1017 lines
81 KiB
XML
1017 lines
81 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6107715" ID-GBIF-Dataset="1291cc93-8e68-4549-aa01-707303d302bf" ID-GBIF-Taxon="114917768" ID-Zenodo-Dep="6107715" checkinTime="1441192651534" checkinUser="donat" docAuthor="Joe A. Macgown, Terence L. Schiefer & Michael G. Branstetter" docDate="2015" docId="FA729970A22EFFAE189F86F3C13ACB24" docLanguage="en" docName="10.11646_zootaxa.4006.2.10.pdf" docOrigin="Zootaxa 4006 (2)" docTitle="Leptanilloides chihuahuaensis" docType="treatment" docVersion="6" lastPageId="7" lastPageNumber="399" masterDocId="064BE108A22CFFA91808862EC445CF10" masterDocTitle="First record of the genus Leptanilloides (Hymenoptera: Formicidae: Dorylinae) from the United States" masterLastPageNumber="400" masterPageNumber="392" pageId="2" pageNumber="394" updateTime="1645022311421" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>First record of the genus Leptanilloides (Hymenoptera: Formicidae: Dorylinae) from the United States</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Joe A. Macgown</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Terence L. Schiefer</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Michael G. Branstetter</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2015</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>4006</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>392</mods:start>
|
||
<mods:end>400</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.11646/zootaxa.4006.2.10</mods:identifier>
|
||
<mods:identifier type="ZooBank">C9E0FAA3-324C-404D-8A1C-3B4AC6DE663E</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6107715" ID-GBIF-Taxon="114917768" ID-Zenodo-Dep="6107715" LSID="urn:lsid:plazi:treatment:FA729970A22EFFAE189F86F3C13ACB24" httpUri="http://treatment.plazi.org/id/FA729970A22EFFAE189F86F3C13ACB24" lastPageId="7" lastPageNumber="399" pageId="2" pageNumber="394">
|
||
<subSubSection box="[151,521,221,247]" pageId="2" pageNumber="394" type="nomenclature">
|
||
<paragraph box="[151,521,221,247]" pageId="2" pageNumber="394">
|
||
<heading bold="true" box="[151,521,221,247]" fontSize="11" level="1" pageId="2" pageNumber="394" reason="0">
|
||
<taxonomicName box="[151,521,221,247]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="chihuahuaensis">
|
||
<emphasis bold="true" box="[151,521,221,247]" italics="true" pageId="2" pageNumber="394">Leptanilloides chihuahuaensis</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[151,274,255,280]" pageId="2" pageNumber="394" type="description">
|
||
<paragraph box="[151,274,255,280]" pageId="2" pageNumber="394">
|
||
(
|
||
<figureCitation box="[159,236,255,280]" captionStart="FIGURE 1" captionStartId="3.[151,249,1527,1548]" captionTargetBox="[151,1436,193,1505]" captionTargetId="figure@3.[151,1436,193,1505]" captionTargetPageId="3" captionText="FIGURE 1. Leptanilloides chihuahuaensis, male, holotype (MEM 207971): (A) head in full-face view, (B) petiole in dorsal view, (C) tergite VIII and exposed male genitalia in dorsal view, (D) sternite IX and exposed male genitalia in ventral view, (E) lateral habitus, and (F) dorsal habitus. Abbreviations: ASIX = ninth abdominal sternite, ATVIII = eighth abdominal tergite, Ax = axilla, Msp = mesopleuron, MsScl = mesoscutellum, MsSct = mesoscutum, Mtn = metanotum, P = petiole, Ped = Pedicel, Pm = paramere, Prnt = pronotum, Prpd = propodeum, Pv = Penivalva, Sc = Scape, and Vol = volsella." pageId="2" pageNumber="394">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation box="[250,265,255,280]" captionStart="FIGURE 2" captionStartId="4.[151,249,1911,1933]" captionTargetBox="[195,1388,193,1886]" captionTargetId="figure@4.[195,1388,193,1886]" captionTargetPageId="4" captionText="FIGURE 2. Leptanilloides chihuahuaensis, male, holotype (MEM 207971): (A) lateral habitus and (B) forewing and hindwing showing labeled veins. Abbreviations: 2 r-rs = second radial sector cross-vein, A = anal vein, C = costal vein, Cu = cubital vein, cu-a = cubital-anal cross-vein, ha = hamuli, M = medial vein, M + Cu = merged medial-cubital veins, Rs + M = merged medial and radial sector plus medial veins, Rs = radial sector, Sc + R = merged subcostal and radial veins." pageId="2" pageNumber="394">2</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="394" type="materials_examined">
|
||
<paragraph pageId="2" pageNumber="394">
|
||
<materialsCitation ID-GBIF-Occurrence="1227780304" collectingDate="2014-07-22" collectingDateMax="2014-07-25" collectingDateMin="2014-07-22" collectionCode="MCZC" collectorName="T. L. Schiefer & W. H. Cross Expedition" country="United States of America" county="Jeff Davis Co." latitude="30.590834" location="Malaise" longLatPrecision="20" longitude="-103.94056" municipality="Davis Mts. St. Pk." pageId="2" pageNumber="394" specimenCode="MEM 207971" specimenCount="2" specimenCount-male="1" stateProvince="TEXAS" typeStatus="holotype">
|
||
<emphasis bold="true" box="[151,334,328,352]" pageId="2" pageNumber="394">
|
||
<typeStatus box="[151,260,328,352]" pageId="2" pageNumber="394">Holotype</typeStatus>
|
||
<specimenCount box="[267,329,328,352]" pageId="2" pageNumber="394" type="male">male</specimenCount>
|
||
:
|
||
</emphasis>
|
||
[
|
||
<collectingCountry box="[347,503,327,352]" name="United States of America" pageId="2" pageNumber="394" value="United States of America">United States</collectingCountry>
|
||
]
|
||
<collectingRegion box="[517,610,328,352]" pageId="2" pageNumber="394">TEXAS</collectingRegion>
|
||
,
|
||
<collectingCounty box="[621,785,327,352]" pageId="2" pageNumber="394">Jeff Davis Co.</collectingCounty>
|
||
,
|
||
<collectingMunicipality box="[797,1004,327,352]" pageId="2" pageNumber="394">Davis Mts. St. Pk.</collectingMunicipality>
|
||
,
|
||
<number box="[1016,1075,328,352]" pageId="2" pageNumber="394" value="5118.0">5118</number>
|
||
',
|
||
<geoCoordinate box="[1091,1223,328,352]" direction="north" orientation="latitude" pageId="2" pageNumber="394" precision="15" value="30.590834">
|
||
<number box="[1091,1120,328,352]" pageId="2" pageNumber="394" value="30.0">30</number>
|
||
°
|
||
<number box="[1130,1158,328,352]" pageId="2" pageNumber="394" value="35.0">35</number>
|
||
'
|
||
<number box="[1161,1191,328,352]" pageId="2" pageNumber="394" value="27.0">27</number>
|
||
"N
|
||
</geoCoordinate>
|
||
<geoCoordinate box="[1230,1382,328,352]" direction="west" orientation="longitude" pageId="2" pageNumber="394" precision="15" value="-103.94056">
|
||
<number box="[1230,1272,328,352]" pageId="2" pageNumber="394" value="103.0">103</number>
|
||
°
|
||
<number box="[1281,1310,328,352]" pageId="2" pageNumber="394" value="56.0">56</number>
|
||
'
|
||
<number box="[1313,1343,328,352]" pageId="2" pageNumber="394" value="26.0">26</number>
|
||
"W
|
||
</geoCoordinate>
|
||
,
|
||
<collectingDate pageId="2" pageNumber="394" value="2014-07-22" valueMax="2014-07-25" valueMin="2014-07-22">
|
||
<date pageId="2" pageNumber="394" value="2014-07-22" valueMax="2014-07-25" valueMin="2014-07-22">
|
||
<number pageId="2" pageNumber="394" value="23.5" valueMax="25.0" valueMin="22.0">22– 25</number>
|
||
July
|
||
<number box="[246,305,364,388]" pageId="2" pageNumber="394" value="2014.0">2014</number>
|
||
</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName box="[320,471,364,388]" pageId="2" pageNumber="394">T.L. Schiefer</collectorName>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:FA729970A22EFFAE189F86F3C13ACB24:77047EBDA22EFFAB19ED8742C67BCE94" box="[485,574,364,388]" country="United States of America" county="Jeff Davis Co." latitude="30.590834" longLatPrecision="20" longitude="-103.94056" municipality="Davis Mts. St. Pk." name="Malaise" pageId="2" pageNumber="394" stateProvince="TEXAS">Malaise</location>
|
||
trap on hillside on highland grassland,
|
||
<collectorName box="[1040,1309,364,388]" pageId="2" pageNumber="394">W.H. Cross Expedition</collectorName>
|
||
. [
|
||
<typeStatus box="[1329,1436,364,388]" pageId="2" pageNumber="394">Holotype</typeStatus>
|
||
deposited in
|
||
<location LSID="urn:lsid:plazi:treatment:FA729970A22EFFAE189F86F3C13ACB24:77047EBDA22EFFAB192D87A1C741CEB8" box="[293,772,399,424]" country="United States of America" county="Jeff Davis Co." latitude="30.590834" longLatPrecision="20" longitude="-103.94056" municipality="Davis Mts. St. Pk." name="Harvard Museum of Comparative Zoology" pageId="2" pageNumber="394" stateProvince="TEXAS">Harvard Museum of Comparative Zoology</location>
|
||
(
|
||
<collectionCode box="[786,870,399,424]" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" pageId="2" pageNumber="394">MCZC</collectionCode>
|
||
),
|
||
<location LSID="urn:lsid:plazi:treatment:FA729970A22EFFAE189F86F3C13ACB24:77047EBDA22EFFAB1B7187A1C7BCCEB8" box="[889,1017,399,424]" country="United States of America" county="Jeff Davis Co." latitude="30.590834" longLatPrecision="20" longitude="-103.94056" municipality="Davis Mts. St. Pk." name="Cambridge" pageId="2" pageNumber="394" stateProvince="TEXAS">Cambridge</location>
|
||
, MA].
|
||
<specimenCount box="[1096,1206,399,424]" pageId="2" pageNumber="394" type="generic">Specimen</specimenCount>
|
||
code
|
||
<specimenCode box="[1272,1433,400,424]" pageId="2" pageNumber="394">
|
||
MEM
|
||
<number box="[1345,1433,400,424]" pageId="2" pageNumber="394" value="207971.0">207971</number>
|
||
</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="1227780303" collectionCode="MEM" country="United States of America" location="USA" pageId="2" pageNumber="394" specimenCode="MEM 207972, MEM 207973" specimenCount="4" specimenCount-male="1" typeStatus="paratype">
|
||
<emphasis box="[151,325,436,460]" italics="true" pageId="2" pageNumber="394">
|
||
<typeStatus box="[151,253,437,460]" pageId="2" pageNumber="394">Paratype</typeStatus>
|
||
<specimenCount box="[260,325,436,460]" pageId="2" pageNumber="394" type="male">males</specimenCount>
|
||
</emphasis>
|
||
: Two point-mounted
|
||
<specimenCount box="[567,684,436,460]" pageId="2" pageNumber="394" type="generic">specimens</specimenCount>
|
||
with the same data as the holotype but with unique
|
||
<specimenCount box="[1263,1369,436,460]" pageId="2" pageNumber="394" type="generic">specimen</specimenCount>
|
||
code:
|
||
<specimenCode box="[151,313,472,496]" pageId="2" pageNumber="394">
|
||
MEM
|
||
<number box="[229,313,472,496]" pageId="2" pageNumber="394" value="207972.0">207972</number>
|
||
</specimenCode>
|
||
and
|
||
<specimenCode box="[376,538,472,496]" pageId="2" pageNumber="394">
|
||
MEM
|
||
<number box="[454,538,472,496]" pageId="2" pageNumber="394" value="207973.0">207973</number>
|
||
</specimenCode>
|
||
[both
|
||
<specimenCount box="[620,736,471,496]" pageId="2" pageNumber="394" type="generic">specimens</specimenCount>
|
||
in Mississippi Entomological Museum (
|
||
<collectionCode box="[1213,1285,471,496]" collectionName="USA, Mississippi, Mississippi, Mississippi State University" pageId="2" pageNumber="394">MEM</collectionCode>
|
||
), Mississippi State, MS,
|
||
<collectingCountry box="[277,339,508,532]" name="United States of America" pageId="2" pageNumber="394" value="United States of America">USA</collectingCountry>
|
||
]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="1227780302" collectingDate="2014-07-21" collectingDateMax="2014-07-25" collectingDateMin="2014-07-21" collectionCode="MEM" collectorName="T. L. Schiefer & W. H. Cross Expedition" country="United States of America" county="Jeff Davis Co." latitude="30.602222" location="Malaise" longLatPrecision="20" longitude="-103.915276" municipality="Davis Mts. St. Pk." pageId="2" pageNumber="394" specimenCode="MEM 207974, MEM 207975, MEM 207976" specimenCount="3" stateProvince="TEXAS">
|
||
and three
|
||
<specimenCount box="[470,590,508,532]" pageId="2" pageNumber="394" type="generic">specimens</specimenCount>
|
||
, one point-mounted and two in
|
||
<number box="[952,980,508,532]" pageId="2" pageNumber="394" value="95.0">95</number>
|
||
% ethanol, with the following collection data: [
|
||
<collectingCountry box="[222,381,543,568]" name="United States of America" pageId="2" pageNumber="394" value="United States of America">United States</collectingCountry>
|
||
]
|
||
<collectingRegion box="[398,491,544,568]" pageId="2" pageNumber="394">TEXAS</collectingRegion>
|
||
,
|
||
<collectingCounty box="[505,675,543,568]" pageId="2" pageNumber="394">Jeff Davis Co.</collectingCounty>
|
||
,
|
||
<collectingMunicipality box="[690,906,543,568]" pageId="2" pageNumber="394">Davis Mts. St. Pk.</collectingMunicipality>
|
||
,
|
||
<number box="[921,982,544,568]" pageId="2" pageNumber="394" value="4915.0">4915</number>
|
||
',
|
||
<geoCoordinate box="[1000,1132,544,568]" direction="north" orientation="latitude" pageId="2" pageNumber="394" precision="15" value="30.602222">
|
||
<number box="[1000,1029,544,568]" pageId="2" pageNumber="394" value="30.0">30</number>
|
||
°
|
||
<number box="[1039,1068,544,568]" pageId="2" pageNumber="394" value="36.0">36</number>
|
||
'08"N
|
||
</geoCoordinate>
|
||
<geoCoordinate box="[1142,1294,544,568]" direction="west" orientation="longitude" pageId="2" pageNumber="394" precision="15" value="-103.915276">
|
||
<number box="[1142,1184,544,568]" pageId="2" pageNumber="394" value="103.0">103</number>
|
||
°
|
||
<number box="[1194,1223,544,568]" pageId="2" pageNumber="394" value="54.0">54</number>
|
||
'
|
||
<number box="[1226,1255,544,568]" pageId="2" pageNumber="394" value="55.0">55</number>
|
||
"W
|
||
</geoCoordinate>
|
||
,
|
||
<collectingDate pageId="2" pageNumber="394" value="2014-07-21" valueMax="2014-07-25" valueMin="2014-07-21">
|
||
<date pageId="2" pageNumber="394" value="2014-07-21" valueMax="2014-07-25" valueMin="2014-07-21">
|
||
<number box="[1309,1379,544,568]" pageId="2" pageNumber="394" value="23.0" valueMax="25.0" valueMin="21.0">21–25</number>
|
||
July
|
||
<number box="[151,209,580,604]" pageId="2" pageNumber="394" value="2014.0">2014</number>
|
||
</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName box="[224,375,580,604]" pageId="2" pageNumber="394">T.L. Schiefer</collectorName>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:FA729970A22EFFAE189F86F3C13ACB24:77047EBDA22EFFAB198D846AC598CD4C" box="[389,477,580,604]" country="United States of America" county="Jeff Davis Co." latitude="30.602222" longLatPrecision="20" longitude="-103.915276" municipality="Davis Mts. St. Pk." name="Malaise" pageId="2" pageNumber="394" stateProvince="TEXAS">Malaise</location>
|
||
trap in riparian area in desert scrub/grassland,
|
||
<collectorName box="[1026,1290,580,604]" pageId="2" pageNumber="394">W.H. Cross Expedition</collectorName>
|
||
with unique
|
||
<specimenCount box="[151,257,615,640]" pageId="2" pageNumber="394" type="generic">specimen</specimenCount>
|
||
codes:
|
||
<specimenCode box="[347,507,616,640]" pageId="2" pageNumber="394">
|
||
MEM
|
||
<number box="[423,507,616,640]" pageId="2" pageNumber="394" value="207974.0">207974</number>
|
||
</specimenCode>
|
||
[point mounted
|
||
<specimenCount box="[701,806,615,640]" pageId="2" pageNumber="394" type="generic">specimen</specimenCount>
|
||
deposited in
|
||
<collectionCode box="[964,1036,616,640]" collectionName="USA, Mississippi, Mississippi, Mississippi State University" pageId="2" pageNumber="394">MEM</collectionCode>
|
||
],
|
||
<specimenCode box="[1056,1217,616,640]" pageId="2" pageNumber="394">
|
||
MEM
|
||
<number box="[1133,1217,616,640]" pageId="2" pageNumber="394" value="207975.0">207975</number>
|
||
</specimenCode>
|
||
[preserved in
|
||
<number box="[1385,1413,616,640]" pageId="2" pageNumber="394" value="95.0">95</number>
|
||
% ethanol for DNA study, deposited in
|
||
<collectionCode box="[602,672,652,676]" collectionName="USA, Mississippi, Mississippi, Mississippi State University" pageId="2" pageNumber="394">MEM</collectionCode>
|
||
] and
|
||
<specimenCode box="[745,910,652,676]" pageId="2" pageNumber="394">
|
||
MEM
|
||
<number box="[825,910,652,676]" pageId="2" pageNumber="394" value="207976.0">207976</number>
|
||
</specimenCode>
|
||
[preserved in
|
||
<number box="[1090,1117,652,676]" pageId="2" pageNumber="394" value="95.0">95</number>
|
||
% ethanol for DNA study, deposited in
|
||
<collectionCode box="[294,367,688,712]" collectionName="USA, Mississippi, Mississippi, Mississippi State University" pageId="2" pageNumber="394">MEM</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="394" type="description">
|
||
<paragraph pageId="2" pageNumber="394">
|
||
<emphasis bold="true" box="[199,645,724,748]" pageId="2" pageNumber="394">
|
||
<typeStatus box="[199,307,725,748]" pageId="2" pageNumber="394">Holotype</typeStatus>
|
||
measurements and indices:
|
||
</emphasis>
|
||
HW
|
||
<number box="[710,762,724,748]" pageId="2" pageNumber="394" value="0.24">0.24</number>
|
||
, HL
|
||
<number box="[822,874,724,748]" pageId="2" pageNumber="394" value="0.24">0.24</number>
|
||
, EL
|
||
<number box="[931,982,724,748]" pageId="2" pageNumber="394" value="0.11">0.11</number>
|
||
, MDL
|
||
<number box="[1067,1119,724,748]" pageId="2" pageNumber="394" value="0.13">0.13</number>
|
||
, SL
|
||
<number box="[1174,1226,724,748]" pageId="2" pageNumber="394" value="0.08">0.08</number>
|
||
, PedL
|
||
<number box="[1308,1360,724,748]" pageId="2" pageNumber="394" value="0.06">0.06</number>
|
||
, FlaLI
|
||
<number box="[151,202,760,784]" pageId="2" pageNumber="394" value="0.04">0.04</number>
|
||
, FlaLII
|
||
<number box="[293,345,760,784]" pageId="2" pageNumber="394" value="0.04">0.04</number>
|
||
, FlaLXI
|
||
<number box="[447,498,760,784]" pageId="2" pageNumber="394" value="0.12">0.12</number>
|
||
, MH
|
||
<number box="[563,614,760,784]" pageId="2" pageNumber="394" value="0.29">0.29</number>
|
||
, ML
|
||
<number box="[676,727,760,784]" pageId="2" pageNumber="394" value="0.46">0.46</number>
|
||
, PrW
|
||
<number box="[798,849,760,784]" pageId="2" pageNumber="394" value="0.2">0.20</number>
|
||
, PW
|
||
<number box="[911,962,760,784]" pageId="2" pageNumber="394" value="0.09">0.09</number>
|
||
, PL
|
||
<number box="[1015,1066,760,784]" pageId="2" pageNumber="394" value="0.07">0.07</number>
|
||
, AIIIW
|
||
<number box="[1160,1212,760,784]" pageId="2" pageNumber="394" value="0.14">0.14</number>
|
||
, AIIIL
|
||
<number box="[1297,1348,760,784]" pageId="2" pageNumber="394" value="0.07">0.07</number>
|
||
, AIVW
|
||
<number box="[151,202,796,820]" pageId="2" pageNumber="394" value="0.2">0.20</number>
|
||
, AIVL
|
||
<number box="[288,339,796,820]" pageId="2" pageNumber="394" value="0.09">0.09</number>
|
||
, FFeW
|
||
<number box="[428,479,796,820]" pageId="2" pageNumber="394" value="0.05">0.05</number>
|
||
, FFeL
|
||
<number box="[559,610,796,820]" pageId="2" pageNumber="394" value="0.23">0.23</number>
|
||
, HFeL
|
||
<number box="[694,745,796,820]" pageId="2" pageNumber="394" value="0.24">0.24</number>
|
||
, HTiL
|
||
<number box="[825,876,796,820]" pageId="2" pageNumber="394" value="0.26">0.26</number>
|
||
, FWL
|
||
<number box="[954,1005,796,820]" pageId="2" pageNumber="394" value="1.46">1.46</number>
|
||
, CI
|
||
<number box="[1052,1096,796,820]" pageId="2" pageNumber="394" value="100.0">100</number>
|
||
, MI
|
||
<number box="[1149,1179,796,820]" pageId="2" pageNumber="394" value="63.0">63</number>
|
||
, PI
|
||
<number box="[1223,1269,796,820]" pageId="2" pageNumber="394" value="128.0">128</number>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="2" pageNumber="394">
|
||
<emphasis box="[199,864,831,856]" italics="true" pageId="2" pageNumber="394">
|
||
Additional
|
||
<typeStatus box="[324,424,832,856]" pageId="2" pageNumber="394">paratype</typeStatus>
|
||
measurements and indices (two males)
|
||
</emphasis>
|
||
: HW
|
||
<number box="[933,984,832,856]" pageId="2" pageNumber="394" value="0.24">0.24</number>
|
||
, HL
|
||
<number box="[1041,1092,832,856]" pageId="2" pageNumber="394" value="0.22">0.22</number>
|
||
, EL
|
||
<number box="[1145,1259,832,856]" pageId="2" pageNumber="394" value="0.1" valueMax="0.11" valueMin="0.09">0.09–0.11</number>
|
||
, MDL
|
||
<number box="[1340,1391,832,856]" pageId="2" pageNumber="394" value="0.13">0.13</number>
|
||
, SL
|
||
<number box="[151,202,868,892]" pageId="2" pageNumber="394" value="0.09">0.09</number>
|
||
, PedL
|
||
<number box="[283,398,868,892]" pageId="2" pageNumber="394" value="0.065" valueMax="0.07" valueMin="0.06">0.06–0.07</number>
|
||
, FlaLI
|
||
<number box="[482,533,868,892]" pageId="2" pageNumber="394" value="0.04">0.04</number>
|
||
, FlaLII
|
||
<number box="[627,678,868,892]" pageId="2" pageNumber="394" value="0.04">0.04</number>
|
||
, FlaLXI
|
||
<number box="[783,896,868,892]" pageId="2" pageNumber="394" value="0.11499999999999999" valueMax="0.12" valueMin="0.11">0.11–0.12</number>
|
||
, MH
|
||
<number box="[963,1078,868,892]" pageId="2" pageNumber="394" value="0.305" valueMax="0.31" valueMin="0.3">0.30–0.31</number>
|
||
, ML
|
||
<number box="[1142,1256,868,892]" pageId="2" pageNumber="394" value="0.455" valueMax="0.46" valueMin="0.45">0.45–0.46</number>
|
||
, PrW
|
||
<number box="[1330,1381,868,892]" pageId="2" pageNumber="394" value="0.18">0.18</number>
|
||
, PW
|
||
<number box="[151,265,904,928]" pageId="2" pageNumber="394" value="0.07500000000000001" valueMax="0.08" valueMin="0.07">0.07–0.08</number>
|
||
, PL
|
||
<number box="[320,435,904,928]" pageId="2" pageNumber="394" value="0.065" valueMax="0.07" valueMin="0.06">0.06–0.07</number>
|
||
, AIIIW
|
||
<number box="[532,583,904,928]" pageId="2" pageNumber="394" value="0.15">0.15</number>
|
||
, AIIIL
|
||
<number box="[671,786,904,928]" pageId="2" pageNumber="394" value="0.085" valueMax="0.1" valueMin="0.07">0.07–0.10</number>
|
||
, AIVW
|
||
<number box="[884,999,904,928]" pageId="2" pageNumber="394" value="0.185" valueMax="0.19" valueMin="0.18">0.18–0.19</number>
|
||
, AIVL
|
||
<geoCoordinate box="[1088,1153,904,928]" pageId="2" pageNumber="394">
|
||
0
|
||
<number box="[1101,1153,904,928]" pageId="2" pageNumber="394" value="0.1">0.10</number>
|
||
</geoCoordinate>
|
||
, FFeW
|
||
<number box="[1246,1297,904,928]" pageId="2" pageNumber="394" value="0.05">0.05</number>
|
||
, FFeL
|
||
<number box="[1380,1431,904,928]" pageId="2" pageNumber="394" value="0.21">0.21</number>
|
||
, HFeL
|
||
<number box="[223,274,940,964]" pageId="2" pageNumber="394" value="0.23">0.23</number>
|
||
, HTiL
|
||
<number box="[354,406,940,964]" pageId="2" pageNumber="394" value="0.26">0.26</number>
|
||
, FWL
|
||
<number box="[484,535,940,964]" pageId="2" pageNumber="394" value="1.38">1.38</number>
|
||
, CI
|
||
<number box="[582,626,940,964]" pageId="2" pageNumber="394" value="109.0">109</number>
|
||
, MI
|
||
<number box="[679,751,940,964]" pageId="2" pageNumber="394" value="67.0" valueMax="69.0" valueMin="65.0">65–69</number>
|
||
, PI
|
||
<number box="[795,894,940,964]" pageId="2" pageNumber="394" value="115.5" valueMax="117.0" valueMin="114.0">114–117</number>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1180,975,1000]" pageId="2" pageNumber="394" type="etymology">
|
||
<paragraph box="[199,1180,975,1000]" pageId="2" pageNumber="394">
|
||
<emphasis bold="true" box="[199,333,976,1000]" pageId="2" pageNumber="394">Etymology.</emphasis>
|
||
Named for the Chihuahuan Desert region where specimens were collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="394" type="diagnosis">
|
||
<paragraph pageId="2" pageNumber="394">
|
||
<emphasis bold="true" box="[199,414,1012,1036]" pageId="2" pageNumber="394">Diagnosis (male).</emphasis>
|
||
Of the six males that have been described in the
|
||
<taxonomicName box="[1072,1233,1012,1036]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1072,1233,1012,1036]" italics="true" pageId="2" pageNumber="394">Leptanilloides</emphasis>
|
||
</taxonomicName>
|
||
genus-group,
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Leptanilloides" genus.bestMatchDistance="502" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="chihuahuaensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="394">
|
||
<taxonomicName box="[1413,1432,1013,1036]" class="Insecta" family="Formicidae" genus="Leptanilloides" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">L</taxonomicName>
|
||
. chihuahuaensis
|
||
</emphasis>
|
||
</taxonomicName>
|
||
is most similar to "Leptanilloidinae male
|
||
<number box="[827,840,1048,1072]" pageId="2" pageNumber="394" value="1.0">1</number>
|
||
" from southern Mexico described by Borowiec & Longino (
|
||
<number box="[262,322,1084,1108]" pageId="2" pageNumber="394" value="2011.0">2011</number>
|
||
).
|
||
<emphasis box="[341,683,1084,1108]" italics="true" pageId="2" pageNumber="394">
|
||
<taxonomicName box="[341,501,1084,1108]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">Leptanilloides</taxonomicName>
|
||
chihuahuaensis
|
||
</emphasis>
|
||
differs from "Leptanilloidinae male
|
||
<number box="[1096,1109,1084,1108]" pageId="2" pageNumber="394" value="1.0">1</number>
|
||
" by having finer and sparser pilosity, in dorsal view the petiole being widened anteriorly and posteriorly, parameres being narrower and incurved, and the hindwing having three hamuli (instead of two). Males of
|
||
<emphasis box="[997,1201,1156,1180]" italics="true" pageId="2" pageNumber="394">
|
||
<taxonomicName box="[997,1201,1156,1180]" class="Insecta" family="Formicidae" genus="Leptanilloides" genus.bestMatchDistance="573" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="chihuahuaensis">L. chihuahuaensis</taxonomicName>
|
||
</emphasis>
|
||
differ from males of
|
||
<taxonomicName authority="Kusnezov, 1953" authorityName="Kusnezov" authorityYear="1953" box="[151,548,1191,1216]" class="Insecta" family="Formicidae" genus="Amyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="golbachi">
|
||
<emphasis box="[151,361,1191,1216]" italics="true" pageId="2" pageNumber="394">Amyrmex golbachi</emphasis>
|
||
Kusnezov,
|
||
<number box="[492,548,1192,1216]" pageId="2" pageNumber="394" value="1953.0">1953</number>
|
||
</taxonomicName>
|
||
by lacking submarginal cells and a stigma in the forewing, possessing a free M vein, and having a larger paramere (about as long as petiole); from
|
||
<emphasis box="[975,1123,1228,1252]" italics="true" pageId="2" pageNumber="394">L. mckennae</emphasis>
|
||
Longino
|
||
<number box="[1244,1300,1228,1252]" pageId="2" pageNumber="394" value="2003.0">2003</number>
|
||
by lacking submarginal cells and a stigma in forewing and having a much shorter paramere (in side view) (Ward
|
||
<number box="[1301,1361,1263,1288]" pageId="2" pageNumber="394" value="2007.0">2007</number>
|
||
); from
|
||
<emphasis box="[151,286,1300,1324]" italics="true" pageId="2" pageNumber="394">L. nubecula</emphasis>
|
||
Donaso, Vieira & Wild,
|
||
<number box="[579,635,1300,1324]" pageId="2" pageNumber="394" value="2006.0">2006</number>
|
||
by being light to medium brown instead of dark brownish-black and being much smaller (
|
||
<collectionCode box="[417,456,1335,1360]" pageId="2" pageNumber="394">HL</collectionCode>
|
||
<quantity box="[473,624,1335,1360]" metricMagnitude="-3" metricUnit="m" metricValue="5.842" metricValueMax="6.096" metricValueMin="5.587999999999999" pageId="2" pageNumber="394" unit="in" value="0.22999999999999998" valueMax="0.24" valueMin="0.22">0.22–0.24 in</quantity>
|
||
<emphasis box="[641,855,1335,1360]" italics="true" pageId="2" pageNumber="394">
|
||
<taxonomicName box="[641,855,1335,1360]" class="Insecta" family="Formicidae" genus="Leptanilloides" genus.bestMatchDistance="677" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="chihuahuaensis">L. chihuahuaensis</taxonomicName>
|
||
</emphasis>
|
||
vs.
|
||
<collectionCode box="[921,959,1337,1360]" pageId="2" pageNumber="394">HL</collectionCode>
|
||
<quantity box="[976,1064,1335,1360]" metricMagnitude="-3" metricUnit="m" metricValue="8.128" pageId="2" pageNumber="394" unit="in" value="0.32">0.32 in</quantity>
|
||
<emphasis box="[1081,1223,1335,1360]" italics="true" pageId="2" pageNumber="394">L. nubecula</emphasis>
|
||
), and from both "Leptanilloidinae male
|
||
<number box="[414,428,1372,1396]" pageId="2" pageNumber="394" value="2.0">2</number>
|
||
and male
|
||
<number box="[544,558,1372,1396]" pageId="2" pageNumber="394" value="3.0">3</number>
|
||
" (Borowiec & Longino
|
||
<number box="[832,888,1372,1396]" pageId="2" pageNumber="394" value="2011.0">2011</number>
|
||
) by the much smaller size (
|
||
<collectionCode box="[1196,1234,1372,1396]" pageId="2" pageNumber="394">HL</collectionCode>
|
||
<number box="[1241,1353,1372,1396]" pageId="2" pageNumber="394" value="0.22999999999999998" valueMax="0.24" valueMin="0.22">0.22–0.24</number>
|
||
vs.
|
||
<collectionCode box="[1399,1436,1373,1396]" pageId="2" pageNumber="394">HL</collectionCode>
|
||
<number box="[151,263,1408,1432]" pageId="2" pageNumber="394" value="0.315" valueMax="0.33" valueMin="0.3">0.30–0.33</number>
|
||
of "Leptanilloidinae male
|
||
<number box="[568,582,1408,1432]" pageId="2" pageNumber="394" value="2.0">2</number>
|
||
and male
|
||
<number box="[701,715,1407,1432]" pageId="2" pageNumber="394" value="3.0">3</number>
|
||
") and by lacking submarginal cells and stigma in the forewing. Five other male morphotypes unassociated with workers were briefly discussed by Ward & Brady (
|
||
<number box="[1363,1424,1444,1468]" pageId="2" pageNumber="394" value="2009.0">2009</number>
|
||
).
|
||
<emphasis box="[151,499,1479,1504]" italics="true" pageId="2" pageNumber="394">
|
||
<taxonomicName box="[151,311,1479,1504]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">Leptanilloides</taxonomicName>
|
||
chihuahuaensis
|
||
</emphasis>
|
||
differs from each of these by lacking submarginal cells and a stigma in the forewing. Geographically,
|
||
<taxonomicName box="[451,654,1516,1540]" class="Insecta" family="Formicidae" genus="Leptanilloides" genus.bestMatchDistance="782" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="chihuahuaensis">
|
||
<emphasis box="[451,654,1516,1540]" italics="true" pageId="2" pageNumber="394">L. chihuahuaensis</emphasis>
|
||
</taxonomicName>
|
||
is the only species thus far know to occur in the United States.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="398" pageId="2" pageNumber="394" type="description">
|
||
<paragraph pageId="2" pageNumber="394">
|
||
<emphasis box="[199,411,1551,1576]" italics="true" pageId="2" pageNumber="394">Description (male)</emphasis>
|
||
: Body size minute. Color light yellowish-brown, head and margins of abdominal segments IV–VII darker, appendages (antennae, mandibles, legs) lighter. Integument mostly smooth and shiny, with simple curved to strongly curved suberect to decumbent setae; pilosity on head, mesosoma, petiole, and first gastral tergite (
|
||
<collectionCode box="[159,209,1660,1684]" pageId="2" pageNumber="394">AIII</collectionCode>
|
||
) scattered and not obscuring the shininess of the integument; pilosity denser on remainder of gaster and appendages.
|
||
</paragraph>
|
||
<paragraph pageId="2" pageNumber="394">Head in full-face view excluding eyes about as long as wide, widest above eyes; posterior corners of head evenly rounded. Eyes large, bulging, occupying almost half the side of head. Ocelli small, protruding slightly, arranged in almost an equilateral triangle, but with distance between lateral ocelli slightly greater than distance between lateral ocellus and median ocellus; distance between median ocellus and eye approximately the length of eye. Clypeus short, tranverse, lacking lamelli; lateroclypeal teeth and hypostomal teeth lacking. Mandible slender, falcate; apex blunt; masticatory margin edentate; external margin of mandible mostly evenly curved along its length; mandible tips overlap at closure.</paragraph>
|
||
<caption pageId="3" pageNumber="395" targetBox="[151,1436,193,1505]" targetPageId="3">
|
||
<paragraph pageId="3" pageNumber="395">
|
||
<emphasis bold="true" box="[151,276,1527,1548]" pageId="3" pageNumber="395">FIGURE 1.</emphasis>
|
||
<emphasis box="[283,593,1526,1549]" italics="true" pageId="3" pageNumber="395">
|
||
<taxonomicName box="[283,428,1526,1549]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="395" phylum="Arthropoda" rank="genus">Leptanilloides</taxonomicName>
|
||
chihuahuaensis
|
||
</emphasis>
|
||
, male, holotype (MEM 207971): (A) head in full-face view, (B) petiole in dorsal view, (C) tergite VIII and exposed male genitalia in dorsal view, (D) sternite IX and exposed male genitalia in ventral view, (E) lateral habitus, and (F) dorsal habitus. Abbreviations: ASIX = ninth abdominal sternite, ATVIII = eighth abdominal tergite, Ax = axilla, Msp = mesopleuron, MsScl = mesoscutellum, MsSct = mesoscutum, Mtn = metanotum, P = petiole, Ped = Pedicel, Pm = paramere, Prnt = pronotum, Prpd = propodeum, Pv = Penivalva, Sc = Scape, and Vol = volsella.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="3" pageNumber="395">
|
||
Antennal sockets circular and exposed, located at the anterior clypeal margin with anterior edge of sockets slightly overlapping anterior clypeal margin. Antenna
|
||
<number box="[782,807,1747,1771]" pageId="3" pageNumber="395" value="13.0">13</number>
|
||
-merous; scape, pedicel, and each flagellomere longer than wide. Scape length about twice the length of the first flagellomere, and about the combined length of the first and second flagellomere; scape subequal to the length of ultimate flagellomere; pedicel thickened, length about
|
||
<number box="[1401,1436,1819,1843]" pageId="3" pageNumber="395" value="1.5">1.5</number>
|
||
times width; flagellomeres
|
||
<number box="[466,516,1855,1879]" pageId="3" pageNumber="395" value="6.0" valueMax="11.0" valueMin="1.0">1-11</number>
|
||
each at least twice as long as wide; first and second flagellomeres subequal in length.
|
||
</paragraph>
|
||
<paragraph lastPageId="5" lastPageNumber="397" pageId="3" pageNumber="395">
|
||
Pronotum U-shaped in dorsal view, reduced anteromedially to a thin horizontal strip, set below the level of the dorsally protruding mesonotum; pronotum triangular in lateral view, with pointed posterior apex directed toward wing base. Mesoscutum lacking notauli; parapsidal lines not discernable. Axillae depressed, not meeting medially, connected by a narrow furrow. Tegula, or tegula-like structure present at wing base, minute, inconspicuous, with
|
||
<number pageId="5" pageNumber="397" value="2.5" valueMax="3.0" valueMin="2.0">2- 3</number>
|
||
erect setae. Mesopleuron lacking oblique transverse sulcus, not divided into anepisternum and katepisternum. Mesoscutellum prominently bulging, as seen in lateral view. Metapleural gland not discernable. Propodeum with dorsal and declivious surfaces not differentiated, evenly rounded. Propodeal spiracle small, ovate, positioned slightly below midheight of propodeum and slightly posterior to the midlength. Legs slender, mesotibia and metatibia each with two simple spurs, pretarsal claw lacking preapical tooth.
|
||
</paragraph>
|
||
<caption pageId="4" pageNumber="396" targetBox="[195,1388,193,1886]" targetPageId="4">
|
||
<paragraph pageId="4" pageNumber="396">
|
||
<emphasis bold="true" box="[151,281,1911,1933]" pageId="4" pageNumber="396">FIGURE 2.</emphasis>
|
||
<emphasis box="[293,607,1911,1933]" italics="true" pageId="4" pageNumber="396">
|
||
<taxonomicName box="[293,438,1911,1933]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">Leptanilloides</taxonomicName>
|
||
chihuahuaensis
|
||
</emphasis>
|
||
, male, holotype (MEM 207971): (A) lateral habitus and (B) forewing and hindwing showing labeled veins. Abbreviations: 2r-rs =second radial sector cross-vein, A = anal vein, C = costal vein, Cu = cubital vein, cu-a = cubital-anal cross-vein, ha = hamuli, M = medial vein, M+Cu = merged medial-cubital veins, Rs+M = merged medial and radial sector plus medial veins, Rs = radial sector, Sc+R = merged subcostal and radial veins.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="5" pageNumber="397">
|
||
Wings with reduced venation; clear; fringed with short to longer fine setae; with numerous short, fine microsetae evenly distributed across wing surfaces and with scattered longer setae present. Fore wing: Costal (
|
||
<collectionCode box="[1408,1427,404,428]" pageId="5" pageNumber="397">C</collectionCode>
|
||
) vein present, about ½ the length of wing, tubular basally for less than ¼ the length of wing, then becoming nebulous before fading out completely. Pterostigma not present. Subcostal and radial veins fused forming Sc+R; located just below and parallel to costal vein, approximately the same length as costal vein and similarly tubular for about ½ of its length before becoming nebulous. Medial (
|
||
<collectionCode box="[830,856,548,572]" pageId="5" pageNumber="397">M</collectionCode>
|
||
) and cubital (Cu) veins fused (
|
||
<collectionCode box="[1227,1253,548,572]" pageId="5" pageNumber="397">M</collectionCode>
|
||
+Cu), nebulous, slightly less than ¼ of the wing length. Anal (
|
||
<collectionCode box="[686,708,583,608]" pageId="5" pageNumber="397">A</collectionCode>
|
||
) vein present beneath M+Cu, about the same length as M+Cu, tubular for slightly more than half of its length before becoming nebulous; apically M+Cu connected to A by nebulous crossvein cu-a; subbasal cell present between M+Cu and A. Branching upward from the top of Cu-a is the tubular abscissa Rs+M (the joined veins M·f
|
||
<number box="[680,690,692,716]" pageId="5" pageNumber="397" value="1.0">1</number>
|
||
and Rs+
|
||
<collectionCode box="[801,827,692,716]" pageId="5" pageNumber="397">M</collectionCode>
|
||
). Radial sector (Rs), tubular, branches upward from Rs+M, then becomes nebulous just past the nebulous/spectral crossvein
|
||
<number box="[975,988,728,752]" pageId="5" pageNumber="397" value="2.0">2</number>
|
||
r-rs (
|
||
<number box="[1045,1053,727,752]" pageId="5" pageNumber="397" value="1.0">1</number>
|
||
r-rs is absent). M branches off of Rs+M just before Rs, appearing spectral basally then continuing as a nebulous vein that does not reach the wing margin. Cu is present as a nebulous vein that branches off near the apex of A; Cu does not reach the wing margin. Posterior margin of fore wing with narrow, conspicuous fold where hamuli attach.
|
||
</paragraph>
|
||
<paragraph pageId="5" pageNumber="397">Hind wing with Sc+R present, tubular basally before becoming nebulous, less than ¼ the length of wing. No other veins present. Anterior margin of hind wing past midlength with a thin, but conspicuous dark stigma. Three hamuli present in the region of the stigma. Jugal lobe absent.</paragraph>
|
||
<paragraph pageId="5" pageNumber="397">
|
||
Petiole (abdominal segment
|
||
<collectionCode box="[530,549,980,1004]" pageId="5" pageNumber="397">II</collectionCode>
|
||
) subquadrate in lateral view, about as long as high or wide, not constricted posteriorly; in dorsal view anterior corners widened, sides slightly constricted, appearing shallowly concave before widening posteriorly; petiole broadly joined to abdominal segment III; petiolar spiracle located on anterior third of the segment, near anterodorsal extremity. Abdominal segment III larger than petiole, not developed as postpetiole, and not separated from abdominal segment IV by a constriction. Abdominal spiracles III, IV, V, and VI located on anterior third of tergites (abdominal spiracle
|
||
<collectionCode box="[682,721,1161,1184]" pageId="5" pageNumber="397">VII</collectionCode>
|
||
not visible in examined specimens). Abdominal tergite VIII (pygidium) small, simple and visible dorsally, not entirely covered by abdominal tergite VII. Pygostyli absent. Abdominal sternite IX (subgenital plate) with posterior margin broadly concave, but with a median subtriangular process. Paramere (basimere + telomere) small (about the length of petiole); basimere roughly triangular, widest at base, narrowing to incurved falcate telomere that terminates in a blunt apex. Volsella triangular, with slightly rounded apex; volsella not differentiated into digitus and cuspis. Penisvalva elongate triangular, extending to about as far as paramere apex.
|
||
</paragraph>
|
||
<caption pageId="5" pageNumber="397" targetBox="[159,1390,1562,1863]" targetIsTable="true" targetPageId="5">
|
||
<paragraph pageId="5" pageNumber="397">
|
||
<emphasis bold="true" box="[151,265,1448,1471]" pageId="5" pageNumber="397">TABLE 1.</emphasis>
|
||
Information for six
|
||
<taxonomicName box="[479,631,1447,1471]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="397" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[479,631,1447,1471]" italics="true" pageId="5" pageNumber="397">Leptanilloides</emphasis>
|
||
</taxonomicName>
|
||
specimens that have available
|
||
<emphasis box="[959,1005,1447,1471]" italics="true" pageId="5" pageNumber="397">COI</emphasis>
|
||
sequence data: species name, country of origin, GenBank and BOLD accession numbers, length of COI sequence fragment, and uncorrected (p) distance as compared to
|
||
<emphasis box="[290,484,1511,1535]" italics="true" pageId="5" pageNumber="397">
|
||
<taxonomicName box="[290,484,1511,1535]" class="Insecta" family="Formicidae" genus="Leptanilloides" genus.bestMatchDistance="2151" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="397" phylum="Arthropoda" rank="species" species="chihuahuaensis">L. chihuahuaensis</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="5" pageNumber="397">
|
||
<table box="[159,1390,1562,1863]" pageId="5" pageNumber="397">
|
||
<tr box="[159,1390,1562,1584]" pageId="5" pageNumber="397">
|
||
<td box="[159,236,1562,1584]" pageId="5" pageNumber="397">Species</td>
|
||
<td box="[396,478,1562,1584]" pageId="5" pageNumber="397">Country</td>
|
||
<td box="[659,767,1562,1584]" pageId="5" pageNumber="397">GenBank#</td>
|
||
<td box="[830,912,1562,1584]" pageId="5" pageNumber="397">BOLD#</td>
|
||
<td box="[1049,1191,1562,1584]" pageId="5" pageNumber="397">COI Size (bp)</td>
|
||
<td box="[1241,1390,1562,1584]" pageId="5" pageNumber="397">p-distance (%)</td>
|
||
</tr>
|
||
<tr box="[159,1390,1604,1656]" pageId="5" pageNumber="397">
|
||
<td box="[159,317,1604,1656]" pageId="5" pageNumber="397">
|
||
<emphasis box="[159,304,1604,1626]" italics="true" pageId="5" pageNumber="397">
|
||
<taxonomicName box="[159,304,1604,1626]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="397" phylum="Arthropoda" rank="genus">Leptanilloides</taxonomicName>
|
||
</emphasis>
|
||
<emphasis box="[159,317,1634,1656]" italics="true" pageId="5" pageNumber="397">chihuahuaensis</emphasis>
|
||
</td>
|
||
<td box="[396,529,1604,1626]" pageId="5" pageNumber="397">USA (Texas)</td>
|
||
<td box="[659,769,1604,1626]" pageId="5" pageNumber="397">KT007966</td>
|
||
<td box="[830,861,1604,1626]" pageId="5" pageNumber="397">n/a</td>
|
||
<td box="[1049,1087,1604,1626]" pageId="5" pageNumber="397">658</td>
|
||
<td box="[1242,1250,1605,1626]" pageId="5" pageNumber="397">-</td>
|
||
</tr>
|
||
<tr box="[159,1390,1676,1698]" pageId="5" pageNumber="397">
|
||
<td box="[159,263,1676,1698]" pageId="5" pageNumber="397">
|
||
<emphasis box="[159,263,1676,1698]" italics="true" pageId="5" pageNumber="397">L. gracilis</emphasis>
|
||
</td>
|
||
<td box="[396,578,1676,1698]" pageId="5" pageNumber="397">Mexico (Chiapas)</td>
|
||
<td box="[659,690,1676,1698]" pageId="5" pageNumber="397">n/a</td>
|
||
<td box="[830,1002,1676,1698]" pageId="5" pageNumber="397">ASLAM1070-11</td>
|
||
<td box="[1049,1087,1676,1698]" pageId="5" pageNumber="397">282</td>
|
||
<td box="[1242,1286,1676,1698]" pageId="5" pageNumber="397">10.1</td>
|
||
</tr>
|
||
<tr box="[159,1390,1717,1739]" pageId="5" pageNumber="397">
|
||
<td box="[159,263,1717,1739]" pageId="5" pageNumber="397">
|
||
<emphasis box="[159,263,1717,1739]" italics="true" pageId="5" pageNumber="397">L. gracilis</emphasis>
|
||
</td>
|
||
<td box="[396,495,1717,1739]" pageId="5" pageNumber="397">Honduras</td>
|
||
<td box="[659,690,1717,1739]" pageId="5" pageNumber="397">n/a</td>
|
||
<td box="[830,1001,1717,1739]" pageId="5" pageNumber="397">ASLAM1144-11</td>
|
||
<td box="[1049,1087,1717,1739]" pageId="5" pageNumber="397">658</td>
|
||
<td box="[1242,1286,1717,1739]" pageId="5" pageNumber="397">10.3</td>
|
||
</tr>
|
||
<tr box="[159,1390,1758,1781]" pageId="5" pageNumber="397">
|
||
<td box="[159,263,1758,1781]" pageId="5" pageNumber="397">
|
||
<emphasis box="[159,263,1758,1781]" italics="true" pageId="5" pageNumber="397">L. gracilis</emphasis>
|
||
</td>
|
||
<td box="[396,494,1758,1781]" pageId="5" pageNumber="397">Honduras</td>
|
||
<td box="[659,758,1758,1781]" pageId="5" pageNumber="397">JF863464</td>
|
||
<td box="[830,989,1758,1781]" pageId="5" pageNumber="397">ASLAM381-11</td>
|
||
<td box="[1049,1087,1758,1781]" pageId="5" pageNumber="397">525</td>
|
||
<td box="[1242,1273,1758,1781]" pageId="5" pageNumber="397">8.2</td>
|
||
</tr>
|
||
<tr box="[159,1390,1800,1822]" pageId="5" pageNumber="397">
|
||
<td box="[159,278,1800,1822]" pageId="5" pageNumber="397">
|
||
<emphasis box="[159,278,1800,1822]" italics="true" pageId="5" pageNumber="397">L. nubecula</emphasis>
|
||
</td>
|
||
<td box="[396,481,1800,1822]" pageId="5" pageNumber="397">Ecuador</td>
|
||
<td box="[659,772,1800,1822]" pageId="5" pageNumber="397">DQ353302</td>
|
||
<td box="[830,986,1800,1822]" pageId="5" pageNumber="397">GBAH2069-06</td>
|
||
<td box="[1049,1087,1800,1822]" pageId="5" pageNumber="397">844</td>
|
||
<td box="[1242,1286,1800,1822]" pageId="5" pageNumber="397">20.1</td>
|
||
</tr>
|
||
<tr box="[159,1390,1841,1863]" pageId="5" pageNumber="397">
|
||
<td box="[159,278,1841,1863]" pageId="5" pageNumber="397">
|
||
<emphasis box="[159,180,1842,1863]" italics="true" pageId="5" pageNumber="397">L.</emphasis>
|
||
MAS001
|
||
</td>
|
||
<td box="[396,507,1841,1863]" pageId="5" pageNumber="397">Costa Rica</td>
|
||
<td box="[659,767,1841,1863]" pageId="5" pageNumber="397">KF371235</td>
|
||
<td box="[830,990,1841,1863]" pageId="5" pageNumber="397">ACGAG099-11</td>
|
||
<td box="[1049,1087,1841,1863]" pageId="5" pageNumber="397">503</td>
|
||
<td box="[1242,1286,1841,1863]" pageId="5" pageNumber="397">10.6</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph pageId="5" pageNumber="397">
|
||
<emphasis bold="true" box="[199,409,1932,1956]" pageId="5" pageNumber="397">Analysis of COI.</emphasis>
|
||
A
|
||
<number box="[455,497,1932,1956]" pageId="5" pageNumber="397" value="658.0">658</number>
|
||
bp region of the mitochondrial gene
|
||
<emphasis box="[955,1010,1932,1956]" italics="true" pageId="5" pageNumber="397">COI,</emphasis>
|
||
the barcode region, was sequenced successfully and the resulting consensus sequence was submitted to GenBank (
|
||
<tableCitation box="[1085,1173,1968,1992]" captionStart="TABLE 1" captionStartId="5.[151,239,1448,1471]" captionTargetBox="[159,1390,1562,1863]" captionTargetPageId="5" captionText="TABLE 1. Information for six Leptanilloides specimens that have available COI sequence data: species name, country of origin, GenBank and BOLD accession numbers, length of COI sequence fragment, and uncorrected (p) distance as compared to L. chihuahuaensis." pageId="5" pageNumber="397">
|
||
Table
|
||
<number box="[1160,1173,1968,1992]" pageId="5" pageNumber="397" value="1.0">1</number>
|
||
</tableCitation>
|
||
). Five additional
|
||
<emphasis box="[1388,1436,1968,1992]" italics="true" pageId="5" pageNumber="397">COI</emphasis>
|
||
sequences for
|
||
<emphasis box="[310,470,2004,2028]" italics="true" pageId="5" pageNumber="397">
|
||
<taxonomicName box="[310,470,2004,2028]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="397" phylum="Arthropoda" rank="genus">Leptanilloides</taxonomicName>
|
||
</emphasis>
|
||
were found and downloaded from the BOLD database (
|
||
<tableCitation box="[1093,1175,2004,2028]" captionStart="TABLE 1" captionStartId="5.[151,239,1448,1471]" captionTargetBox="[159,1390,1562,1863]" captionTargetPageId="5" captionText="TABLE 1. Information for six Leptanilloides specimens that have available COI sequence data: species name, country of origin, GenBank and BOLD accession numbers, length of COI sequence fragment, and uncorrected (p) distance as compared to L. chihuahuaensis." pageId="5" pageNumber="397">
|
||
Table
|
||
<number box="[1163,1175,2004,2028]" pageId="5" pageNumber="397" value="1.0">1</number>
|
||
</tableCitation>
|
||
), including three for
|
||
<emphasis box="[1413,1436,2005,2028]" italics="true" pageId="5" pageNumber="397">L.</emphasis>
|
||
</paragraph>
|
||
<paragraph pageId="6" pageNumber="398">
|
||
<emphasis box="[151,237,151,176]" italics="true" pageId="6" pageNumber="398">gracilis</emphasis>
|
||
Borowiec & Longino (southern Mexico, Honduras), one for
|
||
<emphasis box="[924,1056,151,176]" italics="true" pageId="6" pageNumber="398">L. nubecula</emphasis>
|
||
Donoso
|
||
<emphasis box="[1158,1215,151,176]" italics="true" pageId="6" pageNumber="398">et al.</emphasis>
|
||
(Ecuador), and one for an undescribed species
|
||
<emphasis box="[478,501,189,212]" italics="true" pageId="6" pageNumber="398">L.</emphasis>
|
||
MAS001 (Costa Rica). Comparing
|
||
<emphasis box="[936,1146,188,212]" italics="true" pageId="6" pageNumber="398">
|
||
<taxonomicName box="[936,1146,188,212]" class="Insecta" family="Formicidae" genus="Leptanilloides" genus.bestMatchDistance="2349" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="chihuahuaensis">L. chihuahuaensis</taxonomicName>
|
||
</emphasis>
|
||
to the other specimens, pairwise genetic distances were all above 8.0%, ranging from
|
||
<number box="[849,884,224,248]" pageId="6" pageNumber="398" value="8.2">8.2</number>
|
||
% (
|
||
<emphasis box="[924,1039,223,248]" italics="true" pageId="6" pageNumber="398">L. gracilis</emphasis>
|
||
, Honduras) to
|
||
<number box="[1208,1257,224,248]" pageId="6" pageNumber="398" value="20.1">20.1</number>
|
||
% (
|
||
<emphasis box="[1296,1429,223,248]" italics="true" pageId="6" pageNumber="398">L. nubecula</emphasis>
|
||
, Ecuador).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="398" type="discussion">
|
||
<paragraph box="[151,283,357,383]" pageId="6" pageNumber="398">
|
||
<emphasis bold="true" box="[151,283,357,383]" pageId="6" pageNumber="398">
|
||
<heading bold="true" box="[151,283,357,383]" fontSize="11" level="1" pageId="6" pageNumber="398" reason="0">Discussion</heading>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph pageId="6" pageNumber="398">
|
||
Of the
|
||
<number box="[233,261,428,452]" pageId="6" pageNumber="398" value="16.0">16</number>
|
||
species in the
|
||
<taxonomicName box="[434,594,428,452]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[434,594,428,452]" italics="true" pageId="6" pageNumber="398">Leptanilloides</emphasis>
|
||
</taxonomicName>
|
||
genus-group, males of only three have been given species names, and of those three, only males of
|
||
<emphasis box="[444,576,463,488]" italics="true" pageId="6" pageNumber="398">L. nubecula</emphasis>
|
||
Donoso, Vieira & Wild and
|
||
<emphasis box="[896,1037,463,488]" italics="true" pageId="6" pageNumber="398">L. mckennae</emphasis>
|
||
Longino were associated with their respective worker castes (Donosa
|
||
<emphasis box="[537,594,500,524]" italics="true" pageId="6" pageNumber="398">et al.</emphasis>
|
||
<number box="[601,657,500,524]" pageId="6" pageNumber="398" value="2006.0">2006</number>
|
||
,Ward
|
||
<number box="[732,792,500,524]" pageId="6" pageNumber="398" value="2007.0">2007</number>
|
||
). The description of
|
||
<emphasis box="[1027,1238,500,524]" italics="true" pageId="6" pageNumber="398">
|
||
<taxonomicName authority="Kusnezov, 1953" authorityName="Kusnezov" authorityYear="1953" box="[1027,1238,500,524]" class="Insecta" family="Formicidae" genus="Amyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="golbachi">Amyrmex golbachi</taxonomicName>
|
||
</emphasis>
|
||
was based solely on the male caste (Kusnezov
|
||
<number box="[505,564,535,560]" pageId="6" pageNumber="398" value="1953.0">1953</number>
|
||
). Three other male morphotypes were described by Borowiec & Longino (
|
||
<number box="[159,218,572,596]" pageId="6" pageNumber="398" value="2011.0">2011</number>
|
||
), but, because they were unassociated with workers, they could not be certain of their specific identities. Therefore, Borowiec & Longino (
|
||
<number box="[542,599,607,632]" pageId="6" pageNumber="398" value="2011.0">2011</number>
|
||
) simply referred to these male morphotypes as Leptanilloidinae male
|
||
<number box="[1415,1429,608,632]" pageId="6" pageNumber="398" value="1.0">1</number>
|
||
, male
|
||
<number box="[213,228,644,668]" pageId="6" pageNumber="398" value="2.0">2</number>
|
||
, and male
|
||
<number box="[351,367,644,668]" pageId="6" pageNumber="398" value="3.0">3</number>
|
||
. Borowiec & Longino (
|
||
<number box="[636,693,644,668]" pageId="6" pageNumber="398" value="2011.0">2011</number>
|
||
) stated that "Leptanilloidinae male
|
||
<number box="[1096,1108,644,668]" pageId="6" pageNumber="398" value="1.0">1</number>
|
||
" was likely conspecific with
|
||
<emphasis box="[151,268,679,704]" italics="true" pageId="6" pageNumber="398">L. gracilis</emphasis>
|
||
Borowiec & Longino,
|
||
<number box="[535,590,680,704]" pageId="6" pageNumber="398" value="2011.0">2011</number>
|
||
based on various shared morphological features, the relatively small size, and the largely sympatric distribution of "Leptanilloidinae male
|
||
<number box="[913,925,716,740]" pageId="6" pageNumber="398" value="1.0">1</number>
|
||
" and workers of
|
||
<emphasis box="[1141,1261,716,740]" italics="true" pageId="6" pageNumber="398">L. gracilis</emphasis>
|
||
. The status of Leptanilloidinae males
|
||
<number box="[422,437,752,776]" pageId="6" pageNumber="398" value="2.0">2</number>
|
||
and
|
||
<number box="[500,514,752,776]" pageId="6" pageNumber="398" value="3.0">3</number>
|
||
is less clear. These male morphotypes might represent the males of described species in the group or they may be new species. While searching unidentified material in the Bohart Museum of Entomology collection (
|
||
<collectionCode box="[422,505,823,848]" collectionName="USA, California, Davis, University of California, R.M. Bohart Museum of Entomology" pageId="6" pageNumber="398">UCDC</collectionCode>
|
||
), Ward & Brady (
|
||
<number box="[705,763,823,848]" pageId="6" pageNumber="398" value="2009.0">2009</number>
|
||
) discovered several male
|
||
<taxonomicName box="[1056,1216,823,848]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1056,1216,823,848]" italics="true" pageId="6" pageNumber="398">Leptanilloides</emphasis>
|
||
</taxonomicName>
|
||
genus-group males that represent at least five more species in the group. These appeared to belong to
|
||
<taxonomicName authority="Kusnezov, 1953" authorityName="Kusnezov" authorityYear="1953" box="[1169,1275,860,884]" class="Insecta" family="Formicidae" genus="Amyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1169,1275,860,884]" italics="true" pageId="6" pageNumber="398">Amyrmex</emphasis>
|
||
</taxonomicName>
|
||
and possibly
|
||
<emphasis box="[151,366,895,920]" italics="true" pageId="6" pageNumber="398">
|
||
<taxonomicName box="[151,362,895,920]" class="Insecta" family="Formicidae" genus="Asphinctanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="genus">Asphinctanilloides</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
although it is not clear whether they represented new species or were the males of described species in the group.
|
||
</paragraph>
|
||
<paragraph pageId="6" pageNumber="398">
|
||
All of the described species and even the undescribed males discussed by Ward & Brady (
|
||
<number box="[1289,1347,967,992]" pageId="6" pageNumber="398" value="2009.0">2009</number>
|
||
) in the
|
||
<taxonomicName box="[151,311,1004,1028]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[151,311,1004,1028]" italics="true" pageId="6" pageNumber="398">Leptanilloides</emphasis>
|
||
</taxonomicName>
|
||
genus-group have exclusively Neotropical distributions ranging from Chiapas, Mexico (Borowiec & Longino
|
||
<number box="[253,309,1039,1064]" pageId="6" pageNumber="398" value="2011.0">2011</number>
|
||
) to southeastern Brazil (Silva
|
||
<emphasis box="[652,708,1039,1064]" italics="true" pageId="6" pageNumber="398">et al.</emphasis>
|
||
<number box="[716,776,1039,1064]" pageId="6" pageNumber="398" value="2013.0">2013</number>
|
||
). Based on reported specimen data, members of this genus group have been collected mostly in cloud forests (Borowiec & Longino,
|
||
<number box="[979,1037,1076,1100]" pageId="6" pageNumber="398" value="2011.0">2011</number>
|
||
; Brandão
|
||
<emphasis box="[1151,1208,1076,1100]" italics="true" pageId="6" pageNumber="398">et al.</emphasis>
|
||
<number box="[1214,1273,1076,1100]" pageId="6" pageNumber="398" value="1999.0">1999</number>
|
||
; Donoso
|
||
<emphasis box="[1380,1437,1076,1100]" italics="true" pageId="6" pageNumber="398">et al.</emphasis>
|
||
<number box="[151,210,1112,1136]" pageId="6" pageNumber="398" value="2006.0">2006</number>
|
||
; Longino
|
||
<number box="[333,391,1112,1136]" pageId="6" pageNumber="398" value="2003.0">2003</number>
|
||
, Silva
|
||
<emphasis box="[476,538,1111,1136]" italics="true" pageId="6" pageNumber="398">et al.</emphasis>
|
||
<number box="[549,609,1111,1136]" pageId="6" pageNumber="398" value="2013.0">2013</number>
|
||
). The new species described here, however, was collected in semiarid grassland habitat in the southern foothills of the Davis Mountains in western Texas, approximately
|
||
<quantity box="[1270,1370,1148,1172]" metricMagnitude="6" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="398" unit="km" value="2500.0">2500 km</quantity>
|
||
north of the next closest collection of the genus-group in Chiapas, Mexico.
|
||
</paragraph>
|
||
<paragraph pageId="6" pageNumber="398">
|
||
The specimens of
|
||
<emphasis box="[417,625,1220,1244]" italics="true" pageId="6" pageNumber="398">
|
||
<taxonomicName box="[417,625,1220,1244]" class="Insecta" family="Formicidae" genus="Leptanilloides" genus.bestMatchDistance="2820" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="chihuahuaensis">L. chihuahuaensis</taxonomicName>
|
||
</emphasis>
|
||
were collected in Townes style Malaise traps (Townes
|
||
<number box="[1285,1342,1220,1244]" pageId="6" pageNumber="398" value="1972.0">1972</number>
|
||
) at two locations within Davis Mountains State Park. The first location was on the south (north-facing) slope of Keesey Canyon adjacent to a hillside drainage area. The vegetation at this site was dominated by short grasses but also included scattered forbs and a few low trees and shrubs. The noncalcareous soil, of igneous origin, was shallow, well drained, and extremely rocky (Turner
|
||
<number box="[631,691,1364,1388]" pageId="6" pageNumber="398" value="1977.0">1977</number>
|
||
). The second location was in Limpia Canyon on the dry floodplain of the intermittent Limpia Creek. The soils here were similar to the first site, but the vegetation differed in having taller grasses and forbs, with grasses being less dominant, and with forbs, shrubs, and small trees being more numerous. Many rocky, unvegetated areas were also scattered through this habitat. The Malaise trap at this site was located only
|
||
<quantity box="[295,352,1508,1532]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="398" unit="m" value="20.0">20 m</quantity>
|
||
from the north (south-facing) slope of Limpia Canyon, which had similar habitat to the first site, so it is possible that the ants in this trap originated from the canyon slope rather than the floodplain.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="7" lastPageNumber="399" pageId="6" pageNumber="398" type="biology_ecology">
|
||
<paragraph pageId="6" pageNumber="398">
|
||
Other ant species collected by
|
||
<collectionCode box="[543,610,1581,1604]" collectionName="USA, Mississippi, Mississippi, Mississippi State University" pageId="6" pageNumber="398">MEM</collectionCode>
|
||
researchers in the same localities and during the same time period that
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Leptanilloides" genus.bestMatchDistance="3047" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="chihuahuaensis">
|
||
<emphasis italics="true" pageId="6" pageNumber="398">L. chihuahuaensis</emphasis>
|
||
</taxonomicName>
|
||
was collected were typical species for this arid region. Other species collected in the same Malaise traps included
|
||
<taxonomicName authority="McCook" authorityName="McCook" box="[325,662,1652,1676]" class="Insecta" family="Formicidae" genus="Dorymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="flavus">
|
||
<emphasis box="[325,550,1652,1676]" italics="true" pageId="6" pageNumber="398">Dorymyrmex flavus</emphasis>
|
||
McCook
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Roger" authorityName="Roger" box="[677,994,1652,1676]" class="Insecta" family="Formicidae" genus="Forelius" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="pruinosus">
|
||
<emphasis box="[677,893,1652,1676]" italics="true" pageId="6" pageNumber="398">Forelius pruinosus</emphasis>
|
||
(Roger)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis box="[1010,1187,1653,1676]" italics="true" pageId="6" pageNumber="398">
|
||
<taxonomicName box="[1010,1107,1653,1676]" class="Insecta" family="Formicidae" genus="Formica" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="genus">Formica</taxonomicName>
|
||
gnava
|
||
</emphasis>
|
||
Buckley,
|
||
<taxonomicName authority="Emery" authorityName="Emery" class="Insecta" family="Formicidae" genus="Myrmecina" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="americana">
|
||
<emphasis italics="true" pageId="6" pageNumber="398">Myrmecina americana</emphasis>
|
||
Emery
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authority="Forel" authorityName="Forel" box="[417,764,1687,1712]" class="Insecta" family="Formicidae" genus="Myrmecocystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="melliger">
|
||
<emphasis box="[417,694,1687,1712]" italics="true" pageId="6" pageNumber="398">Myrmecocystus melliger</emphasis>
|
||
Forel
|
||
</taxonomicName>
|
||
, as well as unidentified males of
|
||
<emphasis italics="true" pageId="6" pageNumber="398">
|
||
<taxonomicName box="[1149,1316,1688,1712]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="genus">Crematogaster</taxonomicName>
|
||
,
|
||
<taxonomicName box="[1329,1432,1687,1712]" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
|
||
,
|
||
<taxonomicName box="[151,273,1724,1748]" class="Insecta" family="Formicidae" genus="Solenopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="genus">Solenopsis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<emphasis box="[331,477,1724,1748]" italics="true" pageId="6" pageNumber="398">
|
||
<taxonomicName box="[331,473,1724,1748]" class="Insecta" family="Formicidae" genus="Strumigenys" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="genus">Strumigenys</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
General collections in the same area yielded species including
|
||
<taxonomicName box="[1180,1436,1724,1748]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1180,1436,1724,1748]" italics="true" pageId="6" pageNumber="398">Camponotus festinatus</emphasis>
|
||
</taxonomicName>
|
||
(Buckley),
|
||
<taxonomicName box="[283,582,1759,1784]" pageId="6" pageNumber="398">
|
||
<emphasis box="[283,476,1760,1784]" italics="true" pageId="6" pageNumber="398">C. semitestaceus</emphasis>
|
||
Snelling
|
||
</taxonomicName>
|
||
(found in higher elevations),
|
||
<taxonomicName authority="McCook" authorityName="McCook" box="[946,1279,1759,1784]" class="Insecta" family="Formicidae" genus="Forelius" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="mccooki">
|
||
<emphasis box="[946,1146,1759,1784]" italics="true" pageId="6" pageNumber="398">Forelius mccooki</emphasis>
|
||
(McCook)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Mayr" authorityName="Mayr" class="Insecta" family="Formicidae" genus="Liometopum" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="apiculatum">
|
||
<emphasis italics="true" pageId="6" pageNumber="398">Liometopum apiculatum</emphasis>
|
||
Mayr
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Roger" authorityName="Roger" box="[360,687,1796,1820]" class="Insecta" family="Formicidae" genus="Odontomachus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="clarus">
|
||
<emphasis box="[360,609,1796,1820]" italics="true" pageId="6" pageNumber="398">Odontomachus clarus</emphasis>
|
||
Roger
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Cresson" authorityName="Cresson" box="[700,1049,1796,1820]" class="Insecta" family="Formicidae" genus="Neivamyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis box="[700,929,1797,1820]" italics="true" pageId="6" pageNumber="398">Neivamyrmex minor</emphasis>
|
||
(Cresson)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Watkins" authorityName="Watkins" box="[1062,1285,1796,1820]" class="Insecta" family="Formicidae" genus="Neivamyrmex" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="texanus">
|
||
<emphasis box="[1062,1182,1797,1820]" italics="true" pageId="6" pageNumber="398">N. texanus</emphasis>
|
||
Watkins
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Emery, 1915" authorityName="Emery" authorityYear="1915" class="Insecta" family="Formicidae" genus="Novomessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="6" pageNumber="398">Novomessor cockerelli</emphasis>
|
||
</taxonomicName>
|
||
(André),
|
||
<taxonomicName box="[379,554,1831,1856]" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[379,554,1831,1856]" italics="true" pageId="6" pageNumber="398">Pheidole hyatti</emphasis>
|
||
</taxonomicName>
|
||
Emery,
|
||
<taxonomicName box="[657,760,1832,1856]" pageId="6" pageNumber="398">
|
||
<emphasis box="[657,760,1832,1856]" italics="true" pageId="6" pageNumber="398">P. titanis</emphasis>
|
||
</taxonomicName>
|
||
Wheeler,
|
||
<emphasis box="[884,903,1832,1856]" italics="true" pageId="6" pageNumber="398">P.</emphasis>
|
||
spp.,
|
||
<taxonomicName authority="F. Smith" authorityName="F. Smith" box="[980,1399,1831,1856]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="barbatus">
|
||
<emphasis box="[980,1268,1831,1856]" italics="true" pageId="6" pageNumber="398">Pogonomyrmex barbatus</emphasis>
|
||
(F. Smith)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Wheeler" authorityName="Wheeler" class="Insecta" family="Formicidae" genus="Pogonomyrmex" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="imberbiculus">
|
||
<emphasis italics="true" pageId="6" pageNumber="398">P. imberbiculus</emphasis>
|
||
Wheeler
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Emery" authorityName="Emery" box="[412,611,1869,1892]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="rugosus">
|
||
<emphasis box="[412,527,1869,1892]" italics="true" pageId="6" pageNumber="398">P. rugosus</emphasis>
|
||
Emery
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authority="McCook" authorityName="McCook" box="[669,1136,1868,1892]" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="septentrionalis">
|
||
<emphasis box="[669,1009,1868,1892]" italics="true" pageId="6" pageNumber="398">Trachymyrmex septentrionalis</emphasis>
|
||
(McCook)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph lastPageId="7" lastPageNumber="399" pageId="6" pageNumber="398">
|
||
The climate in the vicinity of Davis Mountains State Park is marked by hot summers and cool, dry winters (Bell
|
||
<emphasis box="[214,263,1940,1964]" italics="true" pageId="6" pageNumber="398">et al</emphasis>
|
||
.
|
||
<number box="[277,337,1940,1964]" pageId="6" pageNumber="398" value="2014.0">2014</number>
|
||
), with over
|
||
<number box="[470,498,1940,1964]" pageId="6" pageNumber="398" value="80.0">80</number>
|
||
% of annual precipitation occurring from May through October (
|
||
<collectionCode box="[1224,1306,1940,1964]" pageId="6" pageNumber="398">NOAA</collectionCode>
|
||
<number box="[1313,1374,1940,1964]" pageId="6" pageNumber="398" value="2014.0">2014</number>
|
||
). The average annual precipitation at nearby Fort Davis is
|
||
<quantity box="[742,836,1976,2000]" metricMagnitude="-1" metricUnit="m" metricValue="4.4399999999999995" pageId="6" pageNumber="398" unit="cm" value="44.4">44.4 cm</quantity>
|
||
, compared to the
|
||
<quantity box="[1037,1214,1976,2000]" metricMagnitude="-1" metricUnit="m" metricValue="2.545" metricValueMax="3.05" metricValueMin="2.04" pageId="6" pageNumber="398" unit="cm" value="25.45" valueMax="30.5" valueMin="20.4">20.4 to 30.5 cm</quantity>
|
||
of rainfall received by the surrounding lower elevation areas of Trans-Pecos Texas (
|
||
<collectionCode box="[883,966,2012,2036]" pageId="6" pageNumber="398">NOAA</collectionCode>
|
||
<number box="[975,1034,2012,2036]" pageId="6" pageNumber="398" value="2014.0">2014</number>
|
||
; Texas State Historical Association
|
||
<number box="[151,211,151,176]" pageId="7" pageNumber="399" value="2010.0">2010</number>
|
||
). This difference in precipitation is a due to the pronounced orographic lifting caused by the Davis Mountains, which results in an approximately
|
||
<quantity box="[671,726,189,212]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="399" unit="cm" value="5.0">5 cm</quantity>
|
||
increase in annual rainfall for each
|
||
<quantity box="[1128,1199,188,212]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="7" pageNumber="399" unit="m" value="305.0">305 m</quantity>
|
||
increase in elevation (DeBaca
|
||
<number box="[256,315,224,248]" pageId="7" pageNumber="399" value="2008.0">2008</number>
|
||
; Turner
|
||
<number box="[409,469,223,248]" pageId="7" pageNumber="399" value="1977.0">1977</number>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="7" pageNumber="399">
|
||
The Davis Mountain range is located within the Chihuahuan Desert ecoregion, which occupies almost
|
||
<number box="[1345,1391,260,284]" pageId="7" pageNumber="399" value="650.0">650</number>
|
||
,
|
||
<geoCoordinate box="[1395,1436,260,284]" pageId="7" pageNumber="399">0 0 0</geoCoordinate>
|
||
square kilometers extending from central Mexico into western Texas, central and southern New Mexico, and southeastern Arizona (Commission for Environmental Cooperation
|
||
<number box="[931,990,332,356]" pageId="7" pageNumber="399" value="2006.0">2006</number>
|
||
; Griffith
|
||
<emphasis box="[1100,1152,332,356]" italics="true" pageId="7" pageNumber="399">et al</emphasis>
|
||
.
|
||
<number box="[1170,1229,332,356]" pageId="7" pageNumber="399" value="2007.0">2007</number>
|
||
; Hoyt
|
||
<number box="[1310,1370,332,356]" pageId="7" pageNumber="399" value="2007.0">2007</number>
|
||
). The isolating effect of the complex topography of this extensive ecoregion, which is characterized by low-lying basins and isolated mountain ranges, contributes to the Chihuanhuan Desert being one of the most diverse arid regions in the world (Bell
|
||
<emphasis box="[334,385,439,464]" italics="true" pageId="7" pageNumber="399">et al</emphasis>
|
||
.
|
||
<number box="[402,461,440,464]" pageId="7" pageNumber="399" value="2014.0">2014</number>
|
||
; Hernández & Gómez-Hinostrosa
|
||
<number box="[857,917,439,464]" pageId="7" pageNumber="399" value="2005.0">2005</number>
|
||
). This diversity is threatened by overgrazing, water diversion, excessive groundwater pumping, introduced species, and overcollecting of animals and plants, such as cacti, for the commercial trade (Hoyt
|
||
<number box="[664,724,511,536]" pageId="7" pageNumber="399" value="2002.0">2002</number>
|
||
). The collection of
|
||
<emphasis box="[942,1102,511,536]" italics="true" pageId="7" pageNumber="399">
|
||
<taxonomicName box="[942,1102,511,536]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="399" phylum="Arthropoda" rank="genus">Leptanilloides</taxonomicName>
|
||
</emphasis>
|
||
from this region is surprising both because of the remarkable range extension it represents for the genus-group as well as the significantly different
|
||
<typeStatus box="[254,302,584,608]" pageId="7" pageNumber="399">type</typeStatus>
|
||
of habitat the specimens were collected in compared to other collections of the genus-group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="7" pageNumber="399" type="discussion">
|
||
<paragraph pageId="7" pageNumber="399">
|
||
Despite the fact that males of all species in the
|
||
<emphasis box="[743,904,620,644]" italics="true" pageId="7" pageNumber="399">
|
||
<taxonomicName box="[743,904,620,644]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="399" phylum="Arthropoda" rank="genus">Leptanilloides</taxonomicName>
|
||
</emphasis>
|
||
genus-group have not been described and that males collected in Texas were unassociated with a colony, we believe there is sufficient evidence for describing
|
||
<emphasis italics="true" pageId="7" pageNumber="399">
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Leptanilloides" genus.bestMatchDistance="3558" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="399" phylum="Arthropoda" rank="species" species="chihuahuaensis">L. chihuahuaensis</taxonomicName>
|
||
</emphasis>
|
||
as a new species. First,
|
||
<emphasis box="[594,797,692,716]" italics="true" pageId="7" pageNumber="399">
|
||
<taxonomicName box="[594,797,692,716]" class="Insecta" family="Formicidae" genus="Leptanilloides" genus.bestMatchDistance="3568" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="399" phylum="Arthropoda" rank="species" species="chihuahuaensis">L. chihuahuaensis</taxonomicName>
|
||
</emphasis>
|
||
has a very disjunct distribution and occurs in a markedly different habitat than other species. Second, there are obvious morphological features that distinguish
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Leptanilloides" genus.bestMatchDistance="3596" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="399" phylum="Arthropoda" rank="species" species="chihuahuaensis">
|
||
<emphasis italics="true" pageId="7" pageNumber="399">L. chihuahuaensis</emphasis>
|
||
</taxonomicName>
|
||
from other described males. And lastly, COI sequence divergence, as compared to several other species with data in GenBank, including
|
||
<emphasis box="[623,740,799,824]" italics="true" pageId="7" pageNumber="399">L. gracilis</emphasis>
|
||
, exceeds
|
||
<number box="[854,868,800,824]" pageId="7" pageNumber="399" value="8.0">8</number>
|
||
%. Although we do not think that this divergence level by itself is sufficient for species status - it has been shown that intraspecific divergence can be quite high (e.g. Meyer and Paulay
|
||
<number box="[364,422,872,896]" pageId="7" pageNumber="399" value="2005.0">2005</number>
|
||
, Wiemers and Fiedler
|
||
<number box="[678,735,871,896]" pageId="7" pageNumber="399" value="2007.0">2007</number>
|
||
) – we do believe that the result strengthens our argument. We also feel confident in using the generic name
|
||
<taxonomicName box="[664,824,908,932]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="399" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[664,824,908,932]" italics="true" pageId="7" pageNumber="399">Leptanilloides</emphasis>
|
||
</taxonomicName>
|
||
because specimens of
|
||
<taxonomicName box="[1083,1287,908,932]" class="Insecta" family="Formicidae" genus="Leptanilloides" genus.bestMatchDistance="3698" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="399" phylum="Arthropoda" rank="species" species="chihuahuaensis">
|
||
<emphasis box="[1083,1287,908,932]" italics="true" pageId="7" pageNumber="399">L. chihuahuaensis</emphasis>
|
||
</taxonomicName>
|
||
most closely match Borowiec & Longino's (
|
||
<number box="[501,558,943,968]" pageId="7" pageNumber="399" value="2011.0">2011</number>
|
||
) description and figures of "Leptanilloidinae male
|
||
<number box="[1138,1150,944,968]" pageId="7" pageNumber="399" value="1.0">1</number>
|
||
," which they believed to be conspecific with
|
||
<emphasis box="[378,501,980,1004]" italics="true" pageId="7" pageNumber="399">L. gracilis.</emphasis>
|
||
Additionally, the taxonomic status of the genera in the
|
||
<taxonomicName box="[1128,1288,980,1004]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="399" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1128,1288,980,1004]" italics="true" pageId="7" pageNumber="399">Leptanilloides</emphasis>
|
||
</taxonomicName>
|
||
genus-group remains unresolved with a strong likelihood that
|
||
<emphasis box="[746,1088,1015,1040]" italics="true" pageId="7" pageNumber="399">
|
||
<taxonomicName authority="Kusnezov, 1953" authorityName="Kusnezov" authorityYear="1953" box="[746,855,1016,1040]" class="Insecta" family="Formicidae" genus="Amyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="399" phylum="Arthropoda" rank="genus">Amyrmex</taxonomicName>
|
||
,
|
||
<taxonomicName box="[873,1084,1015,1040]" class="Insecta" family="Formicidae" genus="Asphinctanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="399" phylum="Arthropoda" rank="genus">Asphinctanilloides</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName box="[1156,1316,1015,1040]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="399" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1156,1316,1015,1040]" italics="true" pageId="7" pageNumber="399">Leptanilloides</emphasis>
|
||
</taxonomicName>
|
||
might be synonymized with one another. If these genera were synonymized, the name
|
||
<emphasis box="[1013,1173,1052,1076]" italics="true" pageId="7" pageNumber="399">
|
||
<taxonomicName box="[1013,1173,1052,1076]" class="Insecta" family="Formicidae" genus="Leptanilloides" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="399" phylum="Arthropoda" rank="genus">Leptanilloides</taxonomicName>
|
||
</emphasis>
|
||
would have priority.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |