="10.3897/zookeys.1158.100518.figures54-57" httpUri="https://binary.pensoft.net/fig/836686" pageId="0" pageNumber="1">56</figureCitation>
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) moderately covered with coarse, round, and shallow punctures; interspaces with fine punctations; pygidial punctations similar to those of propygidium, becoming sparser apically.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Meso- and metaventrite</emphasis>
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: anterior margin of mesoventrite (Fig.
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<figureCitation captionStart="Figures 54–57" captionStartId="F12" captionText="Figures 54 - 57. Atholus bakeri (Bickhardt, 1914) 54 head and pronotum, dorsal view 55 head, frontal view 56 propygidium and pygidium 57 habitus, ventral view." figureDoi="10.3897/zookeys.1158.100518.figures54-57" httpUri="https://binary.pensoft.net/fig/836686" pageId="0" pageNumber="1">57</figureCitation>
|
||
) outwardly arcuate; marginal mesoventral stria crenate and complete; meso-metaventral suture clearly impressed, complete, medially angulate; punctations of intercoxal disk of metaventrite similar to those of mesoventrite; longitudinal suture of metaventrite lightly impressed; lateral disk of metaventrite moderately covered with large, round, shallow punctures.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
|
||
: posterior surface of protibia (Fig.
|
||
<figureCitation captionStart="Figures 54–57" captionStartId="F12" captionText="Figures 54 - 57. Atholus bakeri (Bickhardt, 1914) 54 head and pronotum, dorsal view 55 head, frontal view 56 propygidium and pygidium 57 habitus, ventral view." figureDoi="10.3897/zookeys.1158.100518.figures54-57" httpUri="https://binary.pensoft.net/fig/836686" pageId="0" pageNumber="1">57</figureCitation>
|
||
) flattened and strongly dilated; outer lateral margin with four weak, almost inconspicuous teeth, topped by minute denticles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Endemic to the Philippines (
|
||
<bibRefCitation author="Mazur, S" journalOrPublisher="Warsaw University of Life Sciences SGGW Press, Warsaw" pageId="0" pageNumber="1" refId="B40" refString="Mazur, S, 2011. A Concise Catalogue of the Histeridae (Insecta: Coleoptera). Warsaw University of Life Sciences SGGW Press, Warsaw" title="A Concise Catalogue of the Histeridae (Insecta: Coleoptera)." year="2011">Mazur 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The examined syntype of
|
||
<taxonomicName lsidName="A. bakeri" pageId="0" pageNumber="1" rank="species" species="bakeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">A. bakeri</emphasis>
|
||
</taxonomicName>
|
||
(Bickhardt, 1914) exhibits characters similar to a typical
|
||
<taxonomicName lsidName="A. torquatus" pageId="0" pageNumber="1" rank="species" species="torquatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">A. torquatus</emphasis>
|
||
</taxonomicName>
|
||
(Marseul, 1854). According to
|
||
<bibRefCitation author="Desbordes, H" journalOrPublisher="Annales de la Societe Entomologique de France, Paris" pageId="0" pageNumber="1" pagination="297 - 326" refId="B11" refString="Desbordes, H, 1917. Contribution a la connaissance des Histerides. 2e memoire. Synopsis de divers groupes d' Histeridae. Annales de la Societe Entomologique de France, Paris 85: 297 - 326" title="Contribution a la connaissance des Histerides. 2 e memoire. Synopsis de divers groupes d' Histeridae." volume="85" year="1917">Desbordes (1917)</bibRefCitation>
|
||
,
|
||
<taxonomicName lsidName="A. torquatus" pageId="0" pageNumber="1" rank="species" species="torquatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">A. torquatus</emphasis>
|
||
</taxonomicName>
|
||
(Marseul, 1854) and
|
||
<taxonomicName lsidName="A. bakeri" pageId="0" pageNumber="1" rank="species" species="bakeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">A. bakeri</emphasis>
|
||
</taxonomicName>
|
||
(Bickhardt, 1914) are very similar, being set apart by the pygidial punctation (strong in
|
||
<taxonomicName lsidName="A. bakeri" pageId="0" pageNumber="1" rank="species" species="bakeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">A. bakeri</emphasis>
|
||
</taxonomicName>
|
||
and apically finer in
|
||
<taxonomicName lsidName="A. torquatus" pageId="0" pageNumber="1" rank="species" species="torquatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">A. torquatus</emphasis>
|
||
</taxonomicName>
|
||
). Although the only examined specimen of
|
||
<taxonomicName lsidName="A. bakeri" pageId="0" pageNumber="1" rank="species" species="bakeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">A. bakeri</emphasis>
|
||
</taxonomicName>
|
||
(Bickhardt, 1914) possesses similar pygidial punctations to
|
||
<taxonomicName lsidName="A. torquatus" pageId="0" pageNumber="1" rank="species" species="torquatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">A. torquatus</emphasis>
|
||
</taxonomicName>
|
||
(Bickhardt, 1914), this character remains the primary distinction until further examinations of other types is established. The authors would also encourage a comprehensive observation of both male and female genitalia for future works.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |